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<document id="1CF0C16A2F8F195A5A2C5DC41456AC58" ID-DOI="10.24199/j.mmv.2012.69.05" ID-ISSN="1447-2554" ID-Zenodo-Dep="12212378" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718994422343" checkinUser="felipe" docAuthor="OLoughlin, P. Mark, Barmos, Shari &amp; VandenSpiegel, Didier" docDate="2012" docId="7E4A044D1936FF889938FBA597338DE7" docLanguage="en" docName="MemMusVictoria.69.269-308.pdf" docOrigin="Memoirs of Museum Victoria 69" docSource="https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-69-2012/pages-269-308/" docStyle="DocumentStyle:5B3B5C052A93BFEEE334AD6097A94E45.2:MemMusVictoria.2003-.journal_article.type1" docStyleId="5B3B5C052A93BFEEE334AD6097A94E45" docStyleName="MemMusVictoria.2003-.journal_article.type1" docStyleVersion="2" docTitle="Thyone spenceri O'Loughlin 2012, sp. nov." docType="treatment" docVersion="2" lastPageNumber="306" masterDocId="82737C351914FFAD9A11FFCF956F8E08" masterDocTitle="The phyllophorid sea cucumbers of southern Australia (Echinodermata: Holothuroidea: Dendrochirotida: Phyllophoridae)" masterLastPageNumber="308" masterPageNumber="269" pageNumber="303" updateTime="1719008797983" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="86A6D2DD077736E83B192093E3A74323">The phyllophorid sea cucumbers of southern Australia (Echinodermata: Holothuroidea: Dendrochirotida: Phyllophoridae)</mods:title>
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<mods:namePart id="56F9FCDC48C1B433396F139DA6F4BB95">OLoughlin, P. Mark</mods:namePart>
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<mods:namePart id="CFBA31E8360C0C9812E7139F2B09D08D">Barmos, Shari</mods:namePart>
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<mods:namePart id="4550DE4025E60E3A709BA6AC22C51E4B">VandenSpiegel, Didier</mods:namePart>
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<mods:title id="0537C727B932BBCD3EF602B58236A609">Memoirs of Museum Victoria</mods:title>
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<mods:number id="31FAF2F259C1B65F24D34141E2F4DD35">69</mods:number>
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<treatment id="7E4A044D1936FF889938FBA597338DE7" ID-DOI="http://doi.org/10.5281/zenodo.12212502" ID-Zenodo-Dep="12212502" LSID="urn:lsid:plazi:treatment:7E4A044D1936FF889938FBA597338DE7" httpUri="http://treatment.plazi.org/id/7E4A044D1936FF889938FBA597338DE7" lastPageId="37" lastPageNumber="306" pageId="34" pageNumber="303">
<subSubSection id="BEF9E6D01936FF8F9938FBA591CA8A88" box="[809,1189,1130,1153]" pageId="34" pageNumber="303" type="nomenclature">
<paragraph id="F65CB55B1936FF8F9938FBA591CA8A88" blockId="34.[809,1189,1130,1153]" box="[809,1189,1130,1153]" pageId="34" pageNumber="303">
<heading id="AD1402371936FF8F9938FBA591CA8A88" box="[809,1189,1130,1153]" fontSize="9" level="2" pageId="34" pageNumber="303" reason="5">
<taxonomicName id="31E3CED81936FF8F9938FBA5913C8A89" ID-CoL="56RP8" authority="O'Loughlin" authorityName="O'Loughlin" authorityYear="2012" box="[809,1107,1130,1153]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="34" pageNumber="303" phylum="Echinodermata" rank="species" species="spenceri" status="sp. nov.">
