351 lines
56 KiB
XML
351 lines
56 KiB
XML
<document id="B559233293173DFB91D7DDD52E1DCD6E" ID-CLB-Dataset="294723" ID-DOI="10.4202/app.00415.2017" ID-GBIF-Dataset="e49c83a7-db59-4f74-96ec-ee9a3edb85b3" ID-ISSN="1732-2421" ID-Zenodo-Dep="10980869" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1713284114161" checkinUser="felipe" docAuthor="Baeza-Carratalá, José Francisco, Pérez-Valera, Fernando & Pérez-Valera, Juan Alberto" docDate="2018" docId="7E578795D521A422FF99F8DCFDAAFD84" docLanguage="en" docName="ActaPalaeontolPol.63.1.71-85.pdf" docOrigin="Acta Palaeontologica Polonica 63 (1)" docSource="https://www.mendeley.com/catalogue/6ca80b91-ddf1-3ada-9801-704ee7a80d97/" docStyle="DocumentStyle:E4780F6D34569985EDF9CACFB9BC780E.4:ActaPalaeontolPol.2015-.journal_article" docStyleId="E4780F6D34569985EDF9CACFB9BC780E" docStyleName="ActaPalaeontolPol.2015-.journal_article" docStyleVersion="4" docTitle="Misunithyris goyi Baeza-Carratalá & Pérez-Valera & Pérez-Valera 2018, sp. nov." docType="treatment" docVersion="6" lastPageNumber="80" masterDocId="826EFFEDD524A42BFFFBFFDAFF8AFFF0" masterDocTitle="The oldest post-Paleozoic (Ladinian, Triassic) brachiopods from the Betic Range, SE Spain" masterLastPageNumber="85" masterPageNumber="71" pageNumber="76" updateTime="1713995498171" updateUser="carolina" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B42D29ED91D79A5F6F6517B99E18EE7F">The oldest post-Paleozoic (Ladinian, Triassic) brachiopods from the Betic Range, SE Spain</mods:title>
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<mods:namePart id="D6A907BB06BE35BC0CEB947196525A57">Baeza-Carratalá, José Francisco</mods:namePart>
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<mods:namePart id="E5C1BA116A8BA749147856310A64F269">Pérez-Valera, Fernando</mods:namePart>
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<mods:namePart id="A38CC8636B326AB5D29A43FD6221A33C">Pérez-Valera, Juan Alberto</mods:namePart>
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<mods:typeOfResource id="073C65A35F5CBA777C6D0EF25012614C">text</mods:typeOfResource>
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<mods:title id="DFBE97A80EBBA17718DEA305019B0DC4">Acta Palaeontologica Polonica</mods:title>
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<mods:date id="D3BE77E70D6F0C13B78D96D85319E750">2018</mods:date>
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<treatment id="7E578795D521A422FF99F8DCFDAAFD84" ID-DOI="http://doi.org/10.5281/zenodo.11061685" ID-GBIF-Taxon="226526991" ID-Zenodo-Dep="11061685" LSID="urn:lsid:plazi:treatment:7E578795D521A422FF99F8DCFDAAFD84" httpUri="http://treatment.plazi.org/id/7E578795D521A422FF99F8DCFDAAFD84" lastPageId="9" lastPageNumber="80" pageId="5" pageNumber="76">
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<subSubSection id="BEE46508D521A42EFF99F8DCFE27F8D2" box="[98,429,1798,1826]" pageId="5" pageNumber="76" type="nomenclature">
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<paragraph id="F6413683D521A42EFF99F8DCFE27F8D2" blockId="5.[98,429,1798,1826]" box="[98,429,1798,1826]" pageId="5" pageNumber="76">
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<heading id="AD0981EFD521A42EFF99F8DCFE27F8D2" box="[98,429,1798,1826]" fontSize="12" level="2" pageId="5" pageNumber="76" reason="2">
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<taxonomicName id="31FE4D00D521A42EFF99F8DCFEC8F8D2" authority="Baeza-Carratalá & Pérez-Valera & Pérez-Valera, 2018" authorityName="Baeza-Carratalá & Pérez-Valera & Pérez-Valera" authorityYear="2018" box="[98,322,1798,1826]" class="Rhynchonellata" family="Zeilleriidae" genus="Misunithyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="5" pageNumber="76" phylum="Brachiopoda" rank="species" species="goyi" status="sp. nov.">
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<emphasis id="C48AEA91D521A42EFF99F8DCFEC8F8D2" box="[98,322,1798,1826]" italics="true" pageId="5" pageNumber="76">Misunithyris goyi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="DFB957EAD521A42EFEB1F8DCFE27F8D2" box="[330,429,1798,1826]" pageId="5" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="BEE46508D521A42EFF99F8E8FF14F8B7" box="[98,158,1842,1863]" pageId="5" pageNumber="76" type="description">
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<paragraph id="F6413683D521A42EFF99F8E8FF14F8B7" blockId="5.[98,158,1842,1863]" box="[98,158,1842,1863]" pageId="5" pageNumber="76">
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<figureCitation id="6EC52A06D521A42EFF99F8E8FF11F8B7" box="[98,155,1842,1863]" captionStart="Fig" captionStartId="6.[151,184,1390,1410]" captionTargetBox="[153,1534,160,1365]" captionTargetId="figure-277@6.[153,1535,160,1365]" captionTargetPageId="6" captionText="Fig. 3. Intraspecific variability of zeillerid brachiopod Misunithyris goyi gen. et sp. nov. of the lower Ladinian from the South-Iberian Triassic platform, the Arroyo Hurtado (A, B), Calasparra (C, E), Talave (D, F) sections; in dorsal (A1–F1), anterior (A2–F2), and lateral (A3–F3) views. A. BQ-AH2.1. B. BQ-AH2.2, sectioned in the present work. C. BQ-CL1.1. D. BQ-TA1.1. E. BQ-CL1.2. F. BQ-TA1.2, holotype. All specimens were coated with magnesium oxide." figureDoi="http://doi.org/10.5281/zenodo.10980873" httpUri="https://zenodo.org/record/10980873/files/figure.png" pageId="5" pageNumber="76">Fig. 3</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="BEE46508D521A42EFF99F883FD6FF856" pageId="5" pageNumber="76" type="etymology">
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<paragraph id="F6413683D521A42EFF99F883FD6FF856" blockId="5.[98,770,1881,1958]" pageId="5" pageNumber="76">
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<emphasis id="C48AEA91D521A42EFF99F883FF43F89E" box="[98,201,1881,1902]" italics="true" pageId="5" pageNumber="76">Etymology</emphasis>
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: In a tribute to Antonio Goy (Complutense University of
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<collectingRegion id="343AF861D521A42EFF99F8AFFF27F87A" box="[98,173,1909,1930]" country="Spain" name="Madrid" pageId="5" pageNumber="76">Madrid</collectingRegion>
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), prominent Triassic and Jurassic palaeontologist, to whom the authors are very indebted for long term teaching and collaboration.
