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<document id="B52A737D75CB892057FC72824F2B42DE" ID-DOI="10.24199/j.mmv.2012.69.05" ID-ISSN="1447-2554" ID-Zenodo-Dep="12212378" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718994422343" checkinUser="felipe" docAuthor="OLoughlin, P. Mark, Barmos, Shari &amp; VandenSpiegel, Didier" docDate="2012" docId="7E4A044D1936FF8F9A9DFCE590198A4E" docLanguage="en" docName="MemMusVictoria.69.269-308.pdf" docOrigin="Memoirs of Museum Victoria 69" docSource="https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-69-2012/pages-269-308/" docStyle="DocumentStyle:5B3B5C052A93BFEEE334AD6097A94E45.2:MemMusVictoria.2003-.journal_article.type1" docStyleId="5B3B5C052A93BFEEE334AD6097A94E45" docStyleName="MemMusVictoria.2003-.journal_article.type1" docStyleVersion="2" docTitle="Thyone okeni Bell. We 1884" docType="treatment" docVersion="2" lastPageNumber="303" masterDocId="82737C351914FFAD9A11FFCF956F8E08" masterDocTitle="The phyllophorid sea cucumbers of southern Australia (Echinodermata: Holothuroidea: Dendrochirotida: Phyllophoridae)" masterLastPageNumber="308" masterPageNumber="269" pageNumber="303" updateTime="1719008797983" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="8B6EE2249BBECEA54FA6CDEA1C85C223">The phyllophorid sea cucumbers of southern Australia (Echinodermata: Holothuroidea: Dendrochirotida: Phyllophoridae)</mods:title>
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<mods:namePart id="19C93A8BE82483DC7FE472014CA2A5E8">OLoughlin, P. Mark</mods:namePart>
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<mods:namePart id="55AFE6E1063F2942EA6A27AFD3F756E9">Barmos, Shari</mods:namePart>
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<mods:namePart id="40F34A6DEDE0FE3B610734F67CA6623F">VandenSpiegel, Didier</mods:namePart>
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<treatment id="7E4A044D1936FF8F9A9DFCE590198A4E" ID-DOI="http://doi.org/10.5281/zenodo.12212493" ID-Zenodo-Dep="12212493" LSID="urn:lsid:plazi:treatment:7E4A044D1936FF8F9A9DFCE590198A4E" httpUri="http://treatment.plazi.org/id/7E4A044D1936FF8F9A9DFCE590198A4E" lastPageNumber="303" pageId="34" pageNumber="303">
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<taxonomicName id="31E3CED81936FF8F9A9DFCE594E88D37" ID-CoL="56RMZ" authority="Bell, 1884" authorityName="Bell. We" authorityYear="1884" box="[140,391,810,832]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="34" pageNumber="303" phylum="Echinodermata" rank="species" species="okeni">
<emphasis id="C49769491936FF8F9A9DFCE594758D48" bold="true" box="[140,282,810,832]" italics="true" pageId="34" pageNumber="303">Thyone okeni</emphasis>
<bibRefCitation id="9272C8AA1936FF8F9B31FCE594E88D37" author="Bell, F. J." box="[288,391,810,832]" pageId="34" pageNumber="303" refId="ref22242" refString="Bell, F. J. 1884. Echinodermata. Pp. 117 - 177 in Report on the Zoological Collections made in the Indo-Pacific Ocean during the Voyage of H. M. S. Alert 1881 - 2. Taylor and Francis: London." type="book" year="1884">Bell, 1884</bibRefCitation>
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<figureCitation id="6ED8A9DE1936FF8F9A9DFC99946C8D64" box="[140,259,854,876]" captionStart="Figure 17" captionStartId="27.[147,206,1778,1797]" captionTargetBox="[147,1447,227,1701]" captionTargetId="figure-11@27.[147,1447,227,1701]" captionTargetPageId="27" captionText="Figure 17. Photos of preserved specimens. a, holotype of Thyone flindersi OLoughlin sp. nov. (NMV F151847) (insert: form of the calcareous ring plates); b, specimen of Thyone okeni Bell, 1884 (AM J10868); c, holotype of Thyone joshuai OLoughlin sp. nov. (SAM K2566) (insert: form of the calcareous ring plates); d, holotype of Thyone kerkosa OLoughlin sp. nov. (WAM Z31838); e, holotype of Thyone spenceri OLoughlin sp. nov. (SAM K2562) (insert: form of the calcareous ring plates); f, holotype of Thyone tourvillei OLoughlin sp. nov. (NMV F174902) (insert: form of the calcareous ring plates)." figureDoi="http://doi.org/10.5281/zenodo.12212414" httpUri="https://zenodo.org/record/12212414/files/figure.png" pageId="34" pageNumber="303">Figures 17b</figureCitation>
,