<emphasis id="C49769491936FF8F9938FBA596B98A88" bold="true" box="[809,982,1130,1152]" italics="true" pageId="34" pageNumber="303">Thyone spenceri</emphasis>
OLoughlin
</taxonomicName>
<taxonomicNameLabel id="DFA4D4321936FF8F9E48FBA491CA8A88" box="[1113,1189,1131,1152]" pageId="34" pageNumber="303" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="BEF9E6D01936FF889938FB5897338DE7" lastPageId="37" lastPageNumber="306" pageId="34" pageNumber="303" type="description">
<paragraph id="F65CB55B1936FF8F9938FB5896AD8AA4" blockId="34.[809,962,1175,1197]" box="[809,962,1175,1197]" pageId="34" pageNumber="303">
<figureCitation id="6ED8A9DE1936FF8F9938FB5896F18AA4" box="[809,926,1175,1197]" captionStart="Figure 17" captionStartId="27.[147,206,1778,1797]" captionTargetBox="[147,1447,227,1701]" captionTargetId="figure-11@27.[147,1447,227,1701]" captionTargetPageId="27" captionText="Figure 17. Photos of preserved specimens. a, holotype of Thyone flindersi OLoughlin sp. nov. (NMV F151847) (insert: form of the calcareous ring plates); b, specimen of Thyone okeni Bell, 1884 (AM J10868); c, holotype of Thyone joshuai OLoughlin sp. nov. (SAM K2566) (insert: form of the calcareous ring plates); d, holotype of Thyone kerkosa OLoughlin sp. nov. (WAM Z31838); e, holotype of Thyone spenceri OLoughlin sp. nov. (SAM K2562) (insert: form of the calcareous ring plates); f, holotype of Thyone tourvillei OLoughlin sp. nov. (NMV F174902) (insert: form of the calcareous ring plates)." figureDoi="http://doi.org/10.5281/zenodo.12212414" httpUri="https://zenodo.org/record/12212414/files/figure.png" pageId="34" pageNumber="303">Figures 17e</figureCitation>
,
<figureCitation id="6ED8A9DE1936FF8F99B8FB5896AD8AA4" box="[937,962,1175,1196]" captionStart="Figure 23" captionStartId="36.[140,199,1857,1876]" captionTargetBox="[140,1440,227,1698]" captionTargetId="figure-14@36.[140,1440,227,1698]" captionTargetPageId="36" captionText="Figure 23. SEM images of ossicles from paratype of Thyone spenceri OLoughlin sp. nov. (SAM K2565). Tables from anterior body wall (top); plates and rosettes from tentacles (bottom)." figureDoi="http://doi.org/10.5281/zenodo.12212426" httpUri="https://zenodo.org/record/12212426/files/figure.png" pageId="34" pageNumber="303">23</figureCitation>
</paragraph>
<paragraph id="F65CB55B1936FF8F9938FB0D96BC8B2C" blockId="34.[809,1441,1218,1395]" pageId="34" pageNumber="303">
<emphasis id="C49769491936FF8F9938FB0D96B68ADD" box="[809,985,1218,1237]" italics="true" pageId="34" pageNumber="303">Material examined</emphasis>
.
<materialsCitation id="468BBF061936FF8F99F7FB0D96BC8B2C" collectingDate="1987-09" collectionCode="SAM" collectorName="SA Fisheries Department" elevation="18" latitude="-32.833332" location="Backy Point" longLatPrecision="925" longitude="137.842" municipality="Upper Spencer Gulf" pageId="34" pageNumber="303" specimenCode="K2562" specimenCount="1" stateProvince="South Australia" typeStatus="holotype">
<typeStatus id="29580BF91936FF8F99F7FB0D91528ADD" box="[998,1085,1218,1237]" pageId="34" pageNumber="303" type="holotype">Holotype</typeStatus>
.