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</paragraph>
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</subSubSection>
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<subSubSection id="BEE46508D521A42EFCD3FF7AFAFAFC51" pageId="5" pageNumber="76" type="materials_examined">
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<paragraph id="F6413683D521A42EFCD3FF7AFC65FEFA" blockId="5.[808,1481,160,266]" pageId="5" pageNumber="76">
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<emphasis id="C48AEA91D521A42EFCD3FF7AFC26FF45" box="[808,940,160,181]" italics="true" pageId="5" pageNumber="76">Type material</emphasis>
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:
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<materialsCitation id="46963CDED521A42EFC41FF7AFC6EFF21" location="Talave" pageId="5" pageNumber="76" specimenCode="DCTMA-BQ-TA1.2" specimenCount="3" specimenCount-female="3" typeStatus="holotype">
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<typeStatus id="29458821D521A42EFC41FF7AFB9DFF45" box="[954,1047,160,181]" pageId="5" pageNumber="76" type="holotype">Holotype</typeStatus>
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||
:
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<specimenCode id="A6589EF8D521A42EFBDBFF7AFB6EFF45" box="[1056,1252,160,181]" pageId="5" pageNumber="76">DCTMA-BQ-TA1.2</specimenCode>
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||
(
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||
<figureCitation id="6EC52A06D521A42EFB0AFF7AFAB0FF45" box="[1265,1338,160,181]" captionStart="Fig" captionStartId="6.[151,184,1390,1410]" captionTargetBox="[153,1534,160,1365]" captionTargetId="figure-277@6.[153,1535,160,1365]" captionTargetPageId="6" captionText="Fig. 3. Intraspecific variability of zeillerid brachiopod Misunithyris goyi gen. et sp. nov. of the lower Ladinian from the South-Iberian Triassic platform, the Arroyo Hurtado (A, B), Calasparra (C, E), Talave (D, F) sections; in dorsal (A1–F1), anterior (A2–F2), and lateral (A3–F3) views. A. BQ-AH2.1. B. BQ-AH2.2, sectioned in the present work. C. BQ-CL1.1. D. BQ-TA1.1. E. BQ-CL1.2. F. BQ-TA1.2, holotype. All specimens were coated with magnesium oxide." figureDoi="http://doi.org/10.5281/zenodo.10980873" httpUri="https://zenodo.org/record/10980873/files/figure.png" pageId="5" pageNumber="76">
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Fig.
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<specimenCount id="E0F8FD0AD521A42EFAE4FF7AFAB0FF45" box="[1311,1338,160,181]" pageId="5" pageNumber="76" type="female">3F</specimenCount>
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</figureCitation>
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||
;
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||
<tableCitation id="BB7C0338D521A42EFABEFF7AFA01FF45" box="[1349,1419,160,181]" captionStart="Table 1" captionStartId="5.[808,861,288,309]" captionTargetId="graphics-876@5.[808,1481,415,638]" captionTargetPageId="5" captionText="Table 1. Mainbiometric measurements (in mm) and ratios of the studied specimens of Misunithyris goyi gen. et sp. nov. L, total length; W, total width; T, total thickness; R, ribs on each valve; Rp, ribbing pattern (p, primary rib; s, secondary rib, referred to bifurcate or intercalated ribs)." httpUri="http://table.plazi.org/id/A281660BD521A42EFCD3FEFAFA40FE7D" pageId="5" pageNumber="76" tableUuid="A281660BD521A42EFCD3FEFAFA40FE7D">Table 1</tableCitation>
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) from the
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<location id="F3216058D521A42EFCB4FF66FC05FF21" LSID="urn:lsid:plazi:treatment:7E578795D521A422FF99F8DCFDAAFD84:F3216058D521A42EFCB4FF66FC05FF21" box="[847,911,188,209]" name="Talave" pageId="5" pageNumber="76">Talave</location>
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section.
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</materialsCitation>
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<materialsCitation id="46963CDED521A42EFC16FF66FC65FEFA" location="Intraspecific" pageId="5" pageNumber="76" specimenCode="DCTMA-BQ-AH2.2" specimenCount="1" typeStatus="paratype">
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<typeStatus id="29458821D521A42EFC16FF66FBCFFF21" box="[1005,1093,188,209]" pageId="5" pageNumber="76" type="paratype">Paratype</typeStatus>
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||
:
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<specimenCode id="A6589EF8D521A42EFBA9FF66FA90FF21" box="[1106,1306,188,209]" pageId="5" pageNumber="76">DCTMA-BQ-AH2.2</specimenCode>
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||
(
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<figureCitation id="6EC52A06D521A42EFAD7FF66FA0EFF21" box="[1324,1412,188,209]" captionStart="Fig" captionStartId="6.[151,184,1390,1410]" captionTargetBox="[153,1534,160,1365]" captionTargetId="figure-277@6.[153,1535,160,1365]" captionTargetPageId="6" captionText="Fig. 3. Intraspecific variability of zeillerid brachiopod Misunithyris goyi gen. et sp. nov. of the lower Ladinian from the South-Iberian Triassic platform, the Arroyo Hurtado (A, B), Calasparra (C, E), Talave (D, F) sections; in dorsal (A1–F1), anterior (A2–F2), and lateral (A3–F3) views. A. BQ-AH2.1. B. BQ-AH2.2, sectioned in the present work. C. BQ-CL1.1. D. BQ-TA1.1. E. BQ-CL1.2. F. BQ-TA1.2, holotype. All specimens were coated with magnesium oxide." figureDoi="http://doi.org/10.5281/zenodo.10980873" httpUri="https://zenodo.org/record/10980873/files/figure.png" pageId="5" pageNumber="76">Figs. 3B</figureCitation>
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,
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<figureCitation id="6EC52A06D521A42EFA68FF66FA2BFF21" box="[1427,1441,188,209]" captionStart="Fig" captionStartId="7.[98,131,1824,1844]" captionTargetBox="[140,1436,161,1800]" captionTargetId="figure-118@7.[140,1436,161,1336]" captionTargetPageId="7" captionText="Fig. 4. Internal structure of zeillerid brachiopod Misunithyris goyi gen. et sp. nov. (BQ-AH2.2) from the lower Ladinian of the South-Iberian Triassic platform, the Arroyo Hurtado section. A. Transverse serial sections through the specimen (distance from the beak in mm). B. Microphotographs of acetate peels showing the hinge plates-crural bases relationship: crural bases are located in the ventral part of the hinge plates but a dorsal thickening emerges towards an early dorsal development. Section at 4.10 mm (B1) and 4.30 mm (B2) from the apex. Abbreviations: cb, crural bases; dp, deltidial plates; ld, dental lamellae; ms, median septum; pc, cardinal process; sp, septalium." figureDoi="http://doi.org/10.5281/zenodo.10980875" httpUri="https://zenodo.org/record/10980875/files/figure.png" pageId="5" pageNumber="76">4</figureCitation>
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,
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<figureCitation id="6EC52A06D521A42EFA54FF66FA37FF21" box="[1455,1469,188,209]" captionStart="Fig" captionStartId="8.[151,183,1200,1220]" captionTargetBox="[153,1531,159,1173]" captionTargetId="figure-345@8.[153,1532,159,1173]" captionTargetPageId="8" captionText="Fig. 5. Microphotographs of acetate peels from zeillerid brachiopod Misunithyris goyi gen. et sp. nov. (BQ-AH2.2) from the lower Ladinian of the South- Iberian Triassic platform, the Arroyo Hurtado section. A. Section at 1.00 mm from the apex showing dental lamellae enveloped in a thick-shelled wall. B. Section at 2.60 mm from the apex showing the disposition of deltidial plates.C, D. Sections at 3.30 and 3.50 mm from the apex, respectively, showing the first stages of hinge plates and earlier cardinalia and the evolution of the cardinal process, clearly striated and raised by a high cardinal platform. E, F. Sections at 4.10 and 4.30 mm from the apex, respectively, showing the cardinal area with the position of crural bases and the articulation system.G–I. Partial sections at 4.90, 5.70, and 7.10 mm from the apex, respectively, showing the evolution of the crural architecture and dorsal median septum development." figureDoi="http://doi.org/10.5281/zenodo.10980877" httpUri="https://zenodo.org/record/10980877/files/figure.png" pageId="5" pageNumber="76">5</figureCitation>
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).