<figureCitation id="6ED8A9DE1936FF8F9B1FFC9994488D63" box="[270,295,854,875]" captionStart="Figure 22" captionStartId="35.[147,206,1857,1876]" captionTargetBox="[147,1447,227,1699]" captionTargetId="figure-11@35.[147,1447,227,1699]" captionTargetPageId="35" captionText="Figure 22. SEM images of ossicles from specimen of Thyone okeni Bell, 1884 (AM J7731). Peri-oral body wall rosette and tables with multibranched spires (top); dumbbell-shaped ossicles from tentacles (bottom)." figureDoi="http://doi.org/10.5281/zenodo.12212424" httpUri="https://zenodo.org/record/12212424/files/figure.png" pageId="34" pageNumber="303">22</figureCitation>
</paragraph>
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<treatmentCitation id="7742934A1936FF8F9AA3FC4E949A8D9C" author="Bell, F. J." box="[178,501,897,916]" page="150" pageId="34" pageNumber="303" year="1884">
<taxonomicName id="31E3CED81936FF8F9AA3FC4E949A8D9C" ID-CoL="56RMZ" authority="Bell, 1884: 149 - 150" authorityName="Bell. We" authorityPageNumber="149 - 150" authorityYear="1884" box="[178,501,897,916]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="34" pageNumber="303" phylum="Echinodermata" rank="species" species="okeni">
<emphasis id="C49769491936FF8F9AA3FC4E945F8D9C" box="[178,304,897,916]" italics="true" pageId="34" pageNumber="303">Thyone okeni</emphasis>
<bibRefCitation id="9272C8AA1936FF8F9B28FC4E949A8D9C" author="Bell, F. J." box="[313,501,897,916]" pageId="34" pageNumber="303" refId="ref22242" refString="Bell, F. J. 1884. Echinodermata. Pp. 117 - 177 in Report on the Zoological Collections made in the Indo-Pacific Ocean during the Voyage of H. M. S. Alert 1881 - 2. Taylor and Francis: London." type="book" year="1884">Bell, 1884: 149150</bibRefCitation>
</taxonomicName>
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, pl. 9D.—H. L.
<treatmentCitation id="7742934A1936FF8F9880FC4E95C38DA6" author="Clark, H. L." page="167" pageId="34" pageNumber="303" year="1921">
<bibRefCitation id="9272C8AA1936FF8F9880FC4E95C38DA6" author="Clark, H. L." pageId="34" pageNumber="303" pagination="1 - 223" refId="ref22387" refString="Clark, H. L. 1921. The echinoderm fauna of Torres Strait: its composition and its origin. Publication of the Carnegie Institution of Washington No. 214: i - viii, 1 - 223 pp, 38 pls." type="book chapter" year="1921">Clark, 1921: 167</bibRefCitation>
</treatmentCitation>
.—H. L.
<treatmentCitation id="7742934A1936FF8F9AE9FC5494E08DA6" author="Clark, H. L." box="[248,399,923,942]" page="402" pageId="34" pageNumber="303" year="1946">
<bibRefCitation id="9272C8AA1936FF8F9AE9FC5494E08DA6" author="Clark, H. L." box="[248,399,923,942]" pageId="34" pageNumber="303" refId="ref22469" refString="Clark, H. L. 1946. The Echinoderm Fauna of Australia: Its composition and origin. Publications Carnegie Institute 566. 567 pp." type="book" year="1946">Clark, 1946: 402</bibRefCitation>
</treatmentCitation>
.—
<treatmentCitation id="7742934A1936FF8F9BB8FC5497508DA6" author="Rowe, F. W. E." box="[425,575,923,942]" page="462" pageId="34" pageNumber="303" year="1982">
<bibRefCitation id="9272C8AA1936FF8F9BB8FC5497508DA6" author="Rowe, F. W. E." box="[425,575,923,942]" pageId="34" pageNumber="303" pagination="454 - 476" refId="ref23103" refString="Rowe, F. W. E. 1982. Sea-cucumbers (class Holothurioidea). In Shepherd, S. A. and Thomas, I. M. (eds), Marine Invertebrates of Southern Australia 1: 454 - 476, figs 10: 26 - 10: 37, pls 29 - 32. Adelaide, South Australian Government Printer." type="journal article" year="1982">Rowe, 1982: 462</bibRefCitation>
</treatmentCitation>
(non specimens from Port Phillip and Westernport in
<collectingRegion id="34277BB91936FF8F9BAEFC7A97668DC0" box="[447,521,949,968]" country="Australia" name="Victoria" pageId="34" pageNumber="303">Victoria</collectingRegion>
).—
<treatmentCitation id="7742934A1936FF8F9838FC7A95C48DEA" author="Rowe, F. W. E. &amp; Gates, J." page="316" pageId="34" pageNumber="303" year="1995">
<bibRefCitation id="9272C8AA1936FF8F9838FC7A95C48DEA" author="Rowe, F. W. E. &amp; Gates, J." pageId="34" pageNumber="303" pagination="1 - 510" refId="ref23199" refString="Rowe, F. W. E. and Gates, J. 1995. Echinodermata. In Wells, A. (ed.), Zoological Catalogue of Australia 33: i - xiii, 1 - 510. CSIRO, Melbourne." type="journal article" year="1995">Rowe and Gates, 1995: 316</bibRefCitation>
</treatmentCitation>
(non specimens from southern
<collectingCountry id="8EF4F5CB1936FF8F9BD8FC0097708DEA" box="[457,543,975,994]" name="Australia" pageId="34" pageNumber="303">Australia</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
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<emphasis id="C49769491936FF8F9A9DFC3794538A03" box="[140,316,1016,1035]" italics="true" pageId="34" pageNumber="303">Material examined</emphasis>
.
