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<document id="44AC6F0E45813FF4F305BE10A47813C5" ID-CLB-Dataset="55550" ID-DOI="10.5281/zenodo.6544157" ID-GBIF-Dataset="6cab72d7-5386-4122-8f98-dfd97678ebf6" ID-ISSN="1536-9307" ID-Zenodo-Dep="6544157" ID-ZooBank="6580A0C7-E18D-4B06-A158-522F089CEAF5" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638259586037" checkinUser="felipe" docAuthor="Kovařík, František, Lowe, Graeme &amp; Šťáhlavský, František" docDate="2018" docId="7E4D87D88573FFC80A69DDB6FC314D5E" docLanguage="en" docName="Euscorpius.267-1-41.pdf" docOrigin="Euscorpius 267" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Barbaracurus zambonellii Kovařík &amp; Lowe &amp; Šťáhlavský 2018, comb. n." docType="treatment" docVersion="8" lastPageNumber="36" masterDocId="8274FFA08551FFED0922DE5FFFEC4B65" masterDocTitle="Review of the genus Babycurus Karsch, 1886 (Arachnida, Scorpiones, Buthidae), with descriptions of Barbaracurus gen. n. and two new species from Oman and Yemen" masterLastPageNumber="41" masterPageNumber="1" pageNumber="33" updateTime="1699471299651" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="EC39CBFA5593C23017C406989794613D">Review of the genus Babycurus Karsch, 1886 (Arachnida, Scorpiones, Buthidae), with descriptions of Barbaracurus gen. n. and two new species from Oman and Yemen</mods:title>
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<mods:namePart id="E625BA0D9E73583FDC7A93ACBE86E371">Lowe, Graeme</mods:namePart>
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<mods:namePart id="895504F46B835CAFE0C532C0D0CE3923">Šťáhlavský, František</mods:namePart>
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<treatment id="7E4D87D88573FFC80A69DDB6FC314D5E" ID-DOI="http://doi.org/10.5281/zenodo.5741275" ID-GBIF-Taxon="195303861" ID-Zenodo-Dep="5741275" LSID="urn:lsid:plazi:treatment:7E4D87D88573FFC80A69DDB6FC314D5E" httpUri="http://treatment.plazi.org/id/7E4D87D88573FFC80A69DDB6FC314D5E" lastPageId="37" lastPageNumber="36" pageId="34" pageNumber="33">
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<paragraph id="F65B36CE8573FFCF0A69DDB6FA784F66" blockId="34.[837,1440,1001,1094]" box="[843,1428,1001,1028]" pageId="34" pageNumber="33">
<heading id="AD1381A28573FFCF0A69DDB6FA784F66" box="[843,1428,1001,1028]" centered="true" fontSize="10" level="2" pageId="34" pageNumber="33" reason="4">
<taxonomicName id="31E44D4D8573FFCF0A69DDB6FACB4F66" ID-CoL="KM4Y" authority="(Borelli, 1902)" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2018" baseAuthorityName="Borelli" baseAuthorityYear="1902" box="[843,1319,1001,1028]" class="Arachnida" family="Buthidae" genus="Barbaracurus" kingdom="Animalia" order="Scorpiones" pageId="34" pageNumber="33" phylum="Arthropoda" rank="species" species="zambonellii" status="comb. nov.">
<emphasis id="C490EADC8573FFCF0A69DDB6FB954F61" bold="true" box="[843,1145,1001,1028]" italics="true" pageId="34" pageNumber="33">Barbaracurus zambonellii</emphasis>
(
<bibRefCitation id="92754B3F8573FFCF0DAADDB4FAF24F66" author="BORELLI, A." box="[1160,1310,1003,1027]" journalOrPublisher="Bollettino dei Musei di Zoologia ed Anatomia Comparata della Reale Universita di Torino" pageId="34" pageNumber="33" pagination="1 - 4" part="17" refId="ref16152" refString="BORELLI, A. 1902. Di una nuova specie di scorpione della Colonia Eritrea. Bollettino dei Musei di Zoologia ed Anatomia Comparata della Reale Universita di Torino, 17 (422): 1 - 4." title="Di una nuova specie di scorpione della Colonia Eritrea" type="journal article" year="1902">Borelli, 1902</bibRefCitation>
)
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<emphasis id="C490EADC8573FFCF0C0CDDB6FA834F66" bold="true" box="[1326,1391,1001,1027]" pageId="34" pageNumber="33">comb</emphasis>
.
<emphasis id="C490EADC8573FFCF0C5FDDB6FA614F66" bold="true" box="[1405,1421,1001,1027]" pageId="34" pageNumber="33">n</emphasis>
.
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(
<figureCitation id="6EDF2A4B8573FFCF0A6FDA51FC584F40" box="[845,948,1038,1061]" captionStart="Figures 123" captionStartId="4.[192,278,1544,1568]" captionTargetBox="[200,1432,168,1512]" captionTargetId="figure-16@4.[192,1440,163,1512]" captionTargetPageId="4" captionText="Figures 123: Pedipalp movable finger, comparison of Barbaracurus gen. n. (110) and Babycurus (1123) species. Figures 14: Barbaracurus gen. n., species with 6 non-imbricated almost linear rows of granules. Figure 1. B. ugartei comb. n., female holotype. Figure 2. B. sofomarensis comb. n., male paratype. Figure 3. B. somalicus comb. n., female from Somaliland, locality 17SR. Figure 4. B. subpunctatus comb. n., female from Ethiopia, locality 14EI. Figures 510: Barbaracurus gen. n., species with 7 non-imbricated almost linear rows of granules. Figure 5. B. exquisitus comb. n., male holotype. Figures 67, 10. B. zambonellii comb. n., female (6 and 10) and male (7) from Eritrea, locality 15EH. Figure 8. B. winklerorum sp. n., female paratype. Figure 9. B. yemenensis sp. n., female holotype. Figures 1214: Babycurus, species with 6 imbricated rows of movable fingers. Figure 12. B. ansorgei, female, Congo, Bukama env., FKCP. Figure 13. B. centrurimorphus, female lectotype. Figure 14. B. pictus, female, Congo, Sampwe env., FKCP. Figures 11, 1521: Babycurus, species with 78 imbricated rows of movable fingers. Figure 15. B. buettneri, female, MCSN. Figure 11, 16. B. dunlopi, male paratype. Figure 17. B. jacksoni, male, Tanzania, FKCP, No. 1189. Figure 18. B. kirki, female, Cote dIvoire, Comoe, FKCP. Figure 19. Babycurus melanicus, female holotype. Figure 20. B. taramassoi, female, Somalia, Belet Amin, MCSN. Figure 21. Babycurus wituensis, female lectotype. Figures 2223: Babycurus, species with 9 imbricated rows of movable fingers. Figure 22. B. gigas, female lectotype. Figure 23. B. multisubaculeatus, female holotype. Numbers in red indicate number of granule rows in series of fingers on the left of each number (red arrows)." figureDoi="http://doi.org/10.5281/zenodo.6544159" httpUri="https://zenodo.org/record/6544159/files/figure.png" pageId="34" pageNumber="33">Figures 7</figureCitation>
,