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<location id="F3216058D521A42EFCD3FF03FC2AFF1E" LSID="urn:lsid:plazi:treatment:7E578795D521A422FF99F8DCFDAAFD84:F3216058D521A42EFCD3FF03FC2AFF1E" box="[808,928,217,238]" name="Intraspecific" pageId="5" pageNumber="76">Intraspecific</location>
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variability is shown through the paratypes (
|
||
<figureCitation id="6EC52A06D521A42EFAA1FF03FA1FFF1E" box="[1370,1429,217,238]" captionStart="Fig" captionStartId="6.[151,184,1390,1410]" captionTargetBox="[153,1534,160,1365]" captionTargetId="figure-277@6.[153,1535,160,1365]" captionTargetPageId="6" captionText="Fig. 3. Intraspecific variability of zeillerid brachiopod Misunithyris goyi gen. et sp. nov. of the lower Ladinian from the South-Iberian Triassic platform, the Arroyo Hurtado (A, B), Calasparra (C, E), Talave (D, F) sections; in dorsal (A1–F1), anterior (A2–F2), and lateral (A3–F3) views. A. BQ-AH2.1. B. BQ-AH2.2, sectioned in the present work. C. BQ-CL1.1. D. BQ-TA1.1. E. BQ-CL1.2. F. BQ-TA1.2, holotype. All specimens were coated with magnesium oxide." figureDoi="http://doi.org/10.5281/zenodo.10980873" httpUri="https://zenodo.org/record/10980873/files/figure.png" pageId="5" pageNumber="76">Fig. 3</figureCitation>
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||
) and measured in
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<tableCitation id="BB7C0338D521A42EFC59FF2FFC60FEFA" box="[930,1002,245,266]" captionStart="Table 1" captionStartId="5.[808,861,288,309]" captionTargetId="graphics-876@5.[808,1481,415,638]" captionTargetPageId="5" captionText="Table 1. Mainbiometric measurements (in mm) and ratios of the studied specimens of Misunithyris goyi gen. et sp. nov. L, total length; W, total width; T, total thickness; R, ribs on each valve; Rp, ribbing pattern (p, primary rib; s, secondary rib, referred to bifurcate or intercalated ribs)." httpUri="http://table.plazi.org/id/A281660BD521A42EFCD3FEFAFA40FE7D" pageId="5" pageNumber="76" tableUuid="A281660BD521A42EFCD3FEFAFA40FE7D">
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<location id="F3216058D521A42EFC59FF2FFC5DFEFA" LSID="urn:lsid:plazi:treatment:7E578795D521A422FF99F8DCFDAAFD84:F3216058D521A42EFC59FF2FFC5DFEFA" box="[930,983,245,266]" name="Table" pageId="5" pageNumber="76">Table</location>
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||
1
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</tableCitation>
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||
.
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</materialsCitation>
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</paragraph>
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<paragraph id="F6413683D521A42EFCD3FD45FAF6FD44" blockId="5.[808,1404,671,692]" box="[808,1404,671,692]" pageId="5" pageNumber="76">
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<emphasis id="C48AEA91D521A42EFCD3FD45FC29FD44" box="[808,931,671,692]" italics="true" pageId="5" pageNumber="76">Type locality</emphasis>
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: Talave section (External Betic Zone, SE Spain).
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</paragraph>
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<paragraph id="F6413683D521A42EFCD3FD1BFB64FD03" blockId="5.[808,1480,705,755]" pageId="5" pageNumber="76">
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||
<emphasis id="C48AEA91D521A42EFCD3FD1BFC23FD26" box="[808,937,705,726]" italics="true" pageId="5" pageNumber="76">Type horizon</emphasis>
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: Upper member of Siles Formation, Fassanian (early Ladinian);
|
||
<taxonomicName id="31FE4D00D521A42EFC69FD04FB64FD03" authority="Biochronozone." authorityName="Biochronozone." box="[914,1262,734,755]" class="Cephalopoda" family="Hungaritidae" genus="Gevanites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ceratitida" pageId="5" pageNumber="76" phylum="Mollusca" rank="species" species="epigonus">
|
||
<emphasis id="C48AEA91D521A42EFC69FD04FBC5FD03" box="[914,1103,734,755]" italics="true" pageId="5" pageNumber="76">Gevanites epigonus</emphasis>
|
||
Biochronozone.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="F6413683D521A42EFCD3FCDCFAFAFC51" blockId="5.[808,1482,774,929]" pageId="5" pageNumber="76">
|
||
<emphasis id="C48AEA91D521A42EFCD3FCDCFC02FCEF" box="[808,904,774,799]" italics="true" pageId="5" pageNumber="76">Material</emphasis>
|
||
.—
|
||
<specimenCount id="E0F8FD0AD521A42EFC51FCDCFBC3FCEF" box="[938,1097,774,799]" count="6" pageId="5" pageNumber="76" type="generic">Six specimens</specimenCount>
|
||
(BQ-CL1.1, BQ-CL1.2, BQ-TA1.1, BQ-TA1.2, BQ-AH2.1, and BQ-AH2.2), including
|
||
<typeStatus id="29458821D521A42EFA9FFCFDFA1FFCB0" box="[1380,1429,807,832]" pageId="5" pageNumber="76">type</typeStatus>
|
||
material, with diverse state of preservation. All individuals show articulated but generally fragmented and encrusted valves. Internal recrystallization is visible in
|
||
<specimenCount id="E0F8FD0AD521A42EFB3EFC52FAE7FC51" box="[1221,1389,904,929]" count="2" pageId="5" pageNumber="76" type="generic">two specimens</specimenCount>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BEE46508D521A42EFCD3FC6EFA85FAE2" pageId="5" pageNumber="76" type="diagnosis">
|
||
<paragraph id="F6413683D521A42EFCD3FC6EFA85FAE2" blockId="5.[808,1481,948,1298]" pageId="5" pageNumber="76">
|
||
<emphasis id="C48AEA91D521A42EFCD3FC6EFC11FC3D" box="[808,923,948,973]" italics="true" pageId="5" pageNumber="76">Diagnosis</emphasis>
|
||
.—Medium-sized and ventribiconvex zeilleriid, pyriform in outline with strong sub-labiate beak and epithyrid/permesothyrid foramen. The lateral commissure is straight and the anterior one is clearly sulcate. Sharp ribs (8–13) appearing only near the anterior margin (
|
||
<quantity id="31069B66D521A42EFAAFFBECFA16FBBF" box="[1364,1436,1078,1103]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.27" metricValueMin="0.762" pageId="5" pageNumber="76" unit="in" value="4.0" valueMax="5.0" valueMin="3.0">3–5 in</quantity>
|
||
the median sulcus), often bifurcate and occasionally intercalate. Deltidial plates present and well-developed striate cardinal process. Crural bases are given off dorsally and hinge plates initially fused and subparallel. Dorsal septum long, exceeding the length of the brachidium. Long descending branches, distally with ventral progression.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BEE46508D521A42DFCD3FAFEFA00F858" lastPageId="6" lastPageNumber="77" pageId="5" pageNumber="76" type="description">
|
||
<paragraph id="F6413683D521A42EFCD3FAFEFBB9F914" blockId="5.[808,1482,1316,1960]" pageId="5" pageNumber="76">
|
||
<emphasis id="C48AEA91D521A42EFCD3FAFEFC27FACD" box="[808,941,1316,1341]" italics="true" pageId="5" pageNumber="76">Description</emphasis>
|
||
.—
|
||
<emphasis id="C48AEA91D521A42EFC2BFAFEFB1AFACD" box="[976,1168,1316,1341]" italics="true" pageId="5" pageNumber="76">External features</emphasis>
|
||
(
|
||
<figureCitation id="6EC52A06D521A42EFB66FAFEFB54FACD" box="[1181,1246,1316,1341]" captionStart="Fig" captionStartId="6.[151,184,1390,1410]" captionTargetBox="[153,1534,160,1365]" captionTargetId="figure-277@6.[153,1535,160,1365]" captionTargetPageId="6" captionText="Fig. 3. Intraspecific variability of zeillerid brachiopod Misunithyris goyi gen. et sp. nov. of the lower Ladinian from the South-Iberian Triassic platform, the Arroyo Hurtado (A, B), Calasparra (C, E), Talave (D, F) sections; in dorsal (A1–F1), anterior (A2–F2), and lateral (A3–F3) views. A. BQ-AH2.1. B. BQ-AH2.2, sectioned in the present work. C. BQ-CL1.1. D. BQ-TA1.1. E. BQ-CL1.2. F. BQ-TA1.2, holotype. All specimens were coated with magnesium oxide." figureDoi="http://doi.org/10.5281/zenodo.10980873" httpUri="https://zenodo.org/record/10980873/files/figure.png" pageId="5" pageNumber="76">Fig. 3</figureCitation>
|
||
): Medium-sized ventribiconvex shells (
|
||
<tableCitation id="BB7C0338D521A42EFC01FA9FFBC1FAAE" box="[1018,1099,1349,1374]" captionStart="Table 1" captionStartId="5.[808,861,288,309]" captionTargetId="graphics-876@5.[808,1481,415,638]" captionTargetPageId="5" captionText="Table 1. Mainbiometric measurements (in mm) and ratios of the studied specimens of Misunithyris goyi gen. et sp. nov. L, total length; W, total width; T, total thickness; R, ribs on each valve; Rp, ribbing pattern (p, primary rib; s, secondary rib, referred to bifurcate or intercalated ribs)." httpUri="http://table.plazi.org/id/A281660BD521A42EFCD3FEFAFA40FE7D" pageId="5" pageNumber="76" tableUuid="A281660BD521A42EFCD3FEFAFA40FE7D">Table 1</tableCitation>
|
||
), pyriform to subovoidal in dorsal outline. W/L ratio is rather variable, but all specimens show L> W. Thickness is about 3/5 of the length. Maximum convexity lies in the posterior third while maximum width lies in the anterior third of the shell. Maximum thickness is observed near the mid-length. The beak is massive, sub-labiate, and strong, erect to slightly incurved, with a medium-sized epithyrid to permesothyrid foramen; the beak ridges are well-perceived and blunt, developing short and narrow interareas. The lateral commissure is straight and the anterior one is clearly sulcate, with a wide and arcuate dorsal median sinus. Sulcation is generally shallow, being more pronounced in the larger specimens.
