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<document id="57D94234B08DA33D7B08B5BA71B4238F" ID-DOI="10.5281/zenodo.7666308" ID-ISSN="2305-2562" ID-Zenodo-Dep="7666308" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1677091000000" checkinUser="felipe" docAuthor="Redman, Guy T., Hamer, Michelle L. &amp; Barraclough, David A." docDate="2003" docId="7E3D87A088088007FE7BBA14004FF792" docLanguage="en" docName="AfrInvertebr.44.2.203–277.pdf" docOrigin="African Invertebrates 44 (2)" docStyle="DocumentStyle:C20B10C002986E51083406A4EACF003A.3:AfrInvertebr.2001-2007.journal_article" docStyleId="C20B10C002986E51083406A4EACF003A" docStyleName="AfrInvertebr.2001-2007.journal_article" docStyleVersion="3" docTitle="Harpagophora monodus Attems 1909" docType="treatment" docVersion="2" lastPageNumber="227" masterDocId="8204FFD8881E801FFFECBF750212F119" masterDocTitle="Revision of the Harpagophoridae (Diplopoda, Spirostreptida) of southern Africa, including descriptions of five new species" masterLastPageNumber="277" masterPageNumber="203" pageNumber="225" updateTime="1699455364425" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="1E516E4CEF8E8753FEADB76A76ABC247" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C37CB607524B2A68C7C064144D6DABEE">
<mods:title id="FB285D91EB9F59B1038DF9B60266E626">Revision of the Harpagophoridae (Diplopoda, Spirostreptida) of southern Africa, including descriptions of five new species</mods:title>
</mods:titleInfo>
<mods:name id="D9CF1E4F357028444519AB02485F1C38" type="personal">
<mods:role id="889B31984CE7F318D49D1493362AF324">
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<mods:namePart id="51068D5FBF60CA4483D87C1EE3453C34">Redman, Guy T.</mods:namePart>
</mods:name>
<mods:name id="0FD566AF4E2E4C9BC5753D60BD6506FF" type="personal">
<mods:role id="2825B0CB238A43E41B93E15D04B91B3B">
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<mods:namePart id="98F9B6019E6C2A7E0AB17E1148783C3E">Hamer, Michelle L.</mods:namePart>
</mods:name>
<mods:name id="AE839FECB82B0A377B42A532816F61C0" type="personal">
<mods:role id="D9570D8BE1D233544CC13899911750C7">
<mods:roleTerm id="FA3556AFD5739891CD6A4BD843E41FE6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F40E965645D1B52E0EF9CCF19912A078">Barraclough, David A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="D028B3BF7704A27DC21C14D8969B1C76">text</mods:typeOfResource>
<mods:relatedItem id="1AC1EE868C92F625D31E67B920872BD8" type="host">
<mods:titleInfo id="BC1EAE9685C7066F09D51981476F6B63">
<mods:title id="5059DA15695DB558514A605D4E835650">African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part id="46685C9A71D28E49B5B09C34BD69A343">
<mods:date id="DD20DC5123EAD59C9993FF482EC44C5F">2003</mods:date>
<mods:detail id="6B34D2DF194024661D5F1184302FD982" type="pubDate">
<mods:number id="3D7D6127F0F44E1D369F7D2C750C9AFC">2003-12-31</mods:number>
</mods:detail>
<mods:detail id="BCF93E3E48C199147E94F0E8D16F4239" type="volume">
<mods:number id="7C5A1340D915C5F15213A864EEF1AD36">44</mods:number>
</mods:detail>
<mods:detail id="43106EBFCEF9FD8376BC5A057155C8C9" type="issue">
<mods:number id="2A273BFF46DF7FA61BF0DD634D3124FE">2</mods:number>
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<mods:start id="61854A3F953E79B9F5C53F3B2D8D442C">203</mods:start>
<mods:end id="766222FDA357BE8F9BC0E89FC614B9BA">277</mods:end>
</mods:extent>
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<mods:classification id="8A96DCF8371DE90E22D85D358E29DAB3">journal article</mods:classification>
<mods:identifier id="ABD63012816904B76FA1C9E62A04EA49" type="DOI">10.5281/zenodo.7666308</mods:identifier>
<mods:identifier id="CDE9EEAB5150724FFE21B28213666F0D" type="ISSN">2305-2562</mods:identifier>
<mods:identifier id="9832A316101DA9BEA4C51719A0CF9321" type="Zenodo-Dep">7666308</mods:identifier>
</mods:mods>
<treatment id="7E3D87A088088007FE7BBA14004FF792" LSID="urn:lsid:plazi:treatment:7E3D87A088088007FE7BBA14004FF792" httpUri="http://treatment.plazi.org/id/7E3D87A088088007FE7BBA14004FF792" lastPageId="24" lastPageNumber="227" pageId="22" pageNumber="225">
<subSubSection id="BE8E653D88088009FE7BBA14015AF463" box="[407,840,1377,1402]" pageId="22" pageNumber="225" type="nomenclature">
<paragraph id="F62B36B688088009FE7BBA14015AF463" blockId="22.[144,1104,1377,1674]" box="[407,840,1377,1402]" pageId="22" pageNumber="225">
<heading id="AD6381DA88088009FE7BBA14015AF463" box="[407,840,1377,1402]" centered="true" fontSize="11" level="2" pageId="22" pageNumber="225" reason="3">
<taxonomicName id="31944D3588088009FE7BBA14015AF463" ID-CoL="3JSCD" authority="Attems, 1909" authorityName="Attems" authorityYear="1909" box="[407,840,1377,1402]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="225" phylum="Arthropoda" rank="species" species="monodus">
<emphasis id="C4E0EAA488088009FE7BBA1400B9F460" box="[407,683,1377,1401]" italics="true" pageId="22" pageNumber="225">Harpagophora monodus</emphasis>
<bibRefCitation id="92054B4788088009FD5EBA14015AF463" author="ATTEMS, C." box="[690,840,1377,1402]" pageId="22" pageNumber="225" pagination="1 - 52" refId="ref34241" refString="ATTEMS, C. 1909. Myriopoda. In: Schultze, L., ed., Zoologische und Anthropologische Ergebnisse einer Forschungsreise im Westlichen und Sentralen Sudafrika, ausgefuhrt in den Jahren 1903 - 1905 von Dr Leonard Schultze. Denkschriften Gesellschaft zu Jena 14: 1 - 52." type="journal article" year="1909">Attems, 1909</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BE8E653D88088009FDCFBAFC00AEF4BB" box="[547,700,1417,1442]" pageId="22" pageNumber="225" type="description">
<paragraph id="F62B36B688088009FDCFBAFC00AEF4BB" blockId="22.[144,1104,1377,1674]" box="[547,700,1417,1442]" pageId="22" pageNumber="225">
<figureCitation id="6EAF2A3388088009FDCFBAFC007AF4BB" box="[547,616,1417,1442]" captionStart="Fig" captionStartId="1.[182,215,845,865]" captionTargetBox="[251,1061,203,829]" captionTargetId="figure-7@1.[232,1095,193,840]" captionTargetPageId="1" captionText="Fig. 1. Distribution of Harpagophora, Metaphora and Poratophilus species in southern Africa. G = Gauteng; KZN = KwaZulu-Natal; L = Lesotho; MP = Mpumalanga; S = Swaziland; NW Prov = North West Province; W Cape = Western Cape." figureDoi="http://doi.org/10.5281/zenodo.7666310" httpUri="https://zenodo.org/record/7666310/files/figure.png" pageId="22" pageNumber="225">Figs 1</figureCitation>
,