<figureCitation id="6EDF2A4B8573FFCF0AE2DA51FBE94F40" box="[960,1029,1038,1061]" captionStart="Figures 2439" captionStartId="6.[192,278,1598,1622]" captionTargetBox="[241,1392,199,1576]" captionTargetId="figure-16@6.[239,1405,192,1582]" captionTargetPageId="6" captionText="Figures 2439: Hemispermatophore capsules of Barbaracurus gen. n. and Babycurus species, convex and anterior views. In each figure. left image is convex view with capsule compressed to show shape of lobes, right image anterior view to show profile of basal lobe. Figures 2431. Color images. Figures 2428: Barbaracurus gen. n. Figure 24. B. winklerorum sp. n., holotype. Figure 25. B. exquisitus comb. n., Oman, Jabal Akhdar hotel, 23º04'N 57º38'E. Figure 26. B. sofomarensis comb. n., paratype from the type locality. Figure 27. B. somalicus comb. n., Somaliland, Borama, locality 17SR. Figure 28. B. zambonellii comb. n., Eritrea, Filfil, locality 15EH. Figures 2931: Babycurus. Figure 29: B. jacksoni, Tanzania. Figure 30: B. gigas, Tanzania, Usambara Mts. Figure 31: B. buettneri, Cameroon, Mamba vill., 04°07.311'N 10°11.615'E. Scale bars: 200 µm. Small numbered labels indicate specimen numbers (see lists of Material Examined). Figures 3239. Gray scale images with basal lobes outlined in red. Figures 3236: Barbaracurus gen. n.: B. winklerorum sp. n. (32), B. exquisitus comb. n. (33), B. sofomarensis comb. n. (34), B. somalicus comb. n. (35) and B. zambonellii comb. n. (36). Figures 3739: Babycurus: B. jacksoni (37), B. gigas (38) and B. buettneri (39). Solid red curves indicate either the distal margin of a scoop-like basal lobe (32, 34) or a well developed carina (3537, 39). Dashed red curves indicate either the line of attachment of a scoop-like basal lobe (32, 34) or a weakly developed carina (3536, 38)." figureDoi="http://doi.org/10.5281/zenodo.6544161" httpUri="https://zenodo.org/record/6544161/files/figure.png" pageId="34" pageNumber="33">28, 36</figureCitation>
,
<figureCitation id="6EDF2A4B8573FFCF0D32DA51FBB94F40" box="[1040,1109,1038,1061]" captionStart="Figures 5069" captionStartId="8.[192,278,1229,1253]" captionTargetBox="[195,1434,198,1190]" captionTargetId="figure-48@8.[192,1438,192,1200]" captionTargetPageId="8" captionText="Figures 5069: Pedipalp chela dorsal and external views, comparison of Barbaracurus gen. n. species. Figures 5053: B. sofomarensis comb. n., male (5051) and female (5253) paratypes. Figures 5457: B. somalicus comb. n., male (5455) and female (5657) from Somaliland, locality 17SR. Figures 5861: B. zambonellii comb. n., male (5859) and female (6061) from Eritrea, locality 15EH. Figures 6265: B. exquisitus comb. n., male holotype (6263) and female (6465) paratype. Figures 66 69: B. winklerorum sp. n., male holotype (6667) and female (6869) paratype." figureDoi="http://doi.org/10.5281/zenodo.6544167" httpUri="https://zenodo.org/record/6544167/files/figure.png" pageId="34" pageNumber="33">5861</figureCitation>
,
<figureCitation id="6EDF2A4B8573FFCF0D7DDA51FB494F40" box="[1119,1189,1038,1061]" captionStart="Figures 7080" captionStartId="9.[192,278,1644,1668]" captionTargetBox="[201,1439,170,1605]" captionTargetId="figure-188@9.[192,1439,168,1614]" captionTargetPageId="9" captionText="Figures 7080: Metasoma and telson ventral views, comparison of Barbaracurus gen. n. species. Figures 7071: B. exquisitus comb. n., male holotype (70) and female (71) paratype. Figures 7273: B. sofomarensis comb. n., male (72) and female (73) paratypes. Figures 7474: B. somalicus comb. n., male (74) and female (75) from Somaliland, locality 17SR. Figures 7677: B. winklerorum sp. n., male holotype (76) and female (77) paratype. Figures 78: B. yemenensis sp. n., female holotype. Figures 7980. B. zambonellii comb. n., male (79) and female (80) from Eritrea, locality 15EH." figureDoi="http://doi.org/10.5281/zenodo.6544169" httpUri="https://zenodo.org/record/6544169/files/figure.png" pageId="34" pageNumber="33">7980</figureCitation>
,
<figureCitation id="6EDF2A4B8573FFCF0D92DA51FB184F40" box="[1200,1268,1038,1061]" captionStart="Figures 8189" captionStartId="10.[192,278,1781,1805]" captionTargetBox="[198,1426,218,1748]" captionTargetId="figure-163@10.[192,1439,211,1752]" captionTargetPageId="10" captionText="Figures 8189: Telson, lateral views of both sexes, comparison of Barbaracurus gen. n. species. Figures 8182: B. exquisitus comb. n., male holotype (81) and female (82) paratype. Figures 8384: B. somalicus comb. n., male (83) and female (84) from Somaliland, locality 17SR. Figures 8586: B. winklerorum sp. n., male holotype (85) and female (86) paratype. Figures 87: B. yemenensis sp. n., female holotype. Figures 8889. B. zambonellii comb. n., male (88) and female (89) from Eritrea, locality 15EH." figureDoi="http://doi.org/10.5281/zenodo.6544171" httpUri="https://zenodo.org/record/6544171/files/figure.png" pageId="34" pageNumber="33">8889</figureCitation>
,
<figureCitation id="6EDF2A4B8573FFCF0C22DA51FA8C4F40" box="[1280,1376,1038,1061]" captionStart-0="Figures 219225" captionStart-1="Figures 226238" captionStart-2="Figures 239247" captionStartId-0="31.[192,278,1640,1664]" captionStartId-1="32.[192,278,1770,1794]" captionStartId-2="33.[192,278,1647,1671]" captionTargetBox-0="[191,1440,191,1611]" captionTargetBox-1="[191,1440,191,1742]" captionTargetBox-2="[200,1439,198,1614]" captionTargetId-0="figure-244@31.[192,1439,192,1610]" captionTargetId-1="figure-142@32.[192,1439,192,1741]" captionTargetId-2="figure-6@33.[192,1439,192,1618]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionTargetPageId-2="33" captionText-0="Figures 219225: Barbaracurus zambonellii comb. n. from Eritrea, locality 15EH. Figures 219, 221. Male, dorsal (219) and ventral (221) views. Figures 220, 222. Female, dorsal (220) and ventral (222) views. Figure 223. Right hemispermatophore, convex aspect. Figures 224225. Eritrea, locality 15EH. Scale bars: 10 mm (219222), 1 mm (223)." captionText-1="Figures 226238: Barbaracurus zambonellii comb. n. from Eritrea, locality 15EH. Figures 226, 228, 232238. Male, chelicerae, carapace and tergites IIII (226), sternopectinal region and sternite III (228), pedipalp chela, dorsoexternal (232), external (233), and ventrointernal (234) views, pedipalp patella, dorsal (235) and external (236) views, pedipalp femur and trochanter internal (237) and dorsal (238) views. Figures 227, 229231. Female, chelicerae, carapace and tergites IIII (227), sternopectinal region and sternites IIIIV (229), pedipalp chela, dorsoexternal (230) and external (231) views. The trichobothrial pattern is indicated in Figures 233238 (white circles)." captionText-2="Figures 239247: Figures 239244: Barbaracurus zambonellii comb. n. from Eritrea, locality 15EH. Figures 239241. Male, metasoma and telson, lateral (239), ventral (240), and dorsal (241) views. Figures 242244. Female, metasoma and telson, lateral (242), ventral (243), and dorsal (244) views. Figures 245247: Barbaracurus yemenensis sp. n., Female holotype, metasoma and telson, lateral (245), dorsal (246), and ventral (247) views. Scale bars: 10 mm (239241, 242244, 245247)." figureDoi-0="http://doi.org/10.5281/zenodo.6544201" figureDoi-1="http://doi.org/10.5281/zenodo.6544203" figureDoi-2="http://doi.org/10.5281/zenodo.6544205" httpUri-0="https://zenodo.org/record/6544201/files/figure.png" httpUri-1="https://zenodo.org/record/6544203/files/figure.png" httpUri-2="https://zenodo.org/record/6544205/files/figure.png" pageId="34" pageNumber="33">219244</figureCitation>