|
||
</paragraph>
|
||
<paragraph id="F6413683D521A42DFCB5F936FE2CF957" blockId="5.[808,1482,1316,1960]" lastBlockId="6.[151,825,1517,1960]" lastPageId="6" lastPageNumber="77" pageId="5" pageNumber="76">Ribbing pattern is distinctive of this species. Ribs are absent in the posterior third of the shell; costation consists of 8 to 13 ribs on each valve, 3–5 of which can be present in the median sulcus. Ribs are wide, strong, and square in cross-section, becoming stronger and sharper near the anterior margin. They are often bifurcate; the thicker specimens possess a set of weaker, intercalated ribs. Costation is more evident in the ventral valve and on the flanks of the dorsal one. In the median dorsal sulcus, ribs are only noticed from the mid-length onwards. Concentric, dense and strong growth lines are present on the entire surface giving a reticulate aspect to the shell.</paragraph>
|
||
<caption id="A281660BD522A42DFF6CFAB4FE92FA26" ID-DOI="http://doi.org/10.5281/zenodo.10980873" ID-Zenodo-Dep="10980873" httpUri="https://zenodo.org/record/10980873/files/figure.png" pageId="6" pageNumber="77" startId="6.[151,184,1390,1410]" targetBox="[153,1534,160,1365]" targetPageId="6" targetType="figure">
|
||
<paragraph id="F6413683D522A42DFF6CFAB4FE92FA26" blockId="6.[151,1536,1390,1494]" pageId="6" pageNumber="77">
|
||
Fig. 3. Intraspecific variability of zeillerid brachiopod
|
||
<taxonomicName id="31FE4D00D522A42DFD73FAB4FCA2FA72" authority="Baeza-Carratalá & Pérez-Valera & Pérez-Valera, 2018" authorityName="Baeza-Carratalá & Pérez-Valera & Pérez-Valera" authorityYear="2018" box="[648,808,1390,1410]" class="Rhynchonellata" family="Zeilleriidae" genus="Misunithyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="6" pageNumber="77" phylum="Brachiopoda" rank="species" species="goyi" status="gen. et sp. nov.">
|
||
<emphasis id="C48AEA91D522A42DFD73FAB4FCA2FA72" box="[648,808,1390,1410]" italics="true" pageId="6" pageNumber="77">Misunithyris goyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DFB957EAD522A42DFCD5FAB4FC32FA72" box="[814,952,1390,1410]" pageId="6" pageNumber="77" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
of the lower Ladinian from the South-Iberian Triassic platform, the Arroyo Hurtado (A, B), Calasparra (C, E), Talave (D, F) sections; in dorsal (A
|
||
<subScript id="6A7A34C6D522A42DFC67FA49FC2EFA51" attach="right" box="[924,932,1427,1441]" fontSize="6" pageId="6" pageNumber="77">1</subScript>
|
||
–F
|
||
<subScript id="6A7A34C6D522A42DFC47FA49FC4EFA51" attach="right" box="[956,964,1427,1441]" fontSize="6" pageId="6" pageNumber="77">1</subScript>
|
||
), anterior (A
|
||
<subScript id="6A7A34C6D522A42DFBC4FA49FBCDFA51" attach="left" box="[1087,1095,1427,1441]" fontSize="6" pageId="6" pageNumber="77">2</subScript>
|
||
–F
|
||
<subScript id="6A7A34C6D522A42DFBA4FA49FBEDFA51" attach="left" box="[1119,1127,1427,1441]" fontSize="6" pageId="6" pageNumber="77">2</subScript>
|
||
), and lateral (A
|
||
<subScript id="6A7A34C6D522A42DFB06FA49FA8FFA51" attach="left" box="[1277,1285,1427,1441]" fontSize="6" pageId="6" pageNumber="77">3</subScript>
|
||
–F
|
||
<subScript id="6A7A34C6D522A42DFAE6FA49FAAFFA51" attach="left" box="[1309,1317,1427,1441]" fontSize="6" pageId="6" pageNumber="77">3</subScript>
|
||
) views.
|
||
<emphasis id="C48AEA91D522A42DFA8CFA53FA0DFA6E" bold="true" box="[1399,1415,1417,1438]" pageId="6" pageNumber="77">A</emphasis>
|
||
. BQ-AH2.1.
|
||
<emphasis id="C48AEA91D522A42DFF6CFA7FFF2CFA4A" bold="true" box="[151,166,1445,1466]" pageId="6" pageNumber="77">B</emphasis>
|
||
. BQ-AH2.2, sectioned in the present work.
|
||
<emphasis id="C48AEA91D522A42DFDCDFA7FFDCCFA4B" bold="true" box="[566,582,1445,1467]" pageId="6" pageNumber="77">C</emphasis>
|
||
. BQ-CL1.1.
|
||
<emphasis id="C48AEA91D522A42DFD45FA7FFD44FA4A" bold="true" box="[702,718,1445,1466]" pageId="6" pageNumber="77">D</emphasis>
|
||
. BQ-TA1.1.
|
||
<emphasis id="C48AEA91D522A42DFCBEFA7FFCDEFA4A" bold="true" box="[837,852,1445,1466]" pageId="6" pageNumber="77">E</emphasis>
|
||
. BQ-CL1.2.