<materialsCitation id="468BBF061936FF8F9B58FC3795848A2E" collectionCode="AM" country="Australia" location="Port Jackson" pageId="34" pageNumber="303" specimenCode="J16905" specimenCount="1" stateProvince="New South Wales">
<collectingCountry id="8EF4F5CB1936FF8F9B58FC3794F08A03" box="[329,415,1016,1035]" name="Australia" pageId="34" pageNumber="303">Australia</collectingCountry>
,
<collectingRegion id="34277BB91936FF8F9BBBFC36973E8A03" box="[426,593,1016,1036]" country="Australia" name="New South Wales" pageId="34" pageNumber="303">New South Wales</collectingRegion>
,
<location id="F33CE3801936FF8F984DFC3697BC8A03" LSID="urn:lsid:plazi:treatment:7E4A044D1936FF8F9A9DFCE590198A4E:F33CE3801936FF8F984DFC3697BC8A03" box="[604,723,1016,1036]" country="Australia" name="Port Jackson" pageId="34" pageNumber="303" stateProvince="New South Wales">Port Jackson</location>
,
<collectionCode id="90F22D9E1936FF8F98CFFC36966C8A04" box="[734,771,1017,1036]" country="Australia" name="Australian Museum" pageId="34" pageNumber="303" type="Museum">AM</collectionCode>
<specimenCode id="A6451D201936FF8F9A9DFBDC95A48A2E" box="[140,203,1043,1062]" collectionCode="AM" country="Australia" name="Australian Museum" pageId="34" pageNumber="303" type="Museum">J16905</specimenCode>
(1)
</materialsCitation>
;
<materialsCitation id="468BBF061936FF8F9AE4FBDC942F8A48" collectingDate="1968-11-21" collectionCode="AM" country="Australia" county="Vaucluse" location="Glass Rock" municipality="Bottle" pageId="34" pageNumber="303" specimenCode="J7731" specimenCount="1" stateProvince="New South Wales">
<collectingCounty id="1F3DCDD71936FF8F9AE4FBDC94278A2E" box="[245,328,1043,1062]" pageId="34" pageNumber="303">Vaucluse</collectingCounty>
,
<collectingMunicipality id="16382F211936FF8F9B42FBDC94E48A2E" box="[339,395,1043,1062]" pageId="34" pageNumber="303">Bottle</collectingMunicipality>
and
<location id="F33CE3801936FF8F9BA9FBDC974D8A2E" LSID="urn:lsid:plazi:treatment:7E4A044D1936FF8F9A9DFCE590198A4E:F33CE3801936FF8F9BA9FBDC974D8A2E" box="[440,546,1043,1062]" country="Australia" county="Vaucluse" municipality="Bottle" name="Glass Rock" pageId="34" pageNumber="303" stateProvince="New South Wales">Glass Rock</location>
, under boulders,
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<collectingDate id="92196A731936FF8F98D1FBDC95D58A48" pageId="34" pageNumber="303" value="1968-11-21">21 Nov 1968</collectingDate>
</date>
,
<collectionCode id="90F22D9E1936FF8F9AD2FBE295878A48" box="[195,232,1069,1088]" country="Australia" name="Australian Museum" pageId="34" pageNumber="303" type="Museum">AM</collectionCode>
<specimenCode id="A6451D201936FF8F9AFCFBE2944D8A48" box="[237,290,1069,1088]" collectionCode="AM" country="Australia" name="Australian Museum" pageId="34" pageNumber="303" type="Museum">J7731</specimenCode>
(1)
</materialsCitation>
;
<materialsCitation id="468BBF061936FF8F9B5BFBE297118A48" box="[330,638,1069,1088]" collectingDate="1977-08-29" collectionCode="AM" country="Australia" elevation="5" pageId="34" pageNumber="303" specimenCode="J10868" specimenCount="1" stateProvince="New South Wales">
<quantity id="311B18BE1936FF8F9B5BFBE294018A48" box="[330,366,1069,1088]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="34" pageNumber="303" unit="m" value="5.0">
<elevation id="7DCE52681936FF8F9B5BFBE294018A48" box="[330,366,1069,1088]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="34" pageNumber="303" unit="m" value="5.0">5 m</elevation>
</quantity>
,
<date id="825D939B1936FF8F9B66FBE294848A48" box="[375,491,1069,1088]" pageId="34" pageNumber="303" value="1977-08-29">
<collectingDate id="92196A731936FF8F9B66FBE294848A48" box="[375,491,1069,1088]" pageId="34" pageNumber="303" value="1977-08-29">29 Aug 1977</collectingDate>
</date>
,
<collectionCode id="90F22D9E1936FF8F9BE5FBE297768A48" box="[500,537,1069,1088]" country="Australia" name="Australian Museum" pageId="34" pageNumber="303" type="Museum">AM</collectionCode>
<specimenCode id="A6451D201936FF8F980FFBE297318A48" box="[542,606,1069,1088]" collectionCode="AM" country="Australia" name="Australian Museum" pageId="34" pageNumber="303" type="Museum">J10868</specimenCode>
