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<document id="AFFB3B122CE94920365FD082D10AD597" ID-DOI="10.4202/app.00415.2017" ID-ISSN="1732-2421" ID-Zenodo-Dep="10980869" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1713284114161" checkinUser="felipe" docAuthor="Baeza-Carratalá, José Francisco, Pérez-Valera, Fernando &amp; Pérez-Valera, Juan Alberto" docDate="2018" docId="7E578795D520A42EFF6CFC4FFD90F91E" docLanguage="en" docName="ActaPalaeontolPol.63.1.71-85.pdf" docOrigin="Acta Palaeontologica Polonica 63 (1)" docSource="https://www.mendeley.com/catalogue/6ca80b91-ddf1-3ada-9801-704ee7a80d97/" docStyle="DocumentStyle:E4780F6D34569985EDF9CACFB9BC780E.4:ActaPalaeontolPol.2015-.journal_article" docStyleId="E4780F6D34569985EDF9CACFB9BC780E" docStyleName="ActaPalaeontolPol.2015-.journal_article" docStyleVersion="4" docTitle="Misunithyris Baeza-Carratalá &amp; Pérez-Valera &amp; Pérez-Valera 2018" docType="treatment" docVersion="3" lastPageNumber="76" masterDocId="826EFFEDD524A42BFFFBFFDAFF8AFFF0" masterDocTitle="The oldest post-Paleozoic (Ladinian, Triassic) brachiopods from the Betic Range, SE Spain" masterLastPageNumber="85" masterPageNumber="71" pageNumber="75" updateTime="1713981504003" updateUser="carolina" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1822C80D5B160CD43D16EC6930577ED7">The oldest post-Paleozoic (Ladinian, Triassic) brachiopods from the Betic Range, SE Spain</mods:title>
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<paragraph id="F6413683D520A42FFF6CFC4FFE5BFC41" blockId="4.[151,465,917,945]" box="[151,465,917,945]" pageId="4" pageNumber="75">
<heading id="AD0981EFD520A42FFF6CFC4FFE5BFC41" box="[151,465,917,945]" fontSize="12" level="2" pageId="4" pageNumber="75" reason="2">
Genus
<taxonomicName id="31FE4D00D520A42FFF0AFC4FFE19FC41" authorityName="Baeza-Carratalá &amp; Pérez-Valera &amp; Pérez-Valera" authorityYear="2018" box="[241,403,917,945]" class="Rhynchonellata" family="Zeilleriidae" genus="Misunithyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D520A42FFF0AFC4FFE19FC41" box="[241,403,917,945]" italics="true" pageId="4" pageNumber="75">Misunithyris</emphasis>
</taxonomicName>
nov.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BEE46508D520A42FFF6CFC1BFDBDFBFE" pageId="4" pageNumber="75" type="etymology">
<paragraph id="F6413683D520A42FFF6CFC1BFDBDFBFE" blockId="4.[151,824,961,1038]" pageId="4" pageNumber="75">
<emphasis id="C48AEA91D520A42FFF6CFC1BFF74FC26" box="[151,254,961,982]" italics="true" pageId="4" pageNumber="75">Etymology</emphasis>
: From Mīšūnīš, ancient name of the current Mundo River; in the Mundo River valley, the most significant outcrops of specimens from which this genus is erected are found.
</paragraph>
</subSubSection>
<subSubSection id="BEE46508D520A42FFF6CFBC0FC91FBDF" box="[151,795,1050,1071]" pageId="4" pageNumber="75" type="type_taxon">
<paragraph id="F6413683D520A42FFF6CFBC0FC91FBDF" blockId="4.[151,795,1050,1071]" box="[151,795,1050,1071]" pageId="4" pageNumber="75">
<emphasis id="C48AEA91D520A42FFF6CFBC0FE85FBDF" box="[151,271,1050,1071]" italics="true" pageId="4" pageNumber="75">
<typeStatus id="29458821D520A42FFF6CFBC0FF4EFBDF" box="[151,196,1050,1071]" pageId="4" pageNumber="75">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="31FE4D00D520A42FFEE7FBC0FE4EFBDF" authority="Baeza-Carratalá &amp; Pérez-Valera &amp; Pérez-Valera, 2018" authorityName="Baeza-Carratalá &amp; Pérez-Valera &amp; Pérez-Valera" authorityYear="2018" box="[284,452,1050,1071]" class="Rhynchonellata" family="Zeilleriidae" genus="Misunithyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="species" species="goyi" status="sp. nov.">
<emphasis id="C48AEA91D520A42FFEE7FBC0FE4EFBDF" box="[284,452,1050,1071]" italics="true" pageId="4" pageNumber="75">Misunithyris goyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DFB957EAD520A42FFE30FBC0FD9CFBDF" box="[459,534,1050,1071]" pageId="4" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
(by monotypy); see below.
</paragraph>
</subSubSection>
<subSubSection id="BEE46508D520A42FFF6CFB98FD5AFBAB" box="[151,720,1090,1115]" pageId="4" pageNumber="75" type="diagnosis">
<paragraph id="F6413683D520A42FFF6CFB98FD5AFBAB" blockId="4.[151,720,1090,1115]" box="[151,720,1090,1115]" pageId="4" pageNumber="75">
<emphasis id="C48AEA91D520A42FFF6CFB98FE83FBAB" box="[151,265,1090,1115]" italics="true" pageId="4" pageNumber="75">Diagnosis</emphasis>
.—As for the
<typeStatus id="29458821D520A42FFE58FB98FE5FFBAB" box="[419,469,1090,1115]" pageId="4" pageNumber="75">type</typeStatus>
species by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="BEE46508D520A42EFF6CFBB7FD90F91E" lastPageId="5" lastPageNumber="76" pageId="4" pageNumber="75" type="distribution">
<paragraph id="F6413683D520A42FFF6CFBB7FD92F8D8" blockId="4.[151,826,1133,1960]" pageId="4" pageNumber="75">
<emphasis id="C48AEA91D520A42FFF6CFBB7FF70FB76" box="[151,250,1133,1158]" italics="true" pageId="4" pageNumber="75">Remarks</emphasis>
.—The supra-generic systematic arrangement of
<taxonomicName id="31FE4D00D520A42FFF6CFB57FEACFB56" authorityName="Baeza-Carratalá &amp; Pérez-Valera &amp; Pérez-Valera" authorityYear="2018" box="[151,294,1165,1190]" class="Rhynchonellata" family="Zeilleriidae" genus="Misunithyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D520A42FFF6CFB57FEACFB56" box="[151,294,1165,1190]" italics="true" pageId="4" pageNumber="75">Misunithyris</emphasis>
</taxonomicName>
is debatable, depending on the diagnostic criteria selected. Exceptional concurrence of several internal and external features can make attributable this genus to different groups within
<taxonomicName id="31FE4D00D520A42FFE4CFB37FDDAFAF6" authorityName="Waagen" authorityYear="1883" box="[439,592,1261,1286]" class="Rhynchonellata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="order">Terebratulida</taxonomicName>
. On the one hand, the new genus herein erected shows common features with the superfamily
<taxonomicName id="31FE4D00D520A42FFEA9FAF4FD35FAB7" authority="Schuchert, 1913" authorityName="Schuchert" authorityYear="1913" box="[338,703,1326,1351]" class="Rhynchonellata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="superFamily" superFamily="Dielasmatoidea">Dielasmatoidea Schuchert, 1913</taxonomicName>
, which includes some genera with enveloped dental plates and cardinal process. However, the assignment to
<taxonomicName id="31FE4D00D520A42FFD71FAB4FCB2FA77" authorityName="Schuchert" authorityYear="1913" box="[650,824,1390,1415]" class="Rhynchonellata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="superFamily" superFamily="Dielasmatoidea">Dielasmatoidea</taxonomicName>
can be problematic because