|
||
<emphasis id="C48AEA91D522A42DFC30FA7FFC53FA4A" bold="true" box="[971,985,1445,1466]" pageId="6" pageNumber="77">F</emphasis>
|
||
. BQ-TA1.2, holotype. All specimens were coated with magnesium oxide.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F6413683D522A42DFF46F974FA00F858" blockId="6.[151,825,1517,1960]" lastBlockId="6.[862,1535,1517,1960]" pageId="6" pageNumber="77">
|
||
<emphasis id="C48AEA91D522A42DFF46F974FE09F937" box="[189,387,1710,1735]" italics="true" pageId="6" pageNumber="77">Internal structure</emphasis>
|
||
(
|
||
<figureCitation id="6EC52A06D522A42DFE6FF974FE68F937" box="[404,482,1710,1735]" captionStart="Fig" captionStartId="7.[98,131,1824,1844]" captionTargetBox="[140,1436,161,1800]" captionTargetId="figure-118@7.[140,1436,161,1336]" captionTargetPageId="7" captionText="Fig. 4. Internal structure of zeillerid brachiopod Misunithyris goyi gen. et sp. nov. (BQ-AH2.2) from the lower Ladinian of the South-Iberian Triassic platform, the Arroyo Hurtado section. A. Transverse serial sections through the specimen (distance from the beak in mm). B. Microphotographs of acetate peels showing the hinge plates-crural bases relationship: crural bases are located in the ventral part of the hinge plates but a dorsal thickening emerges towards an early dorsal development. Section at 4.10 mm (B1) and 4.30 mm (B2) from the apex. Abbreviations: cb, crural bases; dp, deltidial plates; ld, dental lamellae; ms, median septum; pc, cardinal process; sp, septalium." figureDoi="http://doi.org/10.5281/zenodo.10980875" httpUri="https://zenodo.org/record/10980875/files/figure.png" pageId="6" pageNumber="77">Figs. 4</figureCitation>
|
||
,
|
||
<figureCitation id="6EC52A06D522A42DFE14F974FE75F937" box="[495,511,1710,1735]" captionStart="Fig" captionStartId="8.[151,183,1200,1220]" captionTargetBox="[153,1531,159,1173]" captionTargetId="figure-345@8.[153,1532,159,1173]" captionTargetPageId="8" captionText="Fig. 5. Microphotographs of acetate peels from zeillerid brachiopod Misunithyris goyi gen. et sp. nov. (BQ-AH2.2) from the lower Ladinian of the South- Iberian Triassic platform, the Arroyo Hurtado section. A. Section at 1.00 mm from the apex showing dental lamellae enveloped in a thick-shelled wall. B. Section at 2.60 mm from the apex showing the disposition of deltidial plates.C, D. Sections at 3.30 and 3.50 mm from the apex, respectively, showing the first stages of hinge plates and earlier cardinalia and the evolution of the cardinal process, clearly striated and raised by a high cardinal platform. E, F. Sections at 4.10 and 4.30 mm from the apex, respectively, showing the cardinal area with the position of crural bases and the articulation system.G–I. Partial sections at 4.90, 5.70, and 7.10 mm from the apex, respectively, showing the evolution of the crural architecture and dorsal median septum development." figureDoi="http://doi.org/10.5281/zenodo.10980877" httpUri="https://zenodo.org/record/10980877/files/figure.png" pageId="6" pageNumber="77">5</figureCitation>
|
||
): This species shows a suboctagonal to ovoidal delthyrial cavity in cross-section where the remains of a short pedicle collar and well-developed deltidial plates are visible. Dental plates are difficult to distinguish as they are enveloped in a thickened-shell wall (
|
||
<figureCitation id="6EC52A06D522A42DFD12F8F4FCA0F8B7" box="[745,810,1838,1863]" captionStart="Fig" captionStartId="8.[151,183,1200,1220]" captionTargetBox="[153,1531,159,1173]" captionTargetId="figure-345@8.[153,1532,159,1173]" captionTargetPageId="8" captionText="Fig. 5. Microphotographs of acetate peels from zeillerid brachiopod Misunithyris goyi gen. et sp. nov. (BQ-AH2.2) from the lower Ladinian of the South- Iberian Triassic platform, the Arroyo Hurtado section. A. Section at 1.00 mm from the apex showing dental lamellae enveloped in a thick-shelled wall. B. Section at 2.60 mm from the apex showing the disposition of deltidial plates.C, D. Sections at 3.30 and 3.50 mm from the apex, respectively, showing the first stages of hinge plates and earlier cardinalia and the evolution of the cardinal process, clearly striated and raised by a high cardinal platform. E, F. Sections at 4.10 and 4.30 mm from the apex, respectively, showing the cardinal area with the position of crural bases and the articulation system.G–I. Partial sections at 4.90, 5.70, and 7.10 mm from the apex, respectively, showing the evolution of the crural architecture and dorsal median septum development." figureDoi="http://doi.org/10.5281/zenodo.10980877" httpUri="https://zenodo.org/record/10980877/files/figure.png" pageId="6" pageNumber="77">Fig. 5</figureCitation>
|
||
). They are short and subparallel. Hinge teeth are massive with crenulations, inserted in broad, shallow, and also crenulated sockets; small denticula are also discernible. It has a noticeable cardinal process, striated and raised by a high cardinal platform where a central cavity is located under the myophore (
|
||
<figureCitation id="6EC52A06D522A42DFC50F9F7FC61F9B6" box="[939,1003,1581,1606]" captionStart="Fig" captionStartId="8.[151,183,1200,1220]" captionTargetBox="[153,1531,159,1173]" captionTargetId="figure-345@8.[153,1532,159,1173]" captionTargetPageId="8" captionText="Fig. 5. Microphotographs of acetate peels from zeillerid brachiopod Misunithyris goyi gen. et sp. nov. (BQ-AH2.2) from the lower Ladinian of the South- Iberian Triassic platform, the Arroyo Hurtado section. A. Section at 1.00 mm from the apex showing dental lamellae enveloped in a thick-shelled wall. B. Section at 2.60 mm from the apex showing the disposition of deltidial plates.C, D. Sections at 3.30 and 3.50 mm from the apex, respectively, showing the first stages of hinge plates and earlier cardinalia and the evolution of the cardinal process, clearly striated and raised by a high cardinal platform. E, F. Sections at 4.10 and 4.30 mm from the apex, respectively, showing the cardinal area with the position of crural bases and the articulation system.G–I. Partial sections at 4.90, 5.70, and 7.10 mm from the apex, respectively, showing the evolution of the crural architecture and dorsal median septum development." figureDoi="http://doi.org/10.5281/zenodo.10980877" httpUri="https://zenodo.org/record/10980877/files/figure.png" pageId="6" pageNumber="77">Fig. 5</figureCitation>
|
||
). Hinge plates are initially horizontal, fused and subparallel. The dorsal median septum is long and well-developed up to 2/3 of the shell. A short elliptical septalium is discernible. Crural bases are located in the ventral area of the hinge plates but an incipient dorsal thickening emerge clearly showing an early dorsal development (
|
||
<figureCitation id="6EC52A06D522A42DFAABF914FA1AF917" box="[1360,1424,1742,1767]" captionStart="Fig" captionStartId="7.[98,131,1824,1844]" captionTargetBox="[140,1436,161,1800]" captionTargetId="figure-118@7.[140,1436,161,1336]" captionTargetPageId="7" captionText="Fig. 4. Internal structure of zeillerid brachiopod Misunithyris goyi gen. et sp. nov. (BQ-AH2.2) from the lower Ladinian of the South-Iberian Triassic platform, the Arroyo Hurtado section. A. Transverse serial sections through the specimen (distance from the beak in mm). B. Microphotographs of acetate peels showing the hinge plates-crural bases relationship: crural bases are located in the ventral part of the hinge plates but a dorsal thickening emerges towards an early dorsal development. Section at 4.10 mm (B1) and 4.30 mm (B2) from the apex. Abbreviations: cb, crural bases; dp, deltidial plates; ld, dental lamellae; ms, median septum; pc, cardinal process; sp, septalium." figureDoi="http://doi.org/10.5281/zenodo.10980875" httpUri="https://zenodo.org/record/10980875/files/figure.png" pageId="6" pageNumber="77">Fig. 4</figureCitation>
|
||
), suggesting a
|
||
<taxonomicName id="31FE4D00D522A42DFC62F934FBB9F8F7" authorityName="Vörös" authorityYear="1983" box="[921,1075,1774,1799]" class="Rhynchonellata" family="Zeilleriidae" genus="Bakonyithyris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Terebratulida" pageId="6" pageNumber="77" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="C48AEA91D522A42DFC62F934FBB9F8F7" box="[921,1075,1774,1799]" italics="true" pageId="6" pageNumber="77">Bakonyithyris</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<typeStatus id="29458821D522A42DFBC0F934FBE4F8F7" box="[1083,1134,1774,1799]" pageId="6" pageNumber="77">type</typeStatus>
|
||
hinge plate/crural base inter-relation (sensu
|
||
<bibRefCitation id="926F4B72D522A42DFC51F8D4FAD3F8D7" author="Baeza-Carratala, J. F. & Garcia Joral, F." box="[938,1369,1806,1831]" pageId="6" pageNumber="77" pagination="651 - 661" refId="ref8777" refString="Baeza-Carratala, J. F. and Garcia Joral, F. 2014. Crural bases position as a structural criterion for supraspecific diagnosis of Early Jurassic zeilleriid brachiopods. Acta Palaeontologica Polonica 59: 651 - 661." type="journal article" year="2014">Baeza-Carratalá and García Joral 2014</bibRefCitation>
|
||
). The descending branches of the brachidium show subparallell and vertical plates and they are developed in the commissural plane in the posterior third of the shell, acquiring a slight ventral development anteriorly. The transverse band is not visible.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="A281660BD523A42CFF99F8FAFD7BF854" ID-DOI="http://doi.org/10.5281/zenodo.10980875" ID-Zenodo-Dep="10980875" httpUri="https://zenodo.org/record/10980875/files/figure.png" pageId="7" pageNumber="78" startId="7.[98,131,1824,1844]" targetBox="[140,1436,161,1800]" targetPageId="7" targetType="figure">
|
||
<paragraph id="F6413683D523A42CFF99F8FAFD7BF854" blockId="7.[98,1481,1824,1956]" pageId="7" pageNumber="78">
|
||
Fig. 4. Internal structure of zeillerid brachiopod
|
||
<taxonomicName id="31FE4D00D523A42CFDD9F8FAFD49F8C4" authority="Baeza-Carratalá & Pérez-Valera & Pérez-Valera, 2018" authorityName="Baeza-Carratalá & Pérez-Valera & Pérez-Valera" authorityYear="2018" box="[546,707,1824,1844]" class="Rhynchonellata" family="Zeilleriidae" genus="Misunithyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="7" pageNumber="78" phylum="Brachiopoda" rank="species" species="goyi" status="gen. et sp. nov.">
|
||
<emphasis id="C48AEA91D523A42CFDD9F8FAFD49F8C4" box="[546,707,1824,1844]" italics="true" pageId="7" pageNumber="78">Misunithyris goyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DFB957EAD523A42CFD31F8FAFCD2F8C4" box="[714,856,1824,1844]" pageId="7" pageNumber="78" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
(BQ-AH2.2) from the lower Ladinian of the South-Iberian Triassic platform, the Arroyo Hurtado section.