,
<figureCitation id="6EDF2A4B8573FFCF0C48DA51FBEE4F23" captionStart-0="Figures 248249" captionStart-1="Figures 250251" captionStartId-0="35.[192,278,1834,1858]" captionStartId-1="36.[192,278,1834,1858]" captionTargetBox-0="[191,1440,191,1806]" captionTargetBox-1="[191,1440,191,1806]" captionTargetId-0="figure-25@35.[192,1439,192,1805]" captionTargetId-1="figure-52@36.[192,1439,192,1805]" captionTargetPageId-0="35" captionTargetPageId-1="36" captionText-0="Figures 248249: Barbaracurus zambonellii comb. n., in vivo habitus. Male (248) and female (249) from Eritrea, locality 15EH." captionText-1="Figures 250251: Barbaracurus zambonellii comb. n. female with newborns (first instar) before first ecdysis (250) and female with juveniles (second instar) after first ecdysis (251) from Eritrea, locality 15EH." figureDoi-0="http://doi.org/10.5281/zenodo.6544207" figureDoi-1="http://doi.org/10.5281/zenodo.6544209" httpUri-0="https://zenodo.org/record/6544207/files/figure.png" httpUri-1="https://zenodo.org/record/6544209/files/figure.png" pageId="34" pageNumber="33">248 251</figureCitation>
,
<figureCitation id="6EDF2A4B8573FFCF0D2FDA70FB814F23" box="[1037,1133,1071,1094]" captionStart="Figures 255264" captionStartId="38.[192,278,1632,1656]" captionTargetBox="[191,1440,191,1603]" captionTargetId="figure-222@38.[192,1439,192,1603]" captionText="Figures 255264: Male postpachytene (255, 257, 261, 263), mitotic metaphase (259), and ideograms (256, 258, 260, 262, 264) (y axis - % of the diploid chromosome length) of Babycurus (255260) and Barbaracurus gen. n. (261264) species. Babycurus jacksoni (1189), Tanzania (2n=16) (255, 256), B. buettneri (1190) from Cameroon, Mamba vill., 04°07.311'N 10°11.615'E (2n=28) (257, 258) and B. gigas (914) from Tanzania (2n=30) (259, 260); Barbaracurus zambonelli comb. n. (876) from Eritrea, locality 15EH (2n=26) (261, 262) and B. somalicus comb. n. (1332), Somaliland, locality 17SR (2n=36) (263, 264). Scale bar: 5 µm (255, 257, 259, 261, 263)." figureDoi="http://doi.org/10.5281/zenodo.6544213" httpUri="https://zenodo.org/record/6544213/files/figure.png" pageId="34" pageNumber="33">261262</figureCitation>
, 265,
<tableCitation id="BB6603758573FFCF0D8FDA70FB104F23" box="[1197,1276,1071,1094]" captionStart="Table 1" captionStartId="21.[192,257,1051,1075]" captionTargetBox="[362,1734,417,1017]" captionTargetId="graphics-6@21.[352,1759,407,1022]" captionText="Table 1: Measurements of Barbaracurus zambonellii comb. n. and B. somalicus comb. n. Abbreviations: length (L), width (W, in carapace depth (D)." httpUri="http://table.plazi.org/id/A29B66468544FFF809E2DA44FEC84F2E" pageId="34" pageNumber="33" tableUuid="A29B66468544FFF809E2DA44FEC84F2E">Table 1</tableCitation>
)
</paragraph>
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<paragraph id="F65B36CE8573FFCF0A1CDA2CFB464E27" blockId="34.[830,1440,1139,1346]" pageId="34" pageNumber="33">
<taxonomicName id="31E44D4D8573FFCF0A1CDA2CFAE24FEF" authority="Borelli, 1902: 1 - 4" authorityName="Borelli" authorityPageNumber="1 - 4" authorityYear="1902" box="[830,1294,1139,1162]" class="Arachnida" family="Buthidae" genus="Babycurus" kingdom="Animalia" order="Scorpiones" pageId="34" pageNumber="33" phylum="Arthropoda" rank="species" species="zambonellii">
<emphasis id="C490EADC8573FFCF0A1CDA2CFBDB4FEF" box="[830,1079,1139,1162]" italics="true" pageId="34" pageNumber="33">Babycurus zambonellii</emphasis>
<bibRefCitation id="92754B3F8573FFCF0D63DA2CFAE24FEF" author="BORELLI, A." box="[1089,1294,1139,1162]" journalOrPublisher="Bollettino dei Musei di Zoologia ed Anatomia Comparata della Reale Universita di Torino" pageId="34" pageNumber="33" pagination="1 - 4" part="17" refId="ref16152" refString="BORELLI, A. 1902. Di una nuova specie di scorpione della Colonia Eritrea. Bollettino dei Musei di Zoologia ed Anatomia Comparata della Reale Universita di Torino, 17 (422): 1 - 4." title="Di una nuova specie di scorpione della Colonia Eritrea" type="journal article" year="1902">Borelli, 1902: 14</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="92754B3F8573FFCF0C3FDA2CFC754FCC" author="HIRST, S." journalOrPublisher="Annals and Magazine of Natural History" pageId="34" pageNumber="33" pagination="208 - 211" part="VII" refId="ref16533" refString="HIRST, S. 1907. Notes on scorpions, with descriptions of two new species. Annals and Magazine of Natural History, VII (19): 208 - 211." title="Notes on scorpions, with descriptions of two new species" type="journal article" year="1907">Hirst, 1907: 209</bibRefCitation>
;
<bibRefCitation id="92754B3F8573FFCF0A8ADACDFB7E4FCC" author="Kraepelin" box="[936,1170,1170,1193]" firstAuthor="Kraepelin" page="181" pageId="34" pageNumber="33" pagination="123 - 196" refId="ref17199" refString="KRAEPELIN, K. 1913. Neue Beitrage zur Systematik der Gliederspinnen. III. A. Bemerkungen zur Skorpionenfauna Indiens. B. Die Skorpione, Pedipalpen und Solifugen Deutsch - Ostafrikas. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 30: 123 - 196." type="journal article" year="1913">Kraepelin, 1913: 181</bibRefCitation>
;
<bibRefCitation id="92754B3F8573FFCF0D82DACDFA704FCC" author="CAPORIACCO, L. DI" box="[1184,1436,1170,1193]" journalOrPublisher="Acta Pontificae Academiae Scientiarum Novi Lyncei" pageId="34" pageNumber="33" pagination="227 - 233" part="11" refId="ref16220" refString="CAPORIACCO, L. DI 1947. Scorpioni dell'Eritrea del Museo zoologici di Firenze. Acta Pontificae Academiae Scientiarum Novi Lyncei, 11 (19): 227 - 233." title="Scorpioni dell'Eritrea del Museo zoologici di Firenze" type="journal article" year="1947">Caporiacco, 1947: 232</bibRefCitation>
;
<bibRefCitation id="92754B3F8573FFCF0A4CDAEFFBDC4FA2" author="Probst" box="[878,1072,1200,1223]" firstAuthor="Probst" page="329" pageId="34" pageNumber="33" pagination="312 - 335" refId="ref17851" refString="PROBST, P. 1973. A review of the scorpions of East Africa with special regard to Kenya and Tanzania. Acta Tropica, 30: 312 - 335." type="journal article" year="1973">Probst, 1973: 329</bibRefCitation>
;
<bibRefCitation id="92754B3F8573FFCF0D1EDAEFFA704FA2" author="LAMORAL, B. H. &amp; S. REYNDERS" box="[1084,1436,1200,1223]" journalOrPublisher="Annals of the Natal Museum" pageId="34" pageNumber="33" pagination="489 - 576" part="22" refId="ref17245" refString="LAMORAL, B. H. &amp; S. REYNDERS 1975. A catalogue of the scorpions described from the Ethiopian Faunal Region up to December 1973. Annals of the Natal Museum, 22: 489 - 576." title="A catalogue of the scorpions described from the Ethiopian Faunal Region up to December 1973" type="journal article" year="1975">Lamoral &amp; Reynders, 1975: 498</bibRefCitation>