<taxonomicName id="31FE4D00D520A42FFE0DFA54FD0DFA57" authorityName="Baeza-Carratalá &amp; Pérez-Valera &amp; Pérez-Valera" authorityYear="2018" box="[502,647,1422,1447]" class="Rhynchonellata" family="Zeilleriidae" genus="Misunithyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D520A42FFE0DFA54FD0DFA57" box="[502,647,1422,1447]" italics="true" pageId="4" pageNumber="75">Misunithyris</emphasis>
</taxonomicName>
does not evidence a dielasmoid-type brachidium (sensu
<bibRefCitation id="926F4B72D520A42FFD6AFA74FC9DFA37" author="Dagys, A. S." box="[657,791,1454,1479]" pageId="4" pageNumber="75" refId="ref9228" refString="Dagys, A. S. 1974. Triasovye brahiopodi (morfologia, sistema, filogenia, stratigraficeskoe znacenie i biogeografia). 387 pp. Izdatel'stvo Nauka (Sibirskoe Otdelenie), Novosibirsk." type="book" year="1974">Dagys 1974</bibRefCitation>
or
<bibRefCitation id="926F4B72D520A42FFF6CFA14FEC2FA17" author="Smirnova, T. N." box="[151,328,1486,1511]" pageId="4" pageNumber="75" pagination="805 - 824" refId="ref12532" refString="Smirnova, T. N. 2008. Ontophylogenetic studies of the brachiopods of the Order Terebratulida. Paleontological Journal 42: 805 - 824." type="journal article" year="2008">Smirnova 2008</bibRefCitation>
). In this sense, supplementary elements such as median ridges or vertical plates (even not forming part in the development of brachidium, as stated by
<bibRefCitation id="926F4B72D520A42FFD7DF9D4FC83F9D7" author="Dagys, A. S." box="[646,777,1550,1575]" pageId="4" pageNumber="75" refId="ref9228" refString="Dagys, A. S. 1974. Triasovye brahiopodi (morfologia, sistema, filogenia, stratigraficeskoe znacenie i biogeografia). 387 pp. Izdatel'stvo Nauka (Sibirskoe Otdelenie), Novosibirsk." type="book" year="1974">Dagys 1974</bibRefCitation>
and
<bibRefCitation id="926F4B72D520A42FFF6CF9F4FEC9F9B7" author="Smirnova, T. N." box="[151,323,1582,1607]" pageId="4" pageNumber="75" pagination="805 - 824" refId="ref12532" refString="Smirnova, T. N. 2008. Ontophylogenetic studies of the brachiopods of the Order Terebratulida. Paleontological Journal 42: 805 - 824." type="journal article" year="2008">Smirnova 2008</bibRefCitation>
) have not been observed in the internal structure of
<emphasis id="C48AEA91D520A42FFF0DF994FE05F997" box="[246,399,1614,1639]" italics="true" pageId="4" pageNumber="75">
<taxonomicName id="31FE4D00D520A42FFF0DF994FE01F997" authorityName="Baeza-Carratalá &amp; Pérez-Valera &amp; Pérez-Valera" authorityYear="2018" box="[246,395,1614,1639]" class="Rhynchonellata" family="Zeilleriidae" genus="Misunithyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="genus">Misunithyris</taxonomicName>
.
</emphasis>
Furthermore, some
<taxonomicName id="31FE4D00D520A42FFD73F994FCB2F997" authorityName="Schuchert" authorityYear="1913" box="[648,824,1614,1639]" class="Rhynchonellata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="superFamily" superFamily="Dielasmatoidea">Dielasmatoidea</taxonomicName>
representatives show septum-supported architectures and often short-looped developments (e.g.,
<taxonomicName id="31FE4D00D520A42FFD88F954FF5BF937" authority="Dagys, 1959" authorityName="Dagys" authorityYear="1959" class="Rhynchonellata" family="Dielasmatidae" genus="Adygella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D520A42FFD88F954FD5DF957" box="[627,727,1678,1703]" italics="true" pageId="4" pageNumber="75">Adygella</emphasis>
Dagys, 1959
</taxonomicName>
;
<taxonomicName id="31FE4D00D520A42FFF26F974FD81F937" authority="Waagen, 1882" authorityName="Waagen" authorityYear="1882" box="[221,523,1710,1735]" class="Rhynchonellata" family="Dielasmatidae" genus="Dielasmina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D520A42FFF26F974FED5F937" box="[221,351,1710,1735]" italics="true" pageId="4" pageNumber="75">Dielasmina</emphasis>
Waagen, 1882
</taxonomicName>
;
<taxonomicName id="31FE4D00D520A42FFDE2F974FDCDF917" authority="Calzada, Peybernes, Kamoun, and Youssef, 1994" authorityName="Calzada, Peybernes, Kamoun, and Youssef" authorityYear="1994" class="Rhynchonellata" family="Dielasmatidae" genus="Tunethyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D520A42FFDE2F974FD1BF937" box="[537,657,1710,1735]" italics="true" pageId="4" pageNumber="75">Tunethyris</emphasis>
Calzada, Peybernes, Kamoun, and Youssef, 1994
</taxonomicName>
). In addition, crural bases are given off dorsally instead the distinctive crural progress revealed in
<emphasis id="C48AEA91D520A42FFE7AF8D5FD92F8D8" box="[385,536,1807,1832]" italics="true" pageId="4" pageNumber="75">
<taxonomicName id="31FE4D00D520A42FFE7AF8D5FD9EF8D8" authorityName="Baeza-Carratalá &amp; Pérez-Valera &amp; Pérez-Valera" authorityYear="2018" box="[385,532,1807,1832]" class="Rhynchonellata" family="Zeilleriidae" genus="Misunithyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="genus">Misunithyris</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="F6413683D520A42FFF46F8F5FB6FFEAA" blockId="4.[151,826,1133,1960]" lastBlockId="4.[862,1535,160,1960]" pageId="4" pageNumber="75">
Within
<taxonomicName id="31FE4D00D520A42FFEE1F8F5FE47F8B8" authorityName="Schuchert" authorityYear="1913" box="[282,461,1839,1864]" class="Rhynchonellata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="superFamily" superFamily="Dielasmatoidea">Dielasmatoidea</taxonomicName>
, higher similarity was expected with the anteriorly multicostate stock attributed to the Permian
<taxonomicName id="31FE4D00D520A42FFEF2F8B5FDC0F878" authority="Waagen, 1882" authorityName="Waagen" authorityYear="1882" box="[265,586,1903,1928]" class="Rhynchonellata" family="Dielasmatidae" genus="Dielasmina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D520A42FFEF2F8B5FE07F878" box="[265,397,1903,1928]" italics="true" pageId="4" pageNumber="75">Dielasmina</emphasis>
Waagen, 1882
</taxonomicName>
and
<taxonomicName id="31FE4D00D520A42FFD6DF8B5FEB3F858" authority="Waagen, 1882" authorityName="Waagen" authorityYear="1882" class="Rhynchonellata" family="Dielasmatidae" genus="Hemiptychina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D520A42FFD6DF8B5FCB2F878" box="[662,824,1903,1928]" italics="true" pageId="4" pageNumber="75">Hemiptychina</emphasis>
Waagen, 1882
</taxonomicName>
and the PermianTriassic
<taxonomicName id="31FE4D00D520A42FFD89F855FBA0FF49" authority="Jin, Sun, and Ye, 1979" authorityName="Jin, Sun, and Ye" authorityYear="1979" box="[626,1066,160,1960]" class="Rhynchonellata" family="Gillediidae" genus="Costoconcha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D520A42FFD89F855FC8CF858" box="[626,774,1935,1960]" italics="true" pageId="4" pageNumber="75">Costoconcha</emphasis>
Jin, Sun, and Ye, 1979
</taxonomicName>
, since besides the anterior ribbed pattern, all genera share quite a few beak features and the presence of cardinal process, but the rest of the internal structure is totally different, mainly referred to the dental plates and the crural development, which is clearly a brachidium-supported structure in the first stages.