(2)
</materialsCitation>
;
<materialsCitation id="468BBF061936FF8F9896FBE294828A52" collectingDate="1968-10-23" collectionCode="NMV" country="Australia" pageId="34" pageNumber="303" specimenCode="F174900" specimenCount="1" stateProvince="New South Wales">
mid-tide rock pools,
<date id="825D939B1936FF8F9AD9FB8894558A52" box="[200,314,1095,1114]" pageId="34" pageNumber="303" value="1968-10-23">
<collectingDate id="92196A731936FF8F9AD9FB8894558A52" box="[200,314,1095,1114]" pageId="34" pageNumber="303" value="1968-10-23">23 Oct 1968</collectingDate>
</date>
,
<collectionCode id="90F22D9E1936FF8F9B55FB8894158A52" box="[324,378,1095,1114]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="34" pageNumber="303" type="Museum">NMV</collectionCode>
<specimenCode id="A6451D201936FF8F9B6EFB8894A18A52" box="[383,462,1095,1114]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="34" pageNumber="303" type="Museum">F174900</specimenCode>
(1)
</materialsCitation>
;
<materialsCitation id="468BBF061936FF8F9BE6FB8894008A7C" collectingDate="1951-01-21" collectionCode="AM" country="Australia" location="Nielsen Park" pageId="34" pageNumber="303" specimenCode="J16869" specimenCount="1" stateProvince="New South Wales">
off
<location id="F33CE3801936FF8F9809FB8897E18A52" LSID="urn:lsid:plazi:treatment:7E4A044D1936FF8F9A9DFCE590198A4E:F33CE3801936FF8F9809FB8897E18A52" box="[536,654,1095,1114]" country="Australia" name="Nielsen Park" pageId="34" pageNumber="303" stateProvince="New South Wales">Nielsen Park</location>
, dredged,
<date id="825D939B1936FF8F98FCFB8895B38A7C" pageId="34" pageNumber="303" value="1951-01-21">
<collectingDate id="92196A731936FF8F98FCFB8895B38A7C" pageId="34" pageNumber="303" value="1951-01-21">21 Jan 1951</collectingDate>
</date>
,
<collectionCode id="90F22D9E1936FF8F9AF7FBAE94648A7C" box="[230,267,1121,1140]" country="Australia" name="Australian Museum" pageId="34" pageNumber="303" type="Museum">AM</collectionCode>
<specimenCode id="A6451D201936FF8F9B00FBAE94208A7C" box="[273,335,1121,1140]" collectionCode="AM" country="Australia" name="Australian Museum" pageId="34" pageNumber="303" type="Museum">J16869</specimenCode>
(1)
</materialsCitation>
;
<materialsCitation id="468BBF061936FF8F9B6BFBAE94438A86" collectingDate="1987-05-21" collectionCode="AM" country="Australia" elevation="5" location="Swansea Channel" municipality="Newcastle" pageId="34" pageNumber="303" specimenCode="J20267" specimenCount="1" stateProvince="New South Wales">
<collectingMunicipality id="16382F211936FF8F9B6BFBAE94B68A7C" box="[378,473,1121,1140]" pageId="34" pageNumber="303">Newcastle</collectingMunicipality>
,
<location id="F33CE3801936FF8F9BF3FBAE97EA8A7C" LSID="urn:lsid:plazi:treatment:7E4A044D1936FF8F9A9DFCE590198A4E:F33CE3801936FF8F9BF3FBAE97EA8A7C" box="[482,645,1121,1140]" country="Australia" municipality="Newcastle" name="Swansea Channel" pageId="34" pageNumber="303" stateProvince="New South Wales">Swansea Channel</location>
,
<quantity id="311B18BE1936FF8F989EFBAE97DB8A7C" box="[655,692,1121,1140]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="34" pageNumber="303" unit="m" value="5.0">
<elevation id="7DCE52681936FF8F989EFBAE97DB8A7C" box="[655,692,1121,1140]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="34" pageNumber="303" unit="m" value="5.0">5 m</elevation>
</quantity>
,
<date id="825D939B1936FF8F98AFFBAE95D78A86" pageId="34" pageNumber="303" value="1987-05-21">
<collectingDate id="92196A731936FF8F98AFFBAE95D78A86" pageId="34" pageNumber="303" value="1987-05-21">21 May 1987</collectingDate>
</date>
,
<collectionCode id="90F22D9E1936FF8F9AD0FBB495898A86" box="[193,230,1147,1166]" country="Australia" name="Australian Museum" pageId="34" pageNumber="303" type="Museum">AM</collectionCode>
<specimenCode id="A6451D201936FF8F9AFAFBB494438A86" box="[235,300,1147,1166]" collectionCode="AM" country="Australia" name="Australian Museum" pageId="34" pageNumber="303" type="Museum">J20267</specimenCode>
</materialsCitation>
(1).