<figureCitation id="6EAF2A3388088009FD9ABAFC00AEF4BB" box="[630,700,1417,1442]" captionStart="Figs 50–58" captionStartId="23.[182,221,1532,1552]" captionTargetBox="[185,1152,412,1511]" captionTargetId="figure-167@23.[181,1146,408,1519]" captionTargetPageId="23" captionText="Figs 50–58. Harpagophora monodus Attems, 1909. 50. Lateral view of collum; arrow indicates anterior end of millipede. 51. Oral view of first pair of male legs. 52. Aboral view of gonopod (NMSA 19102). 53. Aboral view of gonopods (NMSA 16159). 54. Oral view of telopodite. 55. Detail of femoral spines of telopodite. 56–58.Apical elements of telopodite, showing different orientations. bp = black process; f1 = large femoral spine; f2 = small femoral spine; ll = lateral lobe; mrp = median rounded process; p = pectinophore; s = spine branch." figureDoi="http://doi.org/10.5281/zenodo.7666330" httpUri="https://zenodo.org/record/7666330/files/figure.png" pageId="22" pageNumber="225">50–58</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BE8E653D88088009FF7CBADA0137F4C2" pageId="22" pageNumber="225" type="reference_group">
<paragraph id="F62B36B688088009FF7CBADA01E3F4DA" blockId="22.[144,1104,1377,1674]" box="[144,1009,1455,1475]" pageId="22" pageNumber="225">
<treatmentCitationGroup id="D684119888088009FF7CBADA01E3F4DA" box="[144,1009,1455,1475]" pageId="22" pageNumber="225">
<treatmentCitation id="773510A788088009FF7CBADA0001F4DA" author="ATTEMS, C." box="[144,531,1455,1475]" page="43" pageId="22" pageNumber="225" year="1909">
<taxonomicName id="31944D3588088009FF7CBADA0001F4DA" ID-CoL="3JSCD" authority="Attems, 1909: 43" authorityName="Attems" authorityPageNumber="43" authorityYear="1909" box="[144,531,1455,1475]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="225" phylum="Arthropoda" rank="species" species="monodus">
<emphasis id="C4E0EAA488088009FF7CBADA037DF4DA" box="[144,367,1455,1475]" italics="true" pageId="22" pageNumber="225">Harpagophora monodus</emphasis>
<bibRefCitation id="92054B4788088009FE98BADA0001F4DA" author="ATTEMS, C." box="[372,531,1455,1475]" pageId="22" pageNumber="225" pagination="1 - 52" refId="ref34241" refString="ATTEMS, C. 1909. Myriopoda. In: Schultze, L., ed., Zoologische und Anthropologische Ergebnisse einer Forschungsreise im Westlichen und Sentralen Sudafrika, ausgefuhrt in den Jahren 1903 - 1905 von Dr Leonard Schultze. Denkschriften Gesellschaft zu Jena 14: 1 - 52." type="journal article" year="1909">Attems, 1909: 43</bibRefCitation>
</taxonomicName>
</treatmentCitation>
; Attems 1914 1928; Lawrence 1938;
<bibRefCitation id="92054B4788088009FC8BBADA01FFF4DA" author="SCHUBART, O." box="[871,1005,1455,1475]" pageId="22" pageNumber="225" pagination="154 - 172" refId="ref35110" refString="SCHUBART, O. 1966. Diplopoda III. Pselaphognatha, Opisthospermophora, Colobognatha, zugleich Addenda und allgemeine Bemerkungen. In: Hanstrom, B., Brink, P., &amp; Rudebeck, G., eds, South African Animal Life. Stockholm: Swedish Natural Science Research Council 12: 154 - 172." type="journal article" year="1966">Schubart 1966</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="F62B36B688088009FF7CBAB20137F4C2" blockId="22.[144,1104,1377,1674]" box="[144,805,1479,1499]" pageId="22" pageNumber="225">
<taxonomicName id="31944D3588088009FF7CBAB2001EF4C2" ID-CoL="3JSCD" authorityName="Attems" authorityYear="1909" box="[144,524,1479,1499]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="225" phylum="Arthropoda" rank="species" species="monodus" subGenus="Harpagophora">
<emphasis id="C4E0EAA488088009FF7CBAB2030AF4C2" box="[144,280,1479,1499]" italics="true" pageId="22" pageNumber="225">Harpagophora</emphasis>
(
<emphasis id="C4E0EAA488088009FEC9BAB203BFF4C2" box="[293,429,1479,1499]" italics="true" pageId="22" pageNumber="225">Harpagophora</emphasis>
)
<emphasis id="C4E0EAA488088009FE56BAB2001EF4C2" box="[442,524,1479,1499]" italics="true" pageId="22" pageNumber="225">monodus</emphasis>
</taxonomicName>
;
<bibRefCitation id="92054B4788088009FDF4BAB200B6F4C2" author="DEMANGE, J. - M." box="[536,676,1479,1499]" pageId="22" pageNumber="225" pagination="561 - 564" refId="ref34494" refString="DEMANGE, J. - M. 1983. Donnees nouvelles sur la famille des Harpagophoridae (Myriapoda, Diplopoda). Bulletin de Museum National d'Histoire Naturelle Paris (4) 5: 561 - 564." type="journal article" year="1983">Demange 1983</bibRefCitation>
;
<bibRefCitation id="92054B4788088009FD42BAB20133F4C2" author="HAMER, M." box="[686,801,1479,1499]" pageId="22" pageNumber="225" pagination="11 - 82" refId="ref34653" refString="HAMER, M. 1998. Checklist of southern African millipedes. Annals of the Natal Museum 39: 11 - 82." type="journal article" year="1998">Hamer 1998</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BE8E653D88088007FF7CBA9C004FF792" lastPageId="24" lastPageNumber="227" pageId="22" pageNumber="225" type="materials_examined">
<paragraph id="F62B36B688088009FF7CBA9C0625F75B" blockId="22.[144,1104,1377,1674]" pageId="22" pageNumber="225">
<materialsCitation id="46FC3CEB88088009FF7CBA9C0621F75B" location="Swartjies River" municipality="Skilpad Nature Reserve" pageId="22" pageNumber="225" specimenCount="4" stateProvince="Northern Cape" typeStatus="holotype">
<typeStatus id="292F881488088009FF7CBA9C02D5F71B" box="[144,199,1513,1538]" pageId="22" pageNumber="225">Type</typeStatus>
material (not examined): The whereabouts of the
<typeStatus id="292F881488088009FD13BA9C013DF71B" box="[767,815,1513,1538]" pageId="22" pageNumber="225">type</typeStatus>
material is unknown, but the
<typeStatus id="292F881488088009FF55B97C02FBF73B" box="[185,233,1545,1570]" pageId="22" pageNumber="225">type</typeStatus>
locality was given as ‘Cape’ (
<bibRefCitation id="92054B4788088009FDD6B97C00DDF73B" author="ATTEMS, C." box="[570,719,1545,1570]" pageId="22" pageNumber="225" pagination="1 - 52" refId="ref34241" refString="ATTEMS, C. 1909. Myriopoda. In: Schultze, L., ed., Zoologische und Anthropologische Ergebnisse einer Forschungsreise im Westlichen und Sentralen Sudafrika, ausgefuhrt in den Jahren 1903 - 1905 von Dr Leonard Schultze. Denkschriften Gesellschaft zu Jena 14: 1 - 52." type="journal article" year="1909">Attems 1909</bibRefCitation>
). The
<typeStatus id="292F881488088009FCF9B97C0157F73B" box="[789,837,1545,1570]" pageId="22" pageNumber="225">type</typeStatus>
locality is here fixed as
<collectingRegion id="3450F85488088009FF7CB95C032AF75B" box="[144,312,1577,1602]" country="South Africa" name="Northern Cape" pageId="22" pageNumber="225">Northern Cape</collectingRegion>
,
<collectingMunicipality id="164FACCC88088009FEAFB95C0042F75B" box="[323,592,1577,1602]" pageId="22" pageNumber="225">Skilpad Nature Reserve</collectingMunicipality>
,
<location id="F34B606D88088009FDB7B95C011AF75B" LSID="urn:lsid:plazi:treatment:7E3D87A088088007FE7BBA14004FF792:F34B606D88088009FDB7B95C011AF75B" box="[603,776,1577,1602]" municipality="Skilpad Nature Reserve" name="Swartjies River" pageId="22" pageNumber="225" stateProvince="Northern Cape">Swartjies River</location>
(30.16596˚S:17.79866˚E)
</materialsCitation>
.