</paragraph>
<paragraph id="F6413683D520A42FFC7FFEBBFA4EFD4D" blockId="4.[862,1535,160,1960]" pageId="4" pageNumber="75">
Another dielasmatoid morphotype, widely distributed and to some extent contemporary with
<taxonomicName id="31FE4D00D520A42FFAE4FE58FA27FE6B" authorityName="Baeza-Carratalá &amp; Pérez-Valera &amp; Pérez-Valera" authorityYear="2018" box="[1311,1453,386,411]" class="Rhynchonellata" family="Zeilleriidae" genus="Misunithyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D520A42FFAE4FE58FA27FE6B" box="[1311,1453,386,411]" italics="true" pageId="4" pageNumber="75">Misunithyris</emphasis>
</taxonomicName>
is represented by
<taxonomicName id="31FE4D00D520A42FFC1CFE78FAAAFE4B" authority="Douville, 1879" authorityName="Douville" authorityYear="1879" box="[999,1312,418,443]" class="Rhynchonellata" family="Dielasmatidae" genus="Coenothyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D520A42FFC1CFE78FBFAFE4B" box="[999,1136,418,443]" italics="true" pageId="4" pageNumber="75">Coenothyris</emphasis>
Douvillé, 1879
</taxonomicName>
. The species of this Triassic genus display comparable dental plates, undeveloped or fused with the thickened shell wall, evident cardinal process, and long loop. Conversely, it evidences an initial septum-supported structure, notable septalium and, especially, the external features (such as smooth shell, often with strong uniplication) are entirely different (e.g.,
<bibRefCitation id="926F4B72D520A42FFAD9FDBEFBC9FD6D" author="Popiel-Barczyk, E. &amp; Senkowiczowa, H." pageId="4" pageNumber="75" pagination="93 - 111" refId="ref11899" refString="Popiel-Barczyk, E. and Senkowiczowa, H. 1989. Representatives of the genus Coenothyris Douville, 1879 from the Terebratula Bed (Upper Muschelkalk) of the Holy Cross Mts, Central Poland. Acta Geologica Polonica 39: 93 - 111." type="journal article" year="1989">Popiel-Barczyk and Senkowiczowa 1989</bibRefCitation>
;
<bibRefCitation id="926F4B72D520A42FFBB7FD5EFB46FD6D" author="Torok, A." box="[1100,1228,644,669]" pageId="4" pageNumber="75" pagination="161 - 172" refId="ref12660" refString="Torok, A. 1993. Brachiopod beds as indicators of storm events: an example from the Muschelkalk of southern Hungary. In: J. Palfy and A. Voros (eds.), Mesozoic Brachiopods of Alpine Europe, 161 - 172. Hungarian Geological Society, Budapest." type="book chapter" year="1993">Török 1993</bibRefCitation>
;
<bibRefCitation id="926F4B72D520A42FFB2DFD5EFC71FD4D" author="Senkowiczowa, H. &amp; Popiel-Barczyk, E." pageId="4" pageNumber="75" pagination="443 - 466" refId="ref12272" refString="Senkowiczowa, H. and Popiel-Barczyk, E. 1996. Some Terebratulida (Brachiopoda) from the Muschelkalk sediments in the Holy Cross Mts. Geological Quarterly 40: 443 - 466." type="journal article" year="1996">Senkowiczowa and Popiel-Barczyk 1996</bibRefCitation>
;
<bibRefCitation id="926F4B72D520A42FFBFDFD7EFB09FD4D" author="Kaim, A." box="[1030,1155,676,701]" pageId="4" pageNumber="75" pagination="333 - 359" refId="ref10166" refString="Kaim, A. 1997. Brachiopod-bivalve assemblages of the Middle Triassic Terebratula Beds, Upper Silesia, Poland. Acta Palaeontologica Polonica 42: 333 - 359." type="journal article" year="1997">Kaim 1997</bibRefCitation>
;
<bibRefCitation id="926F4B72D520A42FFB74FD7EFA81FD4D" author="Palfy, J." box="[1167,1291,676,701]" pageId="4" pageNumber="75" pagination="139 - 158" refId="ref11002" refString="Palfy, J. 2003. The Pelsonian brachiopod fauna of the Balaton Highland. In: A. Voros (ed.), The Pelsonian Substage on the Balaton Highland (Middle Triassic, Hungary). Geologica Hungarica Serie Palaeontologica 55: 139 - 158." type="journal article" year="2003">Pálfy 2003</bibRefCitation>
;
<bibRefCitation id="926F4B72D520A42FFAEEFD7EFA33FD4D" author="Feldman, H. R." box="[1301,1465,676,701]" pageId="4" pageNumber="75" pagination="1 - 19" refId="ref9669" refString="Feldman, H. R. 2005. Paleoecology, taphonomy, and biogeography of a Coenothyris community (Brachiopoda, Terebratulida) from the Triassic (Upper Anisian - Lower Ladinian) of Israel. American Museum Novitates 3479: 1 - 19." type="journal article" year="2005">Feldman 2005</bibRefCitation>
).
</paragraph>
<paragraph id="F6413683D520A42FFC7FFD1EFBEBFC4F" blockId="4.[862,1535,160,1960]" pageId="4" pageNumber="75">
Arrangement within
<taxonomicName id="31FE4D00D520A42FFB7AFD1EFA27FD2D" authority="Allan, 1940" authorityName="Allan" authorityYear="1940" box="[1153,1453,708,733]" class="Rhynchonellata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="superFamily" superFamily="Zeillerioidea">Zeillerioidea Allan, 1940</taxonomicName>
is the most plausible option, mainly because of the presence of dental lamellae, the well-developed and large dorsal median septum, and a clear zeilleroid-type brachidium (sensu
<bibRefCitation id="926F4B72D520A42FFCA5FC9FFB84FCAE" author="Smirnova, T. N." box="[862,1038,837,862]" pageId="4" pageNumber="75" pagination="805 - 824" refId="ref12532" refString="Smirnova, T. N. 2008. Ontophylogenetic studies of the brachiopods of the Order Terebratulida. Paleontological Journal 42: 805 - 824." type="journal article" year="2008">Smirnova 2008</bibRefCitation>
) with a long-looped development, not connected to the median septum. However, some characters do not fully agree with the various families up to now determined in
<taxonomicName id="31FE4D00D520A42FFC32FC7CFBD7FC4F" authorityName="Allan" authorityYear="1940" box="[969,1117,934,959]" class="Rhynchonellata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="83" phylum="Brachiopoda" rank="superFamily" superFamily="Zeillerioidea">Zeillerioidea</taxonomicName>
.
</paragraph>
<paragraph id="F6413683D520A42FFC7FFC1DFC3AFA73" blockId="4.[862,1535,160,1960]" pageId="4" pageNumber="75">
<taxonomicName id="31FE4D00D520A42FFC7FFC1DFB9FFC10" authorityName="Baeza-Carratalá &amp; Pérez-Valera &amp; Pérez-Valera" authorityYear="2018" box="[900,1045,967,992]" class="Rhynchonellata" family="Zeilleriidae" genus="Misunithyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D520A42FFC7FFC1DFB9FFC10" box="[900,1045,967,992]" italics="true" pageId="4" pageNumber="75">Misunithyris</emphasis>
</taxonomicName>
shares with the family
<taxonomicName id="31FE4D00D520A42FFAC8FC1DFC6DFBF0" authority="Muir-Wood, 1965" authorityName="Muir-Wood" authorityYear="1965" family="Eudessidae" pageId="4" pageNumber="75" rank="family">Eudessidae Muir-Wood, 1965</taxonomicName>
the envelopment of dental plates, the presence of a cardinal process and most of the beak features. The most remarkable difference to
<taxonomicName id="31FE4D00D520A42FFB34FBFDFAC0FBB0" authorityName="Muir-Wood" authorityYear="1965" box="[1231,1354,1063,1088]" family="Eudessidae" pageId="4" pageNumber="75" rank="family">Eudessidae</taxonomicName>
are in a short dorsal median septum and the crural bases given off dorsally, as well as the usual growth of a median cardinal plate of the later group and not perceived in
<taxonomicName id="31FE4D00D520A42FFAE4FB52FA25FB51" authorityName="Baeza-Carratalá &amp; Pérez-Valera &amp; Pérez-Valera" authorityYear="2018" box="[1311,1455,1160,1185]" class="Rhynchonellata" family="Zeilleriidae" genus="Misunithyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D520A42FFAE4FB52FA25FB51" box="[1311,1455,1160,1185]" italics="true" pageId="4" pageNumber="75">Misunithyris</emphasis>
</taxonomicName>
. Some external diagnostic criteria are very different as well, since even showing
<taxonomicName id="31FE4D00D520A42FFBF8FB13FB08FB12" authorityName="Muir-Wood" authorityYear="1965" box="[1027,1154,1225,1250]" family="Eudessidae" pageId="4" pageNumber="75" rank="family">Eudessidae</taxonomicName>
multicostate shells,
<taxonomicName id="31FE4D00D520A42FFA95FB13FA75FB12" authorityName="Baeza-Carratalá &amp; Pérez-Valera &amp; Pérez-Valera" authorityYear="2018" box="[1390,1535,1225,1250]" class="Rhynchonellata" family="Zeilleriidae" genus="Misunithyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D520A42FFA95FB13FA75FB12" box="[1390,1535,1225,1250]" italics="true" pageId="4" pageNumber="75">Misunithyris</emphasis>
</taxonomicName>
shows a marginal ribbing pattern instead the entire multicostate shell-length of
<taxonomicName id="31FE4D00D520A42FFB93FAD3FB61FAD2" authorityName="Muir-Wood" authorityYear="1965" box="[1128,1259,1289,1314]" family="Eudessidae" pageId="4" pageNumber="75" rank="family">Eudessidae</taxonomicName>
. Folding pattern is also unrelated as the conspicuous dorsal sulcus developed by
<taxonomicName id="31FE4D00D520A42FFCA5FA90FC65FA93" authorityName="Baeza-Carratalá &amp; Pérez-Valera &amp; Pérez-Valera" authorityYear="2018" box="[862,1007,1354,1379]" class="Rhynchonellata" family="Zeilleriidae" genus="Misunithyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D520A42FFCA5FA90FC65FA93" box="[862,1007,1354,1379]" italics="true" pageId="4" pageNumber="75">Misunithyris</emphasis>
</taxonomicName>
is not shared with any
<taxonomicName id="31FE4D00D520A42FFAF1FA90FA0DFA93" authorityName="Muir-Wood" authorityYear="1965" box="[1290,1415,1354,1379]" family="Eudessidae" pageId="4" pageNumber="75" rank="family">Eudessidae</taxonomicName>
representatives.