</paragraph>
<paragraph id="F65CB55B1936FF8F9A9DFB69941A8864" blockId="34.[140,771,1189,1903]" pageId="34" pageNumber="303">
<emphasis id="C49769491936FF8F9A9DFB69959D8AB3" box="[140,242,1190,1211]" italics="true" pageId="34" pageNumber="303">Diagnosis</emphasis>
.
<taxonomicName id="31E3CED81936FF8F9B13FB6A94CF8AB3" box="[258,416,1189,1211]" pageId="34" pageNumber="303">
<emphasis id="C49769491936FF8F9B13FB6A94228AB2" box="[258,333,1189,1210]" italics="true" pageId="34" pageNumber="303">Thyone</emphasis>
species
</taxonomicName>
up to
<quantity id="311B18BE1936FF8F9BF8FB6A975C8AB3" box="[489,563,1189,1211]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="34" pageNumber="303" unit="mm" value="75.0">75 mm</quantity>
long, up to
<quantity id="311B18BE1936FF8F98A9FB69966C8AB3" box="[696,771,1190,1211]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="34" pageNumber="303" unit="mm" value="20.0">20 mm</quantity>
diameter (preserved, tentacle withdrawn;
<quantity id="311B18BE1936FF8F9824FB0D97128AD0" box="[565,637,1218,1240]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" pageId="34" pageNumber="303" unit="mm" value="85.0">85 mm</quantity>
long by
<bibRefCitation id="9272C8AA1936FF8F98C8FB0D95AF8AFC" author="Bell, F. J." pageId="34" pageNumber="303" refId="ref22242" refString="Bell, F. J. 1884. Echinodermata. Pp. 117 - 177 in Report on the Zoological Collections made in the Indo-Pacific Ocean during the Voyage of H. M. S. Alert 1881 - 2. Taylor and Francis: London." type="book" year="1884">Bell 1884</bibRefCitation>
); body cylindrical, oral and anal ends tapered, upturned; typically 10 tentacles, 8 large, 2 small ventral; body closely covered with tube feet, about
<quantity id="311B18BE1936FF8F9BACFAD697558B26" box="[445,570,1305,1326]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="34" pageNumber="303" unit="mm" value="0.35" valueMax="0.4" valueMin="0.3">0.30.4 mm</quantity>
diameter, not more concentrated radially; no external evidence of anal scales or papillae; calcareous ring composite, tubular, no evidence of discrete plates for most of length, posterior composite tails not joined distally; radial plates with split anterior point, posterior end divided into two thin composite tails posterior to the end of the inter-radial plates; inter-radial plates tapered to anterior point, truncate posteriorly, about two thirds the length of radial plates; single long tubular polian vesicle; stone canal and madreporite lie anteriorly from attachment along calcareous ring; longitudinal muscles cylindrical, distinctly divided by deep groove mid-body.
</paragraph>
<paragraph id="F65CB55B1936FF8F9AA3F9BC90E18F3D" blockId="34.[140,771,1189,1903]" lastBlockId="34.[809,1440,229,309]" pageId="34" pageNumber="303">
Mid-body wall lacking ossicles; tube feet with endplates only, up to 240
<emphasis id="C49769491936FF8F9B36F95F945A88AD" box="[295,309,1680,1701]" italics="true" pageId="34" pageNumber="303">µ</emphasis>
m diameter lacking any support rods or tables or plates, endplate with irregular perforations, large marginally small centrally, slightly denticulate margin; peri-oral body wall with numerous tables, discs round to slightly oval, up to 192
<emphasis id="C49769491936FF8F9AA6F8CC95AA8910" box="[183,197,1795,1816]" italics="true" pageId="34" pageNumber="303">µ</emphasis>
m diameter/length, up to&gt; 100 small perforations, spires thick short columnar mesh; tentacles branches with stout dumbbell-shaped ossicles, few to about 12 distal perforations, rods up to 320
<emphasis id="C49769491936FF8F9B3EF89594528967" box="[303,317,1882,1903]" italics="true" pageId="34" pageNumber="303">µ</emphasis>
m long, tentacle trunks lacking ossicles, no rosettes seen; introvert ossicles rosettes; anal ossicles rods as in tentacles, rosettes as in introvert, single and multi-layered scale fragments, tube foot endplate diameters up to 200
<emphasis id="C49769491936FF8F9F74FEEF901C8F3D" box="[1381,1395,288,309]" italics="true" pageId="34" pageNumber="303">µ</emphasis>
m.
</paragraph>
<paragraph id="F65CB55B1936FF8F9938FE8490598F93" blockId="34.[809,1440,331,411]" pageId="34" pageNumber="303">
<emphasis id="C49769491936FF8F9938FE8496868F68" box="[809,1001,331,352]" italics="true" pageId="34" pageNumber="303">Colour (preserved)</emphasis>
. Body brown to dark brown to grey-brown; tube feet off-white to grey to speckled to red; tentacle trunks off-white with brown patches, branches dark brown.
