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<document id="AD81EC71FA1E9A1FF4229F56845F2112" ID-CLB-Dataset="45477" ID-DOI="10.11646/zootaxa.3815.3.5" ID-GBIF-Dataset="a135a590-a8a5-4fc1-bb2f-925d6920e48b" ID-ISSN="1175-5326" ID-Zenodo-Dep="225975" ID-ZooBank="CBBEC600-252A-4591-BD83-C754948ED3A3" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460437510317" checkinUser="plazi" docAuthor="Shi, Weifang &amp; Tong, Xiaoli" docDate="2014" docId="7E4287D88858FFC083ECE2D8FD01F1E3" docLanguage="en" docName="zt03815p408.pdf" docOrigin="Zootaxa 3815 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Labiobaetis numeratus Muller-Liebenau 1984, n. comb." docType="treatment" docVersion="9" lastPageNumber="405" masterDocId="827BFFA0885FFFC8837BE434FFEAF32F" masterDocTitle="The genus Labiobaetis (Ephemeroptera: Baetidae) in China, with description of a new species" masterLastPageNumber="408" masterPageNumber="397" pageNumber="404" updateTime="1698332404576" updateUser="ExternalLinkService">
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<mods:title id="2EA5BE3ED382256D0488959F6813898D">The genus Labiobaetis (Ephemeroptera: Baetidae) in China, with description of a new species</mods:title>
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<mods:namePart id="2B75F685A6204990A6F8843ECE6791CB">Shi, Weifang</mods:namePart>
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<mods:namePart id="60B8FB861856D53CDDED0BA3F99F9441">Tong, Xiaoli</mods:namePart>
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<treatment id="7E4287D88858FFC083ECE2D8FD01F1E3" ID-DOI="http://doi.org/10.5281/zenodo.6123026" ID-GBIF-Taxon="119506962" ID-Zenodo-Dep="6123026" LSID="urn:lsid:plazi:treatment:7E4287D88858FFC083ECE2D8FD01F1E3" httpUri="http://treatment.plazi.org/id/7E4287D88858FFC083ECE2D8FD01F1E3" lastPageId="8" lastPageNumber="405" pageId="7" pageNumber="404">
<subSubSection id="BEF165458858FFCF83ECE2D8FEC4F407" pageId="7" pageNumber="404" type="nomenclature">
<paragraph id="F65436CE8858FFCF83ECE2D8FCBCF428" blockId="7.[151,854,1772,1832]" box="[151,854,1772,1799]" pageId="7" pageNumber="404">
<heading id="AD1C81A28858FFCF83ECE2D8FCBCF428" bold="true" box="[151,854,1772,1799]" fontSize="11" level="1" pageId="7" pageNumber="404" reason="1">
<emphasis id="C49FEADC8858FFCF83ECE2D8FCBCF428" bold="true" box="[151,854,1772,1799]" pageId="7" pageNumber="404">
<taxonomicName id="31EB4D4D8858FFCF83ECE2D8FD0EF428" authority="Muller-Liebenau, 1984" authorityName="Muller-Liebenau" authorityYear="1984" box="[151,740,1772,1799]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="404" phylum="Arthropoda" rank="species" species="numeratus" status="comb. nov.">
<emphasis id="C49FEADC8858FFCF83ECE2D8FE46F429" bold="true" box="[151,428,1772,1798]" italics="true" pageId="7" pageNumber="404">Labiobaetis numeratus</emphasis>
(
<bibRefCitation id="927A4B3F8858FFCF82C0E2D9FD36F428" author="Muller-Liebenau" box="[443,732,1773,1799]" pageId="7" pageNumber="404" refString="Muller-Liebenau, I. (1984) New genera and species of the family Baetidae from West-Malaysia (River Gombak). Spixiana, 7, 253 - 284. Available from: http: // biostor. org / reference / 47587 (accessed 3 June 2014)" type="journal article" year="1984">Müller-Liebenau, 1984</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="DFAC57A78858FFCF8197E2D9FCBCF428" box="[748,854,1773,1799]" pageId="7" pageNumber="404">n. comb.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="F65436CE8858FFCF83ECE33BFEC4F407" blockId="7.[151,854,1772,1832]" box="[151,302,1807,1832]" pageId="7" pageNumber="404">
(
<figureCitation id="6ED02A4B8858FFCF83E4E33BFECCF407" box="[159,294,1807,1832]" captionStart="FIGURES 24 – 27" captionStartId="8.[151,264,1954,1977]" captionTargetBox="[160,1428,789,1913]" captionTargetId="figure@8.[151,1436,769,1933]" captionTargetPageId="8" captionText="FIGURES 24 – 27. Labiobaetis numeratus (Müller-Liebenau) 24, right mandible; 25, pronotum; 26, color pattern of abdomen (left: sterna; right: terga); 27, habitus of male nymph." httpUri="https://zenodo.org/record/225980/files/figure.png" pageId="7" pageNumber="404">Figs. 24–27</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="BEF165458858FFC083ECE36DFD0AF232" lastPageId="8" lastPageNumber="405" pageId="7" pageNumber="404" type="reference_group">