<collectingRegion id="34277BB91936FF8F9E57FB0D91B78ADD" box="[1094,1240,1218,1237]" country="Australia" name="South Australia" pageId="34" pageNumber="303">South Australia</collectingRegion>
,
<collectingMunicipality id="16382F211936FF8F9EF3FB0D90F28ADD" box="[1250,1437,1218,1237]" pageId="34" pageNumber="303">Upper Spencer Gulf</collectingMunicipality>
,
<location id="F33CE3801936FF8F9938FB1396CF8AE7" LSID="urn:lsid:plazi:treatment:7E4A044D1936FF889938FBA597338DE7:F33CE3801936FF8F9938FB1396CF8AE7" box="[809,928,1244,1263]" latitude="-32.833332" longLatPrecision="925" longitude="137.842" municipality="Upper Spencer Gulf" name="Backy Point" pageId="34" pageNumber="303" stateProvince="South Australia">Backy Point</location>
West,
<geoCoordinate id="93D7D39C1936FF8F9E19FB1391228AE7" box="[1032,1101,1244,1263]" degrees="32" direction="south" minutes="50" orientation="latitude" pageId="34" pageNumber="303" precision="925" value="-32.833332">32º50'S</geoCoordinate>
<geoCoordinate id="93D7D39C1936FF8F9E4AFB1391A58AE7" box="[1115,1226,1244,1263]" degrees="137" direction="east" minutes="50.52" orientation="longitude" pageId="34" pageNumber="303" precision="9" value="137.842">137º50.52'E</geoCoordinate>
,
<quantity id="311B18BE1936FF8F9ECDFB13907D8AF8" box="[1244,1298,1244,1264]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="34" pageNumber="303" unit="m" value="18.0">
<elevation id="7DCE52681936FF8F9ECDFB13907D8AF8" box="[1244,1298,1244,1264]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="34" pageNumber="303" unit="m" value="18.0">18 m</elevation>
</quantity>
,
<collectorName id="5B16D08D1936FF8F9F35FB1396F88B02" pageId="34" pageNumber="303">SA Fisheries Department</collectorName>
,
<date id="825D939B1936FF8F99B5FB3996948B01" box="[932,1019,1270,1289]" pageId="34" pageNumber="303" value="1987-09">
<collectingDate id="92196A731936FF8F99B5FB3996948B01" box="[932,1019,1270,1289]" pageId="34" pageNumber="303" value="1987-09">Sep 1987</collectingDate>
</date>
,
<collectionCode id="90F22D9E1936FF8F9E19FB3991568B01" box="[1032,1081,1270,1289]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="34" pageNumber="303">SAM</collectionCode>
<specimenCode id="A6451D201936FF8F9E53FB3991108B01" box="[1090,1151,1270,1289]" collectionCode="SAM" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="34" pageNumber="303">K2562</specimenCode>
(ring, tentacle crown, anterior part of body wall).
</materialsCitation>
</paragraph>
<paragraph id="F65CB55B1936FF8F995EFAE491C18B7A" blockId="34.[809,1441,1218,1395]" pageId="34" pageNumber="303">
<materialsCitation id="468BBF061936FF8F995EFAE491C48B7A" collectingDate="1987-09" collectionCode="SAM" collectorName="SA Fisheries Department" elevation="15" latitude="-33.016666" location="Upper Spencer Gulf" longLatPrecision="1207" longitude="137.75" municipality="Port Bonython" pageId="34" pageNumber="303" specimenCode="K2565" specimenCount="1" typeStatus="paratype">
<typeStatus id="29580BF91936FF8F995EFAE496CD8B36" box="[847,930,1323,1342]" pageId="34" pageNumber="303" type="paratype">Paratype</typeStatus>
.
<location id="F33CE3801936FF8F99BDFAE491098B36" LSID="urn:lsid:plazi:treatment:7E4A044D1936FF889938FBA597338DE7:F33CE3801936FF8F99BDFAE491098B36" box="[940,1126,1323,1342]" latitude="-33.016666" longLatPrecision="1207" longitude="137.75" municipality="Port Bonython" name="Upper Spencer Gulf" pageId="34" pageNumber="303">Upper Spencer Gulf</location>
,
<collectingMunicipality id="16382F211936FF8F9E61FAE491978B36" box="[1136,1272,1323,1342]" pageId="34" pageNumber="303">Port Bonython</collectingMunicipality>
,
<geoCoordinate id="93D7D39C1936FF8F9F13FAE4902A8B36" box="[1282,1349,1323,1342]" degrees="33" direction="south" minutes="01" orientation="latitude" pageId="34" pageNumber="303" precision="925" value="-33.016666">33º01'S</geoCoordinate>
<geoCoordinate id="93D7D39C1936FF8F9F5AFAE490F28B36" box="[1355,1437,1323,1342]" degrees="137" direction="east" minutes="45" orientation="longitude" pageId="34" pageNumber="303" precision="925" value="137.75">137º45'E</geoCoordinate>
,
<quantity id="311B18BE1936FF8F9938FA8A96328B51" box="[809,861,1349,1369]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="34" pageNumber="303" unit="m" value="15.0">
<elevation id="7DCE52681936FF8F9938FA8A96328B51" box="[809,861,1349,1369]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="34" pageNumber="303" unit="m" value="15.0">15 m</elevation>
</quantity>
,
<collectorName id="5B16D08D1936FF8F997AFA8A91328B51" box="[875,1117,1349,1369]" pageId="34" pageNumber="303">SA Fisheries Department</collectorName>
,
<date id="825D939B1936FF8F9E7AFA8A91AB8B50" box="[1131,1220,1349,1368]" pageId="34" pageNumber="303" value="1987-09">
<collectingDate id="92196A731936FF8F9E7AFA8A91AB8B50" box="[1131,1220,1349,1368]" pageId="34" pageNumber="303" value="1987-09">Sep 1987</collectingDate>
</date>
,
<collectionCode id="90F22D9E1936FF8F9EC3FA8A906C8B50" box="[1234,1283,1349,1368]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="34" pageNumber="303">SAM</collectionCode>
<specimenCode id="A6451D201936FF8F9F1CFA8A90248B50" box="[1293,1355,1349,1368]" collectionCode="SAM" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="34" pageNumber="303">K2565</specimenCode>
(1, ring, tentacle crown, anterior part of body wall)
</materialsCitation>
.