</paragraph>
<paragraph id="F6413683D520A42FFC7FFA50FBC4F858" blockId="4.[862,1535,160,1960]" pageId="4" pageNumber="75">
The affinities with the family
<taxonomicName id="31FE4D00D520A42FFB11FA50FAE9FA53" box="[1258,1379,1418,1443]" class="Rhynchonellata" family="Zeilleridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="family">Zeilleridae</taxonomicName>
are found in several subfamilies. Some representatives of the subfamily
<taxonomicName id="31FE4D00D520A42FFCA5FA11FBE4FA14" authority="Baker, 2006" authorityName="Baker" authorityYear="2006" box="[862,1134,1483,1508]" class="Rhynchonellata" family="Zeilleridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="subFamily" subFamily="Vectellinae">
Vectellinae
<bibRefCitation id="926F4B72D520A42FFC1FFA11FBE4FA14" author="Baker, P. G." box="[996,1134,1483,1508]" pageId="4" pageNumber="75" pagination="2163 - 2188" refId="ref8816" refString="Baker, P. G. 2006. Zeillerioidea. In: R. L. Kaesler (ed.), Treatise on Invertebrate Paleontology. Part H. Brachiopoda. Revised, 5: Rhynchonelliformea (part), 2163 - 2188. Geological Society of America, Boulder and University of Kansas, Lawrence." type="book chapter" year="2006">Baker, 2006</bibRefCitation>
</taxonomicName>
exhibit a long-looped development and cardinal process as a knob or poorly developed callus. However, MiddleLate Triassic representatives of this subfamily are very different in both external and internal features to the new established genus. It is the case of
<taxonomicName id="31FE4D00D520A42FFA32F996FB13F975" authority="Dagys, 1977" authorityName="Dagys" authorityYear="1977" class="Rhynchonellata" family="Zeilleriidae" genus="Fletcherithyroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D520A42FFA32F996FC74F975" italics="true" pageId="4" pageNumber="75">Fletcherithyroides</emphasis>
Dagys, 1977
</taxonomicName>
,
<taxonomicName id="31FE4D00D520A42FFB52F9B6FA70F975" authority="Dagys, 1965" authorityName="Dagys" authorityYear="1965" box="[1193,1530,1644,1669]" class="Rhynchonellata" family="Zeilleriidae" genus="Aulacothyroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D520A42FFB52F9B6FAD4F975" box="[1193,1374,1644,1669]" italics="true" pageId="4" pageNumber="75">Aulacothyroides</emphasis>
Dagys, 1965
</taxonomicName>
, and
<taxonomicName id="31FE4D00D520A42FFC75F957FA88F956" authority="Xu and Liu, 1983" authorityName="Xu and Liu" authorityYear="1983" box="[910,1282,1677,1702]" class="Rhynchonellata" family="Zeilleriidae" genus="Parantiptychia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="75" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D520A42FFC75F957FBBEF956" box="[910,1076,1677,1702]" italics="true" pageId="4" pageNumber="75">Parantiptychia</emphasis>
Xu and Liu, 1983
</taxonomicName>
, consisting on smooth morphotypes, also showing a long stage of crura supported by septal pillars. Probably the clearest affinity in this subfamily is found in the Upper Triassic?lowermost Jurassic
<taxonomicName id="31FE4D00D520A42FFCA5F8D4FB10F8D7" authority="Seguenza, 1885" authorityName="Seguenza" authorityYear="1885" box="[862,1178,1806,1831]" class="Cephalopoda" family="Hildoceratidae" genus="Tauromenia" kingdom="Animalia" pageId="4" pageNumber="75" phylum="Mollusca" rank="genus">
<emphasis id="C48AEA91D520A42FFCA5F8D4FC6EF8D7" box="[862,996,1806,1831]" italics="true" pageId="4" pageNumber="75">Tauromenia</emphasis>
<bibRefCitation id="926F4B72D520A42FFC11F8D4FB10F8D7" author="Seguenza, G." box="[1002,1178,1806,1831]" pageId="4" pageNumber="75" pagination="377 - 497" refId="ref12207" refString="Seguenza, G. 1885. Monografia delle Spiriferina dei varii piani del Lias Messinese. Bollettino della Societa Geologica Italian a 4: 377 - 497." type="journal article" year="1885">Seguenza, 1885</bibRefCitation>
</taxonomicName>
, due to comparable anterior ribbing pattern and the beak features, as well as the well-developed dorsal median septum (
<bibRefCitation id="926F4B72D520A42FFB3FF894FA2FF897" author="Almeras, Y. &amp; Elmi, S. &amp; Faure, P." box="[1220,1445,1870,1895]" pageId="4" pageNumber="75" pagination="3 - 241" refId="ref8620" refString="Almeras, Y., Elmi, S., and Faure, P. 2007. Les Brachiopodes Liasiques d'Algerie Occidentale. Documents des Laboratoires de Geologie Lyon 163: 3 - 241." type="journal article" year="2007">Alméras et al. 2007</bibRefCitation>
;
<bibRefCitation id="926F4B72D520A42FFA49F894FB31F878" author="Baeza-Carratala, J. F. &amp; Garcia Joral, F." pageId="4" pageNumber="75" pagination="1 - 12" refId="ref8728" refString="Baeza-Carratala, J. F. and Garcia Joral, F. 2012. Multicostate zeillerids (Brachiopoda, Terebratulida) from the Lower Jurassic of the Eastern Subbetic (SE Spain) and their use in correlation and paleobiogeography. Geologica Acta 10: 1 - 12." type="journal article" year="2012">Baeza-Carratalá and García Joral 2012</bibRefCitation>
) and long-looped brachidium (
<bibRefCitation id="926F4B72D520A42FFC9EF855FBC9F858" author="Almeras, Y. &amp; Elmi, S. &amp; Faure, P." box="[869,1091,1935,1960]" pageId="4" pageNumber="75" pagination="3 - 241" refId="ref8620" refString="Almeras, Y., Elmi, S., and Faure, P. 2007. Les Brachiopodes Liasiques d'Algerie Occidentale. Documents des Laboratoires de Geologie Lyon 163: 3 - 241." type="journal article" year="2007">Alméras et al. 2007</bibRefCitation>
).