;
<bibRefCitation id="92754B3F8573FFCF0A4CDA90FBAD4F83" author="KOVARIK, F." bookContentInfo="176 pp" box="[878,1089,1231,1254]" journalOrPublisher="Jihlava (Czech Republic): Publishing House &quot; Madagaskar" pageId="34" pageNumber="33" refId="ref16585" refString="KOVARIK, F. 1998. Stiri [Scorpiones]. Jihlava (Czech Republic): Publishing House &quot; Madagaskar &quot;, 176 pp (in Czech)." title="Stiri [Scorpiones]" type="book" year="1998">Kovařík, 1998: 104</bibRefCitation>
(in part);
<bibRefCitation id="92754B3F8573FFCF0D8DDA90FA4C4F83" author="LOWE" box="[1199,1440,1231,1254]" firstAuthor="Fet" page="80" pageId="34" pageNumber="33" pagination="185 - 192" refId="ref17483" refString="LOWE, G. 2000. A new species of Babycurus (Scorpiones: Buthidae) from northern Oman. Entomological News, 111: 185 - 192." type="journal article" year="2000">Fet &amp; Lowe, 2000: 80</bibRefCitation>
(in part);
<bibRefCitation id="92754B3F8573FFCF0AF8DAB1FB214E60" author="Lowe" box="[986,1229,1262,1285]" firstAuthor="Lowe" page="185 - 191" pageId="34" pageNumber="33" pagination="185 - 192" refId="ref17483" refString="LOWE, G. 2000. A new species of Babycurus (Scorpiones: Buthidae) from northern Oman. Entomological News, 111: 185 - 192." type="journal article" year="2000">Lowe, 2000: 185191</bibRefCitation>
(in part);
<bibRefCitation id="92754B3F8573FFCF0C60DAB1FBFF4E46" author="KOVARIK, F." journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="34" pageNumber="33" pagination="235 - 265" part="64" refId="ref16615" refString="KOVARIK, F. 2000. Revision of Babycurus with descriptions of three new species (Scorpiones: Buthidae). Acta Societatis Zoologicae Bohemicae, 64: 235 - 265." title="Revision of Babycurus with descriptions of three new species (Scorpiones: Buthidae)" type="journal article" year="2000">Kovařík, 2000: 260261</bibRefCitation>
(in part);
<bibRefCitation id="92754B3F8573FFCF0DA8DB53FA8D4E46" author="KOVARIK, F." box="[1162,1377,1292,1315]" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="34" pageNumber="33" pagination="133 - 159" part="67" refId="ref16647" refString="KOVARIK, F. 2003. Scorpions of Djibouti, Eritrea, Ethiopia, and Somalia (Arachnida: Scorpiones) with a key and descriptions of three new species. Acta Societatis Zoologicae Bohemicae, 67: 133 - 159." title="Scorpions of Djibouti, Eritrea, Ethiopia, and Somalia (Arachnida: Scorpiones) with a key and descriptions of three new species" type="journal article" year="2003">Kovařík, 2003: 137</bibRefCitation>
(? in part);
<bibRefCitation id="92754B3F8573FFCF0A8DDB74FB4A4E27" author="CHIARLE, A. &amp; F. KOVARIK &amp; L. LEVI &amp; E. GAVETTI" box="[943,1190,1323,1346]" journalOrPublisher="Arachnologische Mitteilungen" pageId="34" pageNumber="33" pagination="17 - 23" part="43" refId="ref16251" refString="CHIARLE, A., F. KOVARIK, L. LEVI &amp; E. GAVETTI. 2012. The scorpion collections (Arachnida, Scorpiones) held in the Museo Regionale di Scienze Naturali of Turin (Italy). Arachnologische Mitteilungen, 43: 17 - 23." title="The scorpion collections (Arachnida, Scorpiones) held in the Museo Regionale di Scienze Naturali of Turin (Italy)" type="journal article" year="2012">Chiarle et al., 2012: 21</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BEFE65458573FFCF0A1CDB31FB474D1B" pageId="34" pageNumber="33" type="materials_examined">
<paragraph id="F65B36CE8573FFCF0A1CDB31FBFE4EC1" blockId="34.[830,1440,1390,1444]" pageId="34" pageNumber="33">
<collectionCode id="90F5AE0B8573FFCF0A1CDB31FBC74EE0" box="[830,1067,1390,1413]" pageId="34" pageNumber="33">TYPE LOCALITY AND</collectionCode>
<typeStatus id="295F886C8573FFCF0D1ADB2DFB454EE0" box="[1080,1193,1394,1413]" pageId="34" pageNumber="33" type="holotype">HOLOTYPE</typeStatus>
<collectionCode id="90F5AE0B8573FFCF0D94DB2DFAD74EE0" box="[1206,1339,1394,1413]" pageId="34" pageNumber="33">DEPOSITORY</collectionCode>
.
<collectingCountry id="8EF3765E8573FFCF0C73DB31FA704EE0" box="[1361,1436,1390,1413]" name="Eritrea" pageId="34" pageNumber="33">Eritrea</collectingCountry>
,
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;
<collectionCode id="90F5AE0B8573FFCF0A9DDBD2FBE24EC1" box="[959,1038,1421,1444]" country="Italy" httpUri="http://grbio.org/cool/yjj8-is06" name="Museo e Instituto DI Zoologia Sistematica dell' UniversitaDI Torino" pageId="34" pageNumber="33">MZUT</collectionCode>
.
</paragraph>
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<collectionCode id="90F5AE0B8573FFCF0A1CDB91FBD84E81" box="[830,1076,1486,1509]" pageId="34" pageNumber="33">MATERIAL EXAMINED</collectionCode>
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<emphasis id="C490EADC8573FFCF0D6EDB92FB734E80" bold="true" box="[1100,1183,1485,1509]" pageId="34" pageNumber="33">
<collectingCountry id="8EF3765E8573FFCF0D6EDB92FB734E80" box="[1100,1183,1485,1509]" name="Eritrea" pageId="34" pageNumber="33">Eritrea</collectingCountry>
</emphasis>
,
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(
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),
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(
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),
<collectionCode id="90F5AE0B8573FFCF0D75DBB3FB4A4D66" box="[1111,1190,1516,1539]" country="Italy" httpUri="http://grbio.org/cool/yjj8-is06" name="Museo e Instituto DI Zoologia Sistematica dell' UniversitaDI Torino" pageId="34" pageNumber="33">MZUT</collectionCode>
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<location id="F33B60158573FFCF0D90DBB3FB064D66" LSID="urn:lsid:plazi:treatment:7E4D87D88573FFC80A69DDB6FC314D5E:F33B60158573FFCF0D90DBB3FB064D66" box="[1202,1258,1516,1539]" country="Eritrea" latitude="15.6096115" longLatPrecision="1" longitude="38.956333" municipality="Salamuna river" name="Filfil" pageId="34" pageNumber="33">Filfil</location>
,
<collectingMunicipality id="163FACB48573FFCF0DD4DBB3FA704D66" box="[1270,1436,1516,1539]" pageId="34" pageNumber="33">Salamuna river</collectingMunicipality>
,
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<geoCoordinate id="93D050098573FFCF0AF9D854FB804D47" box="[987,1132,1547,1570]" degrees="38" direction="east" minutes="57" orientation="longitude" pageId="34" pageNumber="33" precision="1" seconds="22.8" value="38.956333">38°57'22.8&quot;E</geoCoordinate>
,
<elevation id="7DC9D1FD8573FFCF0D59D854FAEC4D47" box="[1147,1280,1547,1570]" metricMagnitude="2" metricUnit="m" metricValue="8.17" pageId="34" pageNumber="33" unit="m" value="817.0">
<quantity id="311C9B2B8573FFCF0D59D854FB2F4D47" box="[1147,1219,1547,1570]" metricMagnitude="2" metricUnit="m" metricValue="8.17" pageId="34" pageNumber="33" unit="m" value="817.0">817 m</quantity>
a.s.l.