</paragraph>
<paragraph id="F65CB55B1936FF8F9938FE7D91928FCF" blockId="34.[809,1277,433,455]" box="[809,1277,433,455]" pageId="34" pageNumber="303">
<materialsCitation id="468BBF061936FF8F9938FE7D91958FCF" box="[809,1274,433,455]" location="Port Jackson" pageId="34" pageNumber="303" specimenCount="1" stateProvince="New South Wales" typeStatus="holotype">
<emphasis id="C49769491936FF8F9938FE7D96DC8FCE" box="[809,947,433,455]" italics="true" pageId="34" pageNumber="303">
<typeStatus id="29580BF91936FF8F9938FE7D96378FCF" box="[809,856,434,455]" pageId="34" pageNumber="303">Type</typeStatus>
locality.
</emphasis>
<collectingRegion id="34277BB91936FF8F99A9FE7D91028FCF" box="[952,1133,433,455]" country="Australia" name="New South Wales" pageId="34" pageNumber="303">New South Wales</collectingRegion>
,
<location id="F33CE3801936FF8F9E66FE7D91958FCF" LSID="urn:lsid:plazi:treatment:7E4A044D1936FF8F9A9DFCE590198A4E:F33CE3801936FF8F9E66FE7D91958FCF" box="[1143,1274,433,455]" name="Port Jackson" pageId="34" pageNumber="303" stateProvince="New South Wales">Port Jackson</location>
</materialsCitation>
.
</paragraph>
<paragraph id="F65CB55B1936FF8F9938FE12961E8C18" blockId="34.[809,1440,477,528]" pageId="34" pageNumber="303">
<emphasis id="C49769491936FF8F9938FE1296C98FFA" box="[809,934,477,498]" italics="true" pageId="34" pageNumber="303">Distribution</emphasis>
.
<collectingRegion id="34277BB91936FF8F99A9FE1191198FFB" box="[952,1142,477,499]" country="Australia" name="New South Wales" pageId="34" pageNumber="303">New South Wales</collectingRegion>
, Port Jackson to Newcastle;
<quantity id="311B18BE1936FF8F9938FE3596038C18" box="[809,876,506,528]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" pageId="34" pageNumber="303" unit="m" value="2.5" valueMax="5.0" valueMin="0.0">
0
<specimenCount id="E0E57ED21936FF8F9954FE3596038C18" box="[837,876,506,528]" count="5" pageId="34" pageNumber="303" type="generic">5 m</specimenCount>
</quantity>
.
</paragraph>
<paragraph id="F65CB55B1936FF8F9938FDE990198A4E" blockId="34.[809,1443,550,1094]" pageId="34" pageNumber="303">
<emphasis id="C49769491936FF8F9938FDE996EF8C33" box="[809,896,550,571]" italics="true" pageId="34" pageNumber="303">Remarks</emphasis>
. The distribution extensions of
<taxonomicName id="31E3CED81936FF8F9EC9FDE996368C51" authority="Bell, 1884" authorityName="Bell. We" authorityYear="1884" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="34" pageNumber="303" phylum="Echinodermata" rank="species" species="okeni">
<emphasis id="C49769491936FF8F9EC9FDE9900A8C33" box="[1240,1381,550,571]" italics="true" pageId="34" pageNumber="303">Thyone okeni</emphasis>
<bibRefCitation id="9272C8AA1936FF8F9F60FDE996368C51" author="Bell, F. J." pageId="34" pageNumber="303" refId="ref22242" refString="Bell, F. J. 1884. Echinodermata. Pp. 117 - 177 in Report on the Zoological Collections made in the Indo-Pacific Ocean during the Voyage of H. M. S. Alert 1881 - 2. Taylor and Francis: London." type="book" year="1884">Bell, 1884</bibRefCitation>
</taxonomicName>
to Thursday Island off Cape York in northern Queensland (H. L.
<bibRefCitation id="9272C8AA1936FF8F9978FDAF96BA8C7E" author="Clark, H. L." box="[873,981,608,630]" pageId="34" pageNumber="303" pagination="1 - 223" refId="ref22387" refString="Clark, H. L. 1921. The echinoderm fauna of Torres Strait: its composition and its origin. Publication of the Carnegie Institution of Washington No. 214: i - viii, 1 - 223 pp, 38 pls." type="book chapter" year="1921">Clark 1921</bibRefCitation>
) and to
<collectingCountry id="8EF4F5CB1936FF8F9E33FDAE910F8C7E" box="[1058,1120,609,630]" name="Guam" pageId="34" pageNumber="303">Guam</collectingCountry>
in the eastern Pacific (
<bibRefCitation id="9272C8AA1936FF8F9F2DFDAE96F58C9B" author="Rowe, F. W. E. &amp; Doty, J. E." pageId="34" pageNumber="303" pagination="217 - 250" refId="ref23169" refString="Rowe, F. W. E. and Doty, J. E. 1977. The shallow-water holothurians of Guam. Micronesica 13: 217 - 250." type="journal article" year="1977">Rowe and Doty 1977</bibRefCitation>
) are not confirmed.