</paragraph>
<paragraph id="F6413683D521A42EFF7CFF7AFD08FCCD" blockId="5.[98,771,160,1634]" pageId="5" pageNumber="76">
Finally, there are several arguments for and against assigning
<taxonomicName id="31FE4D00D521A42EFF41FF1AFEC2FF29" authorityName="Baeza-Carratalá &amp; Pérez-Valera &amp; Pérez-Valera" authorityYear="2018" box="[186,328,192,217]" class="Rhynchonellata" family="Zeilleriidae" genus="Misunithyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="5" pageNumber="76" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D521A42EFF41FF1AFEC2FF29" box="[186,328,192,217]" italics="true" pageId="5" pageNumber="76">Misunithyris</emphasis>
</taxonomicName>
to the subfamily
<taxonomicName id="31FE4D00D521A42EFDFFFF1AFF12FF09" authority="Schuchert, 1929" authorityName="Schuchert" authorityYear="1929" pageId="5" pageNumber="76" rank="subFamily" subFamily="Zeilleriinae">Zeilleriinae Schuchert, 1929</taxonomicName>
as was defined in Kaesler and Selden (19972007). The septum is clearly well-developed and the brachidium evidences a long-looped progression. Conversely, the presence of cardinal process is atypical in this subfamily, although some genera show primitive lobes as massive callus or small knobs (e.g.,
<taxonomicName id="31FE4D00D521A42EFF16FE5BFD86FE6A" authority="Zittel, 1880" authorityName="Zittel" authorityYear="1880" box="[237,524,385,410]" class="Rhynchonellata" family="Zeilleriidae" genus="Antiptychina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="5" pageNumber="76" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D521A42EFF16FE5BFEF7FE6A" box="[237,381,385,410]" italics="true" pageId="5" pageNumber="76">Antiptychina</emphasis>
Zittel, 1880
</taxonomicName>
;
<taxonomicName id="31FE4D00D521A42EFDE1FE5BFF13FE4A" authority="Dagys, 1965" authorityName="Dagys" authorityYear="1965" class="Rhynchonellata" family="Zeilleriidae" genus="Kolymithyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="5" pageNumber="76" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D521A42EFDE1FE5BFD20FE6A" box="[538,682,385,410]" italics="true" pageId="5" pageNumber="76">Kolymithyris</emphasis>
Dagys, 1965
</taxonomicName>
). On the other hand, some of them also reveal enveloped dental lamellae instead the typical strong and unenveloped dental plates characteristic of this subfamily. As for
<taxonomicName id="31FE4D00D521A42EFF77FDD8FE99FDEB" authorityName="Seguenza" authorityYear="1885" box="[140,275,514,539]" class="Cephalopoda" family="Hildoceratidae" genus="Tauromenia" kingdom="Animalia" pageId="5" pageNumber="76" phylum="Mollusca" rank="genus">
<emphasis id="C48AEA91D521A42EFF77FDD8FE99FDEB" box="[140,275,514,539]" italics="true" pageId="5" pageNumber="76">Tauromenia</emphasis>
</taxonomicName>
, the closer external affinities are found in the anteriorly ribbed representatives of this subfamily, i.e.,
<taxonomicName id="31FE4D00D521A42EFF99FD98FE33FDAB" authority="Owen and Rose, 1997" authorityName="Owen and Rose" authorityYear="1997" box="[98,441,578,603]" class="Rhynchonellata" family="Zeilleriidae" genus="Calpella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="5" pageNumber="76" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D521A42EFF99FD98FF48FDAB" box="[98,194,578,603]" italics="true" pageId="5" pageNumber="76">Calpella</emphasis>
<bibRefCitation id="926F4B72D521A42EFF3CFD98FE33FDAB" author="Owen, E. F. &amp; Rose, E. P. F." box="[199,441,578,603]" pageId="5" pageNumber="76" pagination="497 - 513" refId="ref10854" refString="Owen, E. F. and Rose, E. P. F. 1997. Early Jurassic Brachiopods from Gibraltar, and their Tethyan affinities. Palaeontology 40: 497 - 513." type="journal article" year="1997">Owen and Rose, 1997</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="31FE4D00D521A42EFE15FD98FEB3FD8C" authority="Schuchert and Le Vene, 1929" authorityName="Schuchert and Le Vene" authorityYear="1929" class="Rhynchonellata" family="Zeilleriidae" genus="Parathyridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="5" pageNumber="76" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D521A42EFE15FD98FD06FDAB" box="[494,652,578,603]" italics="true" pageId="5" pageNumber="76">Parathyridina</emphasis>
Schuchert and Le Vene, 1929
</taxonomicName>
, the internal structure of which remains poorly known except for the presence of a prominent median septum (e.g.,
<bibRefCitation id="926F4B72D521A42EFEC4FD79FE5CFD4C" author="Cooper, G. A." box="[319,470,675,700]" pageId="5" pageNumber="76" pagination="1 - 445" refId="ref9167" refString="Cooper, G. A. 1983. The Terebratulacea (Brachiopoda), Triassic to Recent: A study of the Brachidia (Loops). Smithsonian Contributions to Paleobiology 50: 1 - 445." type="journal article" year="1983">Cooper 1983</bibRefCitation>
;
<bibRefCitation id="926F4B72D521A42EFE1DFD79FDFBFD4C" author="Baker, P. G." box="[486,625,675,700]" pageId="5" pageNumber="76" pagination="2163 - 2188" refId="ref8816" refString="Baker, P. G. 2006. Zeillerioidea. In: R. L. Kaesler (ed.), Treatise on Invertebrate Paleontology. Part H. Brachiopoda. Revised, 5: Rhynchonelliformea (part), 2163 - 2188. Geological Society of America, Boulder and University of Kansas, Lawrence." type="book chapter" year="2006">Baker 2006</bibRefCitation>
;
<bibRefCitation id="926F4B72D521A42EFD7AFD79FF44FD2C" author="Almeras Y. &amp; Cougnon, R. &amp; Enay, R. &amp; Mangold, C." pageId="5" pageNumber="76" pagination="1 - 247" refId="ref8567" refString="Almeras Y., Cougnon, R., Enay, R., and Mangold, C. 2010 b. Brachiopodes du Jurassique Inferieur et Moyen d'Arabie Saoudite Centrale. Paleontologie, Biostratigraphie et Paleoenvironnements, echelles chronostratigraphiques. Documents des Laboratoires de Geologie Lyon 168: 1 - 247." type="journal article" year="2010">Alméras et al. 2010b</bibRefCitation>
). In this sense, very close internal and external affinities have been recognized with the recent erected multicostate genus
<taxonomicName id="31FE4D00D521A42EFEE3FCDEFDC5FCED" authority="Feldman, 2013" authorityName="Feldman" authorityYear="2013" box="[280,591,772,797]" class="Rhynchonellata" family="Zeilleriidae" genus="Menathyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="5" pageNumber="76" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D521A42EFEE3FCDEFE12FCED" box="[280,408,772,797]" italics="true" pageId="5" pageNumber="76">Menathyris</emphasis>
<bibRefCitation id="926F4B72D521A42EFE5FFCDEFDC5FCED" author="Feldman, H. R." box="[420,591,772,797]" pageId="5" pageNumber="76" pagination="81 - 85" refId="ref9714" refString="Feldman, H. R. 2013. Menathyris wilsoni (Brachiopoda), new genus and species from the Middle Triassic (Ladinian) of Southern Israel. Annales Societatis Geologorum Poloniae 83: 81 - 85." type="journal article" year="2013">Feldman, 2013</bibRefCitation>
</taxonomicName>
, except for the cardinal process and the anterior folding pattern.
</paragraph>
<caption id="A281660BD521A42EFCD3FEFAFA40FE7D" pageId="5" pageNumber="76" startId="5.[808,861,288,309]" targetBox="[816,1470,423,635]" targetIsTable="true" targetPageId="5" targetType="table">
<paragraph id="F6413683D521A42EFCD3FEFAFA40FE7D" blockId="5.[808,1482,288,397]" pageId="5" pageNumber="76">
Table 1. Main biometric measurements (in mm) and ratios of the studied specimens of
<taxonomicName id="31FE4D00D521A42EFC50FEE7FBD9FEA2" authority="Baeza-Carratalá &amp; Pérez-Valera &amp; Pérez-Valera, 2018" authorityName="Baeza-Carratalá &amp; Pérez-Valera &amp; Pérez-Valera" authorityYear="2018" box="[939,1107,317,338]" class="Rhynchonellata" family="Zeilleriidae" genus="Misunithyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="5" pageNumber="76" phylum="Brachiopoda" rank="species" species="goyi" status="gen. et sp. nov.">
<emphasis id="C48AEA91D521A42EFC50FEE7FBD9FEA2" box="[939,1107,317,338]" italics="true" pageId="5" pageNumber="76">Misunithyris goyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DFB957EAD521A42EFBA2FEE7FB62FEA2" box="[1113,1256,317,338]" pageId="5" pageNumber="76" rank="species">gen. et sp. nov.</taxonomicNameLabel>
L, total length; W, total width; T, total thickness; R, ribs on each valve; Rp, ribbing pattern (p, primary rib; s, secondary rib, referred to bifurcate or intercalated ribs).