|
||
<emphasis id="C48AEA91D523A42CFE41F8E0FE40F8BF" bold="true" box="[442,458,1850,1871]" pageId="7" pageNumber="78">A</emphasis>
|
||
. Transverse serial sections through the specimen (distance from the beak in mm).
|
||
<emphasis id="C48AEA91D523A42CFB57F8E1FB31F8A0" bold="true" box="[1196,1211,1851,1872]" pageId="7" pageNumber="78">B</emphasis>
|
||
. Microphotographs of acetate peels showing the hinge plates-crural bases relationship: crural bases are located in the ventral part of the hinge plates but a dorsal thickening emerges towards an early dorsal development. Section at 4.10 mm (B
|
||
<subScript id="6A7A34C6D523A42CFD77F8A7FD1EF87B" attach="left" box="[652,660,1917,1931]" fontSize="6" pageId="7" pageNumber="78">1</subScript>
|
||
) and 4.30 mm (B
|
||
<subScript id="6A7A34C6D523A42CFCCCF8A7FCB5F87B" attach="left" box="[823,831,1917,1931]" fontSize="6" pageId="7" pageNumber="78">2</subScript>
|
||
) from the apex. Abbreviations: cb, crural bases; dp, deltidial plates; ld, dental lamellae; ms, median septum; pc, cardinal process; sp, septalium.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="A281660BD52CA423FF6CFB6AFAF4FAA0" ID-DOI="http://doi.org/10.5281/zenodo.10980877" ID-Zenodo-Dep="10980877" httpUri="https://zenodo.org/record/10980877/files/figure.png" pageId="8" pageNumber="79" startId="8.[151,183,1200,1220]" targetBox="[153,1531,159,1173]" targetPageId="8" targetType="figure">
|
||
<paragraph id="F6413683D52CA423FF6CFB6AFAF4FAA0" blockId="8.[151,1535,1200,1360]" pageId="8" pageNumber="79">
|
||
Fig. 5. Microphotographs of acetate peels from zeillerid brachiopod
|
||
<taxonomicName id="31FE4D00D52CA423FD05FB6AFC10FB34" authority="Baeza-Carratalá & Pérez-Valera & Pérez-Valera, 2018" authorityName="Baeza-Carratalá & Pérez-Valera & Pérez-Valera" authorityYear="2018" box="[766,922,1200,1220]" class="Rhynchonellata" family="Zeilleriidae" genus="Misunithyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="8" pageNumber="79" phylum="Brachiopoda" rank="species" species="goyi" status="gen. et sp. nov.">
|
||
<emphasis id="C48AEA91D52CA423FD05FB6AFC10FB34" box="[766,922,1200,1220]" italics="true" pageId="8" pageNumber="79">Misunithyris goyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DFB957EAD52CA423FC5AFB6AFBA0FB34" box="[929,1066,1200,1220]" pageId="8" pageNumber="79" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
(BQ-AH2.2) from the lower Ladinian of the South-Iberian Triassic platform, the Arroyo Hurtado section.
|
||
<emphasis id="C48AEA91D52CA423FD7BFB10FD1AFB2F" bold="true" box="[640,656,1226,1247]" pageId="8" pageNumber="79">A</emphasis>
|
||
. Section at 1.00 mm from the apex showing dental lamellae enveloped in a thick-shelled wall.
|
||
<emphasis id="C48AEA91D52CA423FA11FB10FA73FB2F" bold="true" box="[1514,1529,1226,1247]" pageId="8" pageNumber="79">B</emphasis>
|
||
. Section at 2.60 mm from the apex showing the disposition of deltidial plates.
|
||
<emphasis id="C48AEA91D52CA423FCC1FB3CFCC0FB0C" bold="true" box="[826,842,1254,1276]" pageId="8" pageNumber="79">C</emphasis>
|
||
,
|
||
<emphasis id="C48AEA91D52CA423FCAFFB3CFCEEFB0B" bold="true" box="[852,868,1254,1275]" pageId="8" pageNumber="79">D</emphasis>
|
||
. Sections at 3.30 and 3.50 mm from the apex, respectively, showing the first stages of hinge plates and earlier cardinalia and the evolution of the cardinal process, clearly striated and raised by a high cardinal platform.
|
||
<emphasis id="C48AEA91D52CA423FA97FAD8FAF1FAE7" bold="true" box="[1388,1403,1282,1303]" pageId="8" pageNumber="79">E</emphasis>
|
||
,
|
||
<emphasis id="C48AEA91D52CA423FA7DFAD8FA1EFAE7" bold="true" box="[1414,1428,1282,1303]" pageId="8" pageNumber="79">F</emphasis>
|
||
. Sections at 4.10 and 4.30 mm from the apex, respectively, showing the cardinal area with the position of crural bases and the articulation system.
|
||
<emphasis id="C48AEA91D52CA423FAC8FAC4FAD2FAC4" bold="true" box="[1331,1368,1310,1332]" pageId="8" pageNumber="79">G–I</emphasis>
|
||
. Partial sections at 4.90, 5.70, and 7.10 mm from the apex, respectively, showing the evolution of the crural architecture and dorsal median septum development.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="BEE46508D52CA422FF6CFAB4FDAAFD84" lastPageId="9" lastPageNumber="80" pageId="8" pageNumber="79" type="discussion">
|
||
<paragraph id="F6413683D52CA423FF6CFAB4FD8FF8D7" blockId="8.[151,825,1390,1959]" pageId="8" pageNumber="79">
|
||
<emphasis id="C48AEA91D52CA423FF6CFAB4FF72FA77" box="[151,248,1390,1415]" italics="true" pageId="8" pageNumber="79">Remarks</emphasis>
|
||
.—In addition to the main biometric ratios (
|
||
<tableCitation id="BB7C0338D52CA423FD23FAB4FCA2FA77" box="[728,808,1390,1415]" captionStart="Table 1" captionStartId="5.[808,861,288,309]" captionTargetId="graphics-876@5.[808,1481,415,638]" captionTargetPageId="5" captionText="Table 1. Mainbiometric measurements (in mm) and ratios of the studied specimens of Misunithyris goyi gen. et sp. nov. L, total length; W, total width; T, total thickness; R, ribs on each valve; Rp, ribbing pattern (p, primary rib; s, secondary rib, referred to bifurcate or intercalated ribs)." httpUri="http://table.plazi.org/id/A281660BD521A42EFCD3FEFAFA40FE7D" pageId="8" pageNumber="79" tableUuid="A281660BD521A42EFCD3FEFAFA40FE7D">Table 1</tableCitation>
|
||
), intraspecific variability mainly lies in the number of ribs present in each valve, depending on the bifurcation or intercalation. The smooth posterior stage is also variable between ½ and ¼ of the shell. A single specimen shows an exceptionally wide sulcus giving to the anterior commissure an almost rectimarginate aspect, but analyzing in detail this individual the sinus can be noticed from the early stages in the posterior areas of the shell. Thus, even width of sulcus is rather stable, it ranges from mid-width in the BQ-TA1.2 specimen up to wider developments (BQ-AH2.1 and BQ-CL1.1) occupying nearly the entire anterior margin. On the other hand, maximum width can be shifted toward the anterior third, resulting in a more trigonal dorsal outline.