<bibRefCitation id="9272C8AA1936FF8F9E6EFDB2907D8C9B" author="Thandar, A. S." box="[1151,1298,637,659]" pageId="34" pageNumber="303" pagination="207 - 223" refId="ref23494" refString="Thandar, A. S. 1990. The phyllophorid holothurians of southern Africa with the erection of a new genus. South Africa Journal of Zoology 25 (4): 207 - 223." type="journal article" year="1990">Thandar 1990</bibRefCitation>
followed the comment by
<bibRefCitation id="9272C8AA1936FF8F99BFFD5491EB8CA7" author="Rowe, F. W. E. &amp; Doty, J. E." box="[942,1156,666,688]" pageId="34" pageNumber="303" pagination="217 - 250" refId="ref23169" refString="Rowe, F. W. E. and Doty, J. E. 1977. The shallow-water holothurians of Guam. Micronesica 13: 217 - 250." type="journal article" year="1977">Rowe and Doty 1977</bibRefCitation>
that
<taxonomicName id="31E3CED81936FF8F9EA8FD55901C8CB8" authority="Bell" authorityName="Bell. We" authorityYear="1884" box="[1209,1395,666,688]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="34" pageNumber="303" phylum="Echinodermata" rank="species" species="okeni">
<emphasis id="C49769491936FF8F9EA8FD55902D8CA7" box="[1209,1346,666,687]" italics="true" pageId="34" pageNumber="303">Thyone okeni</emphasis>
Bell
</taxonomicName>
was “probably distributed throughout the western Pacific area”, and without examining specimens was “inclined to think that
<taxonomicName id="31E3CED81936FF8F9F9FFD1A91688D0F" authority="Selenka, 1868" authorityName="Selenka" authorityYear="1868" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="34" pageNumber="303" phylum="Echinodermata" rank="species" species="venusta">
<emphasis id="C49769491936FF8F9F9FFD1A96198D0F" italics="true" pageId="34" pageNumber="303">T. venusta</emphasis>
<bibRefCitation id="9272C8AA1936FF8F996DFD3E91688D0F" author="Selenka, E." box="[892,1031,753,775]" pageId="34" pageNumber="303" pagination="109 - 119" refId="ref23385" refString="Selenka, E. 1868. Nachtrag zu den Beitragen zur Anatomie und Systematik der Holothurien. Zeitschrift fur Wissenschaftliche Zoologie 18: 109 - 119 pl. 8." type="journal article" year="1868">Selenka, 1868</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="31E3CED81936FF8F9E26FD3E91D08D0E" authorityName="Bell. We" authorityYear="1884" box="[1079,1215,753,774]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="34" pageNumber="303" phylum="Echinodermata" rank="species" species="okeni">
<emphasis id="C49769491936FF8F9E26FD3E91D08D0E" box="[1079,1215,753,774]" italics="true" pageId="34" pageNumber="303">Thyone okeni</emphasis>
</taxonomicName>
were conspecific” and then formally relegated
<taxonomicName id="31E3CED81936FF8F9E3CFCC190598D2B" authority="Bell, 1884" authorityName="Bell. We" authorityYear="1884" box="[1069,1334,782,804]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="34" pageNumber="303" phylum="Echinodermata" rank="species" species="okeni">
<emphasis id="C49769491936FF8F9E3CFCC191D38D2B" box="[1069,1212,782,803]" italics="true" pageId="34" pageNumber="303">Thyone okeni</emphasis>
<bibRefCitation id="9272C8AA1936FF8F9ED8FCC190598D2B" author="Bell, F. J." box="[1225,1334,782,804]" pageId="34" pageNumber="303" refId="ref22242" refString="Bell, F. J. 1884. Echinodermata. Pp. 117 - 177 in Report on the Zoological Collections made in the Indo-Pacific Ocean during the Voyage of H. M. S. Alert 1881 - 2. Taylor and Francis: London." type="book" year="1884">Bell, 1884</bibRefCitation>
</taxonomicName>
to junior synonymy with
<taxonomicName id="31E3CED81936FF8F99D6FCE4919C8D49" authority="Selenka, 1868" authorityName="Selenka" authorityYear="1868" box="[967,1267,811,833]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="34" pageNumber="303" phylum="Echinodermata" rank="species" species="venusta">
<emphasis id="C49769491936FF8F99D6FCE4910B8D49" box="[967,1124,811,833]" italics="true" pageId="34" pageNumber="303">Thyone venusta</emphasis>
<bibRefCitation id="9272C8AA1936FF8F9E78FCE4919C8D49" author="Selenka, E." box="[1129,1267,811,833]" pageId="34" pageNumber="303" pagination="109 - 119" refId="ref23385" refString="Selenka, E. 1868. Nachtrag zu den Beitragen zur Anatomie und Systematik der Holothurien. Zeitschrift fur Wissenschaftliche Zoologie 18: 109 - 119 pl. 8." type="journal article" year="1868">Selenka, 1868</bibRefCitation>
</taxonomicName>
(
<typeStatus id="29580BF91936FF8F9EEEFCE490448D48" box="[1279,1323,811,832]" pageId="34" pageNumber="303">type</typeStatus>