<paragraph id="F65436CE8858FFCF83ECE36DFD93F442" blockId="7.[151,714,1879,1963]" box="[151,633,1879,1902]" pageId="7" pageNumber="404">
<treatmentCitationGroup id="D6FB11E08858FFCF83ECE36DFD93F442" box="[151,633,1879,1902]" pageId="7" pageNumber="404">
<taxonomicName id="31EB4D4D8858FFCF83ECE36DFD9FF442" authority="Muller-Liebenau, 1984: 268" authorityName="Muller-Liebenau" authorityPageNumber="268" authorityYear="1984" box="[151,629,1879,1902]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="404" phylum="Arthropoda" rank="species" species="numeratus">
<emphasis id="C49FEADC8858FFCF83ECE36DFEADF441" box="[151,327,1881,1902]" italics="true" pageId="7" pageNumber="404">Baetis numeratus</emphasis>
:
<treatmentCitation id="774A10DF8858FFCF822FE363FD9FF442" author="Muller-Liebenau" box="[340,629,1879,1901]" page="268" pageId="7" pageNumber="404" year="1984">
<bibRefCitation id="927A4B3F8858FFCF822FE363FDA9F442" author="Muller-Liebenau" box="[340,579,1879,1901]" pageId="7" pageNumber="404" refString="Muller-Liebenau, I. (1984) New genera and species of the family Baetidae from West-Malaysia (River Gombak). Spixiana, 7, 253 - 284. Available from: http: // biostor. org / reference / 47587 (accessed 3 June 2014)" type="journal article" year="1984">Müller-Liebenau, 1984</bibRefCitation>
: 268
</treatmentCitation>
</taxonomicName>
;
</treatmentCitationGroup>
</paragraph>
<paragraph id="F65436CE8858FFCF83ECE342FD5EF485" blockId="7.[151,714,1879,1963]" pageId="7" pageNumber="404">
<treatmentCitationGroup id="D6FB11E08858FFCF83ECE342FD5EF485" pageId="7" pageNumber="404">
<taxonomicName id="31EB4D4D8858FFCF83ECE342FD2CF4A3" authority="McCafferty &amp; Waltz, 1995: 21" authorityName="McCafferty &amp; Waltz" authorityPageNumber="21" authorityYear="1995" box="[151,710,1910,1932]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="404" phylum="Arthropoda" rank="species" species="numeratus">
<emphasis id="C49FEADC8858FFCF83ECE342FE6AF4A3" box="[151,384,1910,1932]" italics="true" pageId="7" pageNumber="404">Labiobaetis numeratus</emphasis>
:
<treatmentCitation id="774A10DF8858FFCF82F6E342FD2CF4A3" author="McCafferty" box="[397,710,1910,1932]" page="21" pageId="7" pageNumber="404" year="1995">
<bibRefCitation id="927A4B3F8858FFCF82F6E342FD4AF4A3" author="McCafferty" box="[397,672,1910,1932]" pageId="7" pageNumber="404" refString="McCafferty, W. P. &amp; Waltz, R. D. (1995) Labiobaetis (Ephemeroptera: Baetidae): new status, new North American species, and related new genus. Entomological News, 106, 19 - 28." type="journal article" year="1995">McCafferty &amp; Waltz, 1995</bibRefCitation>
: 21
</treatmentCitation>
</taxonomicName>
;
<taxonomicName id="31EB4D4D8858FFCF83ECE3A0FD7BF485" ID-CoL="6WDPG" authority="Lugo-Ortiz" authorityName="Lugo-Ortiz" authorityYear="1999" box="[151,657,1940,1963]" class="Insecta" family="Baetidae" genus="Pseudocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="404" phylum="Arthropoda" rank="species" species="numeratum">
<emphasis id="C49FEADC8858FFCF83ECE3A0FE74F484" box="[151,414,1940,1963]" italics="true" pageId="7" pageNumber="404">Pseudocloeon numeratum</emphasis>
:
<treatmentCitation id="774A10DF8858FFCF82D7E3A0FD7BF485" author="Lugo-Ortiz" box="[428,657,1940,1962]" page="25" pageId="7" pageNumber="404" year="1999">
<bibRefCitation id="927A4B3F8858FFCF82D7E3A0FD7BF485" author="Lugo-Ortiz" box="[428,657,1940,1962]" pageId="7" pageNumber="404" refString="Lugo-Ortiz, C. R., McCafferty, W. P. &amp; Waltz, R. D. (1999) Definition and reorganization of the genus Pseudocloeon (Ephemeroptera: Baetidae) with new species description and combinations. Transactions of the American Entomological Society, 125, 1 - 37. Available from: http: // www. jstor. org / stable / 25078672 (accessed 3 June 2014)" type="journal article" year="1999">Lugo-Ortiz et al, 1999</bibRefCitation>
</treatmentCitation>
</taxonomicName>
: 25
</treatmentCitationGroup>
</paragraph>
<paragraph id="F65436CE8858FFC083ECE3E3FD0AF232" blockId="7.[151,1436,2007,2032]" lastBlockId="8.[151,1436,151,717]" lastPageId="8" lastPageNumber="405" pageId="7" pageNumber="404">
<emphasis id="C49FEADC8858FFCF83ECE3E3FD10F4DF" bold="true" box="[151,762,2007,2032]" pageId="7" pageNumber="404">
Specimens examined.