</elevation>
,
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<collectingDate id="921EE9E68573FFCF0C32D854FA704D47" box="[1296,1436,1547,1570]" pageId="34" pageNumber="33" value="2015-11-03" valueMax="2015-11-04" valueMin="2015-11-03">3.-4.XI.2015</collectingDate>
</date>
, (
<figureCitation id="6EDF2A4B8573FFCF0A64D875FC084D24" box="[838,996,1578,1601]" captionStart="Figures 219225" captionStartId="31.[192,278,1640,1664]" captionTargetBox="[191,1440,191,1611]" captionTargetId="figure-244@31.[192,1439,192,1610]" captionTargetPageId="31" captionText="Figures 219225: Barbaracurus zambonellii comb. n. from Eritrea, locality 15EH. Figures 219, 221. Male, dorsal (219) and ventral (221) views. Figures 220, 222. Female, dorsal (220) and ventral (222) views. Figure 223. Right hemispermatophore, convex aspect. Figures 224225. Eritrea, locality 15EH. Scale bars: 10 mm (219222), 1 mm (223)." figureDoi="http://doi.org/10.5281/zenodo.6544201" httpUri="https://zenodo.org/record/6544201/files/figure.png" pageId="34" pageNumber="33">Figs. 224225</figureCitation>
,
<collectorName id="5B1153188573FFCF0AD2D875FB954D24" box="[1008,1145,1578,1601]" pageId="34" pageNumber="33">Locality No.</collectorName>
<emphasis id="C490EADC8573FFCF0DA2D876FB2E4D24" bold="true" box="[1152,1218,1577,1601]" pageId="34" pageNumber="33">
15
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),
<specimenCount id="E0E2FD478573FFCF0DFBD875FB164D24" box="[1241,1274,1578,1601]" pageId="34" pageNumber="33" type="male">3♂</specimenCount>
<specimenCount id="E0E2FD478573FFCF0DD8D875FAD14D24" box="[1274,1341,1578,1601]" pageId="34" pageNumber="33" type="female">3♀1♀</specimenCount>
im.3juvs. (
<location id="F33B60158573FFCF0A64D817FC834D3A" LSID="urn:lsid:plazi:treatment:7E4D87D88573FFC80A69DDB6FC314D5E:F33B60158573FFCF0A64D817FC834D3A" box="[838,879,1608,1631]" country="Eritrea" name="No." pageId="34" pageNumber="33">No.</location>
876,
<figureCitation id="6EDF2A4B8573FFCF0A9BD817FBAB4D3A" box="[953,1095,1608,1631]" captionStart="Figures 2439" captionStartId="6.[192,278,1598,1622]" captionTargetBox="[241,1392,199,1576]" captionTargetId="figure-16@6.[239,1405,192,1582]" captionTargetPageId="6" captionText="Figures 2439: Hemispermatophore capsules of Barbaracurus gen. n. and Babycurus species, convex and anterior views. In each figure. left image is convex view with capsule compressed to show shape of lobes, right image anterior view to show profile of basal lobe. Figures 2431. Color images. Figures 2428: Barbaracurus gen. n. Figure 24. B. winklerorum sp. n., holotype. Figure 25. B. exquisitus comb. n., Oman, Jabal Akhdar hotel, 23º04'N 57º38'E. Figure 26. B. sofomarensis comb. n., paratype from the type locality. Figure 27. B. somalicus comb. n., Somaliland, Borama, locality 17SR. Figure 28. B. zambonellii comb. n., Eritrea, Filfil, locality 15EH. Figures 2931: Babycurus. Figure 29: B. jacksoni, Tanzania. Figure 30: B. gigas, Tanzania, Usambara Mts. Figure 31: B. buettneri, Cameroon, Mamba vill., 04°07.311'N 10°11.615'E. Scale bars: 200 µm. Small numbered labels indicate specimen numbers (see lists of Material Examined). Figures 3239. Gray scale images with basal lobes outlined in red. Figures 3236: Barbaracurus gen. n.: B. winklerorum sp. n. (32), B. exquisitus comb. n. (33), B. sofomarensis comb. n. (34), B. somalicus comb. n. (35) and B. zambonellii comb. n. (36). Figures 3739: Babycurus: B. jacksoni (37), B. gigas (38) and B. buettneri (39). Solid red curves indicate either the distal margin of a scoop-like basal lobe (32, 34) or a well developed carina (3537, 39). Dashed red curves indicate either the line of attachment of a scoop-like basal lobe (32, 34) or a weakly developed carina (3536, 38)." figureDoi="http://doi.org/10.5281/zenodo.6544161" httpUri="https://zenodo.org/record/6544161/files/figure.png" pageId="34" pageNumber="33">Figs. 28, 36</figureCitation>
,
<figureCitation id="6EDF2A4B8573FFCF0D7AD817FB544D3A" box="[1112,1208,1608,1631]" captionStart="Figures 219225" captionStartId="31.[192,278,1640,1664]" captionTargetBox="[191,1440,191,1611]" captionTargetId="figure-244@31.[192,1439,192,1610]" captionTargetPageId="31" captionText="Figures 219225: Barbaracurus zambonellii comb. n. from Eritrea, locality 15EH. Figures 219, 221. Male, dorsal (219) and ventral (221) views. Figures 220, 222. Female, dorsal (220) and ventral (222) views. Figure 223. Right hemispermatophore, convex aspect. Figures 224225. Eritrea, locality 15EH. Scale bars: 10 mm (219222), 1 mm (223)." figureDoi="http://doi.org/10.5281/zenodo.6544201" httpUri="https://zenodo.org/record/6544201/files/figure.png" pageId="34" pageNumber="33">219223</figureCitation>
,
<figureCitation id="6EDF2A4B8573FFCF0DE8D817FAC64D3A" box="[1226,1322,1608,1631]" captionStart-0="Figures 226238" captionStart-1="Figures 239247" captionStartId-0="32.[192,278,1770,1794]" captionStartId-1="33.[192,278,1647,1671]" captionTargetBox-0="[191,1440,191,1742]" captionTargetBox-1="[200,1439,198,1614]" captionTargetId-0="figure-142@32.[192,1439,192,1741]" captionTargetId-1="figure-6@33.[192,1439,192,1618]" captionTargetPageId-0="32" captionTargetPageId-1="33" captionText-0="Figures 226238: Barbaracurus zambonellii comb. n. from Eritrea, locality 15EH. Figures 226, 228, 232238. Male, chelicerae, carapace and tergites IIII (226), sternopectinal region and sternite III (228), pedipalp chela, dorsoexternal (232), external (233), and ventrointernal (234) views, pedipalp patella, dorsal (235) and external (236) views, pedipalp femur and trochanter internal (237) and dorsal (238) views. Figures 227, 229231. Female, chelicerae, carapace and tergites IIII (227), sternopectinal region and sternites IIIIV (229), pedipalp chela, dorsoexternal (230) and external (231) views. The trichobothrial pattern is indicated in Figures 233238 (white circles)." captionText-1="Figures 239247: Figures 239244: Barbaracurus zambonellii comb. n. from Eritrea, locality 15EH. Figures 239241. Male, metasoma and telson, lateral (239), ventral (240), and dorsal (241) views. Figures 242244. Female, metasoma and telson, lateral (242), ventral (243), and dorsal (244) views. Figures 245247: Barbaracurus yemenensis sp. n., Female holotype, metasoma and telson, lateral (245), dorsal (246), and ventral (247) views. Scale bars: 10 mm (239241, 242244, 245247)." figureDoi-0="http://doi.org/10.5281/zenodo.6544203" figureDoi-1="http://doi.org/10.5281/zenodo.6544205" httpUri-0="https://zenodo.org/record/6544203/files/figure.png" httpUri-1="https://zenodo.org/record/6544205/files/figure.png" pageId="34" pageNumber="33">226244</figureCitation>
,
<figureCitation id="6EDF2A4B8573FFCF0C1ED817FC4C4D1B" captionStart="Figures 255264" captionStartId="38.[192,278,1632,1656]" captionTargetBox="[191,1440,191,1603]" captionTargetId="figure-222@38.[192,1439,192,1603]" captionText="Figures 255264: Male postpachytene (255, 257, 261, 263), mitotic metaphase (259), and ideograms (256, 258, 260, 262, 264) (y axis - % of the diploid chromosome length) of Babycurus (255260) and Barbaracurus gen. n. (261264) species. Babycurus jacksoni (1189), Tanzania (2n=16) (255, 256), B. buettneri (1190) from Cameroon, Mamba vill., 04°07.311'N 10°11.615'E (2n=28) (257, 258) and B. gigas (914) from Tanzania (2n=30) (259, 260); Barbaracurus zambonelli comb. n. (876) from Eritrea, locality 15EH (2n=26) (261, 262) and B. somalicus comb. n. (1332), Somaliland, locality 17SR (2n=36) (263, 264). Scale bar: 5 µm (255, 257, 259, 261, 263)." figureDoi="http://doi.org/10.5281/zenodo.6544213" httpUri="https://zenodo.org/record/6544213/files/figure.png" pageId="34" pageNumber="33">258251, 258259</figureCitation>
), leg.
<collectorName id="5B1153188573FFCF0AC2D838FBBA4D1B" box="[992,1110,1639,1662]" pageId="34" pageNumber="33">F. Kovařík</collectorName>
,
<collectionCode id="90F5AE0B8573FFCF0D43D838FB4B4D1B" box="[1121,1191,1639,1662]" pageId="34" pageNumber="33">FKCP</collectionCode>
.