</paragraph>
</caption>
<paragraph id="F6413683D521A42EFCCEFE7DFA34FD8B" pageId="5" pageNumber="76">
<table id="84FEC423D5215BD4FCCBFE7DFA34FD8B" box="[816,1470,423,635]" gridcols="9" gridrows="7" pageId="5" pageNumber="76">
<tr id="48CE34C1D5215BD4FCCBFE7DFA34FE4C" box="[816,1470,423,444]" gridrow="0" pageId="5" pageNumber="76">
<th id="0B1F5DBDD5215BD4FCCBFE7DFC10FE4C" box="[816,922,423,444]" gridcol="0" gridrow="0" pageId="5" pageNumber="76">Specimen</th>
<th id="0B1F5DBDD5215BD4FC51FE7DFC6AFE4C" box="[938,992,423,444]" gridcol="1" gridrow="0" pageId="5" pageNumber="76">L</th>
<th id="0B1F5DBDD5215BD4FC0FFE7DFBA0FE4C" box="[1012,1066,423,444]" gridcol="2" gridrow="0" pageId="5" pageNumber="76">W</th>
<th id="0B1F5DBDD5215BD4FBC0FE7DFBF4FE4C" box="[1083,1150,423,444]" gridcol="3" gridrow="0" pageId="5" pageNumber="76">T</th>
<th id="0B1F5DBDD5215BD4FB6BFE7DFB36FE4C" box="[1168,1212,423,444]" gridcol="4" gridrow="0" pageId="5" pageNumber="76">W/L</th>
<th id="0B1F5DBDD5215BD4FB2CFE7DFA8BFE4C" box="[1239,1281,423,444]" gridcol="5" gridrow="0" pageId="5" pageNumber="76">T/L</th>
<th id="0B1F5DBDD5215BD4FAE5FE7DFAC0FE4C" box="[1310,1354,423,444]" gridcol="6" gridrow="0" pageId="5" pageNumber="76">T/W</th>
<th id="0B1F5DBDD5215BD4FAA5FE7DFAFCFE4C" box="[1374,1398,423,444]" gridcol="7" gridrow="0" pageId="5" pageNumber="76">R</th>
<th id="0B1F5DBDD5215BD4FA71FE7DFA34FE4C" box="[1418,1470,423,444]" gridcol="8" gridrow="0" pageId="5" pageNumber="76">Rp</th>
</tr>
<tr id="48CE34C1D5215BD4FCCBFE1DFA34FE2C" box="[816,1470,455,476]" gridrow="1" pageId="5" pageNumber="76">
<th id="0B1F5DBDD5215BD4FCCBFE1DFC10FE2C" box="[816,922,455,476]" gridcol="0" gridrow="1" pageId="5" pageNumber="76">BQ-CL1.1</th>
<td id="0B1F5DBDD5215BD4FC51FE1DFC6AFE2C" box="[938,992,455,476]" gridcol="1" gridrow="1" pageId="5" pageNumber="76">17.56</td>
<td id="0B1F5DBDD5215BD4FC0FFE1DFBA0FE2C" box="[1012,1066,455,476]" gridcol="2" gridrow="1" pageId="5" pageNumber="76">12.78</td>
<td id="0B1F5DBDD5215BD4FBC0FE1DFBF4FE2C" box="[1083,1150,455,476]" gridcol="3" gridrow="1" pageId="5" pageNumber="76">10.83</td>
<td id="0B1F5DBDD5215BD4FB6BFE1DFB36FE2C" box="[1168,1212,455,476]" gridcol="4" gridrow="1" pageId="5" pageNumber="76">0.73</td>
<td id="0B1F5DBDD5215BD4FB2CFE1DFA8BFE2C" box="[1239,1281,455,476]" gridcol="5" gridrow="1" pageId="5" pageNumber="76">0.62</td>
<td id="0B1F5DBDD5215BD4FAE5FE1DFAC0FE2C" box="[1310,1354,455,476]" gridcol="6" gridrow="1" pageId="5" pageNumber="76">0.85</td>
<td id="0B1F5DBDD5215BD4FAA5FE1DFAFCFE2C" box="[1374,1398,455,476]" gridcol="7" gridrow="1" pageId="5" pageNumber="76">8</td>
<td id="0B1F5DBDD5215BD4FA71FE1DFA34FE2C" box="[1418,1470,455,476]" gridcol="8" gridrow="1" pageId="5" pageNumber="76">7p 2s</td>
</tr>
<tr id="48CE34C1D5215BD4FCCBFE3DFA34FE0C" box="[816,1470,487,508]" gridrow="2" pageId="5" pageNumber="76">
<th id="0B1F5DBDD5215BD4FCCBFE3DFC10FE0C" box="[816,922,487,508]" gridcol="0" gridrow="2" pageId="5" pageNumber="76">BQ-CL1.2</th>
<td id="0B1F5DBDD5215BD4FC51FE3DFC6AFE0C" box="[938,992,487,508]" gridcol="1" gridrow="2" pageId="5" pageNumber="76">13.00</td>
<td id="0B1F5DBDD5215BD4FC0FFE3DFBA0FE0C" box="[1012,1066,487,508]" gridcol="2" gridrow="2" pageId="5" pageNumber="76">12.44</td>
<td id="0B1F5DBDD5215BD4FBC0FE3DFBF4FE0C" box="[1083,1150,487,508]" gridcol="3" gridrow="2" pageId="5" pageNumber="76">8.20</td>
<td id="0B1F5DBDD5215BD4FB6BFE3DFB36FE0C" box="[1168,1212,487,508]" gridcol="4" gridrow="2" pageId="5" pageNumber="76">0.96</td>
<td id="0B1F5DBDD5215BD4FB2CFE3DFA8BFE0C" box="[1239,1281,487,508]" gridcol="5" gridrow="2" pageId="5" pageNumber="76">0.63</td>
<td id="0B1F5DBDD5215BD4FAE5FE3DFAC0FE0C" box="[1310,1354,487,508]" gridcol="6" gridrow="2" pageId="5" pageNumber="76">0.66</td>
<td id="0B1F5DBDD5215BD4FAA5FE3DFAFCFE0C" box="[1374,1398,487,508]" gridcol="7" gridrow="2" pageId="5" pageNumber="76">9</td>
<td id="0B1F5DBDD5215BD4FA71FE3DFA34FE0C" box="[1418,1470,487,508]" gridcol="8" gridrow="2" pageId="5" pageNumber="76">6p 2s</td>
</tr>
<tr id="48CE34C1D5215BD4FCCBFDDDFA34FDEC" box="[816,1470,519,540]" gridrow="3" pageId="5" pageNumber="76">
<th id="0B1F5DBDD5215BD4FCCBFDDDFC10FDEC" box="[816,922,519,540]" gridcol="0" gridrow="3" pageId="5" pageNumber="76">BQ-TA1.1</th>
<td id="0B1F5DBDD5215BD4FC51FDDDFC6AFDEC" box="[938,992,519,540]" gridcol="1" gridrow="3" pageId="5" pageNumber="76">17.41</td>
<td id="0B1F5DBDD5215BD4FC0FFDDDFBA0FDEC" box="[1012,1066,519,540]" gridcol="2" gridrow="3" pageId="5" pageNumber="76">16.60</td>
<td id="0B1F5DBDD5215BD4FBC0FDDDFBF4FDEC" box="[1083,1150,519,540]" gridcol="3" gridrow="3" pageId="5" pageNumber="76">~8.44</td>
<td id="0B1F5DBDD5215BD4FB6BFDDDFB36FDEC" box="[1168,1212,519,540]" gridcol="4" gridrow="3" pageId="5" pageNumber="76">0.95</td>
<td id="0B1F5DBDD5215BD4FB2CFDDDFA8BFDEC" box="[1239,1281,519,540]" gridcol="5" gridrow="3" pageId="5" pageNumber="76">0.48</td>
<td id="0B1F5DBDD5215BD4FAE5FDDDFAC0FDEC" box="[1310,1354,519,540]" gridcol="6" gridrow="3" pageId="5" pageNumber="76">0.51</td>
<td id="0B1F5DBDD5215BD4FAA5FDDDFAFCFDEC" box="[1374,1398,519,540]" gridcol="7" gridrow="3" pageId="5" pageNumber="76">9</td>
<td id="0B1F5DBDD5215BD4FA71FDDDFA34FDEC" box="[1418,1470,519,540]" gridcol="8" gridrow="3" pageId="5" pageNumber="76">9p</td>
</tr>
<tr id="48CE34C1D5215BD4FCCBFDFCFA34FDCB" box="[816,1470,550,571]" gridrow="4" pageId="5" pageNumber="76">
<th id="0B1F5DBDD5215BD4FCCBFDFCFC10FDCB" box="[816,922,550,571]" gridcol="0" gridrow="4" pageId="5" pageNumber="76">BQ-TA1.2</th>
<td id="0B1F5DBDD5215BD4FC51FDFCFC6AFDCB" box="[938,992,550,571]" gridcol="1" gridrow="4" pageId="5" pageNumber="76">28.34</td>
<td id="0B1F5DBDD5215BD4FC0FFDFCFBA0FDCB" box="[1012,1066,550,571]" gridcol="2" gridrow="4" pageId="5" pageNumber="76">24.43</td>
<td id="0B1F5DBDD5215BD4FBC0FDFCFBF4FDCB" box="[1083,1150,550,571]" gridcol="3" gridrow="4" pageId="5" pageNumber="76">17.01</td>
<td id="0B1F5DBDD5215BD4FB6BFDFCFB36FDCB" box="[1168,1212,550,571]" gridcol="4" gridrow="4" pageId="5" pageNumber="76">0.86</td>
<td id="0B1F5DBDD5215BD4FB2CFDFCFA8BFDCB" box="[1239,1281,550,571]" gridcol="5" gridrow="4" pageId="5" pageNumber="76">0.60</td>
<td id="0B1F5DBDD5215BD4FAE5FDFCFAC0FDCB" box="[1310,1354,550,571]" gridcol="6" gridrow="4" pageId="5" pageNumber="76">0.70</td>
<td id="0B1F5DBDD5215BD4FAA5FDFCFAFCFDCB" box="[1374,1398,550,571]" gridcol="7" gridrow="4" pageId="5" pageNumber="76">13</td>