<collectingCountry id="8EFC765E8858FFCF82E6E3E3FE02F4DF" box="[413,488,2007,2032]" name="China" pageId="7" pageNumber="404">China</collectingCountry>
, Guangdong Province:
</emphasis>
1 nymph on slide and 1 nymph, Wuhua County, Shuanghua Town, near Junying Hydropower Station (
<geoCoordinate id="93DF50098857FFC0810DE4A3FD33F39F" box="[630,729,151,176]" direction="north" orientation="latitude" pageId="8" pageNumber="405" precision="555" value="23.75">23.75°N</geoCoordinate>
,
<geoCoordinate id="93DF50098857FFC0819EE4A3FCBAF39F" box="[741,848,151,176]" direction="east" orientation="longitude" pageId="8" pageNumber="405" precision="555" value="115.85">115.85°E</geoCoordinate>
, alt.
<quantity id="31139B2B8857FFC080F2E4ACFC26F39F" box="[905,972,152,176]" metricMagnitude="2" metricUnit="m" metricValue="2.41" pageId="8" pageNumber="405" unit="m" value="241.0">241m</quantity>
),
<date id="8255100E8857FFC08099E4A3FB8AF39F" box="[994,1120,151,176]" pageId="8" pageNumber="405" value="2011-11-14">14-xi-2011</date>
, coll. Shi Weifang;
<emphasis id="C49FEADC8857FFC08638E4A3FEE3F3FA" bold="true" pageId="8" pageNumber="405">Hainan Province:</emphasis>
1 nymph on slide and 2 nymphs, Baisha County, Yinggeling National Nature Reserve, Daoyin (
<geoCoordinate id="93DF50098857FFC0864DE488FA72F3FB" box="[1334,1432,188,212]" direction="north" orientation="latitude" pageId="8" pageNumber="405" precision="555" value="18.99">18.99°N</geoCoordinate>
,
<geoCoordinate id="93DF50098857FFC083ECE4EBFEEBF3D7" box="[151,257,223,248]" direction="east" orientation="longitude" pageId="8" pageNumber="405" precision="555" value="109.58">109.58°E</geoCoordinate>
),
<date id="8255100E8857FFC08265E4EBFE7DF3D7" box="[286,407,223,248]" pageId="8" pageNumber="405" value="2005-12-05">5-xii-2005</date>
, coll. Yao Haiyuan; 1 nymph on slide, Wanning City, Nanqiao Town, Taiyang River (
<geoCoordinate id="93DF50098857FFC083E4E530FEEBF232" box="[159,257,260,285]" direction="north" orientation="latitude" pageId="8" pageNumber="405" precision="555" value="18.68">18.68°N</geoCoordinate>
,
<geoCoordinate id="93DF50098857FFC08277E530FE92F232" box="[268,376,260,285]" direction="east" orientation="longitude" pageId="8" pageNumber="405" precision="555" value="110.15">110.15°E</geoCoordinate>
),
<date id="8255100E8857FFC082F7E530FDE0F232" box="[396,522,260,285]" pageId="8" pageNumber="405" value="2011-04-23">23-iv-2011</date>
, coll. Shi Weifang.
</paragraph>
</subSubSection>
<subSubSection id="BEF165458857FFC083BCE513FCD4F26F" box="[199,830,295,320]" pageId="8" pageNumber="405" type="distribution">
<paragraph id="F65436CE8857FFC083BCE513FCD4F26F" blockId="8.[151,1436,151,717]" box="[199,830,295,320]" pageId="8" pageNumber="405">
<emphasis id="C49FEADC8857FFC083BCE513FCD4F26F" bold="true" box="[199,830,295,320]" pageId="8" pageNumber="405">
Distribution.
<collectingCountry id="8EFC765E8857FFC0821CE513FE5AF26F" box="[359,432,295,320]" name="China" pageId="8" pageNumber="405">China</collectingCountry>
(Guangdong, Hainan),
<collectingCountry id="8EFC765E8857FFC081B1E513FCD3F26F" box="[714,825,295,320]" name="Malaysia" pageId="8" pageNumber="405">Malaysia</collectingCountry>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="BEF165458857FFC083BCE578FD01F1E3" pageId="8" pageNumber="405" type="discussion">
<paragraph id="F65436CE8857FFC083BCE578FD01F1E3" blockId="8.[151,1436,151,717]" pageId="8" pageNumber="405">
<emphasis id="C49FEADC8857FFC083BCE578FED1F24A" bold="true" box="[199,315,332,357]" pageId="8" pageNumber="405">Remarks.</emphasis>
<taxonomicName id="31EB4D4D8857FFC0823FE579FD5EF24A" authority="Muller-Liebenau" authorityName="Muller-Liebenau" box="[324,692,332,357]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="405" phylum="Arthropoda" rank="species" species="numeratus">
<emphasis id="C49FEADC8857FFC0823FE579FE31F24B" box="[324,475,333,356]" italics="true" pageId="8" pageNumber="405">L. numeratus</emphasis>
(Müller-Liebenau)
</taxonomicName>
is closely similar to
<taxonomicName id="31EB4D4D8857FFC080D7E579FA71F24A" authority="Muller-Liebenau &amp; Hubbard" authorityName="Muller-Liebenau &amp; Hubbard" box="[940,1435,332,357]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="405" phylum="Arthropoda" rank="species" species="ordinatus">
<emphasis id="C49FEADC8857FFC080D7E579FBDCF24B" box="[940,1078,332,356]" italics="true" pageId="8" pageNumber="405">L. ordinatus</emphasis>
(Müller-Liebenau &amp; Hubbard)
</taxonomicName>
from
<collectingCountry id="8EFC765E8857FFC083ADE55BFEACF2A7" box="[214,326,367,392]" name="Sri Lanka" pageId="8" pageNumber="405">Sri Lanka</collectingCountry>
in mouthpart structure because both possess a quite distinctive characteristic within the Oriental species, i.e. having a distinct protuberance between incisor base and molar region on right mandible (
<figureCitation id="6ED02A4B8857FFC08643E5A0FA7AF283" box="[1336,1424,404,429]" captionStart="FIGURES 24 – 27" captionStartId="8.[151,264,1954,1977]" captionTargetBox="[160,1428,789,1913]" captionTargetId="figure@8.[151,1436,769,1933]" captionTargetPageId="8" captionText="FIGURES 24 – 27. Labiobaetis numeratus (Müller-Liebenau) 24, right mandible; 25, pronotum; 26, color pattern of abdomen (left: sterna; right: terga); 27, habitus of male nymph." httpUri="https://zenodo.org/record/225980/files/figure.png" pageId="8" pageNumber="405">Fig. 24</figureCitation>
). However, the color pattern of pronotum and abdomen (
<figureCitation id="6ED02A4B8857FFC0805BE583FC4BF2FF" box="[800,929,439,464]" captionStart="FIGURES 24 – 27" captionStartId="8.[151,264,1954,1977]" captionTargetBox="[160,1428,789,1913]" captionTargetId="figure@8.[151,1436,769,1933]" captionTargetPageId="8" captionText="FIGURES 24 – 27. Labiobaetis numeratus (Müller-Liebenau) 24, right mandible; 25, pronotum; 26, color pattern of abdomen (left: sterna; right: terga); 27, habitus of male nymph." httpUri="https://zenodo.org/record/225980/files/figure.png" pageId="8" pageNumber="405">Fig. 25–27</figureCitation>
), the shape of posterior marginal spines on abdominal terga can be used for differentiating these two species. Similar protuberance on right mandible also exists in Australian genus
<taxonomicName id="31EB4D4D8857FFC082C7E634FB2CF137" authority="Lugo-Ortiz &amp; McCafferty (Lugo-Ortiz &amp; McCafferty 1998)" authorityName="Lugo-Ortiz &amp; McCafferty (Lugo-Ortiz &amp; McCafferty" authorityYear="1998" box="[444,1222,511,536]" class="Insecta" family="Baetidae" genus="Offadens" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="405" phylum="Arthropoda" rank="genus">
<emphasis id="C49FEADC8857FFC082C7E634FDCBF137" box="[444,545,512,536]" italics="true" pageId="8" pageNumber="405">Offadens</emphasis>
<bibRefCitation id="927A4B3F8857FFC0815CE5CBFB2CF137" box="[551,1222,511,536]" pageId="8" pageNumber="405" refString="Lugo-Ortiz, C. R., McCafferty, W. P. &amp; Waltz, R. D. (1998) Offadens, a new genus of small minnow mayflies (Ephemeroptera: Baetidae) from Australia. Proceedings of the Entomological Society of Washington, 100 (2), 306 - 309. http: // dx. doi. org / 10.1051 / limn / 1996022" type="journal article">Lugo-Ortiz &amp; McCafferty (Lugo-Ortiz &amp; McCafferty 1998)</bibRefCitation>
</taxonomicName>
, however, it can be distinguished from
<taxonomicName id="31EB4D4D8857FFC08208E610FE32F113" box="[371,472,548,572]" class="Insecta" family="Baetidae" genus="Offadens" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="405" phylum="Arthropoda" rank="genus">
<emphasis id="C49FEADC8857FFC08208E610FE32F113" box="[371,472,548,572]" italics="true" pageId="8" pageNumber="405">Offadens</emphasis>
</taxonomicName>
by (1) the right mandible with robust prostheca, whereas in
<taxonomicName id="31EB4D4D8857FFC087F0E610FB1AF113" box="[1163,1264,548,572]" class="Insecta" family="Baetidae" genus="Offadens" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="405" phylum="Arthropoda" rank="genus">
<emphasis id="C49FEADC8857FFC087F0E610FB1AF113" box="[1163,1264,548,572]" italics="true" pageId="8" pageNumber="405">Offadens</emphasis>
</taxonomicName>
it is slender or forked (
<bibRefCitation id="927A4B3F8857FFC08389E673FE0AF14F" author="Webb" box="[242,480,583,608]" pageId="8" pageNumber="405" refString="Webb, J. M. &amp; Suter, P. J. (2011) Identification of Larvae of Australian Baetidae. Museum Victoria Science Reports, 15, 1 - 24. Available from: http: // www. museum. vic. gov. au / sciencereports / (accessed 3 June 2014)" type="journal article" year="2011">Webb &amp; Suter 2011</bibRefCitation>
); (2) terminal segment of maxillary palpus with a slightly concave near apex (vs. terminal segment without any concave or notch in
<taxonomicName id="31EB4D4D8857FFC081AAE658FCDCF1AB" box="[721,822,620,644]" class="Insecta" family="Baetidae" genus="Offadens" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="405" phylum="Arthropoda" rank="genus">
<emphasis id="C49FEADC8857FFC081AAE658FCDCF1AB" box="[721,822,620,644]" italics="true" pageId="8" pageNumber="405">Offadens</emphasis>
</taxonomicName>
) and (3) femur with rudimentary villopore, in contrast to the villopore being entirely absent in
<taxonomicName id="31EB4D4D8857FFC08112E6A4FD24F187" box="[617,718,656,680]" class="Insecta" family="Baetidae" genus="Offadens" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="405" phylum="Arthropoda" rank="genus">
<emphasis id="C49FEADC8857FFC08112E6A4FD24F187" box="[617,718,656,680]" italics="true" pageId="8" pageNumber="405">Offadens</emphasis>
</taxonomicName>
which means it can not be considered to belong to the
<taxonomicName id="31EB4D4D8857FFC0862DE6A4FA76F188" box="[1366,1436,656,679]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="405" phylum="Arthropoda" rank="genus">
<emphasis id="C49FEADC8857FFC0862DE6A4FA76F188" box="[1366,1436,656,679]" italics="true" pageId="8" pageNumber="405">Baetis</emphasis>
</taxonomicName>
complex of genera (
<bibRefCitation id="927A4B3F8857FFC0820DE680FD34F1E3" author="Lugo-Ortiz" box="[374,734,692,717]" pageId="8" pageNumber="405" refString="Lugo-Ortiz, C. R., McCafferty, W. P. &amp; Waltz, R. D. (1998) Offadens, a new genus of small minnow mayflies (Ephemeroptera: Baetidae) from Australia. Proceedings of the Entomological Society of Washington, 100 (2), 306 - 309. http: // dx. doi. org / 10.1051 / limn / 1996022" type="journal article" year="1998">Lugo-Ortiz &amp; McCafferty 1998</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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