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</paragraph>
</subSubSection>
<subSubSection id="BEFE65458573FFC80A1CD8F8FBD94F8C" lastPageId="37" lastPageNumber="36" pageId="34" pageNumber="33" type="diagnosis">
<paragraph id="F65B36CE8573FFC80A1CD8F8FD714E98" blockId="34.[830,1441,1703,1910]" lastBlockId="37.[192,802,1112,1533]" lastPageId="37" lastPageNumber="36" pageId="34" pageNumber="33">
DIAGNOSIS. Total length of adult males
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, adult females
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. Coloration yellowish brown to grey with darker markings, chelicerae yellow without or with traces of reticulation. Pedipalp chela manus much wider in male than female, chela length/width ratio
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males and
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female; proximal margins of pedipalp fingers of female straight (
<figureCitation id="6EDF2A4B8573FFCF0D4CD900FB224C13" box="[1134,1230,1887,1910]" captionStart="Figures 5069" captionStartId="8.[192,278,1229,1253]" captionTargetBox="[195,1434,198,1190]" captionTargetId="figure-48@8.[192,1438,192,1200]" captionTargetPageId="8" captionText="Figures 5069: Pedipalp chela dorsal and external views, comparison of Barbaracurus gen. n. species. Figures 5053: B. sofomarensis comb. n., male (5051) and female (5253) paratypes. Figures 5457: B. somalicus comb. n., male (5455) and female (5657) from Somaliland, locality 17SR. Figures 5861: B. zambonellii comb. n., male (5859) and female (6061) from Eritrea, locality 15EH. Figures 6265: B. exquisitus comb. n., male holotype (6263) and female (6465) paratype. Figures 66 69: B. winklerorum sp. n., male holotype (6667) and female (6869) paratype." figureDoi="http://doi.org/10.5281/zenodo.6544167" httpUri="https://zenodo.org/record/6544167/files/figure.png" pageId="34" pageNumber="33">Figs. 61</figureCitation>
,
<figureCitation id="6EDF2A4B8573FFCF0DFBD900FAE84C13" box="[1241,1284,1887,1910]" captionStart="Figures 226238" captionStartId="32.[192,278,1770,1794]" captionTargetBox="[191,1440,191,1742]" captionTargetId="figure-142@32.[192,1439,192,1741]" captionTargetPageId="32" captionText="Figures 226238: Barbaracurus zambonellii comb. n. from Eritrea, locality 15EH. Figures 226, 228, 232238. Male, chelicerae, carapace and tergites IIII (226), sternopectinal region and sternite III (228), pedipalp chela, dorsoexternal (232), external (233), and ventrointernal (234) views, pedipalp patella, dorsal (235) and external (236) views, pedipalp femur and trochanter internal (237) and dorsal (238) views. Figures 227, 229231. Female, chelicerae, carapace and tergites IIII (227), sternopectinal region and sternites IIIIV (229), pedipalp chela, dorsoexternal (230) and external (231) views. The trichobothrial pattern is indicated in Figures 233238 (white circles)." figureDoi="http://doi.org/10.5281/zenodo.6544203" httpUri="https://zenodo.org/record/6544203/files/figure.png" pageId="34" pageNumber="33">231</figureCitation>
), of male undulate so as to leave a gap with fingers closed (
<figureCitation id="6EDF2A4B8574FFC80BE1DA07FCF14F0A" box="[707,797,1112,1135]" captionStart="Figures 5069" captionStartId="8.[192,278,1229,1253]" captionTargetBox="[195,1434,198,1190]" captionTargetId="figure-48@8.[192,1438,192,1200]" captionTargetPageId="8" captionText="Figures 5069: Pedipalp chela dorsal and external views, comparison of Barbaracurus gen. n. species. Figures 5053: B. sofomarensis comb. n., male (5051) and female (5253) paratypes. Figures 5457: B. somalicus comb. n., male (5455) and female (5657) from Somaliland, locality 17SR. Figures 5861: B. zambonellii comb. n., male (5859) and female (6061) from Eritrea, locality 15EH. Figures 6265: B. exquisitus comb. n., male holotype (6263) and female (6465) paratype. Figures 66 69: B. winklerorum sp. n., male holotype (6667) and female (6869) paratype." figureDoi="http://doi.org/10.5281/zenodo.6544167" httpUri="https://zenodo.org/record/6544167/files/figure.png" pageId="37" pageNumber="36">Figs. 59</figureCitation>
,
<figureCitation id="6EDF2A4B8574FFC809E3DA29FF044FE8" box="[193,232,1142,1165]" captionStart="Figures 226238" captionStartId="32.[192,278,1770,1794]" captionTargetBox="[191,1440,191,1742]" captionTargetId="figure-142@32.[192,1439,192,1741]" captionTargetPageId="32" captionText="Figures 226238: Barbaracurus zambonellii comb. n. from Eritrea, locality 15EH. Figures 226, 228, 232238. Male, chelicerae, carapace and tergites IIII (226), sternopectinal region and sternite III (228), pedipalp chela, dorsoexternal (232), external (233), and ventrointernal (234) views, pedipalp patella, dorsal (235) and external (236) views, pedipalp femur and trochanter internal (237) and dorsal (238) views. Figures 227, 229231. Female, chelicerae, carapace and tergites IIII (227), sternopectinal region and sternites IIIIV (229), pedipalp chela, dorsoexternal (230) and external (231) views. The trichobothrial pattern is indicated in Figures 233238 (white circles)." figureDoi="http://doi.org/10.5281/zenodo.6544203" httpUri="https://zenodo.org/record/6544203/files/figure.png" pageId="37" pageNumber="36">233</figureCitation>
); dentate margin of movable finger armed with 7 rows of granules, and a short apical row of 34 denticles (
<figureCitation id="6EDF2A4B8574FFC809EADAEBFEE14FAE" box="[200,269,1204,1227]" captionStart="Figures 123" captionStartId="4.[192,278,1544,1568]" captionTargetBox="[200,1432,168,1512]" captionTargetId="figure-16@4.[192,1440,163,1512]" captionTargetPageId="4" captionText="Figures 123: Pedipalp movable finger, comparison of Barbaracurus gen. n. (110) and Babycurus (1123) species. Figures 14: Barbaracurus gen. n., species with 6 non-imbricated almost linear rows of granules. Figure 1. B. ugartei comb. n., female holotype. Figure 2. B. sofomarensis comb. n., male paratype. Figure 3. B. somalicus comb. n., female from Somaliland, locality 17SR. Figure 4. B. subpunctatus comb. n., female from Ethiopia, locality 14EI. Figures 510: Barbaracurus gen. n., species with 7 non-imbricated almost linear rows of granules. Figure 5. B. exquisitus comb. n., male holotype. Figures 67, 10. B. zambonellii comb. n., female (6 and 10) and male (7) from Eritrea, locality 15EH. Figure 8. B. winklerorum sp. n., female paratype. Figure 9. B. yemenensis sp. n., female holotype. Figures 1214: Babycurus, species with 6 imbricated rows of movable fingers. Figure 12. B. ansorgei, female, Congo, Bukama env., FKCP. Figure 13. B. centrurimorphus, female lectotype. Figure 14. B. pictus, female, Congo, Sampwe env., FKCP. Figures 11, 1521: Babycurus, species with 78 imbricated rows of movable fingers. Figure 15. B. buettneri, female, MCSN. Figure 11, 16. B. dunlopi, male paratype. Figure 17. B. jacksoni, male, Tanzania, FKCP, No. 1189. Figure 18. B. kirki, female, Cote dIvoire, Comoe, FKCP. Figure 19. Babycurus melanicus, female holotype. Figure 20. B. taramassoi, female, Somalia, Belet Amin, MCSN. Figure 21. Babycurus wituensis, female lectotype. Figures 2223: Babycurus, species with 9 imbricated rows of movable fingers. Figure 22. B. gigas, female lectotype. Figure 23. B. multisubaculeatus, female holotype. Numbers in red indicate number of granule rows in series of fingers on the left of each number (red arrows)." figureDoi="http://doi.org/10.5281/zenodo.6544159" httpUri="https://zenodo.org/record/6544159/files/figure.png" pageId="37" pageNumber="36">Fig. 7</figureCitation>
); most proximal granule row with one external accessory granule. Pectines with 1719 teeth in both sexes. Hemispermatophore basal lobe a weak, oblique carina (