<td id="0B1F5DBDD5215BD4FA71FDFCFA34FDCB" box="[1418,1470,550,571]" gridcol="8" gridrow="4" pageId="5" pageNumber="76">9p 2s</td>
</tr>
<tr id="48CE34C1D5215BD4FCCBFD9CFA34FDAB" box="[816,1470,582,603]" gridrow="5" pageId="5" pageNumber="76">
<th id="0B1F5DBDD5215BD4FCCBFD9CFC10FDAB" box="[816,922,582,603]" gridcol="0" gridrow="5" pageId="5" pageNumber="76">BQ-AH2.1</th>
<td id="0B1F5DBDD5215BD4FC51FD9CFC6AFDAB" box="[938,992,582,603]" gridcol="1" gridrow="5" pageId="5" pageNumber="76">21.74</td>
<td id="0B1F5DBDD5215BD4FC0FFD9CFBA0FDAB" box="[1012,1066,582,603]" gridcol="2" gridrow="5" pageId="5" pageNumber="76">15.77</td>
<td id="0B1F5DBDD5215BD4FBC0FD9CFBF4FDAB" box="[1083,1150,582,603]" gridcol="3" gridrow="5" pageId="5" pageNumber="76">15.51</td>
<td id="0B1F5DBDD5215BD4FB6BFD9CFB36FDAB" box="[1168,1212,582,603]" gridcol="4" gridrow="5" pageId="5" pageNumber="76">0.72</td>
<td id="0B1F5DBDD5215BD4FB2CFD9CFA8BFDAB" box="[1239,1281,582,603]" gridcol="5" gridrow="5" pageId="5" pageNumber="76">0.71</td>
<td id="0B1F5DBDD5215BD4FAE5FD9CFAC0FDAB" box="[1310,1354,582,603]" gridcol="6" gridrow="5" pageId="5" pageNumber="76">0.98</td>
<td id="0B1F5DBDD5215BD4FAA5FD9CFAFCFDAB" box="[1374,1398,582,603]" gridcol="7" gridrow="5" pageId="5" pageNumber="76">11</td>
<td id="0B1F5DBDD5215BD4FA71FD9CFA34FDAB" box="[1418,1470,582,603]" gridcol="8" gridrow="5" pageId="5" pageNumber="76">9p 4s</td>
</tr>
<tr id="48CE34C1D5215BD4FCCBFDBCFA34FD8B" box="[816,1470,614,635]" gridrow="6" pageId="5" pageNumber="76">
<th id="0B1F5DBDD5215BD4FCCBFDBCFC10FD8B" box="[816,922,614,635]" gridcol="0" gridrow="6" pageId="5" pageNumber="76">BQ-AH2.2</th>
<td id="0B1F5DBDD5215BD4FC51FDBCFC6AFD8B" box="[938,992,614,635]" gridcol="1" gridrow="6" pageId="5" pageNumber="76">16.64</td>
<td id="0B1F5DBDD5215BD4FC0FFDBCFBA0FD8B" box="[1012,1066,614,635]" gridcol="2" gridrow="6" pageId="5" pageNumber="76">13.54</td>
<td id="0B1F5DBDD5215BD4FBC0FDBCFBF4FD8B" box="[1083,1150,614,635]" gridcol="3" gridrow="6" pageId="5" pageNumber="76">~10.90</td>
<td id="0B1F5DBDD5215BD4FB6BFDBCFB36FD8B" box="[1168,1212,614,635]" gridcol="4" gridrow="6" pageId="5" pageNumber="76">0.81</td>
<td id="0B1F5DBDD5215BD4FB2CFDBCFA8BFD8B" box="[1239,1281,614,635]" gridcol="5" gridrow="6" pageId="5" pageNumber="76">0.65</td>
<td id="0B1F5DBDD5215BD4FAE5FDBCFAC0FD8B" box="[1310,1354,614,635]" gridcol="6" gridrow="6" pageId="5" pageNumber="76">0.80</td>
<td id="0B1F5DBDD5215BD4FAA5FDBCFAFCFD8B" box="[1374,1398,614,635]" gridcol="7" gridrow="6" pageId="5" pageNumber="76">11</td>
<td id="0B1F5DBDD5215BD4FA71FDBCFA34FD8B" box="[1418,1470,614,635]" gridcol="8" gridrow="6" pageId="5" pageNumber="76">9p 2s</td>
</tr>
</table>
</paragraph>
<paragraph id="F6413683D521A42EFF7CFC9EFDCDFB4F" blockId="5.[98,771,160,1634]" pageId="5" pageNumber="76">
It must be kept in mind that the subfamily
<taxonomicName id="31FE4D00D521A42EFD84FC9EFE92FC8D" authority="Schuchert, 1929" authorityName="Schuchert" authorityYear="1929" pageId="5" pageNumber="76" rank="subFamily" subFamily="Zeilleriinae">Zeilleriinae Schuchert, 1929</taxonomicName>
was recently split into several subfamilies by
<bibRefCitation id="926F4B72D521A42EFF79FC5EFDB6FC6D" author="Baeza-Carratala, J. F. &amp; Garcia Joral, F." box="[130,572,900,925]" pageId="5" pageNumber="76" pagination="651 - 661" refId="ref8777" refString="Baeza-Carratala, J. F. and Garcia Joral, F. 2014. Crural bases position as a structural criterion for supraspecific diagnosis of Early Jurassic zeilleriid brachiopods. Acta Palaeontologica Polonica 59: 651 - 661." type="journal article" year="2014">Baeza-Carratalá and García Joral (2014)</bibRefCitation>
on the basis of the hinge plates-crural bases relationship, thus resulting three subfamilies (Aulacothyrinae Babanova, 1964,
<taxonomicName id="31FE4D00D521A42EFD85FC1FFEAEFC0E" authority="Schuchert, 1929" authorityName="Schuchert" authorityYear="1929" pageId="5" pageNumber="76" rank="subFamily" subFamily="Zeilleriinae">Zeilleriinae Schuchert, 1929</taxonomicName>
, and Securininae
<bibRefCitation id="926F4B72D521A42EFDF6FC3FFEBAFBEE" author="Baeza-Carratala, J. F. &amp; Garcia Joral, F." pageId="5" pageNumber="76" pagination="651 - 661" refId="ref8777" refString="Baeza-Carratala, J. F. and Garcia Joral, F. 2014. Crural bases position as a structural criterion for supraspecific diagnosis of Early Jurassic zeilleriid brachiopods. Acta Palaeontologica Polonica 59: 651 - 661." type="journal article" year="2014">Baeza-Carratalá and García Joral, 2014</bibRefCitation>
). Attending to this criterion,
<taxonomicName id="31FE4D00D521A42EFD88FBDFFC88FBEE" authorityName="Baeza-Carratalá &amp; Pérez-Valera &amp; Pérez-Valera" authorityYear="2018" box="[627,770,1029,1054]" class="Rhynchonellata" family="Zeilleriidae" genus="Misunithyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="5" pageNumber="76" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D521A42EFD88FBDFFC88FBEE" box="[627,770,1029,1054]" italics="true" pageId="5" pageNumber="76">Misunithyris</emphasis>
</taxonomicName>
clearly shows a
<taxonomicName id="31FE4D00D521A42EFED8FBFFFE4BFBCE" authorityName="Vörös" authorityYear="1983" box="[291,449,1061,1086]" class="Rhynchonellata" family="Zeilleriidae" genus="Bakonyithyris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Terebratulida" pageId="5" pageNumber="76" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D521A42EFED8FBFFFE4BFBCE" box="[291,449,1061,1086]" italics="true" pageId="5" pageNumber="76">Bakonyithyris</emphasis>
</taxonomicName>
-
<typeStatus id="29458821D521A42EFE32FBFFFE77FBCE" box="[457,509,1061,1086]" pageId="5" pageNumber="76">type</typeStatus>
pattern and might be arranged into Aulacothyrinae. However, the representatives of Aulacothyrinae (sensu
<bibRefCitation id="926F4B72D521A42EFE7CFBBCFF12FB6F" author="Baeza-Carratala, J. F. &amp; Garcia Joral, F." pageId="5" pageNumber="76" pagination="651 - 661" refId="ref8777" refString="Baeza-Carratala, J. F. and Garcia Joral, F. 2014. Crural bases position as a structural criterion for supraspecific diagnosis of Early Jurassic zeilleriid brachiopods. Acta Palaeontologica Polonica 59: 651 - 661." type="journal article" year="2014">Baeza-Carratalá and García Joral 2014</bibRefCitation>
) have not evidenced cardinal process so far, and the progress of dental plates is rather different.