</paragraph>
<paragraph id="F65CB55B1936FF8F9938FA4591D488C9" blockId="34.[809,1441,1418,1903]" pageId="34" pageNumber="303">
<emphasis id="C49769491936FF8F9938FA4596F88B97" box="[809,919,1418,1439]" italics="true" pageId="34" pageNumber="303">Diagnosis.</emphasis>
Anterior body and calcareous ring
<quantity id="311B18BE1936FF8F9F0CFA4590068B97" box="[1309,1385,1418,1439]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="34" pageNumber="303" unit="mm" value="13.0">13 mm</quantity>
long (preserved), body wall soft; 10 tentacles, 8 large 2 ventral small; complete close cover of small tube feet; calcareous ring composite, tubular, posterior ends of composite plates not joined; radial plates tapered anterior with terminal split, long paired composite posterior prolongations separated for most of length, narrowing and further divided distally for about a quarter of total plate length, not joined; inter-radial plates composite, tapered to anterior point, truncate posteriorly; stone canal attached to alimentary canal mesentery, split-pea madreporite form; single polian vesicle.
</paragraph>
<paragraph id="F65CB55B1936FF88995EF90797D78C7B" blockId="34.[809,1441,1418,1903]" lastBlockId="37.[147,779,229,628]" lastPageId="37" lastPageNumber="306" pageId="34" pageNumber="303">
Anterior body wall with abundant tables, tables regular, discs round to oval to triangular, up to 96
<emphasis id="C49769491936FF8F9EE0F929919088F3" box="[1265,1279,1766,1787]" italics="true" pageId="34" pageNumber="303">µ</emphasis>
m long, 2 large central perforations, smaller perforations nearer margin, predominantly 4050 small perforations, spires with 2 pillars, 24
<emphasis id="C49769491936FF8F995AF8F29636895A" box="[843,857,1853,1874]" italics="true" pageId="34" pageNumber="303">µ</emphasis>
m long, 2 tapered ends distally; some table discs rosettelike; some table discs curved for tube foot support; tentacles with abundant perforated plates, rosettes; plates variable in shape, some plates narrow, elongate with large perforations and distal blunt teeth, sometimes small branches along margin of plate, plates up to 280
<emphasis id="C49769491931FF889B69FEF294E98F5A" box="[376,390,317,338]" italics="true" pageId="37" pageNumber="306">µ</emphasis>
m long, some plates cross-shaped with perforate arms, some plates oval with four large central perforations, smaller surrounding ones and distal blunt teeth, plates up to 160
<emphasis id="C49769491931FF889B2CFE5B94248FA1" box="[317,331,404,425]" italics="true" pageId="37" pageNumber="306">µ</emphasis>
m long, some plates curved with denticulate margin and numerous small perforations, plates up to 120
<emphasis id="C49769491931FF8898F8FE7E97988FCE" box="[745,759,433,454]" italics="true" pageId="37" pageNumber="306">µ</emphasis>
m long; rosettes oval, up to 88
<emphasis id="C49769491931FF889BA3FE0194AF8FEB" box="[434,448,462,483]" italics="true" pageId="37" pageNumber="306">µ</emphasis>
m long; peri-oral body wall with tables and rods; tables with 2 spires, discs with 2 central large perforations, small perforations between central large ones and disc margin, each disc with up to more than 30 small perforations, discs about 104
<emphasis id="C49769491931FF889B58FD8E94388C5E" box="[329,343,577,598]" italics="true" pageId="37" pageNumber="306">µ</emphasis>
m long; peri-oral ossicles irregular wide perforated rods, up to 272
<emphasis id="C49769491931FF889B8AFD9194C68C7B" box="[411,425,606,627]" italics="true" pageId="37" pageNumber="306">µ</emphasis>
m long, sometimes crosses.