<figureCitation id="6EDF2A4B8574FFC80834DB4FFE484E42" box="[278,420,1296,1319]" captionStart="Figures 2439" captionStartId="6.[192,278,1598,1622]" captionTargetBox="[241,1392,199,1576]" captionTargetId="figure-16@6.[239,1405,192,1582]" captionTargetPageId="6" captionText="Figures 2439: Hemispermatophore capsules of Barbaracurus gen. n. and Babycurus species, convex and anterior views. In each figure. left image is convex view with capsule compressed to show shape of lobes, right image anterior view to show profile of basal lobe. Figures 2431. Color images. Figures 2428: Barbaracurus gen. n. Figure 24. B. winklerorum sp. n., holotype. Figure 25. B. exquisitus comb. n., Oman, Jabal Akhdar hotel, 23º04'N 57º38'E. Figure 26. B. sofomarensis comb. n., paratype from the type locality. Figure 27. B. somalicus comb. n., Somaliland, Borama, locality 17SR. Figure 28. B. zambonellii comb. n., Eritrea, Filfil, locality 15EH. Figures 2931: Babycurus. Figure 29: B. jacksoni, Tanzania. Figure 30: B. gigas, Tanzania, Usambara Mts. Figure 31: B. buettneri, Cameroon, Mamba vill., 04°07.311'N 10°11.615'E. Scale bars: 200 µm. Small numbered labels indicate specimen numbers (see lists of Material Examined). Figures 3239. Gray scale images with basal lobes outlined in red. Figures 3236: Barbaracurus gen. n.: B. winklerorum sp. n. (32), B. exquisitus comb. n. (33), B. sofomarensis comb. n. (34), B. somalicus comb. n. (35) and B. zambonellii comb. n. (36). Figures 3739: Babycurus: B. jacksoni (37), B. gigas (38) and B. buettneri (39). Solid red curves indicate either the distal margin of a scoop-like basal lobe (32, 34) or a well developed carina (3537, 39). Dashed red curves indicate either the line of attachment of a scoop-like basal lobe (32, 34) or a weakly developed carina (3536, 38)." figureDoi="http://doi.org/10.5281/zenodo.6544161" httpUri="https://zenodo.org/record/6544161/files/figure.png" pageId="37" pageNumber="36">Figs. 28, 36</figureCitation>
). Metasoma narrow, metasoma V length/width ratio is
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males and
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females (
<figureCitation id="6EDF2A4B8574FFC80867DB12FE244E01" box="[325,456,1357,1380]" captionStart="Figures 7080" captionStartId="9.[192,278,1644,1668]" captionTargetBox="[201,1439,170,1605]" captionTargetId="figure-188@9.[192,1439,168,1614]" captionTargetPageId="9" captionText="Figures 7080: Metasoma and telson ventral views, comparison of Barbaracurus gen. n. species. Figures 7071: B. exquisitus comb. n., male holotype (70) and female (71) paratype. Figures 7273: B. sofomarensis comb. n., male (72) and female (73) paratypes. Figures 7474: B. somalicus comb. n., male (74) and female (75) from Somaliland, locality 17SR. Figures 7677: B. winklerorum sp. n., male holotype (76) and female (77) paratype. Figures 78: B. yemenensis sp. n., female holotype. Figures 7980. B. zambonellii comb. n., male (79) and female (80) from Eritrea, locality 15EH." figureDoi="http://doi.org/10.5281/zenodo.6544169" httpUri="https://zenodo.org/record/6544169/files/figure.png" pageId="37" pageNumber="36">Figs. 7677</figureCitation>
); metasoma I with 10 carinae, IIIV with 8 carinae. Telson setose, bearing numerous long macrosetae and short, pointed subaculear tubercle; vesicle smooth, elongate, ellipsoidal, slightly bulbous, telson length/depth ratio
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both sexes; aculeus slender, curved, shorter than vesicle.
</paragraph>
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<paragraph id="F65B36CE8572FFCE09E2D975FF134C3E" blockId="35.[192,1440,1834,1883]" pageId="35" pageNumber="34">
<emphasis id="C490EADC8572FFCE09E2D975FE6A4C27" bold="true" box="[192,390,1834,1858]" pageId="35" pageNumber="34">Figures 248249:</emphasis>
<taxonomicName id="31E44D4D8572FFCE08ADD972FD634C27" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2018" baseAuthorityName="Borelli" baseAuthorityYear="1902" box="[399,655,1837,1858]" class="Arachnida" family="Buthidae" genus="Barbaracurus" kingdom="Animalia" order="Scorpiones" pageId="35" pageNumber="34" phylum="Arthropoda" rank="species" species="zambonellii" status="comb. nov.">
<emphasis id="C490EADC8572FFCE08ADD972FD634C27" box="[399,655,1837,1858]" italics="true" pageId="35" pageNumber="34">Barbaracurus zambonellii</emphasis>
</taxonomicName>
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<emphasis id="C490EADC8572FFCE0BBAD973FD3C4C24" bold="true" box="[664,720,1836,1857]" pageId="35" pageNumber="34">comb</emphasis>
.
<emphasis id="C490EADC8572FFCE0BFDD973FD004C24" bold="true" box="[735,748,1836,1857]" pageId="35" pageNumber="34">n</emphasis>
.
</taxonomicNameLabel>
, in vivo habitus. Male (248) and female (249) from Eritrea, locality 15EH.
</paragraph>
</caption>
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<paragraph id="F65B36CE8575FFC909E2D975FC394C3E" blockId="36.[192,1440,1834,1883]" pageId="36" pageNumber="35">
<emphasis id="C490EADC8575FFC909E2D975FE6F4C27" bold="true" box="[192,387,1834,1858]" pageId="36" pageNumber="35">Figures 250251:</emphasis>
<taxonomicName id="31E44D4D8575FFC908ABD972FD6A4C27" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2018" baseAuthorityName="Borelli" baseAuthorityYear="1902" box="[393,646,1837,1858]" class="Arachnida" family="Buthidae" genus="Barbaracurus" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="35" phylum="Arthropoda" rank="species" species="zambonellii" status="comb. nov.">
<emphasis id="C490EADC8575FFC908ABD972FD6A4C27" box="[393,646,1837,1858]" italics="true" pageId="36" pageNumber="35">Barbaracurus zambonellii</emphasis>
</taxonomicName>
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<emphasis id="C490EADC8575FFC90BAED973FD284C24" bold="true" box="[652,708,1836,1857]" pageId="36" pageNumber="35">comb</emphasis>
.
<emphasis id="C490EADC8575FFC90BF3D973FD324C24" bold="true" box="[721,734,1836,1857]" pageId="36" pageNumber="35">n</emphasis>
.
</taxonomicNameLabel>
female with newborns (first instar) before first ecdysis (250) and female with juveniles (second instar) after first ecdysis (251) from Eritrea, locality 15EH.
</paragraph>
</caption>
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<paragraph id="F65B36CE8574FFC809E2DDADFC274F46" blockId="37.[192,1440,1010,1059]" pageId="37" pageNumber="36">
<emphasis id="C490EADC8574FFC809E2DDADFE914F6F" bold="true" box="[192,381,1010,1034]" pageId="37" pageNumber="36">Figures 252254</emphasis>
:
<taxonomicName id="31E44D4D8574FFC808ACDDAAFD924F6F" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2018" baseAuthorityName="Lowe" baseAuthorityYear="2000" box="[398,638,1013,1034]" class="Arachnida" family="Buthidae" genus="Barbaracurus" kingdom="Animalia" order="Scorpiones" pageId="37" pageNumber="36" phylum="Arthropoda" rank="species" species="exquisitus" status="comb. nov.">
<emphasis id="C490EADC8574FFC808ACDDAAFD924F6F" box="[398,638,1013,1034]" italics="true" pageId="37" pageNumber="36">Barbaracurus exquisitus</emphasis>
</taxonomicName>
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<emphasis id="C490EADC8574FFC80BA5DDABFD534F6C" bold="true" box="[647,703,1012,1033]" pageId="37" pageNumber="36">comb</emphasis>
.
<emphasis id="C490EADC8574FFC80BECDDABFD374F6C" bold="true" box="[718,731,1012,1033]" pageId="37" pageNumber="36">n</emphasis>
.
</taxonomicNameLabel>
Right chelicera of male, dorsal (252), dorsointernal (253) and ventral (254) views. Paratype male, Jabal Shams, Oman, 14.X.1993. Scale bar: 500 µm.