</paragraph>
<paragraph id="F62B36B688088007FF7CB92402CDF316" blockId="22.[144,1104,1377,1674]" lastBlockId="24.[144,1104,206,1675]" lastPageId="24" lastPageNumber="227" pageId="22" pageNumber="225">
<materialsCitation id="46FC3CEB88088009FF7CB9240123F793" collectingDate="1962-06-04" collectionCode="NMSA" collectorName="W. J. Lawson" country="South Africa" location="Port Nolloth" municipality="Material" pageId="22" pageNumber="225" specimenCode="NMSA18922" specimenCount="1" stateProvince="Northern Cape">
<collectingMunicipality id="164FACCC88088009FF7CB92402FDF773" box="[144,239,1617,1642]" pageId="22" pageNumber="225">Material</collectingMunicipality>
examined:
<collectingCountry id="8E83762688088009FE9CB924002FF773" box="[368,573,1617,1642]" name="South Africa" pageId="22" pageNumber="225">SOUTH AFRICA</collectingCountry>
:
<emphasis id="C4E0EAA488088009FDAAB92400FCF770" box="[582,750,1617,1641]" italics="true" pageId="22" pageNumber="225">
<collectingRegion id="3450F85488088009FDAAB92400FCF770" box="[582,750,1617,1641]" country="South Africa" name="Northern Cape" pageId="22" pageNumber="225">Northern Cape</collectingRegion>
</emphasis>
: 1ơ2^,
<quantity id="316C9B5388088009FCBEB92401A3F773" box="[850,945,1617,1642]" metricMagnitude="4" metricUnit="m" metricValue="2.2530816" pageId="22" pageNumber="225" unit="mi" value="14.0">14 miles</quantity>
inland of
<location id="F34B606D88088009FBCDB92402F4F793" LSID="urn:lsid:plazi:treatment:7E3D87A088088007FE7BBA14004FF792:F34B606D88088009FBCDB92402F4F793" country="South Africa" municipality="Material" name="Port Nolloth" pageId="22" pageNumber="225" stateProvince="Northern Cape">Port Nolloth</location>
[2916BD],
<date id="822A107688088009FE80B90403CBF793" box="[364,473,1649,1674]" pageId="22" pageNumber="225" value="1962-06-04">
<collectingDate id="926EE99E88088009FE80B90403CBF793" box="[364,473,1649,1674]" pageId="22" pageNumber="225" value="1962-06-04">4.vi.1962</collectingDate>
</date>
,
<collectorName id="5B61536088088009FE08B904006EF793" box="[484,636,1649,1674]" pageId="22" pageNumber="225">W. J. Lawson</collectorName>
(
<specimenCode id="A6329ECD88088009FD67B904013BF793" box="[651,809,1649,1674]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="22" pageNumber="225" type="Museum">NMSA18922</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="46FC3CEB88088008FCD0B90401C3F1FE" collectingDate="1962-11" collectionCode="NMSA" collectorName="W. Haacke" country="South Africa" county="Little Namaqualand" lastPageId="23" lastPageNumber="226" location="Anenous Pass" municipality="Springbok" pageId="22" pageNumber="225" specimenCode="NMSA 18943" specimenCount="1" stateProvince="Northern Cape">
2ơ,
<collectingCounty id="1F4A4E3A88088009FC85B9040659F793" box="[873,1099,1649,1674]" pageId="22" pageNumber="225">Little Namaqualand</collectingCounty>
,
<collectingMunicipality id="164FACCC88098008FF59BFBB0338F1FE" box="[181,298,206,231]" pageId="23" pageNumber="226">Springbok</collectingMunicipality>
,
<location id="F34B606D88098008FEDEBFBB03D8F1FE" LSID="urn:lsid:plazi:treatment:7E3D87A088088007FE7BBA14004FF792:F34B606D88098008FEDEBFBB03D8F1FE" box="[306,458,206,231]" country="South Africa" county="Little Namaqualand" municipality="Springbok" name="Anenous Pass" pageId="23" pageNumber="226" stateProvince="Northern Cape">Anenous Pass</location>
[2917BA],
<date id="822A107688098008FDA6BFBB00B2F1FE" box="[586,672,206,231]" pageId="23" pageNumber="226" value="1962-11">
<collectingDate id="926EE99E88098008FDA6BFBB00B2F1FE" box="[586,672,206,231]" pageId="23" pageNumber="226" value="1962-11">xi.1962</collectingDate>
</date>
,
<collectorName id="5B61536088098008FD44BFBB010DF1FE" box="[680,799,206,231]" pageId="23" pageNumber="226">W. Haacke</collectorName>
(
<specimenCode id="A6329ECD88098008FCC6BFBB01DAF1FE" box="[810,968,206,231]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="23" pageNumber="226" type="Museum">NMSA 18943</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="46FC3CEB88098008FC36BFBB03EBF05E" collectingDate="1986-08-29" collectionCode="NMSA" collectorName="P. Croeser &amp; J. Londt &amp; Stuckenberg" country="South Africa" location="Studers Pass" municipality="Garies" pageId="23" pageNumber="226" specimenCode="NMSA 16159" specimenCount="1" stateProvince="Northern Cape">
1ơ3^,
<collectingMunicipality id="164FACCC88098008FBC2BFBB0664F1FE" box="[1070,1142,206,231]" pageId="23" pageNumber="226">Garies</collectingMunicipality>
[3018CA],
<quantity id="316C9B5388098008FEDBBF9B0363F01E" box="[311,369,238,263]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="23" pageNumber="226" unit="km" value="6.0">6 km</quantity>
NE of and south of road to
<location id="F34B606D88098008FD5BBF9B015AF01E" LSID="urn:lsid:plazi:treatment:7E3D87A088088007FE7BBA14004FF792:F34B606D88098008FD5BBF9B015AF01E" box="[695,840,238,263]" country="South Africa" municipality="Garies" name="Studers Pass" pageId="23" pageNumber="226" stateProvince="Northern Cape">Studers Pass</location>
, on hill slope facing north under stones and walking around on ground
<date id="822A107688098008FD55BE7B0151F03E" box="[697,835,270,295]" pageId="23" pageNumber="226" value="1986-08-29">
<collectingDate id="926EE99E88098008FD55BE7B0151F03E" box="[697,835,270,295]" pageId="23" pageNumber="226" value="1986-08-29">29.viii.1986</collectingDate>
</date>
,
<collectorName id="5B61536088098008FCBDBE7B01D5F03E" box="[849,967,270,295]" pageId="23" pageNumber="226">P. Croeser</collectorName>
,
<collectorName id="5B61536088098008FC39BE7B0621F03E" box="[981,1075,270,295]" pageId="23" pageNumber="226">J. Londt</collectorName>
&amp; B.
<collectorName id="5B61536088098008FF59BE5B0352F05E" box="[181,320,302,327]" pageId="23" pageNumber="226">Stuckenberg</collectorName>
(
<specimenCode id="A6329ECD88098008FEA3BE5B03E3F05E" box="[335,497,302,327]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="23" pageNumber="226" type="Museum">NMSA 16159</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="46FC3CEB88098008FDE8BE5B0083F07E" collectingDate="1994-09-28" collectionCode="NMSA" collectorName="P. Reavell" country="South Africa" location="Hardeveld" municipality="Kamiesberg Pass" pageId="23" pageNumber="226" specimenCode="NMSA 15632" specimenCount="1" stateProvince="Northern Cape">
1ơ,
<collectingMunicipality id="164FACCC88098008FDDCBE5B00E2F05E" box="[560,752,302,327]" pageId="23" pageNumber="226">Kamiesberg Pass</collectingMunicipality>
[3018AC],
<location id="F34B606D88098008FC99BE5B01FFF05E" LSID="urn:lsid:plazi:treatment:7E3D87A088088007FE7BBA14004FF792:F34B606D88098008FC99BE5B01FFF05E" box="[885,1005,302,327]" country="South Africa" municipality="Kamiesberg Pass" name="Hardeveld" pageId="23" pageNumber="226" stateProvince="Northern Cape">Hardeveld</location>
, under rock,
<date id="822A107688098008FF59BE3B032AF07E" box="[181,312,334,359]" pageId="23" pageNumber="226" value="1994-09-28">
<collectingDate id="926EE99E88098008FF59BE3B032AF07E" box="[181,312,334,359]" pageId="23" pageNumber="226" value="1994-09-28">28.ix.1994</collectingDate>
</date>
,
<collectorName id="5B61536088098008FEA5BE3B03D7F07E" box="[329,453,334,359]" pageId="23" pageNumber="226">P. Reavell</collectorName>
(
<specimenCode id="A6329ECD88098008FE35BE3B009AF07E" box="[473,648,334,359]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="23" pageNumber="226" type="Museum">NMSA 15632</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="46FC3CEB88098007FD4EBE3B017FF1FE" collectingDate="1994-09-29" collectionCode="NMSA" collectorName="P. Reavell" country="South Africa" elevation="900" lastPageId="24" lastPageNumber="227" location="Vlieholteberg" municipality="South Richtersveld" pageId="23" pageNumber="226" specimenCode="NMSA 16489" specimenCount="1" stateProvince="Northern Cape">
1ơ,
<collectingMunicipality id="164FACCC88098008FD3ABE3B01D0F07E" box="[726,962,334,359]" pageId="23" pageNumber="226">South Richtersveld</collectingMunicipality>
,
<location id="F34B606D88098008FC3DBE3B0667F07E" LSID="urn:lsid:plazi:treatment:7E3D87A088088007FE7BBA14004FF792:F34B606D88098008FC3DBE3B0667F07E" box="[977,1141,334,359]" country="South Africa" municipality="South Richtersveld" name="Vlieholteberg" pageId="23" pageNumber="226" stateProvince="Northern Cape">Vlieholteberg</location>
(29˚10'S:17˚37'E),
<quantity id="316C9B5388068007FE8ABFBB03BDF1FE" box="[358,431,206,231]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="24" pageNumber="227" unit="m" value="900.0">
<elevation id="7DB9D18588068007FE8ABFBB03BDF1FE" box="[358,431,206,231]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="24" pageNumber="227" unit="m" value="900.0">900 m</elevation>
</quantity>
,
<date id="822A107688068007FE50BFBB0024F1FE" box="[444,566,206,231]" pageId="24" pageNumber="227" value="1994-09-29">
<collectingDate id="926EE99E88068007FE50BFBB0024F1FE" box="[444,566,206,231]" pageId="24" pageNumber="227" value="1994-09-29">29.ix.1994</collectingDate>
</date>
,
<collectorName id="5B61536088068007FDAEBFBB00A1F1FE" box="[578,691,206,231]" pageId="24" pageNumber="227">P. Reavell</collectorName>
(
<specimenCode id="A6329ECD88068007FD2DBFBB0177F1FE" box="[705,869,206,231]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="24" pageNumber="227" type="Museum">NMSA 16489</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="46FC3CEB88068007FC95BFBB003BF03E" collectingDate="2001-09-10" collectionCode="NMSA" collectorName="R. Slotow &amp; M. Hamer" country="South Africa" county="Skilpad Nature Reserve" location="Succulent Karoo" municipality="Swartjies River" pageId="24" pageNumber="227" specimenCode="NMSA 19101" specimenCount="1" stateProvince="Northern Cape">
2ơ,
<collectingCounty id="1F4A4E3A88068007FC4BBFBB02FFF01E" pageId="24" pageNumber="227">Skilpad Nature Reserve</collectingCounty>
,
<collectingMunicipality id="164FACCC88068007FF15BF9B03BBF01E" box="[249,425,238,263]" pageId="24" pageNumber="227">Swartjies River</collectingMunicipality>
, (30.16596˚S:17.79866˚E),
<location id="F34B606D88068007FD08BF9B01B1F01E" LSID="urn:lsid:plazi:treatment:7E3D87A088088007FE7BBA14004FF792:F34B606D88068007FD08BF9B01B1F01E" box="[740,931,238,263]" country="South Africa" county="Skilpad Nature Reserve" municipality="Swartjies River" name="Succulent Karoo" pageId="24" pageNumber="227" stateProvince="Northern Cape">Succulent Karoo</location>
,
<date id="822A107688068007FC43BF9B063BF01E" box="[943,1065,238,263]" pageId="24" pageNumber="227" value="2001-09-10">
<collectingDate id="926EE99E88068007FC43BF9B063BF01E" box="[943,1065,238,263]" pageId="24" pageNumber="227" value="2001-09-10">10.ix.2001</collectingDate>
</date>
,
<collectorName id="5B61536088068007FBDABF9B02CDF03E" pageId="24" pageNumber="227">R. Slotow</collectorName>
&amp;
<collectorName id="5B61536088068007FEECBE7B0360F03E" box="[256,370,270,295]" pageId="24" pageNumber="227">M. Hamer</collectorName>
(
<specimenCode id="A6329ECD88068007FE93BE7B0033F03E" box="[383,545,270,295]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="24" pageNumber="227" type="Museum">NMSA 19101</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="46FC3CEB88068007FDD8BE7B00AAF05E" collectionCode="SAMC" country="South Africa" location="Little Namaqualand" municipality="Garies" pageId="24" pageNumber="227" specimenCode="C2102" specimenCount="1" stateProvince="Northern Cape">
1ơ 4^,
<collectingMunicipality id="164FACCC88068007FD63BE7B00CAF03E" box="[655,728,270,295]" pageId="24" pageNumber="227">Garies</collectingMunicipality>
[3018CA],
<location id="F34B606D88068007FCB0BE7B062EF03E" LSID="urn:lsid:plazi:treatment:7E3D87A088088007FE7BBA14004FF792:F34B606D88068007FCB0BE7B062EF03E" box="[860,1084,270,295]" country="South Africa" municipality="Garies" name="Little Namaqualand" pageId="24" pageNumber="227" stateProvince="Northern Cape">Little Namaqualand</location>
, - no original label with specimen lot (
<collectionCode id="9085AE7388068007FDE1BE5B004FF05E" box="[525,605,302,327]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="24" pageNumber="227" type="Museum">SAMC</collectionCode>
<specimenCode id="A6329ECD88068007FD8EBE5B00BDF05E" box="[610,687,302,327]" collectionCode="SAMC" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="24" pageNumber="227" type="Museum">C2102</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="46FC3CEB88068007FD53BE5B065EF05E" box="[703,1100,302,327]" collectionCode="SAMC" country="South Africa" pageId="24" pageNumber="227" specimenCode="B7674" specimenCount="1" stateProvince="Northern Cape">
<taxonomicName id="31944D3588068007FD53BE5B017EF05F" box="[703,876,302,326]" genus="No" pageId="24" pageNumber="227" rank="subSpecies" species="locality" subSpecies="data">
<emphasis id="C4E0EAA488068007FD53BE5B017EF05F" box="[703,876,302,326]" italics="true" pageId="24" pageNumber="227">No locality data</emphasis>
</taxonomicName>
: 2ơ (
<collectionCode id="9085AE7388068007FC4EBE5B01E0F05E" box="[930,1010,302,327]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="24" pageNumber="227" type="Museum">SAMC</collectionCode>
<specimenCode id="A6329ECD88068007FC1BBE5B0656F05E" box="[1015,1092,302,327]" collectionCode="SAMC" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="24" pageNumber="227" type="Museum">B7674</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="46FC3CEB88068007FF7CBE2302C9F316" collectorName="Telocoxite &amp; Lateral &amp; Telocoxal" country="South Africa" county="Diagnosis" location="Spine" municipality="Distinct" pageId="24" pageNumber="227" specimenCount="1" stateProvince="Northern Cape">
<emphasis id="C4E0EAA488068007FF7CBE230310F077" box="[144,258,342,366]" italics="true" pageId="24" pageNumber="227">
<collectingCounty id="1F4A4E3A88068007FF7CBE230310F077" box="[144,258,342,366]" pageId="24" pageNumber="227">Diagnosis</collectingCounty>
</emphasis>
:
<collectingMunicipality id="164FACCC88068007FEFCBE23037BF076" box="[272,361,342,367]" pageId="24" pageNumber="227">Distinct</collectingMunicipality>
black projection at apex of larger femoral spine, directed laterally at right angle to major part of spine (
<figureCitation id="6EAF2A3388068007FDF1BE030062F096" box="[541,624,374,399]" captionStart="Figs 50–58" captionStartId="23.[182,221,1532,1552]" captionTargetBox="[185,1152,412,1511]" captionTargetId="figure-167@23.[181,1146,408,1519]" captionTargetPageId="23" captionText="Figs 50–58. Harpagophora monodus Attems, 1909. 50. Lateral view of collum; arrow indicates anterior end of millipede. 51. Oral view of first pair of male legs. 52. Aboral view of gonopod (NMSA 19102). 53. Aboral view of gonopods (NMSA 16159). 54. Oral view of telopodite. 55. Detail of femoral spines of telopodite. 56–58.Apical elements of telopodite, showing different orientations. bp = black process; f1 = large femoral spine; f2 = small femoral spine; ll = lateral lobe; mrp = median rounded process; p = pectinophore; s = spine branch." figureDoi="http://doi.org/10.5281/zenodo.7666330" httpUri="https://zenodo.org/record/7666330/files/figure.png" pageId="24" pageNumber="227">Fig. 55</figureCitation>
).
<location id="F34B606D88068007FD6DBE0300D3F096" LSID="urn:lsid:plazi:treatment:7E3D87A088088007FE7BBA14004FF792:F34B606D88068007FD6DBE0300D3F096" box="[641,705,374,399]" country="South Africa" county="Diagnosis" municipality="Distinct" name="Spine" pageId="24" pageNumber="227" stateProvince="Northern Cape">Spine</location>
branch a laminate plate, broadening apically with smooth margin and one large, curved, black tooth at one end (
<figureCitation id="6EAF2A3388068007FC02BEE30651F0B6" box="[1006,1091,406,431]" captionStart="Figs 50–58" captionStartId="23.[182,221,1532,1552]" captionTargetBox="[185,1152,412,1511]" captionTargetId="figure-167@23.[181,1146,408,1519]" captionTargetPageId="23" captionText="Figs 50–58. Harpagophora monodus Attems, 1909. 50. Lateral view of collum; arrow indicates anterior end of millipede. 51. Oral view of first pair of male legs. 52. Aboral view of gonopod (NMSA 19102). 53. Aboral view of gonopods (NMSA 16159). 54. Oral view of telopodite. 55. Detail of femoral spines of telopodite. 56–58.Apical elements of telopodite, showing different orientations. bp = black process; f1 = large femoral spine; f2 = small femoral spine; ll = lateral lobe; mrp = median rounded process; p = pectinophore; s = spine branch." figureDoi="http://doi.org/10.5281/zenodo.7666330" httpUri="https://zenodo.org/record/7666330/files/figure.png" pageId="24" pageNumber="227">Fig. 57</figureCitation>
).
<collectorName id="5B61536088068007FF7CBEC30314F0D6" box="[144,262,438,463]" pageId="24" pageNumber="227">Telocoxite</collectorName>
rounded with short, blunt protuberance on medial distal margin.
<collectorName id="5B61536088068007FBECBEC3065DF0D6" box="[1024,1103,438,463]" pageId="24" pageNumber="227">Lateral</collectorName>
margin of telocoxite forming obscure, broadly rounded lobe (
<figureCitation id="6EAF2A3388068007FCD6BEA3019FF0F6" box="[826,909,470,495]" captionStart="Figs 50–58" captionStartId="23.[182,221,1532,1552]" captionTargetBox="[185,1152,412,1511]" captionTargetId="figure-167@23.[181,1146,408,1519]" captionTargetPageId="23" captionText="Figs 50–58. Harpagophora monodus Attems, 1909. 50. Lateral view of collum; arrow indicates anterior end of millipede. 51. Oral view of first pair of male legs. 52. Aboral view of gonopod (NMSA 19102). 53. Aboral view of gonopods (NMSA 16159). 54. Oral view of telopodite. 55. Detail of femoral spines of telopodite. 56–58.Apical elements of telopodite, showing different orientations. bp = black process; f1 = large femoral spine; f2 = small femoral spine; ll = lateral lobe; mrp = median rounded process; p = pectinophore; s = spine branch." figureDoi="http://doi.org/10.5281/zenodo.7666330" httpUri="https://zenodo.org/record/7666330/files/figure.png" pageId="24" pageNumber="227">Fig. 53</figureCitation>
).
<collectorName id="5B61536088068007FC4CBEA3061CF0F6" box="[928,1038,470,495]" pageId="24" pageNumber="227">Telocoxal</collectorName>
spine absent
</materialsCitation>
.
</paragraph>
<caption id="A2EB663E88098008FF5ABA89012DF791" ID-DOI="http://doi.org/10.5281/zenodo.7666330" ID-Zenodo-Dep="7666330" httpUri="https://zenodo.org/record/7666330/files/figure.png" pageId="23" pageNumber="226" startId="23.[182,221,1532,1552]" targetBox="[185,1152,412,1511]" targetPageId="23">
<paragraph id="F62B36B688098008FF5ABA89012DF791" blockId="23.[182,1143,1532,1672]" pageId="23" pageNumber="226">
Figs 50–58.
<taxonomicName id="31944D3588098008FEC5BA89009CF709" authority="Attems, 1909" authorityName="Attems" authorityYear="1909" box="[297,654,1532,1552]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="226" phylum="Arthropoda" rank="species" species="monodus">
<emphasis id="C4E0EAA488098008FEC5BA890018F709" box="[297,522,1532,1552]" italics="true" pageId="23" pageNumber="226">Harpagophora monodus</emphasis>
<bibRefCitation id="92054B4788098008FDFCBA89009CF709" author="ATTEMS, C." box="[528,654,1532,1552]" pageId="23" pageNumber="226" pagination="1 - 52" refId="ref34241" refString="ATTEMS, C. 1909. Myriopoda. In: Schultze, L., ed., Zoologische und Anthropologische Ergebnisse einer Forschungsreise im Westlichen und Sentralen Sudafrika, ausgefuhrt in den Jahren 1903 - 1905 von Dr Leonard Schultze. Denkschriften Gesellschaft zu Jena 14: 1 - 52." type="journal article" year="1909">Attems, 1909</bibRefCitation>
</taxonomicName>
. 50. Lateral view of collum; arrow indicates anterior end of millipede. 51. Oral view of first pair of male legs. 52. Aboral view of gonopod (NMSA 19102). 53. Aboral view of gonopods (NMSA 16159). 54. Oral view of telopodite. 55. Detail of femoral spines of telopodite. 56–58.Apical elements of telopodite, showing different orientations. bp = black process; f1 = large femoral spine; f2 = small femoral spine; ll = lateral lobe; mrp = median rounded process; p = pectinophore; s = spine branch.
</paragraph>
</caption>
<paragraph id="F62B36B688068007FF7CBD680309F32F" blockId="24.[144,1104,206,1675]" box="[144,283,541,566]" pageId="24" pageNumber="227">Description:</paragraph>
<paragraph id="F62B36B688068007FF7CBD3102A8F3A4" blockId="24.[144,1104,206,1675]" pageId="24" pageNumber="227">
<emphasis id="C4E0EAA488068007FF7CBD310306F345" box="[144,276,580,604]" italics="true" pageId="24" pageNumber="227">Dimensions</emphasis>
: Males, n = 8; females, n = 2. Body width 5.7–8.3 [7.3], female 9.0–11.0; collum width 7.0–8.7 [6.9], female 7.8–9.0; body length 65.0–100.0 [81.3], female 85.0–86.0; leg length 2.6–5.0, female 2.5–4.0; antenna length 5.0–5.4 [5.1], female 5.0.
</paragraph>
<paragraph id="F62B36B688068007FF7CBDB10091F3C4" blockId="24.[144,1104,206,1675]" box="[144,643,708,733]" pageId="24" pageNumber="227">
<emphasis id="C4E0EAA488068007FF7CBDB10366F3C5" box="[144,372,708,732]" italics="true" pageId="24" pageNumber="227">Number of segments</emphasis>
: 47–51 [49], female 49.
</paragraph>
<paragraph id="F62B36B688068007FF7CBD9103E6F224" blockId="24.[144,1104,206,1675]" pageId="24" pageNumber="227">
<emphasis id="C4E0EAA488068007FF7CBD9102CDF3E5" box="[144,223,740,764]" italics="true" pageId="24" pageNumber="227">Colour</emphasis>
: Head, clypeus and collum light brown. Prozonites brown, mesozonites light brown, metazonites dark brown. Anal valve and caudal spine light brown. Legs and antennae light brown to orange.
</paragraph>
<paragraph id="F62B36B688068007FF7CBC3103A1F244" blockId="24.[144,1104,206,1675]" box="[144,435,836,861]" pageId="24" pageNumber="227">
<emphasis id="C4E0EAA488068007FF7CBC310334F245" box="[144,294,836,860]" italics="true" pageId="24" pageNumber="227">First ozopore</emphasis>
: Segment 6.
</paragraph>
<paragraph id="F62B36B688068007FF7CBC110127F284" blockId="24.[144,1104,206,1675]" pageId="24" pageNumber="227">
<emphasis id="C4E0EAA488068007FF7CBC1102F0F265" box="[144,226,868,892]" italics="true" pageId="24" pageNumber="227">Collum</emphasis>
: Anterior corner with approximately 90˚ angle, posterior corner passing a little beyond posterior margin (
<figureCitation id="6EAF2A3388068007FE5EBCF10014F284" box="[434,518,900,925]" captionStart="Figs 50–58" captionStartId="23.[182,221,1532,1552]" captionTargetBox="[185,1152,412,1511]" captionTargetId="figure-167@23.[181,1146,408,1519]" captionTargetPageId="23" captionText="Figs 50–58. Harpagophora monodus Attems, 1909. 50. Lateral view of collum; arrow indicates anterior end of millipede. 51. Oral view of first pair of male legs. 52. Aboral view of gonopod (NMSA 19102). 53. Aboral view of gonopods (NMSA 16159). 54. Oral view of telopodite. 55. Detail of femoral spines of telopodite. 56–58.Apical elements of telopodite, showing different orientations. bp = black process; f1 = large femoral spine; f2 = small femoral spine; ll = lateral lobe; mrp = median rounded process; p = pectinophore; s = spine branch." figureDoi="http://doi.org/10.5281/zenodo.7666330" httpUri="https://zenodo.org/record/7666330/files/figure.png" pageId="24" pageNumber="227">Fig. 50</figureCitation>
). One submarginal groove.
</paragraph>
<paragraph id="F62B36B688068007FF7CBCD1000FF5E4" blockId="24.[144,1104,206,1675]" pageId="24" pageNumber="227">
<emphasis id="C4E0EAA488068007FF7CBCD10313F2A5" box="[144,257,932,956]" italics="true" pageId="24" pageNumber="227">Gonopods</emphasis>
: Larger femoral spine directed distally, apex broad with black spine positioned at right angle and pointing towards lateral margin of telocoxite. Margin of area at base of black spine serrated (
<figureCitation id="6EAF2A3388068007FE49BC9103EEF2E4" box="[421,508,996,1021]" captionStart="Figs 50–58" captionStartId="23.[182,221,1532,1552]" captionTargetBox="[185,1152,412,1511]" captionTargetId="figure-167@23.[181,1146,408,1519]" captionTargetPageId="23" captionText="Figs 50–58. Harpagophora monodus Attems, 1909. 50. Lateral view of collum; arrow indicates anterior end of millipede. 51. Oral view of first pair of male legs. 52. Aboral view of gonopod (NMSA 19102). 53. Aboral view of gonopods (NMSA 16159). 54. Oral view of telopodite. 55. Detail of femoral spines of telopodite. 56–58.Apical elements of telopodite, showing different orientations. bp = black process; f1 = large femoral spine; f2 = small femoral spine; ll = lateral lobe; mrp = median rounded process; p = pectinophore; s = spine branch." figureDoi="http://doi.org/10.5281/zenodo.7666330" httpUri="https://zenodo.org/record/7666330/files/figure.png" pageId="24" pageNumber="227">Fig. 55</figureCitation>
). Smaller femoral spine slender and partially curled round base of larger femoral spine (
<figureCitation id="6EAF2A3388068007FDFEBB710071F504" box="[530,611,1028,1053]" captionStart="Figs 50–58" captionStartId="23.[182,221,1532,1552]" captionTargetBox="[185,1152,412,1511]" captionTargetId="figure-167@23.[181,1146,408,1519]" captionTargetPageId="23" captionText="Figs 50–58. Harpagophora monodus Attems, 1909. 50. Lateral view of collum; arrow indicates anterior end of millipede. 51. Oral view of first pair of male legs. 52. Aboral view of gonopod (NMSA 19102). 53. Aboral view of gonopods (NMSA 16159). 54. Oral view of telopodite. 55. Detail of femoral spines of telopodite. 56–58.Apical elements of telopodite, showing different orientations. bp = black process; f1 = large femoral spine; f2 = small femoral spine; ll = lateral lobe; mrp = median rounded process; p = pectinophore; s = spine branch." figureDoi="http://doi.org/10.5281/zenodo.7666330" httpUri="https://zenodo.org/record/7666330/files/figure.png" pageId="24" pageNumber="227">Fig. 55</figureCitation>
). Pectinophore with laminate plate expanded apically, hollow, saucer-like and curved inwards towards other apical elements. Surface of pectinophore with several setiform processes along entire margin of branch (
<figureCitation id="6EAF2A3388068007FC1ABB3102A2F564" captionStart="Figs 50–58" captionStartId="23.[182,221,1532,1552]" captionTargetBox="[185,1152,412,1511]" captionTargetId="figure-167@23.[181,1146,408,1519]" captionTargetPageId="23" captionText="Figs 50–58. Harpagophora monodus Attems, 1909. 50. Lateral view of collum; arrow indicates anterior end of millipede. 51. Oral view of first pair of male legs. 52. Aboral view of gonopod (NMSA 19102). 53. Aboral view of gonopods (NMSA 16159). 54. Oral view of telopodite. 55. Detail of femoral spines of telopodite. 56–58.Apical elements of telopodite, showing different orientations. bp = black process; f1 = large femoral spine; f2 = small femoral spine; ll = lateral lobe; mrp = median rounded process; p = pectinophore; s = spine branch." figureDoi="http://doi.org/10.5281/zenodo.7666330" httpUri="https://zenodo.org/record/7666330/files/figure.png" pageId="24" pageNumber="227">Figs 56, 57</figureCitation>
). Spine branch a laminate plate with apical black spine (
<figureCitation id="6EAF2A3388068007FCC0BB110193F564" box="[812,897,1124,1149]" captionStart="Figs 50–58" captionStartId="23.[182,221,1532,1552]" captionTargetBox="[185,1152,412,1511]" captionTargetId="figure-167@23.[181,1146,408,1519]" captionTargetPageId="23" captionText="Figs 50–58. Harpagophora monodus Attems, 1909. 50. Lateral view of collum; arrow indicates anterior end of millipede. 51. Oral view of first pair of male legs. 52. Aboral view of gonopod (NMSA 19102). 53. Aboral view of gonopods (NMSA 16159). 54. Oral view of telopodite. 55. Detail of femoral spines of telopodite. 56–58.Apical elements of telopodite, showing different orientations. bp = black process; f1 = large femoral spine; f2 = small femoral spine; ll = lateral lobe; mrp = median rounded process; p = pectinophore; s = spine branch." figureDoi="http://doi.org/10.5281/zenodo.7666330" httpUri="https://zenodo.org/record/7666330/files/figure.png" pageId="24" pageNumber="227">Fig. 57</figureCitation>
). Telocoxal spines absent. Telocoxite rounded with short, blunt protuberance on medial distal margin. Surface of telocoxite raised between medial margin and larger femoral spine, such that first femoral spine partially concealed. Outer margin of telocoxite formed into obscure, broadly rounded lobe (
<figureCitation id="6EAF2A3388068007FE62BB910002F5E4" box="[398,528,1252,1277]" captionStart="Figs 50–58" captionStartId="23.[182,221,1532,1552]" captionTargetBox="[185,1152,412,1511]" captionTargetId="figure-167@23.[181,1146,408,1519]" captionTargetPageId="23" captionText="Figs 50–58. Harpagophora monodus Attems, 1909. 50. Lateral view of collum; arrow indicates anterior end of millipede. 51. Oral view of first pair of male legs. 52. Aboral view of gonopod (NMSA 19102). 53. Aboral view of gonopods (NMSA 16159). 54. Oral view of telopodite. 55. Detail of femoral spines of telopodite. 56–58.Apical elements of telopodite, showing different orientations. bp = black process; f1 = large femoral spine; f2 = small femoral spine; ll = lateral lobe; mrp = median rounded process; p = pectinophore; s = spine branch." figureDoi="http://doi.org/10.5281/zenodo.7666330" httpUri="https://zenodo.org/record/7666330/files/figure.png" pageId="24" pageNumber="227">Figs 52, 53</figureCitation>
).
</paragraph>
<paragraph id="F62B36B688068007FF7CBA7102FAF424" blockId="24.[144,1104,206,1675]" pageId="24" pageNumber="227">
<emphasis id="C4E0EAA488068007FF7CBA710333F405" box="[144,289,1284,1308]" italics="true" pageId="24" pageNumber="227">Pre-anal ring</emphasis>
: Caudal spine short, just reaching margin of anal valve and sharply upturned distally.