</paragraph>
<caption id="A29CE5D31937FF8E9A82F88E964A8966" ID-DOI="http://doi.org/10.5281/zenodo.12212424" ID-Zenodo-Dep="12212424" httpUri="https://zenodo.org/record/12212424/files/figure.png" pageId="35" pageNumber="304" startId="35.[147,206,1857,1876]" targetBox="[147,1447,227,1699]" targetPageId="35" targetType="figure">
<paragraph id="F65CB55B1937FF8E9A82F88E964A8966" blockId="35.[147,1447,1857,1902]" pageId="35" pageNumber="304">
Figure 22. SEM images of ossicles from specimen of
<taxonomicName id="31E3CED81937FF8E9892F88E960C895C" authority="Bell, 1884" authorityName="Bell. We" authorityYear="1884" box="[643,867,1857,1876]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="35" pageNumber="304" phylum="Echinodermata" rank="species" species="okeni">
<emphasis id="C49769491937FF8E9892F88E9790895C" box="[643,767,1857,1876]" italics="true" pageId="35" pageNumber="304">Thyone okeni</emphasis>
<bibRefCitation id="9272C8AA1937FF8E9917F88E960C895C" author="Bell, F. J." box="[774,867,1857,1876]" pageId="35" pageNumber="304" refId="ref22242" refString="Bell, F. J. 1884. Echinodermata. Pp. 117 - 177 in Report on the Zoological Collections made in the Indo-Pacific Ocean during the Voyage of H. M. S. Alert 1881 - 2. Taylor and Francis: London." type="book" year="1884">Bell, 1884</bibRefCitation>
</taxonomicName>
(AM J7731). Peri-oral body wall rosette and tables with multibranched spires (top); dumbbell-shaped ossicles from tentacles (bottom).
</paragraph>
</caption>
<caption id="A29CE5D31930FF899A9DF88E977B8966" ID-DOI="http://doi.org/10.5281/zenodo.12212426" ID-Zenodo-Dep="12212426" httpUri="https://zenodo.org/record/12212426/files/figure.png" pageId="36" pageNumber="305" startId="36.[140,199,1857,1876]" targetBox="[140,1440,227,1698]" targetPageId="36" targetType="figure">
<paragraph id="F65CB55B1930FF899A9DF88E977B8966" blockId="36.[140,1440,1857,1902]" pageId="36" pageNumber="305">
Figure 23. SEM images of ossicles from paratype of
<taxonomicName id="31E3CED81930FF89987DF88E961B895C" authority="O'Loughlin" authorityName="O'Loughlin" authorityYear="2012" box="[620,884,1857,1876]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="36" pageNumber="305" phylum="Echinodermata" rank="species" species="spenceri" status="sp. nov.">
<emphasis id="C49769491930FF89987DF88E966B895B" box="[620,772,1857,1876]" italics="true" pageId="36" pageNumber="305">Thyone spenceri</emphasis>
OLoughlin
</taxonomicName>
<taxonomicNameLabel id="DFA4D4321930FF89996BF88E96D0895C" box="[890,959,1857,1876]" pageId="36" pageNumber="305" rank="species">sp. nov.</taxonomicNameLabel>
(SAM K2565). Tables from anterior body wall (top); plates and rosettes from tentacles (bottom).
</paragraph>
</caption>
<paragraph id="F65CB55B1931FF889A82FD4594FA8CB5" blockId="37.[147,778,650,701]" pageId="37" pageNumber="306">
<emphasis id="C49769491931FF889A82FD45943C8C97" box="[147,339,650,671]" italics="true" pageId="37" pageNumber="306">Colour (preserved)</emphasis>
. Body off-white; tentacles brown with dark brown to black markings.
</paragraph>
<paragraph id="F65CB55B1931FF889A82FD1C97948CE0" blockId="37.[147,763,723,745]" box="[147,763,723,745]" pageId="37" pageNumber="306">
<emphasis id="C49769491931FF889A82FD1C947F8CE0" box="[147,272,723,744]" italics="true" pageId="37" pageNumber="306">Distribution</emphasis>
.