|
||
</paragraph>
|
||
<paragraph id="F6413683D52CA423FF46F8F4FA27F917" blockId="8.[151,825,1390,1959]" lastBlockId="8.[862,1537,1390,1959]" pageId="8" pageNumber="79">
|
||
As it is discussed above in the supraspecific assignment, this species shows high singularity in the external and internal traits, probably because it is an endemic taxon. The closer affinity is shown with the Lower Jurassic (mostly anteriorly) multicostulate zeilleriids
|
||
<taxonomicName id="31FE4D00D52CA423FB3DFAB4FA51FA77" authorityName="Seguenza" authorityYear="1855" box="[1222,1499,1390,1415]" class="Cephalopoda" family="Hildoceratidae" genus="Tauromenia" kingdom="Animalia" pageId="8" pageNumber="79" phylum="Mollusca" rank="species" species="polymorpha">
|
||
<emphasis id="C48AEA91D52CA423FB3DFAB4FA51FA77" box="[1222,1499,1390,1415]" italics="true" pageId="8" pageNumber="79">Tauromenia polymorpha</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="31FE4D00D52CA423FA11FAB4FC6DFA57" baseAuthorityName="Dubar" baseAuthorityYear="1942" class="Cephalopoda" family="Hildoceratidae" genus="Tauromenia" kingdom="Animalia" pageId="8" pageNumber="79" phylum="Mollusca" rank="species" species="brevicostata">
|
||
<emphasis id="C48AEA91D52CA423FA11FAB4FC6DFA57" italics="true" pageId="8" pageNumber="79">T. brevicostata</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="31FE4D00D52CA423FBDEFA54FB7EFA57" box="[1061,1268,1422,1447]" class="Rhynchonellata" family="Zeilleriidae" genus="Fimbriothyris" kingdom="Animalia" order="Terebratulida" pageId="8" pageNumber="79" phylum="Brachiopoda" rank="species" species="undetermined">
|
||
<emphasis id="C48AEA91D52CA423FBDEFA54FB35FA57" box="[1061,1215,1422,1447]" italics="true" pageId="8" pageNumber="79">Fimbriothyris</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="926F4B72D52CA423FAF8FA54FA27FA57" author="Seguenza, G." box="[1283,1453,1422,1447]" pageId="8" pageNumber="79" pagination="377 - 497" refId="ref12207" refString="Seguenza, G. 1885. Monografia delle Spiriferina dei varii piani del Lias Messinese. Bollettino della Societa Geologica Italian a 4: 377 - 497." type="journal article" year="1885">Seguenza 1885</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="926F4B72D52CA423FA42FA54FC1CFA37" author="Dubar, G." pageId="8" pageNumber="79" pagination="25 - 53" refId="ref9420" refString="Dubar, G. 1942. Etudes paleontologiques sur le Lias du Maroc. Brachiopodes. Terebratules et Zeilleries multiplisees. Notes et Memoires du Service Geologique du Maroc 57: 25 - 53." type="journal article" year="1942">Dubar 1942</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="926F4B72D52CA423FC5AFA74FBDFFA37" author="Elmi, S. & Almeras, Y. & Benhamou, M. & Mekahli, L. & Marok, A." box="[929,1109,1454,1479]" pageId="8" pageNumber="79" pagination="695 - 706" refId="ref9525" refString="Elmi, S., Almeras, Y., Benhamou, M., Mekahli, L., and Marok, A. 2003. Biostratigraphie des brachiopodes et age Carixien (Pliensbachien inferieur) des calcaries a grands bivalves en Algerie occidentale. Geobios 36: 695 - 706." type="journal article" year="2003">Elmi et al. 2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="926F4B72D52CA423FB9AFA74FAB4FA37" author="Almeras, Y. & Elmi, S. & Faure, P." box="[1121,1342,1454,1479]" pageId="8" pageNumber="79" pagination="3 - 241" refId="ref8620" refString="Almeras, Y., Elmi, S., and Faure, P. 2007. Les Brachiopodes Liasiques d'Algerie Occidentale. Documents des Laboratoires de Geologie Lyon 163: 3 - 241." type="journal article" year="2007">Alméras et al. 2007</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="926F4B72D52CA423FAB1FA74FBEDFA17" author="Baeza-Carratala, J. F. & Garcia Joral, F." pageId="8" pageNumber="79" pagination="1 - 12" refId="ref8728" refString="Baeza-Carratala, J. F. and Garcia Joral, F. 2012. Multicostate zeillerids (Brachiopoda, Terebratulida) from the Lower Jurassic of the Eastern Subbetic (SE Spain) and their use in correlation and paleobiogeography. Geologica Acta 10: 1 - 12." type="journal article" year="2012">Baeza-Carratalá and García Joral 2012</bibRefCitation>
|
||
), mainly recorded in the southern part of Western Tethys and even with the Middle Jurassic
|
||
<taxonomicName id="31FE4D00D52CA423FCA5F9D4FC5DF9D7" authorityName="Muir-Wood" authorityYear="1965" box="[862,983,1550,1575]" class="Rhynchonellata" family="Eudesiidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Terebratulida" pageId="8" pageNumber="79" phylum="Brachiopoda" rank="family">Eudesiidae</taxonomicName>
|
||
representatives recorded in both Tethyan margins and the Mid-East platforms (e.g.,
|
||
<bibRefCitation id="926F4B72D52CA423FB3CF9F4FAE8F9B7" author="Almeras, Y." box="[1223,1378,1582,1607]" pageId="8" pageNumber="79" pagination="261 - 268" refId="ref8440" refString="Almeras, Y. 1987. Origin and evolution of the Jurassic Multicostate Zeileriid Brachiopods Eudesia King and Flabellothyris Deslongchamps. Palaeogeography, Paleoclimatology, Palaeoecology, 59: 261 - 268." type="journal article" year="1987">Alméras 1987</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="926F4B72D52CA423FA97F9F4FA70F9B7" author="Cooper, G. A." box="[1388,1530,1582,1607]" pageId="8" pageNumber="79" pagination="1 - 213" refId="ref9204" refString="Cooper, G. A. 1989. Jurassic brachiopods of Saudi Arabia. Smithsonian Contributions to Paleobiology 65: 1 - 213." type="journal article" year="1989">Cooper 1989</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="926F4B72D52CA423FCA5F994FBDEF997" author="Mukherjee, D. & Bardhan, S. & Ghosh, D. N." box="[862,1108,1614,1639]" pageId="8" pageNumber="79" pagination="347 - 364" refId="ref10800" refString="Mukherjee, D., Bardhan, S., and Ghosh, D. N. 2000. Evolution and migration of zeillerid brachiopods: A new record of Eudesia King from the Middle Jurassic of Kutch, India. Neues Jahrbuch fur Geologie und Palaontologie Abhandlungen 215: 347 - 364." type="journal article" year="2000">Mukherjee et al. 2000</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="926F4B72D52CA423FBA4F994FACFF997" author="Almeras Y. & Cougnon, R. & Enay, R. & Mangold, C." box="[1119,1349,1614,1639]" pageId="8" pageNumber="79" pagination="1 - 247" refId="ref8567" refString="Almeras Y., Cougnon, R., Enay, R., and Mangold, C. 2010 b. Brachiopodes du Jurassique Inferieur et Moyen d'Arabie Saoudite Centrale. Paleontologie, Biostratigraphie et Paleoenvironnements, echelles chronostratigraphiques. Documents des Laboratoires de Geologie Lyon 168: 1 - 247." type="journal article" year="2010">Alméras et al. 2010b</bibRefCitation>
|
||
), having in common with these last taxa several internal features (cardinal process present in several taxa, long median septum) but not sharing the brachidium architecture and the ribs covering the whole surface of the shell in the Middle Jurassic taxa.