</paragraph>
</caption>
<paragraph id="F65B36CE8574FFC809E2D875FBD94F8C" blockId="37.[192,802,1578,1907]" lastBlockId="37.[830,1442,1112,1258]" pageId="37" pageNumber="36">
NOTE. In his original description,
<bibRefCitation id="92754B3F8574FFC80B79D875FCCE4D24" author="BORELLI, A." box="[603,802,1578,1601]" journalOrPublisher="Bollettino dei Musei di Zoologia ed Anatomia Comparata della Reale Universita di Torino" pageId="37" pageNumber="36" pagination="1 - 4" part="17" refId="ref16152" refString="BORELLI, A. 1902. Di una nuova specie di scorpione della Colonia Eritrea. Bollettino dei Musei di Zoologia ed Anatomia Comparata della Reale Universita di Torino, 17 (422): 1 - 4." title="Di una nuova specie di scorpione della Colonia Eritrea" type="journal article" year="1902">Borelli (1902: 3)</bibRefCitation>
assumed that the
<typeStatus id="295F886C8574FFC808AFD817FE064D3A" box="[397,490,1608,1631]" pageId="37" pageNumber="36" type="holotype">holotype</typeStatus>
was a male, and this was accepted by most subsequent authors. Only
<bibRefCitation id="92754B3F8574FFC80BBAD838FECA4DF8" author="Lowe" firstAuthor="Lowe" page="190 - 191" pageId="37" pageNumber="36" pagination="185 - 192" refId="ref17483" refString="LOWE, G. 2000. A new species of Babycurus (Scorpiones: Buthidae) from northern Oman. Entomological News, 111: 185 - 192." type="journal article" year="2000">Lowe (2000: 190191)</bibRefCitation>
questioned this assumption, observing that the
<typeStatus id="295F886C8574FFC809E3D8FBFEF14DDE" box="[193,285,1700,1723]" pageId="37" pageNumber="36" type="holotype">holotype</typeStatus>
exhibited some characters more consistent with females of other species of
<taxonomicName id="31E44D4D8574FFC808DFD89CFD834DBF" authorityName=": Kraepelin" authorityYear="1913" box="[509,623,1731,1754]" class="Arachnida" family="Buthidae" genus="Babycurus" kingdom="Animalia" order="Scorpiones" pageId="37" pageNumber="36" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="C490EADC8574FFC808DFD89CFD834DBF" box="[509,623,1731,1754]" italics="true" pageId="37" pageNumber="36">Babycurus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DFA357A78574FFC80B58D89CFD054DBF" box="[634,745,1731,1754]" pageId="37" pageNumber="36" sensu="lato">sensu lato</taxonomicNameLabel>
, and noted that “Study of additional material is needed to clarify variation and sexual dimorphism in
<taxonomicName id="31E44D4D8574FFC80BE5D95FFEFD4C53" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2018" baseAuthorityName="Borelli" baseAuthorityYear="1902" class="Arachnida" family="Buthidae" genus="Barbaracurus" kingdom="Animalia" order="Scorpiones" pageId="37" pageNumber="36" phylum="Arthropoda" rank="species" species="zambonellii">
<emphasis id="C490EADC8574FFC80BE5D95FFEFD4C53" italics="true" pageId="37" pageNumber="36">B. zambonellii</emphasis>
</taxonomicName>
”. Recently, the first author (F.K.) was able to finally settle this question by collecting additional material representing both sexes of
<taxonomicName id="31E44D4D8574FFC80B69D903FD044C16" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2018" baseAuthorityName="Borelli" baseAuthorityYear="1902" box="[587,744,1884,1907]" class="Arachnida" family="Buthidae" genus="Barbaracurus" kingdom="Animalia" order="Scorpiones" pageId="37" pageNumber="36" phylum="Arthropoda" rank="species" species="zambonellii">
<emphasis id="C490EADC8574FFC80B69D903FD044C16" box="[587,744,1884,1907]" italics="true" pageId="37" pageNumber="36">B. zambonellii</emphasis>
</taxonomicName>
. The new specimens clearly demonstrated that the
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is indeed female, not male.
<taxonomicName id="31E44D4D8574FFC80D7CDA29FB124FE8" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2018" baseAuthorityName="Borelli" baseAuthorityYear="1902" box="[1118,1278,1142,1165]" class="Arachnida" family="Buthidae" genus="Barbaracurus" kingdom="Animalia" order="Scorpiones" pageId="37" pageNumber="36" phylum="Arthropoda" rank="species" species="zambonellii">
<emphasis id="C490EADC8574FFC80D7CDA29FB124FE8" box="[1118,1278,1142,1165]" italics="true" pageId="37" pageNumber="36">B. zambonellii</emphasis>
</taxonomicName>
is an Eritrean endemic and the female which
<bibRefCitation id="92754B3F8574FFC80DACDACAFAC64FC9" author="Sissom" box="[1166,1322,1173,1196]" firstAuthor="Sissom" pageId="37" pageNumber="36" refId="ref18158" refString="SISSOM, W. D. 1994. Descriptions of new and poorly known scorpions of Yemen (Scorpiones: Buthidae," type="book" year="1994">Sissom (1994)</bibRefCitation>
cited from
<collectingCountry id="8EF3765E8574FFC80A1DDAEBFC644FAE" box="[831,904,1204,1227]" name="Yemen" pageId="37" pageNumber="36">Yemen</collectingCountry>
represents a new species which we describe here as
<taxonomicName id="31E44D4D8574FFC80A79DA8DFC1E4F8C" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2018" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2018" box="[859,1010,1234,1257]" class="Arachnida" family="Buthidae" genus="Barbaracurus" kingdom="Animalia" order="Scorpiones" pageId="37" pageNumber="36" phylum="Arthropoda" rank="species" species="yemenensis" status="sp. nov.">
<emphasis id="C490EADC8574FFC80A79DA8DFC1E4F8C" box="[859,1010,1234,1257]" italics="true" pageId="37" pageNumber="36">B. yemenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DFA357A78574FFC80ADBDA8DFBD94F8C" box="[1017,1077,1234,1258]" pageId="37" pageNumber="36" rank="species">
<emphasis id="C490EADC8574FFC80ADBDA8DFBFE4F8F" bold="true" box="[1017,1042,1234,1258]" pageId="37" pageNumber="36">sp</emphasis>
.
<emphasis id="C490EADC8574FFC80D02DA8DFBC34F8F" bold="true" box="[1056,1071,1234,1258]" pageId="37" pageNumber="36">n</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="BEFE65458574FFC80A1CDB4FFC314D5E" pageId="37" pageNumber="36" type="discussion">
<paragraph id="F65B36CE8574FFC80A1CDB4FFC314D5E" blockId="37.[830,1441,1296,1595]" pageId="37" pageNumber="36">
COMMENTS ON LOCALITY AND LIFE STRATEGY. The first author (F.K.) visited the locality 15EH (
<figureCitation id="6EDF2A4B8574FFC80DD7DB71FA784E20" box="[1269,1428,1326,1349]" captionStart="Figures 219225" captionStartId="31.[192,278,1640,1664]" captionTargetBox="[191,1440,191,1611]" captionTargetId="figure-244@31.[192,1439,192,1610]" captionTargetPageId="31" captionText="Figures 219225: Barbaracurus zambonellii comb. n. from Eritrea, locality 15EH. Figures 219, 221. Male, dorsal (219) and ventral (221) views. Figures 220, 222. Female, dorsal (220) and ventral (222) views. Figure 223. Right hemispermatophore, convex aspect. Figures 224225. Eritrea, locality 15EH. Scale bars: 10 mm (219222), 1 mm (223)." figureDoi="http://doi.org/10.5281/zenodo.6544201" httpUri="https://zenodo.org/record/6544201/files/figure.png" pageId="37" pageNumber="36">Figs. 224225</figureCitation>
), a montane, forested habitat along a riverbed of an occasional river, on
<date id="825A100E8574FFC80D08DB33FAFA4EE6" box="[1066,1302,1388,1411]" pageId="37" pageNumber="36" value="2015-11-03" valueMax="2015-11-04" valueMin="2015-11-03">34 November 2015</date>
. At this locality, the author recorded a maximum daytime temperature of 30.3 ºC, and minimum nighttime temperatures of 19.6 ºC. The recorded humidity was between 46% and 92%. In addition to
<taxonomicName id="31E44D4D8574FFC80DB9DBB9FAD74E98" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2018" baseAuthorityName="Borelli" baseAuthorityYear="1902" box="[1179,1339,1510,1533]" class="Arachnida" family="Buthidae" genus="Barbaracurus" kingdom="Animalia" order="Scorpiones" pageId="37" pageNumber="36" phylum="Arthropoda" rank="species" species="zambonellii">
<emphasis id="C490EADC8574FFC80DB9DBB9FAD74E98" box="[1179,1339,1510,1533]" italics="true" pageId="37" pageNumber="36">B. zambonellii</emphasis>
</taxonomicName>
the first author also recorded
<taxonomicName id="31E44D4D8574FFC80D05D85AFA4C4D79" authority="(L. Koch, 1875)" baseAuthorityName="L. Koch" baseAuthorityYear="1875" box="[1063,1440,1541,1564]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="37" pageNumber="36" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="C490EADC8574FFC80D05D85AFB044D79" box="[1063,1256,1541,1564]" italics="true" pageId="37" pageNumber="36">Hottentotta minax</emphasis>
(L. Koch, 1875)
</taxonomicName>
at this locality.
</paragraph>
</subSubSection>
</treatment>
</document>