<collectingRegion id="34277BB91931FF889B0CFD1C94D28CE1" box="[285,445,723,745]" country="Australia" name="South Australia" pageId="37" pageNumber="306">South Australia</collectingRegion>
, Upper Spencer Gulf,
<quantity id="311B18BE1931FF88988FFD1C97988CE0" box="[670,759,723,745]" metricMagnitude="1" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="37" pageNumber="306" unit="m" value="16.5" valueMax="18.0" valueMin="15.0">
15
<specimenCount id="E0E57ED21931FF8898D2FD1C97988CE0" box="[707,759,723,745]" count="18" pageId="37" pageNumber="306" type="generic">18 m</specimenCount>
</quantity>
.
</paragraph>
<paragraph id="F65CB55B1931FF889A82FD3097F48D3A" blockId="37.[147,778,767,818]" pageId="37" pageNumber="306">
<emphasis id="C49769491931FF889A82FD3094688D1C" box="[147,263,767,788]" italics="true" pageId="37" pageNumber="306">Etymology.</emphasis>
Named with reference to Spencer Gulf in
<collectingRegion id="34277BB91931FF8898DEFD30959F8D3A" country="Australia" name="South Australia" pageId="37" pageNumber="306">South Australia</collectingRegion>
where the
<typeStatus id="29580BF91931FF889B4EFCD394E48D39" box="[351,395,796,817]" pageId="37" pageNumber="306">type</typeStatus>
specimens were collected.
</paragraph>
<paragraph id="F65CB55B1931FF889A82FC8797338DE7" blockId="37.[147,779,840,1007]" pageId="37" pageNumber="306">
<emphasis id="C49769491931FF889A82FC87959C8D55" box="[147,243,840,861]" italics="true" pageId="37" pageNumber="306">Remarks.</emphasis>
Both the
<typeStatus id="29580BF91931FF889B48FC8794DE8D56" box="[345,433,840,862]" pageId="37" pageNumber="306" type="holotype">holotype</typeStatus>
and
<typeStatus id="29580BF91931FF889BF4FC8797528D55" box="[485,573,840,861]" pageId="37" pageNumber="306" type="paratype">paratype</typeStatus>
of
<taxonomicName id="31E3CED81931FF889870FC8794658D73" authority="O'Loughlin" authorityName="O'Loughlin" authorityYear="2012" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="37" pageNumber="306" phylum="Echinodermata" rank="species" species="spenceri" status="sp. nov.">
<emphasis id="C49769491931FF889870FC8796658D55" box="[609,778,840,861]" italics="true" pageId="37" pageNumber="306">Thyone spenceri</emphasis>
OLoughlin
</taxonomicName>
<taxonomicNameLabel id="DFA4D4321931FF889B1EFCAA94358D72" box="[271,346,869,890]" pageId="37" pageNumber="306" rank="species">sp. nov.</taxonomicNameLabel>
are damaged and most of the body wall has been lost. But there are good morphological systematic characters for the erection of this new
<taxonomicName id="31E3CED81931FF889838FC5096648DBD" box="[553,779,927,949]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="37" pageNumber="306" phylum="Echinodermata" rank="species" species="undetermined">
<emphasis id="C49769491931FF889838FC50971B8DBC" box="[553,628,927,948]" italics="true" pageId="37" pageNumber="306">Thyone</emphasis>
Oken species.
</taxonomicName>
<taxonomicName id="31E3CED81931FF889A82FC7394258DD9" authorityName="O'Loughlin" authorityYear="2012" box="[147,330,956,977]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="37" pageNumber="306" phylum="Echinodermata" rank="species" species="spenceri">
<emphasis id="C49769491931FF889A82FC7394258DD9" box="[147,330,956,977]" italics="true" pageId="37" pageNumber="306">Thyone spenceri</emphasis>
</taxonomicName>
is distinguished from other southern Australian
<taxonomicName id="31E3CED81931FF889B15FC1694F18DE7" box="[260,414,985,1007]" pageId="37" pageNumber="306">
<emphasis id="C49769491931FF889B15FC1694208DE6" box="[260,335,985,1006]" italics="true" pageId="37" pageNumber="306">Thyone</emphasis>
species
</taxonomicName>
in the key (above).
</paragraph>
</subSubSection>
</treatment>
</document>