</paragraph>
<paragraph id="F6413683D521A42EFF7CFB1DFC89F992" blockId="5.[98,771,160,1634]" pageId="5" pageNumber="76">
Summarizing, the arrangement of
<taxonomicName id="31FE4D00D521A42EFDCFFB1DFD4FFB10" authorityName="Baeza-Carratalá &amp; Pérez-Valera &amp; Pérez-Valera" authorityYear="2018" box="[564,709,1223,1248]" class="Rhynchonellata" family="Zeilleriidae" genus="Misunithyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="5" pageNumber="76" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D521A42EFDCFFB1DFD4FFB10" box="[564,709,1223,1248]" italics="true" pageId="5" pageNumber="76">Misunithyris</emphasis>
</taxonomicName>
into family
<taxonomicName id="31FE4D00D521A42EFF4DFB3DFEBBFAF0" box="[182,305,1255,1280]" class="Rhynchonellata" family="Zeilleridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="5" pageNumber="76" phylum="Brachiopoda" rank="family">Zeilleridae</taxonomicName>
is the best plausible determination, but does not fully agree with the currently defined subfamilies. Combining external and internal features, it seems to be closer to the multicostate zeilleriid stock, such as
<taxonomicName id="31FE4D00D521A42EFF99FAB2FE19FA71" authority="Feldman, 2013" authorityName="Feldman" authorityYear="2013" box="[98,403,1384,1409]" class="Rhynchonellata" family="Zeilleriidae" genus="Menathyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="5" pageNumber="76" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D521A42EFF99FAB2FF69FA71" box="[98,227,1384,1409]" italics="true" pageId="5" pageNumber="76">Menathyris</emphasis>
<bibRefCitation id="926F4B72D521A42EFF11FAB2FE19FA71" author="Feldman, H. R." box="[234,403,1384,1409]" pageId="5" pageNumber="76" pagination="81 - 85" refId="ref9714" refString="Feldman, H. R. 2013. Menathyris wilsoni (Brachiopoda), new genus and species from the Middle Triassic (Ladinian) of Southern Israel. Annales Societatis Geologorum Poloniae 83: 81 - 85." type="journal article" year="2013">Feldman, 2013</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="31FE4D00D521A42EFE65FAB2FD74FA71" authority="Owen and Rose, 1997" authorityName="Owen and Rose" authorityYear="1997" box="[414,766,1384,1409]" class="Rhynchonellata" family="Zeilleriidae" genus="Calpella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="5" pageNumber="76" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D521A42EFE65FAB2FE75FA71" box="[414,511,1384,1409]" italics="true" pageId="5" pageNumber="76">Calpella</emphasis>
<bibRefCitation id="926F4B72D521A42EFDFEFAB2FD74FA71" author="Owen, E. F. &amp; Rose, E. P. F." box="[517,766,1384,1409]" pageId="5" pageNumber="76" pagination="497 - 513" refId="ref10854" refString="Owen, E. F. and Rose, E. P. F. 1997. Early Jurassic Brachiopods from Gibraltar, and their Tethyan affinities. Palaeontology 40: 497 - 513." type="journal article" year="1997">Owen and Rose, 1997</bibRefCitation>
</taxonomicName>
, or
<taxonomicName id="31FE4D00D521A42EFF7AFA52FE4CFA51" authority="Seguenza, 1885" authorityName="Seguenza" authorityYear="1885" box="[129,454,1416,1441]" class="Cephalopoda" family="Hildoceratidae" genus="Tauromenia" kingdom="Animalia" pageId="5" pageNumber="76" phylum="Mollusca" rank="genus">
<emphasis id="C48AEA91D521A42EFF7AFA52FE83FA51" box="[129,265,1416,1441]" italics="true" pageId="5" pageNumber="76">Tauromenia</emphasis>
<bibRefCitation id="926F4B72D521A42EFEEAFA52FE4CFA51" author="Seguenza, G." box="[273,454,1416,1441]" pageId="5" pageNumber="76" pagination="377 - 497" refId="ref12207" refString="Seguenza, G. 1885. Monografia delle Spiriferina dei varii piani del Lias Messinese. Bollettino della Societa Geologica Italian a 4: 377 - 497." type="journal article" year="1885">Seguenza, 1885</bibRefCitation>
</taxonomicName>
. A new subfamily might be erected, either with
<taxonomicName id="31FE4D00D521A42EFEA8FA72FE68FA31" authorityName="Baeza-Carratalá &amp; Pérez-Valera &amp; Pérez-Valera" authorityYear="2018" box="[339,482,1448,1473]" class="Rhynchonellata" family="Zeilleriidae" genus="Misunithyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="5" pageNumber="76" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D521A42EFEA8FA72FE68FA31" box="[339,482,1448,1473]" italics="true" pageId="5" pageNumber="76">Misunithyris</emphasis>
</taxonomicName>
as monotypic taxon, or together with the aforementioned genera, but further studies of internal structures in their representatives will be required to establish the validity of this approach, emphasizing examination of cardinal process and brachidium architecture in addition to the already known shared features.
</paragraph>
<paragraph id="F6413683D521A42EFF99F9AEFD90F91E" blockId="5.[98,770,1652,1774]" pageId="5" pageNumber="76">
<emphasis id="C48AEA91D521A42EFF99F9AEFD81F97D" box="[98,523,1652,1677]" italics="true" pageId="5" pageNumber="76">Stratigraphic and geographic range</emphasis>
.—
<taxonomicName id="31FE4D00D521A42EFDD5F9AEFD37F97D" authorityName="Baeza-Carratalá &amp; Pérez-Valera &amp; Pérez-Valera" authorityYear="2018" box="[558,701,1652,1677]" class="Rhynchonellata" family="Zeilleriidae" genus="Misunithyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="5" pageNumber="76" phylum="Brachiopoda" rank="genus">
<emphasis id="C48AEA91D521A42EFDD5F9AEFD37F97D" box="[558,701,1652,1677]" italics="true" pageId="5" pageNumber="76">Misunithyris</emphasis>
</taxonomicName>
is so far a monospecific genus recorded in the
<taxonomicName id="31FE4D00D521A42EFDBBF94EFE96F93D" authority="Chronozone" authorityName="Chronozone" class="Cephalopoda" family="Hungaritidae" genus="Gevanites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ceratitida" pageId="5" pageNumber="76" phylum="Mollusca" rank="species" species="epigonus">
<emphasis id="C48AEA91D521A42EFDBBF94EFF00F93D" italics="true" pageId="5" pageNumber="76">Gevanites epigonus</emphasis>
Chronozone
</taxonomicName>
of the Fassanian (lower Ladinian, Middle Triassic) from the Betic Range (Fig. 2).
</paragraph>
</subSubSection>
</treatment>
</document>