locality the Red Sea). In the key in his paper
<bibRefCitation id="9272C8AA1936FF8F9E69FC87907C8D55" author="Thandar, A. S." box="[1144,1299,840,862]" pageId="34" pageNumber="303" pagination="207 - 223" refId="ref23494" refString="Thandar, A. S. 1990. The phyllophorid holothurians of southern Africa with the erection of a new genus. South Africa Journal of Zoology 25 (4): 207 - 223." type="journal article" year="1990">Thandar (1990)</bibRefCitation>
described the body of
<taxonomicName id="31E3CED81936FF8F9991FCAA91DB8D73" authority="Selenka, 1868" authorityName="Selenka" authorityYear="1868" box="[896,1204,869,891]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="34" pageNumber="303" phylum="Echinodermata" rank="species" species="venusta">
<emphasis id="C49769491936FF8F9991FCAA914F8D73" box="[896,1056,869,891]" italics="true" pageId="34" pageNumber="303">Thyone venusta</emphasis>
<bibRefCitation id="9272C8AA1936FF8F9E39FCAA91DB8D73" author="Selenka, E." box="[1064,1204,869,891]" pageId="34" pageNumber="303" pagination="109 - 119" refId="ref23385" refString="Selenka, E. 1868. Nachtrag zu den Beitragen zur Anatomie und Systematik der Holothurien. Zeitschrift fur Wissenschaftliche Zoologie 18: 109 - 119 pl. 8." type="journal article" year="1868">Selenka, 1868</bibRefCitation>
</taxonomicName>
as being violet at both ends, and the tube feet as lacking ossicles. Neither character is true of
<taxonomicName id="31E3CED81936FF8F9961FC50910D8DBC" authority="Bell, 1884" authorityName="Bell. We" authorityYear="1884" box="[880,1122,927,949]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="34" pageNumber="303" phylum="Echinodermata" rank="species" species="okeni">
<emphasis id="C49769491936FF8F9961FC5096998DBC" box="[880,1014,927,948]" italics="true" pageId="34" pageNumber="303">Thyone okeni</emphasis>
<bibRefCitation id="9272C8AA1936FF8F99EAFC50910D8DBC" author="Bell, F. J." box="[1019,1122,927,949]" pageId="34" pageNumber="303" refId="ref22242" refString="Bell, F. J. 1884. Echinodermata. Pp. 117 - 177 in Report on the Zoological Collections made in the Indo-Pacific Ocean during the Voyage of H. M. S. Alert 1881 - 2. Taylor and Francis: London." type="book" year="1884">Bell, 1884</bibRefCitation>
</taxonomicName>
. There are distinctive dumbbell-shaped endplate support rods in the tube feet of
<taxonomicName id="31E3CED81936FF8F9F06FC7396E88DE6" authority="Bell. We" authorityName="Bell. We" authorityYear="1884" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="34" pageNumber="303" phylum="Echinodermata" rank="species" species="okeni">
<emphasis id="C49769491936FF8F9F06FC7390CF8DD9" box="[1303,1440,956,977]" italics="true" pageId="34" pageNumber="303">Thyone okeni</emphasis>
Bell. We
</taxonomicName>
reject this synonymy on the basis of inadequate systematic evidence for these two species with widely separated
<typeStatus id="29580BF91936FF8F9938FBDC963A8A20" box="[809,853,1043,1064]" pageId="34" pageNumber="303">type</typeStatus>
localities.
<taxonomicName id="31E3CED81936FF8F99D7FBDC91D08A20" authority="Bell, 1884" authorityName="Bell. We" authorityYear="1884" box="[966,1215,1043,1065]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="34" pageNumber="303" phylum="Echinodermata" rank="species" species="okeni">
<emphasis id="C49769491936FF8F99D7FBDC913F8A20" box="[966,1104,1043,1064]" italics="true" pageId="34" pageNumber="303">Thyone okeni</emphasis>
<bibRefCitation id="9272C8AA1936FF8F9E49FBDC91D08A20" author="Bell, F. J." box="[1112,1215,1043,1065]" pageId="34" pageNumber="303" refId="ref22242" refString="Bell, F. J. 1884. Echinodermata. Pp. 117 - 177 in Report on the Zoological Collections made in the Indo-Pacific Ocean during the Voyage of H. M. S. Alert 1881 - 2. Taylor and Francis: London." type="book" year="1884">Bell, 1884</bibRefCitation>
</taxonomicName>
is distinguished from other southern
<collectingCountry id="8EF4F5CB1936FF8F99D1FBFF91738A4E" box="[960,1052,1072,1094]" name="Australia" pageId="34" pageNumber="303">Australia</collectingCountry>
<taxonomicName id="31E3CED81936FF8F9E33FBFF91D58A4E" box="[1058,1210,1072,1094]" pageId="34" pageNumber="303">
<emphasis id="C49769491936FF8F9E33FBFF91038A4D" box="[1058,1132,1072,1093]" italics="true" pageId="34" pageNumber="303">Thyone</emphasis>
species
</taxonomicName>
in the key (above).
</paragraph>
</subSubSection>
</treatment>
</document>