</paragraph>
<paragraph id="F62B36B688068007FF7CBA3F0130F49A" blockId="24.[144,1104,206,1675]" pageId="24" pageNumber="227">
Distribution: The species is endemic to the
<collectingRegion id="3450F85488068007FD64BA3F0122F47A" box="[648,816,1354,1379]" country="South Africa" name="Northern Cape" pageId="24" pageNumber="227">Northern Cape</collectingRegion>
(
<figureCitation id="6EAF2A3388068007FCAEBA3F0194F47A" box="[834,902,1354,1379]" captionStart="Fig" captionStartId="1.[182,215,845,865]" captionTargetBox="[251,1061,203,829]" captionTargetId="figure-7@1.[232,1095,193,840]" captionTargetPageId="1" captionText="Fig. 1. Distribution of Harpagophora, Metaphora and Poratophilus species in southern Africa. G = Gauteng; KZN = KwaZulu-Natal; L = Lesotho; MP = Mpumalanga; S = Swaziland; NW Prov = North West Province; W Cape = Western Cape." figureDoi="http://doi.org/10.5281/zenodo.7666310" httpUri="https://zenodo.org/record/7666310/files/figure.png" pageId="24" pageNumber="227">Fig. 1</figureCitation>
). It occurs in the western part of the province in the succulent Karoo biome.
</paragraph>
<paragraph id="F62B36B688068007FF7CBAE701E5F732" blockId="24.[144,1104,206,1675]" pageId="24" pageNumber="227">
Remarks: In terms of gonopod structure, the species is most similar to
<taxonomicName id="31944D3588068007FC79BAE60631F4B2" baseAuthorityName="Brandt" baseAuthorityYear="1841" box="[917,1059,1426,1451]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="24" pageNumber="227" phylum="Arthropoda" rank="species" species="attenuata">
<emphasis id="C4E0EAA488068007FC79BAE601BBF4B2" box="[917,937,1427,1451]" italics="true" pageId="24" pageNumber="227">H</emphasis>
.
<emphasis id="C4E0EAA488068007FC5ABAE60631F4B2" box="[950,1059,1427,1451]" italics="true" pageId="24" pageNumber="227">attenuata</emphasis>
</taxonomicName>
<emphasis id="C4E0EAA488068007FBCEBAE60634F4B2" box="[1058,1062,1427,1451]" italics="true" pageId="24" pageNumber="227">,</emphasis>
but
<taxonomicName id="31944D3588068007FF7CBAC6030AF4D2" authorityName="Attems" authorityYear="1909" box="[144,280,1459,1483]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="24" pageNumber="227" phylum="Arthropoda" rank="species" species="monodus">
<emphasis id="C4E0EAA488068007FF7CBAC6030AF4D2" box="[144,280,1459,1483]" italics="true" pageId="24" pageNumber="227">H. monodus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="31944D3588068007FE5DBAC6002DF4D2" baseAuthorityName="Brandt" baseAuthorityYear="1841" box="[433,575,1458,1483]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="24" pageNumber="227" phylum="Arthropoda" rank="species" species="attenuata">
<emphasis id="C4E0EAA488068007FE5DBAC603D7F4D2" box="[433,453,1459,1483]" italics="true" pageId="24" pageNumber="227">H</emphasis>
.
<emphasis id="C4E0EAA488068007FE39BAC6002DF4D2" box="[469,575,1459,1483]" italics="true" pageId="24" pageNumber="227">attenuata</emphasis>
</taxonomicName>
in the lack of telocoxal spines along the inner margin of the telocoxite. In
<taxonomicName id="31944D3588068007FE29BAA6005EF4F2" authorityName="Attems" authorityYear="1909" box="[453,588,1490,1515]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="24" pageNumber="227" phylum="Arthropoda" rank="species" species="monodus">
<emphasis id="C4E0EAA488068007FE29BAA603CBF4F2" box="[453,473,1491,1515]" italics="true" pageId="24" pageNumber="227">H</emphasis>
.
<emphasis id="C4E0EAA488068007FE0BBAA6005EF4F2" box="[487,588,1491,1515]" italics="true" pageId="24" pageNumber="227">monodus</emphasis>
</taxonomicName>
the protuberances on the medial distal margin of the telocoxite are not as long as in
<taxonomicName id="31944D3588068007FDBCBA8600CDF712" baseAuthorityName="Brandt" baseAuthorityYear="1841" box="[592,735,1522,1547]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="24" pageNumber="227" phylum="Arthropoda" rank="species" species="attenuata">
<emphasis id="C4E0EAA488068007FDBCBA860076F712" box="[592,612,1523,1547]" italics="true" pageId="24" pageNumber="227">H</emphasis>
.
<emphasis id="C4E0EAA488068007FD99BA8600CDF712" box="[629,735,1523,1547]" italics="true" pageId="24" pageNumber="227">attenuata</emphasis>
</taxonomicName>
, and are more bluntly rounded. Specimens of
<taxonomicName id="31944D3588068007FEC2B96603A4F732" authorityName="Attems" authorityYear="1909" box="[302,438,1555,1579]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="24" pageNumber="227" phylum="Arthropoda" rank="species" species="monodus">
<emphasis id="C4E0EAA488068007FEC2B96603A4F732" box="[302,438,1555,1579]" italics="true" pageId="24" pageNumber="227">H. monodus</emphasis>
</taxonomicName>
are much shorter and less robust than
<taxonomicName id="31944D3588068007FC88B96601E3F732" baseAuthorityName="Brandt" baseAuthorityYear="1841" box="[868,1009,1554,1579]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="24" pageNumber="227" phylum="Arthropoda" rank="species" species="attenuata">
<emphasis id="C4E0EAA488068007FC88B966016AF732" box="[868,888,1555,1579]" italics="true" pageId="24" pageNumber="227">H</emphasis>
.
<emphasis id="C4E0EAA488068007FC6BB96601E3F732" box="[903,1009,1555,1579]" italics="true" pageId="24" pageNumber="227">attenuata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="F62B36B688068007FF46B94701B6F752" blockId="24.[144,1104,206,1675]" box="[170,932,1586,1611]" pageId="24" pageNumber="227">In some specimens the suture on the syncoxosternum is incomplete.</paragraph>
<paragraph id="F62B36B688068007FF46B927004FF792" blockId="24.[144,1104,206,1675]" pageId="24" pageNumber="227">
The origin of the name
<taxonomicName id="31944D3588068007FE43B9260006F772" authorityName="Attems" authorityYear="1909" box="[431,532,1619,1643]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="24" pageNumber="227" phylum="Arthropoda" rank="species" species="monodus">
<emphasis id="C4E0EAA488068007FE43B9260006F772" box="[431,532,1619,1643]" italics="true" pageId="24" pageNumber="227">monodus</emphasis>
</taxonomicName>
is unclear and it has therefore not been modified to agree with the gender of the genus name.
</paragraph>
</subSubSection>
</treatment>
</document>
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