|
||
</paragraph>
|
||
<paragraph id="F6413683D52CA422FC7FF934FEE8FE33" blockId="8.[862,1537,1390,1959]" lastBlockId="9.[98,771,160,451]" lastPageId="9" lastPageNumber="80" pageId="8" pageNumber="79">
|
||
Among the Triassic representatives, the closer affinities are found in
|
||
<taxonomicName id="31FE4D00D52CA423FC15F8D4FAF9F8D7" authority="Feldman, 2013" authorityName="Feldman" authorityYear="2013" box="[1006,1395,1806,1831]" class="Rhynchonellata" family="Zeilleriidae" genus="Menathyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="8" pageNumber="79" phylum="Brachiopoda" rank="species" species="wilsoni">
|
||
<emphasis id="C48AEA91D52CA423FC15F8D4FB4DF8D7" box="[1006,1223,1806,1831]" italics="true" pageId="8" pageNumber="79">Menathyris wilsoni</emphasis>
|
||
<bibRefCitation id="926F4B72D52CA423FB35F8D4FAF9F8D7" author="Feldman, H. R." box="[1230,1395,1806,1831]" pageId="8" pageNumber="79" pagination="81 - 85" refId="ref9714" refString="Feldman, H. R. 2013. Menathyris wilsoni (Brachiopoda), new genus and species from the Middle Triassic (Ladinian) of Southern Israel. Annales Societatis Geologorum Poloniae 83: 81 - 85." type="journal article" year="2013">Feldman, 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="8EE97613D52CA423FA42F8D4FA71F8D7" box="[1465,1531,1806,1831]" name="Israel" pageId="8" pageNumber="79">Israel</collectingCountry>
|
||
, virtually contemporary with the Betic material, sharing the sulcate shape of the shell, the marginal ribbing pattern and comparable internal structure, except for the cardinal process, absent in
|
||
<taxonomicName id="31FE4D00D52CA423FBF3F854FBF4F857" authorityName="Feldman" authorityYear="2013" box="[1032,1150,1934,1959]" class="Rhynchonellata" family="Zeilleriidae" genus="Menathyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="8" pageNumber="79" phylum="Brachiopoda" rank="species" species="wilsoni">
|
||
<emphasis id="C48AEA91D52CA423FBF3F854FBF4F857" box="[1032,1150,1934,1959]" italics="true" pageId="8" pageNumber="79">M. wilsoni</emphasis>
|
||
</taxonomicName>
|
||
; however, sulcus is shallower and narrower, anterior commissure is generally rectimarginate and ribs are simple without bifurcation or intercalation in such species. Curious similarity concerning the internal structure is observed in
|
||
<specimenCount id="E0F8FD0AD52DA422FE94FED9FD83FEEC" box="[367,521,259,284]" count="1" pageId="9" pageNumber="80" type="generic">one specimen</specimenCount>
|
||
of
|
||
<taxonomicName id="31FE4D00D52DA422FDD7FED9FD3DFEEC" authorityName="Douville" authorityYear="1879" box="[556,695,259,284]" class="Rhynchonellata" family="Dielasmatidae" genus="Coenothyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="9" pageNumber="80" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="C48AEA91D52DA422FDD7FED9FD3DFEEC" box="[556,695,259,284]" italics="true" pageId="9" pageNumber="80">Coenothyris</emphasis>
|
||
</taxonomicName>
|
||
with a central umbonal chamber in the cardinal process, similarly to the Betic material, and several
|
||
<taxonomicName id="31FE4D00D52DA422FE15FE9CFC88FEAF" box="[494,770,326,351]" pageId="9" pageNumber="80">
|
||
<emphasis id="C48AEA91D52DA422FE15FE9CFD2DFEAF" box="[494,679,326,351]" italics="true" pageId="9" pageNumber="80">Aulacothyroides</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
developing wide cardinal process, all of them depicted by
|
||
<bibRefCitation id="926F4B72D52DA422FF99FE53FDC9FE52" author="Senkowiczowa, H. & Popiel-Barczyk, E." box="[98,579,393,418]" pageId="9" pageNumber="80" pagination="443 - 466" refId="ref12272" refString="Senkowiczowa, H. and Popiel-Barczyk, E. 1996. Some Terebratulida (Brachiopoda) from the Muschelkalk sediments in the Holy Cross Mts. Geological Quarterly 40: 443 - 466." type="journal article" year="1996">Senkowiczowa and Popiel-Barczyk (1996)</bibRefCitation>
|
||
in the Anisian– Ladinian from
|
||
<collectingCountry id="8EE97613D52DA422FEF6FE70FED4FE33" box="[269,350,426,451]" name="Poland" pageId="9" pageNumber="80">Poland</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="F6413683D52DA422FF99FE0CFDAAFD84" blockId="9.[98,770,470,628]" pageId="9" pageNumber="80">
|
||
<emphasis id="C48AEA91D52DA422FF99FE0CFD8AFE1F" box="[98,512,470,495]" italics="true" pageId="9" pageNumber="80">Stratigraphic and geographic range</emphasis>
|
||
.—In the Betic Range, this new species has been recorded together with biostratigraphical marker of the
|
||
<taxonomicName id="31FE4D00D52DA422FE8AFDC2FEDCFDA2" authority="Biochronozone Perez-Valera F. 2005" authorityName="Biochronozone Perez-Valera F." authorityYear="2005" class="Cephalopoda" family="Hungaritidae" genus="Gevanites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ceratitida" pageId="9" pageNumber="80" phylum="Mollusca" rank="species" species="epigonus">
|
||
<emphasis id="C48AEA91D52DA422FE8AFDC2FDDAFDC1" box="[369,592,536,561]" italics="true" pageId="9" pageNumber="80">Gevanites epigonus</emphasis>
|
||
Biochronozone Pérez-Valera F. 2005
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="926F4B72D52DA422FE9AFDE3FDE0FDA2" author="Perez-Valera, J. A. & Goy, A. & Perez-Valera, F. & Perez-Lopez, A." box="[353,618,569,594]" pageId="9" pageNumber="80" pagination="23 - 24" refId="ref11661" refString="Perez-Valera, J. A., Goy, A., Perez-Valera, F., and Perez-Lopez, A. 2011. Ammonoid biostratigraphy of Muschelkalk carbonates (Ladinian) ofthe South-Iberian Triassic (Betic External Zones, S. Spain). In: A. Maate (ed.), Livret des Resumes du Sixieme Reunion du Groupe Marocain du Permien et du Trias, Tetouan, 20 - 21 May, 2011, 23 - 24. University of Tetouan, Tetouan." type="book chapter" year="2011">Pérez-Valera et al. 2011</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="926F4B72D52DA422FD8FFDE3FF13FD84" author="Perez-Valera, J. A." pageId="9" pageNumber="80" refId="ref11570" refString="Perez-Valera, J. A. 2015. Ammonoideos y bioestratigrafia del Triasico medio (Ladiniense) del sector oriental de la Cordillera Betica. 489 pp. Unpublished Ph. D. Thesis, Universidad Complutense de Madrid, Madrid." type="book" year="2015">Pérez-Valera 2015</bibRefCitation>
|
||
) of the Fassanian (early Ladinian).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |