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<mods:title id="6A37EA54F0A13F7C67750E863947EE03">Revision of Eccoptolonthus Bernhauer (Coleoptera: Staphylininae: Philonthina) with descriptions of four new species from China</mods:title>
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<mods:namePart id="D871961B682B5E411ECD16D35FB57C3B">Fei, Xu-Dong</mods:namePart>
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<emphasis id="C48FEADFE3592F74FF2EFA38FD5E077E" bold="true" box="[151,697,1510,1537]" pageId="1" pageNumber="474">
The Genus 
<taxonomicName id="31FB4D4EE3592F74FE9EFA39FD5E077E" ID-CoL="8X7C9" authority="Bernhauer, 1912" authorityName="Bernhauer" authorityYear="1912" box="[295,697,1510,1537]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3592F74FE9EFA39FE03077F" bold="true" box="[295,484,1510,1537]" italics="true" pageId="1" pageNumber="474">Eccoptolonthus</emphasis>
<bibRefCitation id="926A4B3CE3592F74FE52FA38FD5E077E" author="Bernhauer, M." box="[491,697,1510,1537]" pageId="1" pageNumber="474" pagination="177 - 183" refId="ref14451" refString="Bernhauer, M. (1912) Beitrag zur Staphylinidenfauna von Afrika (Col.). Entomologische Mitteilungen, 1 (6 &amp; 7), 177 - 183 &amp; 203 - 209." type="journal article" year="1912">Bernhauer, 1912</bibRefCitation>
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<typeStatus id="2940886FE3592F74FF2EF9F4FF2B073D" box="[151,204,1579,1603]" pageId="1" pageNumber="474">Type</typeStatus>
species:
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<taxonomicName id="31FB4D4EE3592F74FE95F9F4FCBB073D" authority="Bernhauer, 1912" authorityName="Bernhauer" authorityYear="1912" box="[300,860,1579,1603]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="species" species="conradti" subGenus="Eccoptolonthus">
<emphasis id="C48FEADFE3592F74FE95F9F4FD4C073D" box="[300,683,1579,1603]" italics="true" pageId="1" pageNumber="474">Philonthus (Eccoptolonthus) conradti</emphasis>
<bibRefCitation id="926A4B3CE3592F74FD08F9F4FCBB073D" author="Bernhauer, M." box="[689,860,1579,1603]" pageId="1" pageNumber="474" pagination="177 - 183" refId="ref14451" refString="Bernhauer, M. (1912) Beitrag zur Staphylinidenfauna von Afrika (Col.). Entomologische Mitteilungen, 1 (6 &amp; 7), 177 - 183 &amp; 203 - 209." type="journal article" year="1912">Bernhauer, 1912</bibRefCitation>
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.
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<paragraph id="F64436CDE3592F74FF2EF9B1FCED0623" blockId="1.[151,1437,1646,2038]" pageId="1" pageNumber="474">
<bibRefCitation id="926A4B3CE3592F74FF2EF9B1FE9007F8" author="Bernhauer, M." box="[151,375,1646,1670]" pageId="1" pageNumber="474" pagination="177 - 183" refId="ref14451" refString="Bernhauer, M. (1912) Beitrag zur Staphylinidenfauna von Afrika (Col.). Entomologische Mitteilungen, 1 (6 &amp; 7), 177 - 183 &amp; 203 - 209." type="journal article" year="1912">Bernhauer, 1912: 206</bibRefCitation>
(subgenus of 
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<emphasis id="C48FEADFE3592F74FDB4F9B1FD9C07F8" box="[525,635,1646,1670]" italics="true" pageId="1" pageNumber="474">Philonthus</emphasis>
</taxonomicName>
, species included: 
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<emphasis id="C48FEADFE3592F74FCFBF9B1FC7F07F8" box="[834,920,1646,1670]" italics="true" pageId="1" pageNumber="474">conradti</emphasis>
</taxonomicName>
); 
<bibRefCitation id="926A4B3CE3592F74FC09F9B1FAF707F8" author="Bernhauer, M. &amp; Schubert, K." box="[944,1296,1646,1670]" pageId="1" pageNumber="474" pagination="289 - 408" refId="ref14553" refString="Bernhauer, M. &amp; Schubert, K. (1914) Staphylinidae IV. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus. Vol. 5. Pars 57. Junk, Berlin, pp. 289 - 408." type="book chapter" year="1914">Bernhauer &amp; Schubert, 1914: 326</bibRefCitation>
(synonym of 
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<emphasis id="C48FEADFE3592F74FF7EF953FED207DA" box="[199,309,1676,1700]" italics="true" pageId="1" pageNumber="474">Philonthus</emphasis>
</taxonomicName>
); 
<bibRefCitation id="926A4B3CE3592F74FEF2F952FDEA07DA" author="Cameron, M." box="[331,525,1676,1701]" pageId="1" pageNumber="474" pagination="1 - 443" refId="ref14719" refString="Cameron, M. (1932) The Fauna of British India including Ceylon and Burma. Coleoptera. Staphylinidae. Vol. III. Taylor &amp; Francis London, 1 - 443." type="book chapter" year="1932">Cameron, 1932: 62</bibRefCitation>
(subgenus of 
<taxonomicName id="31FB4D4EE3592F74FD25F953FCED07DA" ID-CoL="8VRYR" authorityName="Stephens" authorityYear="1829" box="[668,778,1676,1700]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3592F74FD25F953FCED07DA" box="[668,778,1676,1700]" italics="true" pageId="1" pageNumber="474">Philonthus</emphasis>
</taxonomicName>
); 
<bibRefCitation id="926A4B3CE3592F74FC99F953FBF607DB" author="Blackwelder, W." box="[800,1041,1676,1701]" pageId="1" pageNumber="474" pagination="1 - 658" refId="ref14604" refString="Blackwelder, W. (1943) Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, Washington, 182, 1 - 658. https: // doi. org / 10.5479 / si. 03629236.182. i" type="journal article" year="1943">Blackwelder, 1943: 399</bibRefCitation>
(subgenus of 
<taxonomicName id="31FB4D4EE3592F74FB19F953FAE907DA" ID-CoL="8VRYR" authorityName="Stephens" authorityYear="1829" box="[1184,1294,1676,1700]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3592F74FB19F953FAE907DA" box="[1184,1294,1676,1700]" italics="true" pageId="1" pageNumber="474">Philonthus</emphasis>
</taxonomicName>
, 
<typeStatus id="2940886FE3592F74FAA3F952FAA107DB" box="[1306,1350,1677,1701]" pageId="1" pageNumber="474">type</typeStatus>
species: 
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<emphasis id="C48FEADFE3592F74FF7EF974FEFA07BD" box="[199,285,1707,1731]" italics="true" pageId="1" pageNumber="474">conradti</emphasis>
</taxonomicName>
); 
<bibRefCitation id="926A4B3CE3592F74FE8DF974FDCE07BD" author="Blackwelder, R. E." box="[308,553,1707,1731]" pageId="1" pageNumber="474" pagination="1 - 483" refId="ref14652" refString="Blackwelder, R. E. (1952) The generic names of the beetle family Staphylinidae, with an essay on genotypy. United States National Museum Bulletin, 200, 1 - 483." type="journal article" year="1952">Blackwelder, 1952: 138</bibRefCitation>
(subgenus of 
<taxonomicName id="31FB4D4EE3592F74FD04F974FCCC07BD" ID-CoL="8VRYR" authorityName="Stephens" authorityYear="1829" box="[701,811,1707,1731]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3592F74FD04F974FCCC07BD" box="[701,811,1707,1731]" italics="true" pageId="1" pageNumber="474">Philonthus</emphasis>
</taxonomicName>
, 
<typeStatus id="2940886FE3592F74FC80F973FC8207BA" box="[825,869,1708,1732]" pageId="1" pageNumber="474">type</typeStatus>
species: 
<taxonomicName id="31FB4D4EE3592F74FC7DF974FBFD07BD" authorityName="Bernhauer" authorityYear="1912" box="[964,1050,1707,1731]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="species" species="conradti">
<emphasis id="C48FEADFE3592F74FC7DF974FBFD07BD" box="[964,1050,1707,1731]" italics="true" pageId="1" pageNumber="474">conradti</emphasis>
</taxonomicName>
); 
<bibRefCitation id="926A4B3CE3592F74FB88F973FAF607BD" author="Herman, L. H." box="[1073,1297,1707,1731]" pageId="1" pageNumber="474" pagination="2819 - 2949" refId="ref15354" refString="Herman, L. H. (2001 a) Catalog of the Staphylinidae (Insecta: Coleoptera): 1758 to the end of the second millennium. V. Staphylinine group (Part 2). Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina). Bulletin of the American Museum of Natural History, 265, 2819 - 2949. https: // doi. org / 10.1206 / 0003 - 0090.265.1.5" type="journal article" year="2001">Herman, 2001a: 2733</bibRefCitation>
(subgenus of 
<taxonomicName id="31FB4D4EE3592F74FF7EF915FED2079C" ID-CoL="8VRYR" authorityName="Stephens" authorityYear="1829" box="[199,309,1738,1762]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3592F74FF7EF915FED2079C" box="[199,309,1738,1762]" italics="true" pageId="1" pageNumber="474">Philonthus</emphasis>
</taxonomicName>
); 
<bibRefCitation id="926A4B3CE3592F74FEF5F915FDE1079C" author="Newton, A. F. &amp; New Nomenclatural and Taxonomic Acts &amp; Lobl, I. &amp; Lobl, D." box="[332,518,1738,1762]" pageId="1" pageNumber="474" pagination="1" refId="ref15999" refString="Newton, A. F. (2015) New Nomenclatural and Taxonomic Acts, and Comments. In: Lobl, I. &amp; Lobl, D. (2015) Catalogue of Palaearctic Coleoptera. Vol. 2 / 1. Hydrophiloidea-Staphylinoidea Revised and Updated Edition. Vol. 1. Brill, Leiden and Boston, pp. 12 - 24. https: // doi. org / 10.1163 / 9789004296855 _ 002" type="book chapter" year="2015">Newton, 2015: 13</bibRefCitation>
(as valid genus, not subgenus or synonym of 
<taxonomicName id="31FB4D4EE3592F74FC5FF915FB7B079C" ID-CoL="8VRYR" authority="Curtis" authorityName="Curtis" authorityYear="1829" box="[998,1180,1738,1762]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3592F74FC5FF915FBB3079C" box="[998,1108,1738,1762]" italics="true" pageId="1" pageNumber="474">Philonthus</emphasis>
Curtis
</taxonomicName>
, catalog); 
<bibRefCitation id="926A4B3CE3592F74FAB0F915FEC6067F" author="Schillhammer, H. &amp; Staphylinidae, New Synonyms &amp; In &amp; Lobl, I. &amp; Lobl, D." pageId="1" pageNumber="474" pagination="1" refId="ref16408" refString="Schillhammer, H. (2015) Staphylinidae, New Synonyms. In: Lobl, I. &amp; Lobl, D. (2015) Catalogue of Palaearctic Coleoptera. Vol. 2 / 1. Hydrophiloidea-Staphylinoidea Revised and Updated Edition. Vol. 1. Brill, Leiden and Boston, pp. 23 - 24." type="book chapter" year="2015">Schillhammer, 2015: 23</bibRefCitation>
, 24 (new synonyms and new combinations of 
<taxonomicName id="31FB4D4EE3592F74FD4BF937FC77067E" ID-CoL="8X7C9" authorityName="Bernhauer" authorityYear="1912" box="[754,912,1768,1792]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3592F74FD4BF937FC77067E" box="[754,912,1768,1792]" italics="true" pageId="1" pageNumber="474">Eccoptolonthus</emphasis>
</taxonomicName>
); 
<bibRefCitation id="926A4B3CE3592F74FC1CF937FB0C067F" author="Schulke, M. &amp; Smetana, A." box="[933,1259,1768,1793]" pageId="1" pageNumber="474" pagination="1" refId="ref16467" refString="Schulke, M. &amp; Smetana, A. (2015) Staphylinidae, subtribe Philonthina. In: Lobl, I. &amp; Lobl, D. (2015) Catalogue of Palaearctic Coleoptera. Vol. 2 / 1. Hydrophiloidea-Staphylinoidea Revised and Updated Edition. Vol. 1. Brill, Leiden and Boston, pp. 1017 - 1054." type="book chapter" year="2015">Schülke &amp; Smetana, 2015: 1021</bibRefCitation>
(
<taxonomicName id="31FB4D4EE3592F74FB41F937FA71067E" ID-CoL="8X7C9" authorityName="Bernhauer" authorityYear="1912" box="[1272,1430,1768,1792]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3592F74FB41F937FA71067E" box="[1272,1430,1768,1792]" italics="true" pageId="1" pageNumber="474">Eccoptolonthus</emphasis>
</taxonomicName>
, catalog); Chani-Posse, 
<emphasis id="C48FEADFE3592F74FE0DF8D7FE050661" box="[436,482,1799,1823]" italics="true" pageId="1" pageNumber="474">et al</emphasis>
. 2018: 34 (two species of 
<taxonomicName id="31FB4D4EE3592F74FD4EF8D7FCB00661" ID-CoL="8VRQP" authorityName="Fauvel" authorityYear="1874" box="[759,855,1800,1823]" class="Insecta" family="Staphylinidae" genus="Hesperus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3592F74FD4EF8D7FCB00661" box="[759,855,1800,1823]" italics="true" pageId="1" pageNumber="474">Hesperus</emphasis>
</taxonomicName>
transferred to 
<taxonomicName id="31FB4D4EE3592F74FC49F8D8FB690661" ID-CoL="8X7C9" authorityName="Bernhauer" authorityYear="1912" box="[1008,1166,1799,1823]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3592F74FC49F8D8FB690661" box="[1008,1166,1799,1823]" italics="true" pageId="1" pageNumber="474">Eccoptolonthus</emphasis>
</taxonomicName>
); 
<bibRefCitation id="926A4B3CE3592F74FB1DF8D8FAD60661" author="Li, L. Z." box="[1188,1329,1799,1823]" pageId="1" pageNumber="474" refId="ref15835" refString="Li, L. Z. (2019) s. n. In: Catalogue of Chinese Coleoptera. Vol. 3. Staphylinidae. Science Press, Beijing, pp. 421 - 445." type="book" year="2019">Li, 2019: 424</bibRefCitation>
(
<taxonomicName id="31FB4D4EE3592F74FAF8F8D7FEF50640" ID-CoL="8X7C9" authorityName="Bernhauer" authorityYear="1912" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3592F74FAF8F8D7FEF50640" italics="true" pageId="1" pageNumber="474">Eccoptolonthus</emphasis>
</taxonomicName>
, catalog of Chinese); 
<bibRefCitation id="926A4B3CE3592F74FE41F8F9FD280640" author="Mikatova, S. &amp; Machackova, L. &amp; Hajek, J." box="[504,719,1830,1854]" pageId="1" pageNumber="474" pagination="353 - 389" refId="ref15872" refString="Mikatova, S., Machackova, L. &amp; Hajek, J. (2020) Catalogue of type specimens of beetles (Coleoptera) deposited in the National Museum, Prague, Czech Republic). Acta Entomolgica Muse Nationalis Pragae, 60 (1), 353 - 389. https: // doi. org / 10.37520 / aemnp. 2020.021" type="journal article" year="2020">Mikátová, 2020: 386</bibRefCitation>
(catalogue of 
<typeStatus id="2940886FE3592F74FCDFF8F9FC750640" box="[870,914,1830,1854]" pageId="1" pageNumber="474">type</typeStatus>
specimens of 
<taxonomicName id="31FB4D4EE3592F74FB91F8F9FB210640" ID-CoL="8X7C9" authorityName="Bernhauer" authorityYear="1912" box="[1064,1222,1830,1854]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3592F74FB91F8F9FB210640" box="[1064,1222,1830,1854]" italics="true" pageId="1" pageNumber="474">Eccoptolonthus</emphasis>
</taxonomicName>
in 
<collectingCountry id="8EEC765DE3592F74FB53F8F9FA760640" box="[1258,1425,1830,1854]" name="Czech Republic" pageId="1" pageNumber="474">Czech Republic</collectingCountry>
); 
<bibRefCitation id="926A4B3CE3592F74FF7EF89BFE6A0622" author="Hayashi, Y." box="[199,397,1860,1885]" pageId="1" pageNumber="474" pagination="135 - 142" refId="ref15304" refString="Hayashi, Y. (2020) A New Subgenus of the Genus Eccoptolonthus Bernhauer, with Descriptions of a New Species and a New Subspecies (Coleoptera, Staphylinidae, Philonthina). Elytra, Tokyo, New Series, 10 (1), 135 - 142." type="journal article" year="2020">Hayashi, 2020: 135</bibRefCitation>
(a new subgenus of 
<taxonomicName id="31FB4D4EE3592F74FDE7F89BFD1B0622" ID-CoL="8X7C9" authorityName="Bernhauer" authorityYear="1912" box="[606,764,1860,1884]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3592F74FDE7F89BFD1B0622" box="[606,764,1860,1884]" italics="true" pageId="1" pageNumber="474">Eccoptolonthus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="F64436CDE3592F74FF2EF8BCFE960688" blockId="1.[151,1437,1646,2038]" pageId="1" pageNumber="474">
<emphasis id="C48FEADFE3592F74FF2EF8BCFF280605" bold="true" box="[151,207,1891,1915]" pageId="1" pageNumber="474">Syn.:</emphasis>
<taxonomicName id="31FB4D4EE3592F74FF6FF8BCFDAC0605" ID-CoL="9JMLR" authority="Hayashi, 2008: 146" authorityName="Hayashi" authorityPageNumber="146" authorityYear="2008" box="[214,587,1891,1915]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3592F74FF6FF8BCFE9C0605" box="[214,379,1891,1915]" italics="true" pageId="1" pageNumber="474">Pseudohesperus</emphasis>
<bibRefCitation id="926A4B3CE3592F74FE3AF8BCFDAC0605" author="Hayashi, Y." box="[387,587,1891,1915]" pageId="1" pageNumber="474" pagination="145 - 153" refId="ref15256" refString="Hayashi, Y. (2008) Pseudohesperus, a new genus of Philonthina (Coleoptera: Staphylinidae) from East Asia, with redescription of its type species Philonthus rutiliventris Sharp. The Entomological Review of Japan, 63 (2), 145 - 153." type="journal article" year="2008">Hayashi, 2008: 146</bibRefCitation>
</taxonomicName>
(new genus, 
<typeStatus id="2940886FE3592F74FD6FF8BBFCE50602" box="[726,770,1892,1916]" pageId="1" pageNumber="474">type</typeStatus>
species: 
<taxonomicName id="31FB4D4EE3592F74FCDBF8BCFB300605" authority="Sharp, 1874" authorityName="Sharp" authorityYear="1874" box="[866,1239,1891,1915]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="species" species="rutiliventris">
<emphasis id="C48FEADFE3592F74FCDBF8BCFBA80605" box="[866,1103,1891,1915]" italics="true" pageId="1" pageNumber="474">Philonthus rutiliventris</emphasis>
<bibRefCitation id="926A4B3CE3592F74FBEEF8BCFB300605" author="Sharp, D." box="[1111,1239,1891,1915]" pageId="1" pageNumber="474" pagination="47 - 48" refId="ref16531" refString="Sharp, D. (1874) The Staphylinidae of Japan. Transactions of the Entomological Society of London, 1, 47 - 48. https: // doi. org / 10.1111 / j. 1365 - 2311.1874. tb 00159. x" type="journal article" year="1874">Sharp, 1874</bibRefCitation>
</taxonomicName>
); 
<bibRefCitation id="926A4B3CE3592F74FB53F8BCFF0A06E4" author="Hromadka, L." pageId="1" pageNumber="474" pagination="495 - 516" refId="ref15506" refString="Hromadka, L. (2010) Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae, 50 (2), 495 - 516." type="journal article" year="2010">Hromádka, 2010: 495</bibRefCitation>
(revised the Afrotropical fauna of 
<taxonomicName id="31FB4D4EE3592F74FDE9F85DFD1206E4" ID-CoL="9JMLR" authorityName="Hayashi" authorityYear="2008" box="[592,757,1922,1946]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3592F74FDE9F85DFD1206E4" box="[592,757,1922,1946]" italics="true" pageId="1" pageNumber="474">Pseudohesperus</emphasis>
</taxonomicName>
); 
<bibRefCitation id="926A4B3CE3592F74FCB2F85DFC0C06E4" author="Li, L. &amp; Zhou, H. Z." box="[779,1003,1922,1946]" pageId="1" pageNumber="474" pagination="679 - 722" refId="ref15762" refString="Li, L. &amp; Zhou, H. Z. (2011) Revision and phylogenetic assessment of the rove beetle genus Pseudohesperus Hayashi, with broad reference to the subtribe Philonthina (Coleoptera: Staphylinidae: Staphylinini). Zoological Journal of the Linnean Society, 163 (3), 679 - 722. https: // doi. org / 10.1111 / j. 1096 - 3642.2011.00731. x" type="journal article" year="2011">Li &amp; Zhou, 2011: 679</bibRefCitation>
(revised 
<collectingCountry id="8EEC765DE3592F74FBF3F85DFB6006E4" box="[1098,1159,1922,1946]" name="China" pageId="1" pageNumber="474">China</collectingCountry>
fauna of 
<taxonomicName id="31FB4D4EE3592F74FB51F85DFA6A06E4" ID-CoL="9JMLR" authorityName="Hayashi" authorityYear="2008" box="[1256,1421,1922,1946]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3592F74FB51F85DFA6A06E4" box="[1256,1421,1922,1946]" italics="true" pageId="1" pageNumber="474">Pseudohesperus</emphasis>
</taxonomicName>
); 
<bibRefCitation id="926A4B3CE3592F74FF7EF87FFE2206C7" author="Schillhammer, H." box="[199,453,1952,1977]" pageId="1" pageNumber="474" pagination="133 - 163" refId="ref16374" refString="Schillhammer, H. (2011) Old and new Staphylinini from the Palearctic and Oriental Regions (Coleoptera: Staphylinidae: Staphylininae). Koleopterologische Rundschau, 81, 133 - 163." type="journal article" year="2011">Schillhammer, 2011: 147</bibRefCitation>
(a new species of 
<taxonomicName id="31FB4D4EE3592F74FDC7F87FFCC406C6" ID-CoL="9JMLR" authorityName="Hayashi" authorityYear="2008" box="[638,803,1952,1976]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3592F74FDC7F87FFCC406C6" box="[638,803,1952,1976]" italics="true" pageId="1" pageNumber="474">Pseudohesperus</emphasis>
</taxonomicName>
); 
<bibRefCitation id="926A4B3CE3592F74FC8EF87EFC0B06C7" author="Newton, A. F. &amp; New Nomenclatural and Taxonomic Acts &amp; Lobl, I. &amp; Lobl, D." box="[823,1004,1952,1977]" pageId="1" pageNumber="474" pagination="1" refId="ref15999" refString="Newton, A. F. (2015) New Nomenclatural and Taxonomic Acts, and Comments. In: Lobl, I. &amp; Lobl, D. (2015) Catalogue of Palaearctic Coleoptera. Vol. 2 / 1. Hydrophiloidea-Staphylinoidea Revised and Updated Edition. Vol. 1. Brill, Leiden and Boston, pp. 12 - 24. https: // doi. org / 10.1163 / 9789004296855 _ 002" type="book chapter" year="2015">Newton, 2015: 12</bibRefCitation>
(synonym of 
<taxonomicName id="31FB4D4EE3592F74FBC1F87FFAF106C6" ID-CoL="8X7C9" authorityName="Bernhauer" authorityYear="1912" box="[1144,1302,1952,1976]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3592F74FBC1F87FFAF106C6" box="[1144,1302,1952,1976]" italics="true" pageId="1" pageNumber="474">Eccoptolonthus</emphasis>
</taxonomicName>
); 
<bibRefCitation id="926A4B3CE3592F74FA92F87FFEC806A9" author="Hromadka, L." pageId="1" pageNumber="474" pagination="371 - 381" refId="ref15555" refString="Hromadka, L. (2016) Six new Philonthini from the Afrotropical Region (Coleoptera: Staphylinidae: Philonthina). Studies and Reports, Taxonomical Series, 12 (2), 371 - 381." type="journal article" year="2016">Hromádka, 2016: 376</bibRefCitation>
(a new species of 
<taxonomicName id="31FB4D4EE3592F74FE4DF860FD7E06A9" ID-CoL="9JMLR" authorityName="Hayashi" authorityYear="2008" box="[500,665,1983,2007]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3592F74FE4DF860FD7E06A9" box="[500,665,1983,2007]" italics="true" pageId="1" pageNumber="474">Pseudohesperus</emphasis>
</taxonomicName>
); 
<bibRefCitation id="926A4B3CE3592F74FD08F860FC6F06A9" author="Mikatova, S. &amp; Machackova, L. &amp; Hajek, J." box="[689,904,1983,2007]" pageId="1" pageNumber="474" pagination="353 - 389" refId="ref15872" refString="Mikatova, S., Machackova, L. &amp; Hajek, J. (2020) Catalogue of type specimens of beetles (Coleoptera) deposited in the National Museum, Prague, Czech Republic). Acta Entomolgica Muse Nationalis Pragae, 60 (1), 353 - 389. https: // doi. org / 10.37520 / aemnp. 2020.021" type="journal article" year="2020">Mikátová, 2020: 386</bibRefCitation>
(catalogue of 
<typeStatus id="2940886FE3592F74FB99F81FFBAB06A6" box="[1056,1100,1984,2008]" pageId="1" pageNumber="474">type</typeStatus>
specimens of 
<taxonomicName id="31FB4D4EE3592F74FB5AF860FA6606A9" ID-CoL="8X7C9" authorityName="Bernhauer" authorityYear="1912" box="[1251,1409,1983,2007]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3592F74FB5AF860FA6606A9" box="[1251,1409,1983,2007]" italics="true" pageId="1" pageNumber="474">Eccoptolonthus</emphasis>
</taxonomicName>
in 
<collectingCountry id="8EEC765DE3592F74FF7EF801FE8D0688" box="[199,362,2014,2038]" name="Czech Republic" pageId="1" pageNumber="474">Czech Republic</collectingCountry>
)
</paragraph>
<paragraph id="F64436CDE35A2F77FF2EFF49FA8D01D0" blockId="2.[151,1386,150,174]" box="[151,1386,150,174]" pageId="2" pageNumber="475">
<emphasis id="C48FEADFE35A2F77FF2EFF49FEEF01D0" bold="true" box="[151,264,150,174]" pageId="2" pageNumber="475">Subgenus:</emphasis>
<taxonomicName id="31FB4D4EE35A2F77FEB7FF48FD8801D0" ID-CoL="8XRDQ" authority="Hayashi, 2020: 136" authorityName="Hayashi" authorityYear="2020" box="[270,623,150,174]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="496" phylum="Arthropoda" rank="subGenus" subGenus="Eumorimotous">
<emphasis id="C48FEADFE35A2F77FEB7FF48FE4401D0" box="[270,419,151,174]" italics="true" pageId="2" pageNumber="475">Eumorimotous</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FE13FF49FD8801D0" author="Hayashi, Y." box="[426,623,150,174]" pageId="2" pageNumber="475" pagination="135 - 142" refId="ref15304" refString="Hayashi, Y. (2020) A New Subgenus of the Genus Eccoptolonthus Bernhauer, with Descriptions of a New Species and a New Subspecies (Coleoptera, Staphylinidae, Philonthina). Elytra, Tokyo, New Series, 10 (1), 135 - 142." type="journal article" year="2020">Hayashi, 2020: 136</bibRefCitation>
</taxonomicName>
(
<typeStatus id="2940886FE35A2F77FDC4FF49FD4D01D0" box="[637,682,150,174]" pageId="2" pageNumber="475">type</typeStatus>
species: 
<taxonomicName id="31FB4D4EE35A2F77FCBEFF49FA8101D0" ID-CoL="8XZX4" authority="(Fauvel, 1895))" authorityName="" baseAuthorityName="Fauvel" baseAuthorityYear="1895" box="[775,1382,150,174]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="laevigatus" subGenus="Eumorimotous">
<emphasis id="C48FEADFE35A2F77FCBEFF49FC4201D0" box="[775,933,150,174]" italics="true" pageId="2" pageNumber="475">Eccoptolonthus</emphasis>
(
<emphasis id="C48FEADFE35A2F77FC0AFF48FBAF01D0" box="[947,1096,151,174]" italics="true" pageId="2" pageNumber="475">Eumorimotous</emphasis>
) 
<emphasis id="C48FEADFE35A2F77FBEEFF49FB5801D0" box="[1111,1215,150,174]" italics="true" pageId="2" pageNumber="475">laevigatus</emphasis>
(
<bibRefCitation id="926A4B3CE35A2F77FB74FF49FAB201D0" author="Fauvel, A." box="[1229,1365,150,174]" pageId="2" pageNumber="475" pagination="180 - 286" refId="ref15110" refString="Fauvel, A. (1895) Staphylinides nouveaux de l'Inde et de la Malaisie. Revue d'Entomologie, 14, 180 - 286. https: // doi. org / 10.1017 / S 102688120005892 X" type="journal article" year="1895">Fauvel, 1895</bibRefCitation>
))
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="BEE16546E35A2F77FF2EFF03FD91008C" pageId="2" pageNumber="475" type="diagnosis">
<paragraph id="F64436CDE35A2F77FF2EFF03FD91008C" blockId="2.[151,1437,220,1795]" pageId="2" pageNumber="475">
<emphasis id="C48FEADFE35A2F77FF2EFF03FEED0188" bold="true" box="[151,266,220,246]" pageId="2" pageNumber="475">Diagnosis</emphasis>
: The genus 
<taxonomicName id="31FB4D4EE35A2F77FE2EFF03FDA20188" authorityName="Bernhauer" authorityYear="1912" box="[407,581,220,246]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE35A2F77FE2EFF03FDA20188" box="[407,581,220,246]" italics="true" pageId="2" pageNumber="475">Eccoptolonthus</emphasis>
</taxonomicName>
was characterized in great detail by 
<bibRefCitation id="926A4B3CE35A2F77FC5CFF03FB710189" author="Hayashi, Y." box="[997,1174,220,247]" pageId="2" pageNumber="475" pagination="145 - 153" refId="ref15256" refString="Hayashi, Y. (2008) Pseudohesperus, a new genus of Philonthina (Coleoptera: Staphylinidae) from East Asia, with redescription of its type species Philonthus rutiliventris Sharp. The Entomological Review of Japan, 63 (2), 145 - 153." type="journal article" year="2008">Hayashi (2008)</bibRefCitation>
, 
<bibRefCitation id="926A4B3CE35A2F77FB1AFF03FA8B0189" author="Hromadka, L." box="[1187,1388,220,247]" pageId="2" pageNumber="475" pagination="495 - 516" refId="ref15506" refString="Hromadka, L. (2010) Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae, 50 (2), 495 - 516." type="journal article" year="2010">Hromádka (2010)</bibRefCitation>
and 
<bibRefCitation id="926A4B3CE35A2F77FF2EFEDFFE860065" author="Li, L. &amp; Zhou, H. Z." box="[151,353,256,283]" pageId="2" pageNumber="475" pagination="679 - 722" refId="ref15762" refString="Li, L. &amp; Zhou, H. Z. (2011) Revision and phylogenetic assessment of the rove beetle genus Pseudohesperus Hayashi, with broad reference to the subtribe Philonthina (Coleoptera: Staphylinidae: Staphylinini). Zoological Journal of the Linnean Society, 163 (3), 679 - 722. https: // doi. org / 10.1111 / j. 1096 - 3642.2011.00731. x" type="journal article" year="2011">Li &amp; Zhou (2011)</bibRefCitation>
: the genus is closely allied to 
<taxonomicName id="31FB4D4EE35A2F77FD12FEDEFC490065" authority="Stephens, 1829" authorityName="Stephens" authorityYear="1829" box="[683,942,256,283]" class="Insecta" family="Staphylinidae" genus="Bisnius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE35A2F77FD12FEDEFD1A0064" box="[683,765,257,282]" italics="true" pageId="2" pageNumber="475">Bisnius</emphasis>
Stephens, 1829
</taxonomicName>
and 
<taxonomicName id="31FB4D4EE35A2F77FC5DFEDFFB140065" authority="Stephens, 1829" authorityName="Stephens" authorityYear="1829" box="[996,1267,256,283]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE35A2F77FC5DFEDFFBA70064" box="[996,1088,256,282]" italics="true" pageId="2" pageNumber="475">Gabrius</emphasis>
Stephens, 1829
</taxonomicName>
, because of the similar structure of protarsi, but is easily distinguishable from them by the following characters: (1) maxillary and labial palpi with fourth segment rod-like; (2) gular sutures gradually convergent posteriad to the neck constriction, not parallel in posterior (basal) half; (3) chaetotaxy of the pronotum composed of only a pair of anterolateral macrosetae, basolateral setae absent; (4) mesoventrite with apical portion distinctly and abruptly convex ventrad, without transverse carina; (5) sternite IX of males in most species with a pair of long appendages and without any long erect bristles; only a few with short appendages.
</paragraph>
</subSubSection>
<subSubSection id="BEE16546E35A2F77FF7EFE23FC3A067C" pageId="2" pageNumber="475" type="discussion">
<paragraph id="F64436CDE35A2F77FF7EFE23FDE003D9" blockId="2.[151,1437,220,1795]" pageId="2" pageNumber="475">
<emphasis id="C48FEADFE35A2F77FF7EFE23FE490368" bold="true" box="[199,430,508,534]" pageId="2" pageNumber="475">Taxonomic history:</emphasis>
the name 
<taxonomicName id="31FB4D4EE35A2F77FD98FE23FC680369" authority="Bernhauer, 1912" authorityName="Bernhauer" authorityYear="1912" box="[545,911,508,535]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE35A2F77FD98FE23FD280368" box="[545,719,508,534]" italics="true" pageId="2" pageNumber="475">Eccoptolonthus</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FD6FFE23FC680369" author="Bernhauer, M." box="[726,911,508,535]" pageId="2" pageNumber="475" pagination="177 - 183" refId="ref14451" refString="Bernhauer, M. (1912) Beitrag zur Staphylinidenfauna von Afrika (Col.). Entomologische Mitteilungen, 1 (6 &amp; 7), 177 - 183 &amp; 203 - 209." type="journal article" year="1912">Bernhauer, 1912</bibRefCitation>
</taxonomicName>
was treated as a subgenus of the genus 
<taxonomicName id="31FB4D4EE35A2F77FAF1FE23FE9B0345" authority="Stephens, 1829" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE35A2F77FAF1FE23FF210344" italics="true" pageId="2" pageNumber="475">Philonthus</emphasis>
Stephens, 1829
</taxonomicName>
(
<bibRefCitation id="926A4B3CE35A2F77FE35FDFFFD3F0345" author="Bernhauer, M. &amp; Schubert, K." box="[396,728,544,571]" pageId="2" pageNumber="475" pagination="289 - 408" refId="ref14553" refString="Bernhauer, M. &amp; Schubert, K. (1914) Staphylinidae IV. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus. Vol. 5. Pars 57. Junk, Berlin, pp. 289 - 408." type="book chapter" year="1914">Bernhauer &amp; Schubert, 1914</bibRefCitation>
; 
<bibRefCitation id="926A4B3CE35A2F77FD5FFDFEFC700345" author="Cameron, M." box="[742,919,545,571]" pageId="2" pageNumber="475" pagination="1 - 443" refId="ref14719" refString="Cameron, M. (1932) The Fauna of British India including Ceylon and Burma. Coleoptera. Staphylinidae. Vol. III. Taylor &amp; Francis London, 1 - 443." type="book chapter" year="1932">Cameron, 1932</bibRefCitation>
; 
<bibRefCitation id="926A4B3CE35A2F77FC1CFDFFFB9B0345" author="Blackwelder, W." box="[933,1148,544,571]" pageId="2" pageNumber="475" pagination="1 - 658" refId="ref14604" refString="Blackwelder, W. (1943) Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, Washington, 182, 1 - 658. https: // doi. org / 10.5479 / si. 03629236.182. i" type="journal article" year="1943">Blackwelder, 1943</bibRefCitation>
; 
<bibRefCitation id="926A4B3CE35A2F77FB33FDFFFA860345" author="Blackwelder, R. E." box="[1162,1377,544,571]" pageId="2" pageNumber="475" pagination="1 - 483" refId="ref14652" refString="Blackwelder, R. E. (1952) The generic names of the beetle family Staphylinidae, with an essay on genotypy. United States National Museum Bulletin, 200, 1 - 483." type="journal article" year="1952">Blackwelder, 1952</bibRefCitation>
; 
<bibRefCitation id="926A4B3CE35A2F77FAD7FDFEFED40320" author="Tottenham, C. E." pageId="2" pageNumber="475" pagination="230 - 231" refId="ref16769" refString="Tottenham, C. E. (1962) Coleoptera Staphylinidae Staphylininae. Mission zoologique de l'I. R. S. A. C. en Afrique orientale (P. Basilewsky et N. Leleup, 1957). LXXVI. Annales du Musee Royal de l'Afrique Centrale, Tervuren, Serie 8 º: Sciences Zoologiques, 110, 230 - 231." type="journal article" year="1962">Tottenham, 1962</bibRefCitation>
; 
<bibRefCitation id="926A4B3CE35A2F77FE86FD9AFE0A0321" author="Herman, L. H." box="[319,493,581,607]" pageId="2" pageNumber="475" pagination="2819 - 2949" refId="ref15354" refString="Herman, L. H. (2001 a) Catalog of the Staphylinidae (Insecta: Coleoptera): 1758 to the end of the second millennium. V. Staphylinine group (Part 2). Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina). Bulletin of the American Museum of Natural History, 265, 2819 - 2949. https: // doi. org / 10.1206 / 0003 - 0090.265.1.5" type="journal article" year="2001">Herman, 2001a</bibRefCitation>
) and included only a few species: 
<taxonomicName id="31FB4D4EE35A2F77FCCAFD9BFF2803FD" authority="Bernhauer, 1912" authorityName="Bernhauer" authorityYear="1912" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="conradti" subGenus="Eccoptolonthus">
<emphasis id="C48FEADFE35A2F77FCCAFD9BFC0B0320" box="[883,1004,580,606]" italics="true" pageId="2" pageNumber="475">Philonthus</emphasis>
(
<emphasis id="C48FEADFE35A2F77FC44FD9BFB4C0320" box="[1021,1195,580,606]" italics="true" pageId="2" pageNumber="475">Eccoptolonthus</emphasis>
) 
<emphasis id="C48FEADFE35A2F77FB02FD9BFAFD0320" box="[1211,1306,580,606]" italics="true" pageId="2" pageNumber="475">conradti</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FA9BFD9BFF2803FD" author="Bernhauer, M." pageId="2" pageNumber="475" pagination="177 - 183" refId="ref14451" refString="Bernhauer, M. (1912) Beitrag zur Staphylinidenfauna von Afrika (Col.). Entomologische Mitteilungen, 1 (6 &amp; 7), 177 - 183 &amp; 203 - 209." type="journal article" year="1912">Bernhauer, 1912</bibRefCitation>
</taxonomicName>
as the 
<typeStatus id="2940886FE35A2F77FE9CFDB6FEB203FD" box="[293,341,617,643]" pageId="2" pageNumber="475">type</typeStatus>
species, 
<taxonomicName id="31FB4D4EE35A2F77FE79FDB7FBBA03FC" authority="Tottenham, 1962" authorityName="Tottenham" authorityYear="1962" box="[448,1117,616,643]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="eccoptomus" subGenus="Eccoptolonthus">
<emphasis id="C48FEADFE35A2F77FE79FDB7FDDE03FC" box="[448,569,616,642]" italics="true" pageId="2" pageNumber="475">Philonthus</emphasis>
(
<emphasis id="C48FEADFE35A2F77FDF5FDB7FD1D03FC" box="[588,762,616,642]" italics="true" pageId="2" pageNumber="475">Eccoptolonthus</emphasis>
) 
<emphasis id="C48FEADFE35A2F77FCB4FDB6FC7603FC" box="[781,913,617,642]" italics="true" pageId="2" pageNumber="475">eccoptomus</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FC22FDB7FBBA03FC" author="Tottenham, C. E." box="[923,1117,616,642]" pageId="2" pageNumber="475" pagination="230 - 231" refId="ref16769" refString="Tottenham, C. E. (1962) Coleoptera Staphylinidae Staphylininae. Mission zoologique de l'I. R. S. A. C. en Afrique orientale (P. Basilewsky et N. Leleup, 1957). LXXVI. Annales du Musee Royal de l'Afrique Centrale, Tervuren, Serie 8 º: Sciences Zoologiques, 110, 230 - 231." type="journal article" year="1962">Tottenham, 1962</bibRefCitation>
</taxonomicName>
, and another un-named one mentioned by 
<bibRefCitation id="926A4B3CE35A2F77FE8EFD53FDE403D9" author="Tottenham, C. E." box="[311,515,652,679]" pageId="2" pageNumber="475" pagination="230 - 231" refId="ref16769" refString="Tottenham, C. E. (1962) Coleoptera Staphylinidae Staphylininae. Mission zoologique de l'I. R. S. A. C. en Afrique orientale (P. Basilewsky et N. Leleup, 1957). LXXVI. Annales du Musee Royal de l'Afrique Centrale, Tervuren, Serie 8 º: Sciences Zoologiques, 110, 230 - 231." type="journal article" year="1962">Tottenham (1962)</bibRefCitation>
.
</paragraph>
<paragraph id="F64436CDE35A2F77FF7EFD6EFAF0054D" blockId="2.[151,1437,220,1795]" pageId="2" pageNumber="475">
In contrast, the genus 
<taxonomicName id="31FB4D4EE35A2F77FE07FD6FFCFA03B5" authority="Hayashi, 2008" authorityName="Hayashi" authorityYear="2008" box="[446,797,688,715]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE35A2F77FE07FD6FFD9303B4" box="[446,628,688,714]" italics="true" pageId="2" pageNumber="475">Pseudohesperus</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FDC5FD6FFCFA03B5" author="Hayashi, Y." box="[636,797,688,715]" pageId="2" pageNumber="475" pagination="145 - 153" refId="ref15256" refString="Hayashi, Y. (2008) Pseudohesperus, a new genus of Philonthina (Coleoptera: Staphylinidae) from East Asia, with redescription of its type species Philonthus rutiliventris Sharp. The Entomological Review of Japan, 63 (2), 145 - 153." type="journal article" year="2008">Hayashi, 2008</bibRefCitation>
</taxonomicName>
was established as a separate genus and it was treated as valid for a long time after its description. In the original paper, 
<bibRefCitation id="926A4B3CE35A2F77FCEFFD0BFBE40391" author="Hayashi, Y." box="[854,1027,724,751]" pageId="2" pageNumber="475" pagination="145 - 153" refId="ref15256" refString="Hayashi, Y. (2008) Pseudohesperus, a new genus of Philonthina (Coleoptera: Staphylinidae) from East Asia, with redescription of its type species Philonthus rutiliventris Sharp. The Entomological Review of Japan, 63 (2), 145 - 153." type="journal article" year="2008">Hayashi (2008)</bibRefCitation>
transferred the 
<typeStatus id="2940886FE35A2F77FB0AFD0AFB040391" box="[1203,1251,725,751]" pageId="2" pageNumber="475">type</typeStatus>
species, 
<taxonomicName id="31FB4D4EE35A2F77FAF1FD0BFE39026D" authority="Sharp, 1874" authorityName="Sharp" authorityYear="1874" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="rutiliventris">
<emphasis id="C48FEADFE35A2F77FAF1FD0BFEB7026C" italics="true" pageId="2" pageNumber="475">Philonthus rutiliventris</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FEEEFD27FE39026D" author="Sharp, D." box="[343,478,760,787]" pageId="2" pageNumber="475" pagination="47 - 48" refId="ref16531" refString="Sharp, D. (1874) The Staphylinidae of Japan. Transactions of the Entomological Society of London, 1, 47 - 48. https: // doi. org / 10.1111 / j. 1365 - 2311.1874. tb 00159. x" type="journal article" year="1874">Sharp, 1874</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="31FB4D4EE35A2F77FDADFD27FC69026D" authority="Kraatz, 1859" authorityName="Kraatz" authorityYear="1859" box="[532,910,760,787]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="eustilbus">
<emphasis id="C48FEADFE35A2F77FDADFD27FD1F026C" box="[532,760,760,786]" italics="true" pageId="2" pageNumber="475">Philonthus eustilbus</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FD47FD26FC69026D" author="Kraatz, G." box="[766,910,760,787]" pageId="2" pageNumber="475" pagination="98 - 99" refId="ref15650" refString="Kraatz, G. (1859) Die Staphylinen-Fauna von Ostindien, insbesondere der Insel Ceylan. Archiv fur Naturgeschichte, 25 (1), 98 - 99. https: // doi. org / 10.5962 / bhl. title. 66002" type="journal article" year="1859">Kraatz, 1859</bibRefCitation>
</taxonomicName>
from the genus 
<taxonomicName id="31FB4D4EE35A2F77FBFAFD27FB5B026C" authorityName="Stephens" authorityYear="1829" box="[1091,1212,760,786]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE35A2F77FBFAFD27FB5B026C" box="[1091,1212,760,786]" italics="true" pageId="2" pageNumber="475">Philonthus</emphasis>
</taxonomicName>
to 
<taxonomicName id="31FB4D4EE35A2F77FB66FD27FA72026C" authorityName="Hayashi" authorityYear="2008" box="[1247,1429,760,786]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE35A2F77FB66FD27FA72026C" box="[1247,1429,760,786]" italics="true" pageId="2" pageNumber="475">Pseudohesperus</emphasis>
</taxonomicName>
. 
<bibRefCitation id="926A4B3CE35A2F77FF2EFCC3FEB80249" author="Hromadka, L." box="[151,351,796,823]" pageId="2" pageNumber="475" pagination="495 - 516" refId="ref15506" refString="Hromadka, L. (2010) Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae, 50 (2), 495 - 516." type="journal article" year="2010">Hromádka (2010)</bibRefCitation>
revised the Afrotropical fauna, namely, 
<taxonomicName id="31FB4D4EE35A2F77FC9AFCC2FBB30249" authority="Hromadka, 2010" authorityName="Hromadka" authorityYear="2010" box="[803,1108,796,823]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="apsilus">
<emphasis id="C48FEADFE35A2F77FC9AFCC2FC6A0248" box="[803,909,796,822]" italics="true" pageId="2" pageNumber="475">P. apsilus</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FC2CFCC3FBB30249" author="Hromadka, L." box="[917,1108,796,823]" pageId="2" pageNumber="475" pagination="495 - 516" refId="ref15506" refString="Hromadka, L. (2010) Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae, 50 (2), 495 - 516." type="journal article" year="2010">Hromádka, 2010</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="31FB4D4EE35A2F77FBE6FCC2FA7F0249" authority="Hromadka, 2010" authorityName="Hromadka" authorityYear="2010" box="[1119,1432,796,823]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="tauraco">
<emphasis id="C48FEADFE35A2F77FBE6FCC2FB350248" box="[1119,1234,797,822]" italics="true" pageId="2" pageNumber="475">P. tauraco</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FB60FCC3FA7F0249" author="Hromadka, L." box="[1241,1432,796,823]" pageId="2" pageNumber="475" pagination="495 - 516" refId="ref15506" refString="Hromadka, L. (2010) Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae, 50 (2), 495 - 516." type="journal article" year="2010">Hromádka, 2010</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="31FB4D4EE35A2F77FF2EFC9EFE4E0225" authority="Hromadka, 2010" authorityName="Hromadka" authorityYear="2010" box="[151,425,832,859]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="tyto">
<emphasis id="C48FEADFE35A2F77FF2EFC9EFF380224" box="[151,223,833,858]" italics="true" pageId="2" pageNumber="475">P. tyto</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FF51FC9FFE4E0225" author="Hromadka, L." box="[232,425,832,859]" pageId="2" pageNumber="475" pagination="495 - 516" refId="ref15506" refString="Hromadka, L. (2010) Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae, 50 (2), 495 - 516." type="journal article" year="2010">Hromádka, 2010</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="31FB4D4EE35A2F77FE0FFC9EFD1E0225" authority="Hromadka, 2010" authorityName="Hromadka" authorityYear="2010" box="[438,761,832,859]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="varanus">
<emphasis id="C48FEADFE35A2F77FE0FFC9EFDC90224" box="[438,558,833,858]" italics="true" pageId="2" pageNumber="475">P. varanus</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FD8EFC9FFD1E0225" author="Hromadka, L." box="[567,761,832,859]" pageId="2" pageNumber="475" pagination="495 - 516" refId="ref15506" refString="Hromadka, L. (2010) Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae, 50 (2), 495 - 516." type="journal article" year="2010">Hromádka, 2010</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="31FB4D4EE35A2F77FCBFFC9EFBBB0225" authority="(Bernhauer, 1912)" baseAuthorityName="Bernhauer" baseAuthorityYear="1912" box="[774,1116,832,859]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="conradti">
<emphasis id="C48FEADFE35A2F77FCBFFC9EFC650224" box="[774,898,832,858]" italics="true" pageId="2" pageNumber="475">P. conradti</emphasis>
(
<bibRefCitation id="926A4B3CE35A2F77FC2AFC9FFBB30225" author="Bernhauer, M." box="[915,1108,832,859]" pageId="2" pageNumber="475" pagination="177 - 183" refId="ref14451" refString="Bernhauer, M. (1912) Beitrag zur Staphylinidenfauna von Afrika (Col.). Entomologische Mitteilungen, 1 (6 &amp; 7), 177 - 183 &amp; 203 - 209." type="journal article" year="1912">Bernhauer, 1912</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="31FB4D4EE35A2F77FBD3FC9EFF3C0201" authority="(Tottenham, 1962)" baseAuthorityName="Tottenham" baseAuthorityYear="1962" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="eccoptomus">
<emphasis id="C48FEADFE35A2F77FBD3FC9EFAEC0224" box="[1130,1291,833,858]" italics="true" pageId="2" pageNumber="475">P. eccoptomus</emphasis>
(
<bibRefCitation id="926A4B3CE35A2F77FAA5FC9FFF340201" author="Tottenham, C. E." pageId="2" pageNumber="475" pagination="230 - 231" refId="ref16769" refString="Tottenham, C. E. (1962) Coleoptera Staphylinidae Staphylininae. Mission zoologique de l'I. R. S. A. C. en Afrique orientale (P. Basilewsky et N. Leleup, 1957). LXXVI. Annales du Musee Royal de l'Afrique Centrale, Tervuren, Serie 8 º: Sciences Zoologiques, 110, 230 - 231." type="journal article" year="1962">Tottenham, 1962</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="31FB4D4EE35A2F77FF51FCBAFDA80201" authority="(Bernhauer, 1912)" baseAuthorityName="Bernhauer" baseAuthorityYear="1912" box="[232,591,868,895]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="proselytus">
<emphasis id="C48FEADFE35A2F77FF51FCBAFE900200" box="[232,375,868,894]" italics="true" pageId="2" pageNumber="475">P. proselytus</emphasis>
(
<bibRefCitation id="926A4B3CE35A2F77FE3EFCBBFDAF0201" author="Bernhauer, M." box="[391,584,868,895]" pageId="2" pageNumber="475" pagination="177 - 183" refId="ref14451" refString="Bernhauer, M. (1912) Beitrag zur Staphylinidenfauna von Afrika (Col.). Entomologische Mitteilungen, 1 (6 &amp; 7), 177 - 183 &amp; 203 - 209." type="journal article" year="1912">Bernhauer, 1912</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="31FB4D4EE35A2F77FDE4FCBAFC270201" authority="(Levasseur, 1967)" baseAuthorityName="Levasseur" baseAuthorityYear="1967" box="[605,960,868,895]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="bafutensis">
<emphasis id="C48FEADFE35A2F77FDE4FCBAFD0B0200" box="[605,748,868,894]" italics="true" pageId="2" pageNumber="475">P. bafutensis</emphasis>
(
<bibRefCitation id="926A4B3CE35A2F77FD42FCBAFC5E0201" author="Levasseur, L." box="[763,953,868,895]" pageId="2" pageNumber="475" pagination="954 - 977" refId="ref15694" refString="Levasseur, L. (1967) Nouveaux genres et especes de Staphylinidae (Col.) d'Afrique. Bulletin de l'Institut Fondamental d'Afrique Noire, (A), 29 (3), 954 - 977." type="journal article" year="1967">Levasseur, 1967</bibRefCitation>
)
</taxonomicName>
, and 
<taxonomicName id="31FB4D4EE35A2F77FC46FCBAFA920201" authority="(Scheerpeltz, 1956)" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1956" box="[1023,1397,868,895]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="natalensis">
<emphasis id="C48FEADFE35A2F77FC46FCBAFB690200" box="[1023,1166,868,894]" italics="true" pageId="2" pageNumber="475">P. natalensis</emphasis>
(
<bibRefCitation id="926A4B3CE35A2F77FB24FCBBFA8A0201" author="Scheerpeltz, O." box="[1181,1389,868,895]" pageId="2" pageNumber="475" pagination="478 - 516" refId="ref16219" refString="Scheerpeltz, O. (1956) Die von Prof. Dr. K. E. Schedl in Belgisch-Kongo aufgefundenen Staphyliniden (Coleoptera). Revue de Zoologie et de Botanique Africaines, 53 &amp; 54, 478 - 516 &amp; 8 - 46." type="journal article" year="1956">Scheerpeltz, 1956</bibRefCitation>
)
</taxonomicName>
. At the same time 
<taxonomicName id="31FB4D4EE35A2F77FE85FC56FD0502DD" authority="Cameron, 1950" authorityName="Cameron" authorityYear="1950" box="[316,738,904,931]" class="Insecta" family="Staphylinidae" genus="Hesperus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="longicornis">
<emphasis id="C48FEADFE35A2F77FE85FC56FDC902DC" box="[316,558,904,930]" italics="true" pageId="2" pageNumber="475">Hesperus longicornis</emphasis>
Cameron, 1950
</taxonomicName>
was synonymized with 
<taxonomicName id="31FB4D4EE35A2F77FC4DFC57FE5002B9" authority="(Bernhauer, 1912) (Hromadka, 2010)" authorityName="Hromadka" authorityYear="2010" baseAuthorityName="Bernhauer" baseAuthorityYear="1912" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="conradti">
<emphasis id="C48FEADFE35A2F77FC4DFC57FAF602DC" box="[1012,1297,904,930]" italics="true" pageId="2" pageNumber="475">Pseudohesperus conradti</emphasis>
(
<bibRefCitation id="926A4B3CE35A2F77FA99FC57FF3702B9" author="Bernhauer, M." pageId="2" pageNumber="475" pagination="177 - 183" refId="ref14451" refString="Bernhauer, M. (1912) Beitrag zur Staphylinidenfauna von Afrika (Col.). Entomologische Mitteilungen, 1 (6 &amp; 7), 177 - 183 &amp; 203 - 209." type="journal article" year="1912">Bernhauer, 1912</bibRefCitation>
) (
<bibRefCitation id="926A4B3CE35A2F77FF50FC73FE4802B9" author="Hromadka, L." box="[233,431,940,967]" pageId="2" pageNumber="475" pagination="495 - 516" refId="ref15506" refString="Hromadka, L. (2010) Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae, 50 (2), 495 - 516." type="journal article" year="2010">Hromádka, 2010</bibRefCitation>
)
</taxonomicName>
. 
<bibRefCitation id="926A4B3CE35A2F77FE7DFC73FD5302B9" author="Schillhammer, H." box="[452,692,940,967]" pageId="2" pageNumber="475" pagination="133 - 163" refId="ref16374" refString="Schillhammer, H. (2011) Old and new Staphylinini from the Palearctic and Oriental Regions (Coleoptera: Staphylinidae: Staphylininae). Koleopterologische Rundschau, 81, 133 - 163." type="journal article" year="2011">Schillhammer (2011)</bibRefCitation>
described another species, 
<taxonomicName id="31FB4D4EE35A2F77FC48FC73FF360295" authority="Schillhammer, 2011" authorityName="Schillhammer" authorityYear="2011" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="ernsti">
<emphasis id="C48FEADFE35A2F77FC48FC73FB1702B8" box="[1009,1264,940,966]" italics="true" pageId="2" pageNumber="475">Pseudohesperus ernsti</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FB40FC73FF360295" author="Schillhammer, H." pageId="2" pageNumber="475" pagination="133 - 163" refId="ref16374" refString="Schillhammer, H. (2011) Old and new Staphylinini from the Palearctic and Oriental Regions (Coleoptera: Staphylinidae: Staphylininae). Koleopterologische Rundschau, 81, 133 - 163." type="journal article" year="2011">Schillhammer, 2011</bibRefCitation>
</taxonomicName>
. In the same year, 
<bibRefCitation id="926A4B3CE35A2F77FE13FC0FFD9F0295" author="Li, L. &amp; Zhou, H. Z." box="[426,632,976,1003]" pageId="2" pageNumber="475" pagination="679 - 722" refId="ref15762" refString="Li, L. &amp; Zhou, H. Z. (2011) Revision and phylogenetic assessment of the rove beetle genus Pseudohesperus Hayashi, with broad reference to the subtribe Philonthina (Coleoptera: Staphylinidae: Staphylinini). Zoological Journal of the Linnean Society, 163 (3), 679 - 722. https: // doi. org / 10.1111 / j. 1096 - 3642.2011.00731. x" type="journal article" year="2011">Li &amp; Zhou (2011)</bibRefCitation>
revised the Chinese fauna with a phylogenetic analysis of 
<taxonomicName id="31FB4D4EE35A2F77FAA2FC0FFA7B0294" authorityName="Kirby" authorityYear="1837" box="[1307,1436,976,1002]" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="subTribe" subTribe="Philonthina">Philonthina</taxonomicName>
and reported six Chinese species, of which four were described as new: 
<taxonomicName id="31FB4D4EE35A2F77FC01FC2BFA9B0571" authority="Li &amp; Zhou, 2011" authorityName="Li &amp; Zhou" authorityYear="2011" box="[952,1404,1012,1039]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="luteus">
<emphasis id="C48FEADFE35A2F77FC01FC2BFB5F0570" box="[952,1208,1012,1038]" italics="true" pageId="2" pageNumber="475">Pseudohesperus luteus</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FB07FC2BFA9B0571" author="Li, L. &amp; Zhou, H. Z." box="[1214,1404,1012,1039]" pageId="2" pageNumber="475" pagination="679 - 722" refId="ref15762" refString="Li, L. &amp; Zhou, H. Z. (2011) Revision and phylogenetic assessment of the rove beetle genus Pseudohesperus Hayashi, with broad reference to the subtribe Philonthina (Coleoptera: Staphylinidae: Staphylinini). Zoological Journal of the Linnean Society, 163 (3), 679 - 722. https: // doi. org / 10.1111 / j. 1096 - 3642.2011.00731. x" type="journal article" year="2011">Li &amp; Zhou, 2011</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="31FB4D4EE35A2F77FA31FC2AFE0B054D" authority="Li &amp; Zhou, 2011" authorityName="Li &amp; Zhou" authorityYear="2011" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="pedatiformis">
<emphasis id="C48FEADFE35A2F77FA31FC2AFEC2054C" italics="true" pageId="2" pageNumber="475">P. pedatiformis</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FE95FBC7FE0B054D" author="Li, L. &amp; Zhou, H. Z." box="[300,492,1048,1075]" pageId="2" pageNumber="475" pagination="679 - 722" refId="ref15762" refString="Li, L. &amp; Zhou, H. Z. (2011) Revision and phylogenetic assessment of the rove beetle genus Pseudohesperus Hayashi, with broad reference to the subtribe Philonthina (Coleoptera: Staphylinidae: Staphylinini). Zoological Journal of the Linnean Society, 163 (3), 679 - 722. https: // doi. org / 10.1111 / j. 1096 - 3642.2011.00731. x" type="journal article" year="2011">Li &amp; Zhou, 2011</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="31FB4D4EE35A2F77FE4EFBC6FC6F054D" authority="Li &amp; Zhou, 2011" authorityName="Li &amp; Zhou" authorityYear="2011" box="[503,904,1048,1075]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="sparsipunctatus">
<emphasis id="C48FEADFE35A2F77FE4EFBC6FD23054C" box="[503,708,1049,1074]" italics="true" pageId="2" pageNumber="475">P. sparsipunctatus</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FD72FBC7FC6F054D" author="Li, L. &amp; Zhou, H. Z." box="[715,904,1048,1075]" pageId="2" pageNumber="475" pagination="679 - 722" refId="ref15762" refString="Li, L. &amp; Zhou, H. Z. (2011) Revision and phylogenetic assessment of the rove beetle genus Pseudohesperus Hayashi, with broad reference to the subtribe Philonthina (Coleoptera: Staphylinidae: Staphylinini). Zoological Journal of the Linnean Society, 163 (3), 679 - 722. https: // doi. org / 10.1111 / j. 1096 - 3642.2011.00731. x" type="journal article" year="2011">Li &amp; Zhou, 2011</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="31FB4D4EE35A2F77FC07FBC6FAF5054D" authority="Li &amp; Zhou, 2011" authorityName="Li &amp; Zhou" authorityYear="2011" box="[958,1298,1048,1075]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="tripartitus">
<emphasis id="C48FEADFE35A2F77FC07FBC6FBAB054C" box="[958,1100,1049,1074]" italics="true" pageId="2" pageNumber="475">P. tripartitus</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FBEBFBC7FAF5054D" author="Li, L. &amp; Zhou, H. Z." box="[1106,1298,1048,1075]" pageId="2" pageNumber="475" pagination="679 - 722" refId="ref15762" refString="Li, L. &amp; Zhou, H. Z. (2011) Revision and phylogenetic assessment of the rove beetle genus Pseudohesperus Hayashi, with broad reference to the subtribe Philonthina (Coleoptera: Staphylinidae: Staphylinini). Zoological Journal of the Linnean Society, 163 (3), 679 - 722. https: // doi. org / 10.1111 / j. 1096 - 3642.2011.00731. x" type="journal article" year="2011">Li &amp; Zhou, 2011</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="F64436CDE35A2F77FF7EFBE3FCED0731" blockId="2.[151,1437,220,1795]" pageId="2" pageNumber="475">
The taxonomy of 
<taxonomicName id="31FB4D4EE35A2F77FE20FBE3FDA80528" authorityName="Hayashi" authorityYear="2008" box="[409,591,1084,1110]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE35A2F77FE20FBE3FDA80528" box="[409,591,1084,1110]" italics="true" pageId="2" pageNumber="475">Pseudohesperus</emphasis>
</taxonomicName>
was maintained as above until 
<bibRefCitation id="926A4B3CE35A2F77FC78FBE2FB960529" author="Newton, A. F. &amp; New Nomenclatural and Taxonomic Acts &amp; Lobl, I. &amp; Lobl, D." box="[961,1137,1084,1111]" pageId="2" pageNumber="475" pagination="1" refId="ref15999" refString="Newton, A. F. (2015) New Nomenclatural and Taxonomic Acts, and Comments. In: Lobl, I. &amp; Lobl, D. (2015) Catalogue of Palaearctic Coleoptera. Vol. 2 / 1. Hydrophiloidea-Staphylinoidea Revised and Updated Edition. Vol. 1. Brill, Leiden and Boston, pp. 12 - 24. https: // doi. org / 10.1163 / 9789004296855 _ 002" type="book chapter" year="2015">Newton (2015)</bibRefCitation>
, following the concept of 
<bibRefCitation id="926A4B3CE35A2F77FF2EFBBFFE850505" author="Hromadka, L." box="[151,354,1120,1147]" pageId="2" pageNumber="475" pagination="495 - 516" refId="ref15506" refString="Hromadka, L. (2010) Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae, 50 (2), 495 - 516." type="journal article" year="2010">Hromádka (2010)</bibRefCitation>
, recognized that 
<taxonomicName id="31FB4D4EE35A2F77FD99FBBFFC990505" authority="Hayashi, 2008" authorityName="Hayashi" authorityYear="2008" box="[544,894,1120,1147]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE35A2F77FD99FBBFFD310504" box="[544,726,1120,1146]" italics="true" pageId="2" pageNumber="475">Pseudohesperus</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FD64FBBFFC990505" author="Hayashi, Y." box="[733,894,1120,1147]" pageId="2" pageNumber="475" pagination="145 - 153" refId="ref15256" refString="Hayashi, Y. (2008) Pseudohesperus, a new genus of Philonthina (Coleoptera: Staphylinidae) from East Asia, with redescription of its type species Philonthus rutiliventris Sharp. The Entomological Review of Japan, 63 (2), 145 - 153." type="journal article" year="2008">Hayashi, 2008</bibRefCitation>
</taxonomicName>
should be as a synonym of 
<taxonomicName id="31FB4D4EE35A2F77FB0FFBBFFECC05E1" authority="Bernhauer, 1912" authorityName="Bernhauer" authorityYear="1912" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE35A2F77FB0FFBBFFA830504" box="[1206,1380,1120,1146]" italics="true" pageId="2" pageNumber="475">Eccoptolonthus</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FAD2FBBEFECC05E1" author="Bernhauer, M." pageId="2" pageNumber="475" pagination="177 - 183" refId="ref14451" refString="Bernhauer, M. (1912) Beitrag zur Staphylinidenfauna von Afrika (Col.). Entomologische Mitteilungen, 1 (6 &amp; 7), 177 - 183 &amp; 203 - 209." type="journal article" year="1912">Bernhauer, 1912</bibRefCitation>
</taxonomicName>
. As a result, all the species included previously in 
<taxonomicName id="31FB4D4EE35A2F77FCE0FB5BFBE805E0" authorityName="Hayashi" authorityYear="2008" box="[857,1039,1156,1182]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE35A2F77FCE0FB5BFBE805E0" box="[857,1039,1156,1182]" italics="true" pageId="2" pageNumber="475">Pseudohesperus</emphasis>
</taxonomicName>
were transferred to 
<taxonomicName id="31FB4D4EE35A2F77FB57FB5BFDE905BD" authority="Bernhauer, 1912" authorityName="Bernhauer" authorityYear="1912" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE35A2F77FB57FB5BFA7B05E0" box="[1262,1436,1156,1182]" italics="true" pageId="2" pageNumber="475">Eccoptolonthus</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FF2EFB77FEB305BD" author="Bernhauer, M." box="[151,340,1192,1219]" pageId="2" pageNumber="475" pagination="177 - 183" refId="ref14451" refString="Bernhauer, M. (1912) Beitrag zur Staphylinidenfauna von Afrika (Col.). Entomologische Mitteilungen, 1 (6 &amp; 7), 177 - 183 &amp; 203 - 209." type="journal article" year="1912">Bernhauer, 1912</bibRefCitation>
(
<bibRefCitation id="926A4B3CE35A2F77FEDCFB76FDE005BD" author="Newton, A. F. &amp; New Nomenclatural and Taxonomic Acts &amp; Lobl, I. &amp; Lobl, D." box="[357,519,1192,1219]" pageId="2" pageNumber="475" pagination="1" refId="ref15999" refString="Newton, A. F. (2015) New Nomenclatural and Taxonomic Acts, and Comments. In: Lobl, I. &amp; Lobl, D. (2015) Catalogue of Palaearctic Coleoptera. Vol. 2 / 1. Hydrophiloidea-Staphylinoidea Revised and Updated Edition. Vol. 1. Brill, Leiden and Boston, pp. 12 - 24. https: // doi. org / 10.1163 / 9789004296855 _ 002" type="book chapter" year="2015">Newton 2015</bibRefCitation>
)
</taxonomicName>
. In the same year, 
<bibRefCitation id="926A4B3CE35A2F77FD54FB77FC3805BD" author="Schillhammer, H. &amp; Staphylinidae, New Synonyms &amp; In &amp; Lobl, I. &amp; Lobl, D." box="[749,991,1192,1219]" pageId="2" pageNumber="475" pagination="1" refId="ref16408" refString="Schillhammer, H. (2015) Staphylinidae, New Synonyms. In: Lobl, I. &amp; Lobl, D. (2015) Catalogue of Palaearctic Coleoptera. Vol. 2 / 1. Hydrophiloidea-Staphylinoidea Revised and Updated Edition. Vol. 1. Brill, Leiden and Boston, pp. 23 - 24." type="book chapter" year="2015">Schillhammer (2015)</bibRefCitation>
confirmed this generic synonymy and transferred 
<taxonomicName id="31FB4D4EE35A2F77FEA1FB13FD680599" authority="Sharp, 1874" authorityName="Sharp" authorityYear="1874" box="[280,655,1228,1255]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="gastralis">
<emphasis id="C48FEADFE35A2F77FEA1FB13FE1B0598" box="[280,508,1228,1254]" italics="true" pageId="2" pageNumber="475">Philonthus gastralis</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FDBDFB13FD680599" author="Sharp, D." box="[516,655,1228,1255]" pageId="2" pageNumber="475" pagination="47 - 48" refId="ref16531" refString="Sharp, D. (1874) The Staphylinidae of Japan. Transactions of the Entomological Society of London, 1, 47 - 48. https: // doi. org / 10.1111 / j. 1365 - 2311.1874. tb 00159. x" type="journal article" year="1874">Sharp, 1874</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="31FB4D4EE35A2F77FD22FB13FB620599" authority="Bernhauer, 1931" authorityName="Bernhauer" authorityYear="1931" box="[667,1157,1228,1255]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="hongkongensis">
<emphasis id="C48FEADFE35A2F77FD22FB13FC250598" box="[667,962,1228,1254]" italics="true" pageId="2" pageNumber="475">Philonthus hongkongensis</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FC72FB13FB620599" author="Bernhauer, M." box="[971,1157,1228,1255]" pageId="2" pageNumber="475" pagination="125 - 132" refId="ref14527" refString="Bernhauer, M. (1931) Zur Staphylinidenfauna des chinesischen Reiches. Wiener Entomologische Zeitung, 48 (3), 125 - 132." type="journal article" year="1931">Bernhauer, 1931</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="31FB4D4EE35A2F77FB07FB13FED20475" authority="Coiffait, 1981" authorityName="Coiffait" authorityYear="1981" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis id="C48FEADFE35A2F77FB07FB13FA7B0598" box="[1214,1436,1228,1254]" italics="true" pageId="2" pageNumber="475">Philonthus lassallei</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FF2EFB2FFED20475" author="Coiffait, H." box="[151,309,1264,1291]" pageId="2" pageNumber="475" pagination="323 - 335" refId="ref14966" refString="Coiffait, H. (1981) Staphylinides nouveaux du Nepal. Nouvelle Revue d'Entomologie, 11 (4), 323 - 335." type="journal article" year="1981">Coiffait, 1981</bibRefCitation>
</taxonomicName>
to 
<taxonomicName id="31FB4D4EE35A2F77FEE2FB2FFDEE0474" authorityName="Bernhauer" authorityYear="1912" box="[347,521,1264,1290]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE35A2F77FEE2FB2FFDEE0474" box="[347,521,1264,1290]" italics="true" pageId="2" pageNumber="475">Eccoptolonthus</emphasis>
</taxonomicName>
. In the same paper, 
<taxonomicName id="31FB4D4EE35A2F77FD48FB2FFB9C0475" authority="Fauvel, 1875" authorityName="Fauvel" authorityYear="1875" box="[753,1147,1264,1291]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="insignitus">
<emphasis id="C48FEADFE35A2F77FD48FB2FFC380474" box="[753,991,1264,1290]" italics="true" pageId="2" pageNumber="475">Philonthus insignitus</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FC51FB2FFB9C0475" author="Fauvel, A." box="[1000,1147,1264,1291]" pageId="2" pageNumber="475" refId="ref15061" refString="Fauvel, A. (1875) Faune Gallo-Rhenane … Catalogue systematique des staphylinides de la Faune Gallo-Rhenane avec l'addition synonymique des especes europeennes, siberiennes, caucasiques et Mediterraneennes et descriptions nouvelles. Vol. 3. Livraison 6. Le Blanc-Hardel, Caen, xxxviii pp." type="book" year="1875">Fauvel, 1875</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="31FB4D4EE35A2F77FB0DFB2FFE620451" authority="Kashcheev, 1999" authorityName="Kashcheev" authorityYear="1999" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="longipalpis">
<emphasis id="C48FEADFE35A2F77FB0DFB2FFF5F0450" italics="true" pageId="2" pageNumber="475">Philonthus longipalpis</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FF7BFACBFE620451" author="Kashcheev, V. A." box="[194,389,1300,1327]" pageId="2" pageNumber="475" pagination="55 - 60" refId="ref15621" refString="Kashcheev, V. A. (1999) Koprobiontnye stafilinidy (Coleoptera, Staphylinidae) iugo-zapadnogo Altaia. Selevinia, 1998 - 1999, 55 - 60." type="journal article" year="1999">Kashcheev, 1999</bibRefCitation>
</taxonomicName>
were synonymized with 
<taxonomicName id="31FB4D4EE35A2F77FD14FACBFA7E0451" authority="(Sharp, 1874) (Schillhammer, 2015)" authorityName="Schillhammer" authorityYear="2015" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[685,1433,1300,1327]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="rutiliventris">
<emphasis id="C48FEADFE35A2F77FD14FACBFC0D0450" box="[685,1002,1300,1326]" italics="true" pageId="2" pageNumber="475">Eccoptolonthus rutiliventris</emphasis>
(
<bibRefCitation id="926A4B3CE35A2F77FC44FACBFB6D0451" author="Sharp, D." box="[1021,1162,1300,1327]" pageId="2" pageNumber="475" pagination="47 - 48" refId="ref16531" refString="Sharp, D. (1874) The Staphylinidae of Japan. Transactions of the Entomological Society of London, 1, 47 - 48. https: // doi. org / 10.1111 / j. 1365 - 2311.1874. tb 00159. x" type="journal article" year="1874">Sharp, 1874</bibRefCitation>
) (
<bibRefCitation id="926A4B3CE35A2F77FB1DFACBFA760451" author="Schillhammer, H. &amp; Staphylinidae, New Synonyms &amp; In &amp; Lobl, I. &amp; Lobl, D." box="[1188,1425,1300,1327]" pageId="2" pageNumber="475" pagination="1" refId="ref16408" refString="Schillhammer, H. (2015) Staphylinidae, New Synonyms. In: Lobl, I. &amp; Lobl, D. (2015) Catalogue of Palaearctic Coleoptera. Vol. 2 / 1. Hydrophiloidea-Staphylinoidea Revised and Updated Edition. Vol. 1. Brill, Leiden and Boston, pp. 23 - 24." type="book chapter" year="2015">Schillhammer, 2015</bibRefCitation>
)
</taxonomicName>
. 
<bibRefCitation id="926A4B3CE35A2F77FF2EFAE7FE87042D" author="Hromadka, L." box="[151,352,1336,1363]" pageId="2" pageNumber="475" pagination="371 - 381" refId="ref15555" refString="Hromadka, L. (2016) Six new Philonthini from the Afrotropical Region (Coleoptera: Staphylinidae: Philonthina). Studies and Reports, Taxonomical Series, 12 (2), 371 - 381." type="journal article" year="2016">Hromádka (2016)</bibRefCitation>
described another species but under 
<taxonomicName id="31FB4D4EE35A2F77FD46FAE7FC52042C" authorityName="Hayashi" authorityYear="2008" box="[767,949,1336,1362]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE35A2F77FD46FAE7FC52042C" box="[767,949,1336,1362]" italics="true" pageId="2" pageNumber="475">Pseudohesperus</emphasis>
</taxonomicName>
: 
<taxonomicName id="31FB4D4EE35A2F77FC7DFAE7FA7E042C" authority="Hromadka, 2016" authorityName="Hromadka" authorityYear="2016" box="[964,1433,1336,1362]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="terezae">
<emphasis id="C48FEADFE35A2F77FC7DFAE7FB35042C" box="[964,1234,1336,1362]" italics="true" pageId="2" pageNumber="475">Pseudohesperus terezae</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FB60FAE7FA7E042C" author="Hromadka, L." box="[1241,1433,1336,1362]" pageId="2" pageNumber="475" pagination="371 - 381" refId="ref15555" refString="Hromadka, L. (2016) Six new Philonthini from the Afrotropical Region (Coleoptera: Staphylinidae: Philonthina). Studies and Reports, Taxonomical Series, 12 (2), 371 - 381." type="journal article" year="2016">Hromádka, 2016</bibRefCitation>
</taxonomicName>
. The correct combination under 
<taxonomicName id="31FB4D4EE35A2F77FDBDFA83FD550408" authorityName="Bernhauer" authorityYear="1912" box="[516,690,1372,1398]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE35A2F77FDBDFA83FD550408" box="[516,690,1372,1398]" italics="true" pageId="2" pageNumber="475">Eccoptolonthus</emphasis>
</taxonomicName>
was given by 
<bibRefCitation id="926A4B3CE35A2F77FCD9FA83FB8E0409" author="Mikatova, S. &amp; Machackova, L. &amp; Hajek, J." box="[864,1129,1372,1399]" pageId="2" pageNumber="475" pagination="353 - 389" refId="ref15872" refString="Mikatova, S., Machackova, L. &amp; Hajek, J. (2020) Catalogue of type specimens of beetles (Coleoptera) deposited in the National Museum, Prague, Czech Republic). Acta Entomolgica Muse Nationalis Pragae, 60 (1), 353 - 389. https: // doi. org / 10.37520 / aemnp. 2020.021" type="journal article" year="2020">
Mikátová 
<emphasis id="C48FEADFE35A2F77FC6FFA82FBED0408" box="[982,1034,1372,1398]" italics="true" pageId="2" pageNumber="475">et al</emphasis>
. (2020)
</bibRefCitation>
. 
<bibRefCitation id="926A4B3CE35A2F77FBCFFA83FA7B0409" author="Chani-Posse, M. R. &amp; Brunke, A. J. &amp; Chatzimanolis, S. &amp; Schillhammer, H. &amp; Solodovnikov, A." box="[1142,1436,1372,1399]" pageId="2" pageNumber="475" pagination="1 - 40" refId="ref14893" refString="Chani-Posse, M. R., Brunke, A. J., Chatzimanolis, S., Schillhammer, H. &amp; Solodovnikov, A. (2018) Phylogeny of the hyper-diverse rove beetle subtribe Philonthina with implications for classification of the tribe Staphylinini (Coleoptera: Staphylinidae). Cladistics, 34, 1 - 40. https: // doi. org / 10.1111 / cla. 12188" type="journal article" year="2018">
Chani-Posse 
<emphasis id="C48FEADFE35A2F77FAB4FA82FAA60408" box="[1293,1345,1372,1398]" italics="true" pageId="2" pageNumber="475">et al</emphasis>
. (2018)
</bibRefCitation>
reported the results of phylogenetic analyses, which supported the transfer of two 
<taxonomicName id="31FB4D4EE35A2F77FB83FA5EFB4304E4" authorityName="Fauvel" authorityYear="1874" box="[1082,1188,1409,1434]" class="Insecta" family="Staphylinidae" genus="Hesperus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE35A2F77FB83FA5EFB4304E4" box="[1082,1188,1409,1434]" italics="true" pageId="2" pageNumber="475">Hesperus</emphasis>
</taxonomicName>
species to this genus: 
<taxonomicName id="31FB4D4EE35A2F77FF2EFA7BFD8C04C1" authority="(Fauvel, 1895)" authorityName="" baseAuthorityName="Fauvel" baseAuthorityYear="1895" box="[151,619,1444,1471]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="laevigatus">
<emphasis id="C48FEADFE35A2F77FF2EFA7BFE5804C0" box="[151,447,1444,1470]" italics="true" pageId="2" pageNumber="475">Eccoptolonthus laevigatus</emphasis>
(
<bibRefCitation id="926A4B3CE35A2F77FE77FA7BFD8404C1" author="Fauvel, A." box="[462,611,1444,1471]" pageId="2" pageNumber="475" pagination="180 - 286" refId="ref15110" refString="Fauvel, A. (1895) Staphylinides nouveaux de l'Inde et de la Malaisie. Revue d'Entomologie, 14, 180 - 286. https: // doi. org / 10.1017 / S 102688120005892 X" type="journal article" year="1895">Fauvel, 1895</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="31FB4D4EE35A2F77FD1BFA7BFB9804C1" authority="(Bernhauer, 1911)" baseAuthorityName="Bernhauer" baseAuthorityYear="1911" box="[674,1151,1444,1471]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="roepkei">
<emphasis id="C48FEADFE35A2F77FD1BFA7BFC4D04C0" box="[674,938,1444,1470]" italics="true" pageId="2" pageNumber="475">Eccoptolonthus roepkei</emphasis>
(
<bibRefCitation id="926A4B3CE35A2F77FC00FA7BFB9004C1" author="Bernhauer, M." box="[953,1143,1444,1471]" pageId="2" pageNumber="475" pagination="55 - 62" refId="ref14424" refString="Bernhauer, M. (1911) Zur Staphylinidenfauna Ostindiens und der Sundainseln. Entomologische Blatter, 7, 55 - 62 + 86 - 93." type="journal article" year="1911">Bernhauer, 1911</bibRefCitation>
)
</taxonomicName>
. 
<bibRefCitation id="926A4B3CE35A2F77FB33FA7BFADF04C1" author="Hayashi, Y." box="[1162,1336,1444,1471]" pageId="2" pageNumber="475" pagination="135 - 142" refId="ref15304" refString="Hayashi, Y. (2020) A New Subgenus of the Genus Eccoptolonthus Bernhauer, with Descriptions of a New Species and a New Subspecies (Coleoptera, Staphylinidae, Philonthina). Elytra, Tokyo, New Series, 10 (1), 135 - 142." type="journal article" year="2020">Hayashi (2020)</bibRefCitation>
reported the newest findings and erected the subgenus 
<taxonomicName id="31FB4D4EE35A2F77FD23FA16FC0F049D" authority="Hayashi, 2020" authorityName="Hayashi" authorityYear="2020" box="[666,1000,1480,1507]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="subGenus" subGenus="Eumorimotous">
<emphasis id="C48FEADFE35A2F77FD23FA16FCD8049C" box="[666,831,1481,1506]" italics="true" pageId="2" pageNumber="475">Eumorimotous</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FCFFFA17FC0F049D" author="Hayashi, Y." box="[838,1000,1480,1507]" pageId="2" pageNumber="475" pagination="135 - 142" refId="ref15304" refString="Hayashi, Y. (2020) A New Subgenus of the Genus Eccoptolonthus Bernhauer, with Descriptions of a New Species and a New Subspecies (Coleoptera, Staphylinidae, Philonthina). Elytra, Tokyo, New Series, 10 (1), 135 - 142." type="journal article" year="2020">
Hayashi, 
<quantity id="31039B28E35A2F77FC09FA16FC0F049D" box="[944,1000,1481,1507]" metricMagnitude="1" metricUnit="m" metricValue="5.1308" pageId="2" pageNumber="475" unit="in" value="2020.0">2020</quantity>
</bibRefCitation>
</taxonomicName>
in the genus 
<taxonomicName id="31FB4D4EE35A2F77FBC7FA17FACB049C" authorityName="Bernhauer" authorityYear="1912" box="[1150,1324,1480,1506]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE35A2F77FBC7FA17FACB049C" box="[1150,1324,1480,1506]" italics="true" pageId="2" pageNumber="475">Eccoptolonthus</emphasis>
</taxonomicName>
, with one species and one subspecies described: 
<taxonomicName id="31FB4D4EE35A2F77FDF0FA33FB3B0779" authority="Hayashi, 2020" authorityName="Hayashi" authorityYear="2020" box="[585,1244,1516,1543]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="morimotoi" subGenus="Eumorimotous">
<emphasis id="C48FEADFE35A2F77FDF0FA33FD100778" box="[585,759,1516,1542]" italics="true" pageId="2" pageNumber="475">Eccoptolonthus</emphasis>
(
<emphasis id="C48FEADFE35A2F77FCB1FA32FC4A0778" box="[776,941,1517,1542]" italics="true" pageId="2" pageNumber="475">Eumorimotous</emphasis>
) 
<emphasis id="C48FEADFE35A2F77FC07FA32FBD40778" box="[958,1075,1517,1542]" italics="true" pageId="2" pageNumber="475">morimotoi</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FB83FA33FB3B0779" author="Hayashi, Y." box="[1082,1244,1516,1543]" pageId="2" pageNumber="475" pagination="135 - 142" refId="ref15304" refString="Hayashi, Y. (2020) A New Subgenus of the Genus Eccoptolonthus Bernhauer, with Descriptions of a New Species and a New Subspecies (Coleoptera, Staphylinidae, Philonthina). Elytra, Tokyo, New Series, 10 (1), 135 - 142." type="journal article" year="2020">Hayashi, 2020</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="31FB4D4EE35A2F77FAAAFA33FCA50755" authority="Hayashi, 2020" authorityName="Hayashi" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="subSpecies" species="laevigatus" subGenus="Eumorimotous" subSpecies="tiomanensis">
<emphasis id="C48FEADFE35A2F77FAAAFA33FF210754" italics="true" pageId="2" pageNumber="475">Eccoptolonthus</emphasis>
(
<emphasis id="C48FEADFE35A2F77FF61F9CEFE9A0754" box="[216,381,1553,1578]" italics="true" pageId="2" pageNumber="475">Eumorimotous</emphasis>
) 
<emphasis id="C48FEADFE35A2F77FE36F9CFFD760754" box="[399,657,1552,1578]" italics="true" pageId="2" pageNumber="475">laevigatus tiomanensis</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FD23F9CFFCA50755" author="Hayashi, Y." box="[666,834,1552,1579]" pageId="2" pageNumber="475" pagination="135 - 142" refId="ref15304" refString="Hayashi, Y. (2020) A New Subgenus of the Genus Eccoptolonthus Bernhauer, with Descriptions of a New Species and a New Subspecies (Coleoptera, Staphylinidae, Philonthina). Elytra, Tokyo, New Series, 10 (1), 135 - 142." type="journal article" year="2020">Hayashi, 2020</bibRefCitation>
</taxonomicName>
. 
<taxonomicName id="31FB4D4EE35A2F77FCF7F9CFFF3F0731" authority="(Fauvel, 1895)" baseAuthorityName="Fauvel" baseAuthorityYear="1895" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="wasmanni" subGenus="Eumorimotous">
<emphasis id="C48FEADFE35A2F77FCF7F9CFFC1B0754" box="[846,1020,1552,1578]" italics="true" pageId="2" pageNumber="475">Eccoptolonthus</emphasis>
(
<emphasis id="C48FEADFE35A2F77FBB7F9CEFB540754" box="[1038,1203,1553,1578]" italics="true" pageId="2" pageNumber="475">Eumorimotous</emphasis>
) 
<emphasis id="C48FEADFE35A2F77FB7CF9CEFAD00754" box="[1221,1335,1553,1578]" italics="true" pageId="2" pageNumber="475">wasmanni</emphasis>
(
<bibRefCitation id="926A4B3CE35A2F77FAF1F9CFFF370731" author="Fauvel, A." pageId="2" pageNumber="475" pagination="180 - 286" refId="ref15110" refString="Fauvel, A. (1895) Staphylinides nouveaux de l'Inde et de la Malaisie. Revue d'Entomologie, 14, 180 - 286. https: // doi. org / 10.1017 / S 102688120005892 X" type="journal article" year="1895">Fauvel, 1895</bibRefCitation>
)
</taxonomicName>
was also transferred from 
<taxonomicName id="31FB4D4EE35A2F77FDBDF9EAFD890730" authorityName="Fauvel" authorityYear="1874" box="[516,622,1589,1614]" class="Insecta" family="Staphylinidae" genus="Hesperus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE35A2F77FDBDF9EAFD890730" box="[516,622,1589,1614]" italics="true" pageId="2" pageNumber="475">Hesperus</emphasis>
</taxonomicName>
to this genus.
</paragraph>
<paragraph id="F64436CDE35A2F77FF7EF986FC3A067C" blockId="2.[151,1437,220,1795]" pageId="2" pageNumber="475">
In conclusion, there were 24 species and one subspecies included in the genus 
<taxonomicName id="31FB4D4EE35A2F77FB96F987FA7B070D" authority="Bernhauer, 1912" authorityName="Bernhauer" authorityYear="1912" box="[1071,1436,1624,1651]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE35A2F77FB96F987FB3A070C" box="[1071,1245,1624,1650]" italics="true" pageId="2" pageNumber="475">Eccoptolonthus</emphasis>
<bibRefCitation id="926A4B3CE35A2F77FB5AF987FA7B070D" author="Bernhauer, M." box="[1251,1436,1624,1651]" pageId="2" pageNumber="475" pagination="177 - 183" refId="ref14451" refString="Bernhauer, M. (1912) Beitrag zur Staphylinidenfauna von Afrika (Col.). Entomologische Mitteilungen, 1 (6 &amp; 7), 177 - 183 &amp; 203 - 209." type="journal article" year="1912">Bernhauer, 1912</bibRefCitation>
</taxonomicName>
before this study (
<bibRefCitation id="926A4B3CE35A2F77FEE4F9A3FDFD07E9" author="Bernhauer, M." box="[349,538,1660,1687]" pageId="2" pageNumber="475" pagination="177 - 183" refId="ref14451" refString="Bernhauer, M. (1912) Beitrag zur Staphylinidenfauna von Afrika (Col.). Entomologische Mitteilungen, 1 (6 &amp; 7), 177 - 183 &amp; 203 - 209." type="journal article" year="1912">Bernhauer, 1912</bibRefCitation>
; 
<bibRefCitation id="926A4B3CE35A2F77FD9CF9A3FC8107E9" author="Bernhauer, M. &amp; Schubert, K." box="[549,870,1660,1687]" pageId="2" pageNumber="475" pagination="289 - 408" refId="ref14553" refString="Bernhauer, M. &amp; Schubert, K. (1914) Staphylinidae IV. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus. Vol. 5. Pars 57. Junk, Berlin, pp. 289 - 408." type="book chapter" year="1914">Bernhauer &amp; Schubert, 1914</bibRefCitation>
; 
<bibRefCitation id="926A4B3CE35A2F77FCC8F9A2FBF807E9" author="Cameron, M." box="[881,1055,1661,1687]" pageId="2" pageNumber="475" pagination="1 - 443" refId="ref14719" refString="Cameron, M. (1932) The Fauna of British India including Ceylon and Burma. Coleoptera. Staphylinidae. Vol. III. Taylor &amp; Francis London, 1 - 443." type="book chapter" year="1932">Cameron, 1932</bibRefCitation>
; 
<bibRefCitation id="926A4B3CE35A2F77FB93F9A3FB1907E9" author="Blackwelder, W." box="[1066,1278,1660,1687]" pageId="2" pageNumber="475" pagination="1 - 658" refId="ref14604" refString="Blackwelder, W. (1943) Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, Washington, 182, 1 - 658. https: // doi. org / 10.5479 / si. 03629236.182. i" type="journal article" year="1943">Blackwelder, 1943</bibRefCitation>
; 
<bibRefCitation id="926A4B3CE35A2F77FAB0F9A3FF3507C5" author="Blackwelder, R. E." pageId="2" pageNumber="475" pagination="1 - 483" refId="ref14652" refString="Blackwelder, R. E. (1952) The generic names of the beetle family Staphylinidae, with an essay on genotypy. United States National Museum Bulletin, 200, 1 - 483." type="journal article" year="1952">Blackwelder, 1952</bibRefCitation>
; 
<bibRefCitation id="926A4B3CE35A2F77FF67F97FFE7807C4" author="Tottenham, C. E." box="[222,415,1696,1722]" pageId="2" pageNumber="475" pagination="230 - 231" refId="ref16769" refString="Tottenham, C. E. (1962) Coleoptera Staphylinidae Staphylininae. Mission zoologique de l'I. R. S. A. C. en Afrique orientale (P. Basilewsky et N. Leleup, 1957). LXXVI. Annales du Musee Royal de l'Afrique Centrale, Tervuren, Serie 8 º: Sciences Zoologiques, 110, 230 - 231." type="journal article" year="1962">Tottenham, 1962</bibRefCitation>
; 
<bibRefCitation id="926A4B3CE35A2F77FE12F97EFDBB07C5" author="Herman, L. H." box="[427,604,1697,1723]" pageId="2" pageNumber="475" pagination="2819 - 2949" refId="ref15354" refString="Herman, L. H. (2001 a) Catalog of the Staphylinidae (Insecta: Coleoptera): 1758 to the end of the second millennium. V. Staphylinine group (Part 2). Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina). Bulletin of the American Museum of Natural History, 265, 2819 - 2949. https: // doi. org / 10.1206 / 0003 - 0090.265.1.5" type="journal article" year="2001">Herman, 2001a</bibRefCitation>
; 
<bibRefCitation id="926A4B3CE35A2F77FDD1F97FFCEA07C5" author="Hayashi, Y." box="[616,781,1696,1723]" pageId="2" pageNumber="475" pagination="145 - 153" refId="ref15256" refString="Hayashi, Y. (2008) Pseudohesperus, a new genus of Philonthina (Coleoptera: Staphylinidae) from East Asia, with redescription of its type species Philonthus rutiliventris Sharp. The Entomological Review of Japan, 63 (2), 145 - 153." type="journal article" year="2008">Hayashi, 2008</bibRefCitation>
; 
<bibRefCitation id="926A4B3CE35A2F77FCA0F97FFC3D07C5" author="Hromadka, L." box="[793,986,1696,1723]" pageId="2" pageNumber="475" pagination="495 - 516" refId="ref15506" refString="Hromadka, L. (2010) Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae, 50 (2), 495 - 516." type="journal article" year="2010">Hromádka, 2010</bibRefCitation>
; 
<bibRefCitation id="926A4B3CE35A2F77FC5CF97FFB4007C5" author="Li, L. &amp; Zhou, H. Z." box="[997,1191,1696,1723]" pageId="2" pageNumber="475" pagination="679 - 722" refId="ref15762" refString="Li, L. &amp; Zhou, H. Z. (2011) Revision and phylogenetic assessment of the rove beetle genus Pseudohesperus Hayashi, with broad reference to the subtribe Philonthina (Coleoptera: Staphylinidae: Staphylinini). Zoological Journal of the Linnean Society, 163 (3), 679 - 722. https: // doi. org / 10.1111 / j. 1096 - 3642.2011.00731. x" type="journal article" year="2011">Li &amp; Zhou, 2011</bibRefCitation>
; 
<bibRefCitation id="926A4B3CE35A2F77FB0AF97FFA7007C5" author="Schillhammer, H." box="[1203,1431,1696,1723]" pageId="2" pageNumber="475" pagination="133 - 163" refId="ref16374" refString="Schillhammer, H. (2011) Old and new Staphylinini from the Palearctic and Oriental Regions (Coleoptera: Staphylinidae: Staphylininae). Koleopterologische Rundschau, 81, 133 - 163." type="journal article" year="2011">Schillhammer, 2011</bibRefCitation>
; 
<bibRefCitation id="926A4B3CE35A2F77FF2EF91AFEDB07A1" author="Newton, A. F. &amp; New Nomenclatural and Taxonomic Acts &amp; Lobl, I. &amp; Lobl, D." box="[151,316,1732,1759]" pageId="2" pageNumber="475" pagination="1" refId="ref15999" refString="Newton, A. F. (2015) New Nomenclatural and Taxonomic Acts, and Comments. In: Lobl, I. &amp; Lobl, D. (2015) Catalogue of Palaearctic Coleoptera. Vol. 2 / 1. Hydrophiloidea-Staphylinoidea Revised and Updated Edition. Vol. 1. Brill, Leiden and Boston, pp. 12 - 24. https: // doi. org / 10.1163 / 9789004296855 _ 002" type="book chapter" year="2015">Newton, 2015</bibRefCitation>
; 
<bibRefCitation id="926A4B3CE35A2F77FEF2F91BFDD307A1" author="Schillhammer, H. &amp; Staphylinidae, New Synonyms &amp; In &amp; Lobl, I. &amp; Lobl, D." box="[331,564,1732,1759]" pageId="2" pageNumber="475" pagination="1" refId="ref16408" refString="Schillhammer, H. (2015) Staphylinidae, New Synonyms. In: Lobl, I. &amp; Lobl, D. (2015) Catalogue of Palaearctic Coleoptera. Vol. 2 / 1. Hydrophiloidea-Staphylinoidea Revised and Updated Edition. Vol. 1. Brill, Leiden and Boston, pp. 23 - 24." type="book chapter" year="2015">Schillhammer, 2015</bibRefCitation>
; 
<bibRefCitation id="926A4B3CE35A2F77FDFAF91BFCE007A0" author="Hromadka, L." box="[579,775,1732,1758]" pageId="2" pageNumber="475" pagination="371 - 381" refId="ref15555" refString="Hromadka, L. (2016) Six new Philonthini from the Afrotropical Region (Coleoptera: Staphylinidae: Philonthina). Studies and Reports, Taxonomical Series, 12 (2), 371 - 381." type="journal article" year="2016">Hromádka, 2016</bibRefCitation>
; 
<bibRefCitation id="926A4B3CE35A2F77FCACF91BFC0A07A1" author="Chani-Posse, M. R. &amp; Brunke, A. J. &amp; Chatzimanolis, S. &amp; Schillhammer, H. &amp; Solodovnikov, A." box="[789,1005,1732,1759]" pageId="2" pageNumber="475" pagination="1 - 40" refId="ref14893" refString="Chani-Posse, M. R., Brunke, A. J., Chatzimanolis, S., Schillhammer, H. &amp; Solodovnikov, A. (2018) Phylogeny of the hyper-diverse rove beetle subtribe Philonthina with implications for classification of the tribe Staphylinini (Coleoptera: Staphylinidae). Cladistics, 34, 1 - 40. https: // doi. org / 10.1111 / cla. 12188" type="journal article" year="2018">Chani-Posse, 2018</bibRefCitation>
; 
<bibRefCitation id="926A4B3CE35A2F77FC45F91AFB4507A1" author="Newton, A. F." box="[1020,1186,1733,1759]" pageId="2" pageNumber="475" pagination="2000" refId="ref16076" refString="Newton, A. F. (2019) StaphBase: Staphyliniformia World Catalog Database. Version Nov 2018. Online. In: Species 2000 &amp; ITIS Catalogue of Life, 2019 Annual Checklist. Available from: https: // www. catalogueoflife. org / col / details / database / id / 204 (accessed 15 April 2020)." type="journal article" year="2019">Newton, 2019</bibRefCitation>
; 
<bibRefCitation id="926A4B3CE35A2F77FB08F91BFABC07A1" author="Hayashi, Y." box="[1201,1371,1732,1759]" pageId="2" pageNumber="475" pagination="135 - 142" refId="ref15304" refString="Hayashi, Y. (2020) A New Subgenus of the Genus Eccoptolonthus Bernhauer, with Descriptions of a New Species and a New Subspecies (Coleoptera, Staphylinidae, Philonthina). Elytra, Tokyo, New Series, 10 (1), 135 - 142." type="journal article" year="2020">Hayashi, 2020</bibRefCitation>
). Our study contributed new findings after revising the above-mentioned studies.
</paragraph>
</subSubSection>
<subSection id="9F742F26E35A2F76FF2EF88AFA7B05EC" lastPageId="3" lastPageNumber="476" pageId="2" pageNumber="475" type="key">
<subSubSection id="BEE16546E35A2F76FF2EF88AFA7B05EC" lastPageId="3" lastPageNumber="476" pageId="2" pageNumber="475" type="key">
<paragraph id="F64436CDE35A2F77FF2EF88AFCA50611" blockId="2.[151,834,1876,1903]" box="[151,834,1876,1903]" pageId="2" pageNumber="475">
<heading id="AD0C81A1E35A2F77FF2EF88AFCA50611" bold="true" box="[151,834,1876,1903]" fontSize="11" level="1" pageId="2" pageNumber="475" reason="1">
<emphasis id="C48FEADFE35A2F77FF2EF88AFCA50611" bold="true" box="[151,834,1876,1903]" pageId="2" pageNumber="475">
Key to the Chinese species of the genus 
<taxonomicName id="31FB4D4EE35A2F77FD3CF88BFCA50611" ID-CoL="8X7C9" authorityName="Bernhauer" authorityYear="1912" box="[645,834,1876,1903]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE35A2F77FD3CF88BFCA50611" bold="true" box="[645,834,1876,1903]" italics="true" pageId="2" pageNumber="475">Eccoptolonthus</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="F64436CDE35A2F77FF2EF846FA7B06D0" blockId="2.[151,1436,1945,1997]" box="[151,1436,1945,1967]" pageId="2" pageNumber="475">1. Male sternite VIII with a pair of long appendages (Figs. 7D, 9H, 11D)........................................... 2</paragraph>
<paragraph id="F64436CDE35A2F77FF2EF868FA7B06B2" blockId="2.[151,1436,1945,1997]" box="[151,1436,1974,1997]" pageId="2" pageNumber="475">- Male sternite VIII without a pair of long appendages (Figs. 1D, 3E, 5D).......................................... 8</paragraph>
<paragraph id="F64436CDE35A2F77FF2EF80BFA7B0697" blockId="2.[151,1436,2004,2026]" box="[151,1436,2004,2026]" pageId="2" pageNumber="475">2. Head with distinct microsculpture........................................................................ 3</paragraph>
<paragraph id="F64436CDE35B2F76FF2EFF47FA7B01D3" blockId="3.[151,1437,151,1171]" box="[151,1436,151,173]" pageId="3" pageNumber="476">- Head without microsculpture............................................................................ 5</paragraph>
<paragraph id="F64436CDE35B2F76FF2EFF6AFA7B0199" blockId="3.[151,1437,151,1171]" pageId="3" pageNumber="476">
3. Paramere weakly constricted at midlength in ventral view: paramere at narrowest part more than half the corresponding width of median lobe (
<figureCitation id="6EC02A48E35B2F76FEC6FF0DFE330196" box="[383,468,210,232]" captionStart="FIGURE 11" captionStartId="19.[151,250,1900,1924]" captionTargetBox="[169,1422,827,1863]" captionTargetId="figure-267@19.[160,1427,813,1876]" captionTargetPageId="19" captionText="FIGURE 11. Eccoptolonthus xiaolongmenensis Fei &amp; Zhou, sp. nov. A. Habitus; B. aedeagus, dorsal (parameral side) view; C. aedeagus, lateral view; D. sternite IX of male; E. stylus of tergite IX of male; F. sternite VIII of male; G. tergite X of male. Scale bars, A= 1 mm; B, C, D, E, F, G= 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4640505" httpUri="https://zenodo.org/record/4640505/files/figure.png" pageId="3" pageNumber="476">Fig. 11B</figureCitation>
)............................................................................... 4
</paragraph>
<paragraph id="F64436CDE35B2F76FF2EFF2FFA7B005C" blockId="3.[151,1437,151,1171]" pageId="3" pageNumber="476">
- Paramere strongly constricted at midlength in ventral view: paramere at narrowest part less than half the corresponding width of median lobe (
<figureCitation id="6EC02A48E35B2F76FEC6FED2FE2F005C" box="[383,456,269,291]" captionStart="FIGURE 7" captionStartId="12.[151,250,1936,1960]" captionTargetBox="[205,1376,707,1907]" captionTargetId="figure-204@12.[199,1388,705,1912]" captionTargetPageId="12" captionText="FIGURE 7. Eccoptolonthus fuyuensis Fei &amp; Zhou, sp. nov. A. Habitus; B. aedeagus, dorsal (parameral side) view; C. aedeagus, lateral view; D. sternite IX of male; E. tergite X of male; F. sternite VIII of male; g. stylus of tergite IX of male. Scale bars, A, B, C, D, E, F, G, H= 0.2 mm;" figureDoi="http://doi.org/10.5281/zenodo.4640491" httpUri="https://zenodo.org/record/4640491/files/figure.png" pageId="3" pageNumber="476">Fig. 7B</figureCitation>
)..................................................... 
<taxonomicName id="31FB4D4EE35B2F76FBE3FED2FAA1005D" ID-CoL="8XZXB" authority="Fei &amp; Zhou" authorityName="Fei &amp; Zhou" authorityYear="2021" box="[1114,1350,269,291]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="476" phylum="Arthropoda" rank="species" species="fuyuensis" status="sp. nov.">
<emphasis id="C48FEADFE35B2F76FBE3FED2FB2A005C" box="[1114,1229,269,290]" italics="true" pageId="3" pageNumber="476">E. fuyuensis</emphasis>
Fei &amp; Zhou
</taxonomicName>
, 
<emphasis id="C48FEADFE35B2F76FAE9FED2FA7B005C" bold="true" box="[1360,1436,269,290]" pageId="3" pageNumber="476">
<taxonomicNameLabel id="DFBC57A4E35B2F76FAE9FED2FA7B005C" box="[1360,1436,269,290]" pageId="3" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="F64436CDE35B2F76FF2EFEF5FA7B003E" blockId="3.[151,1437,151,1171]" box="[151,1436,298,320]" pageId="3" pageNumber="476">
4. Male tergite X with arcuate medioapical emargination.................................. 
<taxonomicName id="31FB4D4EE35B2F76FB3FFEF4FA7B003E" ID-CoL="8XKW8" authority="(Sharp, 1874)" authorityName="Schillhammer" authorityYear="2015" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[1158,1436,298,320]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="476" phylum="Arthropoda" rank="species" species="rutiliventris">
<emphasis id="C48FEADFE35B2F76FB3FFEF4FAF50041" box="[1158,1298,298,320]" italics="true" pageId="3" pageNumber="476">E. rutiliventris</emphasis>
(
<bibRefCitation id="926A4B3CE35B2F76FAA6FEF5FA72003E" author="Sharp, D." box="[1311,1429,298,320]" pageId="3" pageNumber="476" pagination="47 - 48" refId="ref16531" refString="Sharp, D. (1874) The Staphylinidae of Japan. Transactions of the Entomological Society of London, 1, 47 - 48. https: // doi. org / 10.1111 / j. 1365 - 2311.1874. tb 00159. x" type="journal article" year="1874">Sharp, 1874</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="F64436CDE35B2F76FF2EFE97FA7B0023" blockId="3.[151,1437,151,1171]" box="[151,1436,327,349]" pageId="3" pageNumber="476">
- Male tergite X with acute triangular medioapical emargination................ 
<taxonomicName id="31FB4D4EE35B2F76FBBEFE97FAA10023" ID-CoL="8XZXC" authority="Fei &amp; Zhou" authorityName="Fei &amp; Zhou" authorityYear="2021" box="[1031,1350,327,349]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="476" phylum="Arthropoda" rank="species" species="xiaolongmenensis" status="sp. nov.">
<emphasis id="C48FEADFE35B2F76FBBEFE97FB290022" box="[1031,1230,327,349]" italics="true" pageId="3" pageNumber="476">E. xiaolongmenensis</emphasis>
Fei &amp; Zhou
</taxonomicName>
, 
<emphasis id="C48FEADFE35B2F76FAE9FE97FA7B0023" bold="true" box="[1360,1436,328,349]" pageId="3" pageNumber="476">
<taxonomicNameLabel id="DFBC57A4E35B2F76FAE9FE97FA7B0023" box="[1360,1436,328,349]" pageId="3" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="F64436CDE35B2F76FF2EFEBAFA7B00CB" blockId="3.[151,1437,151,1171]" pageId="3" pageNumber="476">
5. Antennae black, with apical three or four antennomeres yellowish white; abdomen black with reddish yellow posterior margins in 3rd, 4th, 7th and 8th ventrites; 5th and 6th brownish red; inner face of paramere sparingly scattered with about 16 peg-setae along margins in apical third................................ 
<taxonomicName id="31FB4D4EE35B2F76FC39FE7FFA7B00CB" ID-CoL="8YDHD" authority="laevigatus (Fauvel, 1895)" authorityName="laevigatus (Fauvel" authorityYear="1895" baseAuthorityName="Fauvel" baseAuthorityYear="1895" box="[896,1436,415,437]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="476" phylum="Arthropoda" rank="subSpecies" species="laevigatus" subGenus="Eumorimotous" subSpecies="laevigatus">
<emphasis id="C48FEADFE35B2F76FC39FE7FFC7200CB" box="[896,917,416,437]" italics="true" pageId="3" pageNumber="476">E.</emphasis>
(
<emphasis id="C48FEADFE35B2F76FC1BFE7FFBC800CB" box="[930,1071,416,437]" italics="true" pageId="3" pageNumber="476">Eumorimotous</emphasis>
) 
<emphasis id="C48FEADFE35B2F76FB87FE40FAEE00CA" box="[1086,1289,415,436]" italics="true" pageId="3" pageNumber="476">laevigatus laevigatus</emphasis>
(
<bibRefCitation id="926A4B3CE35B2F76FAAFFE40FA7200CB" author="Fauvel, A." box="[1302,1429,415,437]" pageId="3" pageNumber="476" pagination="180 - 286" refId="ref15110" refString="Fauvel, A. (1895) Staphylinides nouveaux de l'Inde et de la Malaisie. Revue d'Entomologie, 14, 180 - 286. https: // doi. org / 10.1017 / S 102688120005892 X" type="journal article" year="1895">Fauvel, 1895</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="F64436CDE35B2F76FF2EFE62FA7B008E" blockId="3.[151,1437,151,1171]" pageId="3" pageNumber="476">- Antennae black, with antennomeres I–III to various extent reddish-brown, antennomere XI or antennomeres X–XI dark brown; abdomen black; paramere with more than 16 peg-setae....................................................... 6</paragraph>
<paragraph id="F64436CDE35B2F76FF2EFE28FA7A0355" blockId="3.[151,1437,151,1171]" pageId="3" pageNumber="476">
6. Eyes very large, 1.75–2.33 times as long as tempora; pronotum with sparse punctures; paramere much wider and underside of paramere with larger number of peg setae..................................... 
<taxonomicName id="31FB4D4EE35B2F76FB8BFDCAFA7A0355" authority="(Li &amp; Zhou, 2011)" baseAuthorityName="Li &amp; Zhou" baseAuthorityYear="2011" box="[1074,1437,533,555]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="476" phylum="Arthropoda" rank="species" species="sparsipunctatus">
<emphasis id="C48FEADFE35B2F76FB8BFDCAFB030354" box="[1074,1252,533,554]" italics="true" pageId="3" pageNumber="476">E. sparsipunctatus</emphasis>
(
<bibRefCitation id="926A4B3CE35B2F76FB48FDCAFA720355" author="Li, L. &amp; Zhou, H. Z." box="[1265,1429,533,555]" pageId="3" pageNumber="476" pagination="679 - 722" refId="ref15762" refString="Li, L. &amp; Zhou, H. Z. (2011) Revision and phylogenetic assessment of the rove beetle genus Pseudohesperus Hayashi, with broad reference to the subtribe Philonthina (Coleoptera: Staphylinidae: Staphylinini). Zoological Journal of the Linnean Society, 163 (3), 679 - 722. https: // doi. org / 10.1111 / j. 1096 - 3642.2011.00731. x" type="journal article" year="2011">Li &amp; Zhou, 2011</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="F64436CDE35B2F76FF2EFDECFA7B0336" blockId="3.[151,1437,151,1171]" box="[151,1436,562,584]" pageId="3" pageNumber="476">- Eyes flat, 1.3 times as long as tempora; pronotum with dense punctures.......................................... 7</paragraph>
<paragraph id="F64436CDE35B2F76FF2EFD8FFA7B03FD" blockId="3.[151,1437,151,1171]" pageId="3" pageNumber="476">
7. Pronotum longer than broad; apex of abdomen dark.................................................................................................................................. 
<taxonomicName id="31FB4D4EE35B2F76FB87FDB2FA7B03FD" authority="(Bernhauer, 1931)" baseAuthorityName="Bernhauer" baseAuthorityYear="1931" box="[1086,1436,621,643]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="476" phylum="Arthropoda" rank="species" species="hongkongensis">
<emphasis id="C48FEADFE35B2F76FB87FDB2FB0F03FC" box="[1086,1256,621,642]" italics="true" pageId="3" pageNumber="476">E. hongkongensis</emphasis>
(
<bibRefCitation id="926A4B3CE35B2F76FB4DFDB2FA7203FD" author="Bernhauer, M." box="[1268,1429,621,643]" pageId="3" pageNumber="476" pagination="125 - 132" refId="ref14527" refString="Bernhauer, M. (1931) Zur Staphylinidenfauna des chinesischen Reiches. Wiener Entomologische Zeitung, 48 (3), 125 - 132." type="journal article" year="1931">Bernhauer, 1931</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="F64436CDE35B2F76FF2EFD54FA7B03DE" blockId="3.[151,1437,151,1171]" box="[151,1436,650,672]" pageId="3" pageNumber="476">
- Pronotum about as long as broad; apex of abdomen reddish-yellow.......................... 
<taxonomicName id="31FB4D4EE35B2F76FB23FD54FA7B03DE" authority="(Kraatz, 1859)" baseAuthorityName="Kraatz" baseAuthorityYear="1859" box="[1178,1436,650,672]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="476" phylum="Arthropoda" rank="species" species="eustilbus">
<emphasis id="C48FEADFE35B2F76FB23FD54FAED03E1" box="[1178,1290,650,672]" italics="true" pageId="3" pageNumber="476">E. eustilbus</emphasis>
(
<bibRefCitation id="926A4B3CE35B2F76FAAEFD55FA7203DE" author="Kraatz, G." box="[1303,1429,650,672]" pageId="3" pageNumber="476" pagination="98 - 99" refId="ref15650" refString="Kraatz, G. (1859) Die Staphylinen-Fauna von Ostindien, insbesondere der Insel Ceylan. Archiv fur Naturgeschichte, 25 (1), 98 - 99. https: // doi. org / 10.5962 / bhl. title. 66002" type="journal article" year="1859">Kraatz, 1859</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="F64436CDE35B2F76FF2EFD77FA7B03C3" blockId="3.[151,1437,151,1171]" box="[151,1436,679,701]" pageId="3" pageNumber="476">8. Paramere trilobed..................................................................................... 9</paragraph>
<paragraph id="F64436CDE35B2F76FF2EFD1AFA7B03A4" blockId="3.[151,1437,151,1171]" box="[151,1436,709,731]" pageId="3" pageNumber="476">- Paramere with single lobe............................................................................. 11</paragraph>
<paragraph id="F64436CDE35B2F76FF2EFD3DFA7A0386" blockId="3.[151,1437,151,1171]" box="[151,1437,738,760]" pageId="3" pageNumber="476">
9. Eyes longer than tempora.................................................... 
<taxonomicName id="31FB4D4EE35B2F76FBE9FD3CFA7A0386" authority="(Li &amp; Zhou, 2011)" baseAuthorityName="Li &amp; Zhou" baseAuthorityYear="2011" box="[1104,1437,738,760]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="476" phylum="Arthropoda" rank="species" species="pedatiformis">
<emphasis id="C48FEADFE35B2F76FBE9FD3CFB030389" box="[1104,1252,738,760]" italics="true" pageId="3" pageNumber="476">E. pedatiformis</emphasis>
(
<bibRefCitation id="926A4B3CE35B2F76FB48FD3DFA720386" author="Li, L. &amp; Zhou, H. Z." box="[1265,1429,738,760]" pageId="3" pageNumber="476" pagination="679 - 722" refId="ref15762" refString="Li, L. &amp; Zhou, H. Z. (2011) Revision and phylogenetic assessment of the rove beetle genus Pseudohesperus Hayashi, with broad reference to the subtribe Philonthina (Coleoptera: Staphylinidae: Staphylinini). Zoological Journal of the Linnean Society, 163 (3), 679 - 722. https: // doi. org / 10.1111 / j. 1096 - 3642.2011.00731. x" type="journal article" year="2011">Li &amp; Zhou, 2011</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="F64436CDE35B2F76FF2EFCDFFA7B026B" blockId="3.[151,1437,151,1171]" box="[151,1436,767,789]" pageId="3" pageNumber="476">- Eyes shorter than tempora............................................................................. 10</paragraph>
<paragraph id="F64436CDE35B2F76FF2EFCC2FA7A022E" blockId="3.[151,1437,151,1171]" pageId="3" pageNumber="476">
10. Middle lobe of paramere distinctly exceeding median lobe, apex of median lobe with discernible terminal angles in ventral view....................................................................... 
<taxonomicName id="31FB4D4EE35B2F76FBCDFCE4FA7A022E" authority="(Schillhammer, 2011)" baseAuthorityName="Schillhammer" baseAuthorityYear="2011" box="[1140,1437,826,848]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="476" phylum="Arthropoda" rank="species" species="ernsti">
<emphasis id="C48FEADFE35B2F76FBCDFCE4FB21022E" box="[1140,1222,827,848]" italics="true" pageId="3" pageNumber="476">E. ernsti</emphasis>
(
<bibRefCitation id="926A4B3CE35B2F76FB6BFCE5FA72022E" author="Schillhammer, H." box="[1234,1429,826,848]" pageId="3" pageNumber="476" pagination="133 - 163" refId="ref16374" refString="Schillhammer, H. (2011) Old and new Staphylinini from the Palearctic and Oriental Regions (Coleoptera: Staphylinidae: Staphylininae). Koleopterologische Rundschau, 81, 133 - 163." type="journal article" year="2011">Schillhammer, 2011</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="F64436CDE35B2F76FF2EFC87FA7A02F5" blockId="3.[151,1437,151,1171]" pageId="3" pageNumber="476">
- Middle lobe of paramere not exceeding median lobe, median lobe distinctly narrowed into subacute apex in ventral view.............................................................................. 
<taxonomicName id="31FB4D4EE35B2F76FBD1FCAAFA7A02F5" authority="(Li &amp; Zhou, 2011)" baseAuthorityName="Li &amp; Zhou" baseAuthorityYear="2011" box="[1128,1437,885,907]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="476" phylum="Arthropoda" rank="species" species="tripartitus">
<emphasis id="C48FEADFE35B2F76FBD1FCAAFB0302F4" box="[1128,1252,885,906]" italics="true" pageId="3" pageNumber="476">E. tripartitus</emphasis>
(
<bibRefCitation id="926A4B3CE35B2F76FB48FCAAFA7202F5" author="Li, L. &amp; Zhou, H. Z." box="[1265,1429,885,907]" pageId="3" pageNumber="476" pagination="679 - 722" refId="ref15762" refString="Li, L. &amp; Zhou, H. Z. (2011) Revision and phylogenetic assessment of the rove beetle genus Pseudohesperus Hayashi, with broad reference to the subtribe Philonthina (Coleoptera: Staphylinidae: Staphylinini). Zoological Journal of the Linnean Society, 163 (3), 679 - 722. https: // doi. org / 10.1111 / j. 1096 - 3642.2011.00731. x" type="journal article" year="2011">Li &amp; Zhou, 2011</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="F64436CDE35B2F76FF2EFC4DFA7B02D9" blockId="3.[151,1437,151,1171]" box="[151,1436,914,936]" pageId="3" pageNumber="476">
11. Sensory peg setae form two circular groups in ventral view (
<figureCitation id="6EC02A48E35B2F76FC9FFC4DFC8802D6" box="[806,879,914,936]" captionStart="FIGURE 3" captionStartId="6.[151,250,1865,1889]" captionTargetBox="[163,1424,786,1827]" captionTargetId="figure-212@6.[151,1436,777,1842]" captionTargetPageId="6" captionText="FIGURE 3. Eccoptolonthus dafoensis Fei &amp; Zhou, sp. nov. A. Habitus; B. aedeagus, dorsal (parameral side) view; C. apex of paramere, enlarged view, dorsal (parameral side) view; D. aedeagus, lateral view; E. sternite IX of male; F. stylus of tergite IX of male; G. sternite VIII of male; H. tergite X of male. Scale bars, A=2 mm, B, C, D, E, F, G, H= 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4640477" httpUri="https://zenodo.org/record/4640477/files/figure.png" pageId="3" pageNumber="476">Fig. 3C</figureCitation>
).................. 
<taxonomicName id="31FB4D4EE35B2F76FBE1FC4CFAA102D6" ID-CoL="8XZX9" authority="Fei &amp; Zhou" authorityName="Fei &amp; Zhou" authorityYear="2021" box="[1112,1350,914,936]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="476" phylum="Arthropoda" rank="species" species="dafoensis" status="sp. nov.">
<emphasis id="C48FEADFE35B2F76FBE1FC4CFB2902D9" box="[1112,1230,914,936]" italics="true" pageId="3" pageNumber="476">E. dafoensis</emphasis>
Fei &amp; Zhou
</taxonomicName>
, 
<emphasis id="C48FEADFE35B2F76FAE9FC4DFA7B02D9" bold="true" box="[1360,1436,914,935]" pageId="3" pageNumber="476">
<taxonomicNameLabel id="DFBC57A4E35B2F76FAE9FC4DFA7B02D9" box="[1360,1436,914,935]" pageId="3" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="F64436CDE35B2F76FF2EFC6FFA7B02BB" blockId="3.[151,1437,151,1171]" box="[151,1436,943,965]" pageId="3" pageNumber="476">- Sensory peg setae uniformly distributed in the apex......................................................... 12</paragraph>
<paragraph id="F64436CDE35B2F76FF2EFC12FA7B029D" blockId="3.[151,1437,151,1171]" box="[151,1436,973,995]" pageId="3" pageNumber="476">
12. Pronotum as long as elytra; elytra obscure-brownish...................................... 
<taxonomicName id="31FB4D4EE35B2F76FB1DFC12FA7B029D" authority="(Sharp, 1874)" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[1188,1436,973,995]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="476" phylum="Arthropoda" rank="species" species="gastralis">
<emphasis id="C48FEADFE35B2F76FB1DFC12FAF5029C" box="[1188,1298,973,994]" italics="true" pageId="3" pageNumber="476">E. gastralis</emphasis>
(
<bibRefCitation id="926A4B3CE35B2F76FAA6FC12FA72029D" author="Sharp, D." box="[1311,1429,973,995]" pageId="3" pageNumber="476" pagination="47 - 48" refId="ref16531" refString="Sharp, D. (1874) The Staphylinidae of Japan. Transactions of the Entomological Society of London, 1, 47 - 48. https: // doi. org / 10.1111 / j. 1365 - 2311.1874. tb 00159. x" type="journal article" year="1874">Sharp, 1874</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="F64436CDE35B2F76FF2EFC34FA7B0281" blockId="3.[151,1437,151,1171]" box="[151,1436,1002,1024]" pageId="3" pageNumber="476">- Pronotum shorter than elytra, elytra black-brown with suture and posterior margin markedly reddish-brown............ 13</paragraph>
<paragraph id="F64436CDE35B2F76FF2EFBD7FA7A0545" blockId="3.[151,1437,151,1171]" pageId="3" pageNumber="476">
13. Eyes about 1.25–1.67 times as long as tempora; paramere conical, weakly, gradually broadening towards base; male tergite X broadly emarginate at apex; male sternite IX with basal portion weakly deflexed laterad........ 
<taxonomicName id="31FB4D4EE35B2F76FB29FBFAFA7A0545" authority="(Li &amp; Zhou, 2011)" baseAuthorityName="Li &amp; Zhou" baseAuthorityYear="2011" box="[1168,1437,1061,1083]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="476" phylum="Arthropoda" rank="species" species="luteus">
<emphasis id="C48FEADFE35B2F76FB29FBFAFB030544" box="[1168,1252,1061,1082]" italics="true" pageId="3" pageNumber="476">E. luteus</emphasis>
(
<bibRefCitation id="926A4B3CE35B2F76FB48FBFAFA720545" author="Li, L. &amp; Zhou, H. Z." box="[1265,1429,1061,1083]" pageId="3" pageNumber="476" pagination="679 - 722" refId="ref15762" refString="Li, L. &amp; Zhou, H. Z. (2011) Revision and phylogenetic assessment of the rove beetle genus Pseudohesperus Hayashi, with broad reference to the subtribe Philonthina (Coleoptera: Staphylinidae: Staphylinini). Zoological Journal of the Linnean Society, 163 (3), 679 - 722. https: // doi. org / 10.1111 / j. 1096 - 3642.2011.00731. x" type="journal article" year="2011">Li &amp; Zhou, 2011</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="F64436CDE35B2F76FF2EFB9CFA7B05EC" blockId="3.[151,1437,151,1171]" pageId="3" pageNumber="476">
- Eyes about 0.8 times as long as tempora; paramere cylindrical, strongly broadened from the middle to base; male tergite X with smaller, triangular medioapical emargination; male sternite IX with basal portion sharply deflexed laterad (
<figureCitation id="6EC02A48E35B2F76FB5CFB80FAC40508" box="[1253,1315,1119,1142]" captionStart="FIGURE 1" captionStartId="4.[151,250,1907,1931]" captionTargetBox="[152,1407,811,1862]" captionTargetId="figure-265@4.[151,1436,810,1881]" captionTargetPageId="4" captionText="FIGURE 1. Eccoptolonthus conaensis Fei &amp; Zhou, sp. nov. A. Habitus; B. aedeagus, dorsal (parameral side) view; C. aedeagus, lateral view; D. sternite IX of male; E. stylus of tergite IX of male; F. tergite X of male; G. sternite VIII of male. Scale bars, A= 1 mm; B, C, D, E, F, G= 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4640471" httpUri="https://zenodo.org/record/4640471/files/figure.png" pageId="3" pageNumber="476">Figs. 1</figureCitation>
F-G).............................................................................. 
<taxonomicName id="31FB4D4EE35B2F76FBE3FBA2FAAD05ED" ID-CoL="8XZX8" authority="Fei &amp; Zhou" authorityName="Fei &amp; Zhou" authorityYear="2021" box="[1114,1354,1149,1171]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="476" phylum="Arthropoda" rank="species" species="conaensis" status="sp. nov.">
<emphasis id="C48FEADFE35B2F76FBE3FBA2FB3305EC" box="[1114,1236,1149,1170]" italics="true" pageId="3" pageNumber="476">E. conaensis</emphasis>
Fei &amp; Zhou
</taxonomicName>
<emphasis id="C48FEADFE35B2F76FAE9FBA2FA7B05EC" bold="true" box="[1360,1436,1149,1170]" pageId="3" pageNumber="476">
<taxonomicNameLabel id="DFBC57A4E35B2F76FAE9FBA2FA7B05EC" box="[1360,1436,1149,1170]" pageId="3" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</subSection>
</treatment>
</document>