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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.271.4062" ID-GBIF-Dataset="c93c3a93-d2e1-4c85-8ece-bf3aa1a5109d" ID-PMC="PMC3652427" ID-Pensoft-Pub="1313-2970-271-1" ID-PubMed="23717185" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-271-1" ModsDocOrigin="ZooKeys 271" ModsDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" checkinTime="1451247655320" checkinUser="pensoft" docAuthor="Caterino, Michael S. &amp; Tishechkin, Alexey K." docDate="2013" docId="7E3A84157DF84BE775EC6B1058DAED8A" docLanguage="en" docName="ZooKeys 271: 1-401" docOrigin="ZooKeys 271" docSource="http://dx.doi.org/10.3897/zookeys.271.4062" docTitle="Operclipygus pustulifer Caterino &amp; Tishechkin, 2013, sp. n." docType="treatment" docVersion="4" lastPageNumber="40" masterDocId="4403FF80FFE7FF95FFDA5D0E29259B2F" masterDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" masterLastPageNumber="401" masterPageNumber="1" pageNumber="40" updateTime="1668155382655" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Caterino, Michael S.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tishechkin, Alexey K.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="volume">
<mods:number>271</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>401</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.271.4062</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.271.4062</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-271-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152040924" LSID="urn:lsid:zoobank.org:act:1423C21B-1A81-487B-97D4-EFA7A03E9863" httpUri="http://treatment.plazi.org/id/7E3A84157DF84BE775EC6B1058DAED8A" lastPageNumber="40" pageId="39" pageNumber="40">
<subSubSection pageId="39" pageNumber="40" type="nomenclature">
<paragraph pageId="39" pageNumber="40">
<taxonomicName LSID="urn:lsid:zoobank.org:act:1423C21B-1A81-487B-97D4-EFA7A03E9863" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus pustulifer" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="pustulifer">Operclipygus pustulifer</taxonomicName>
<taxonomicNameLabel pageId="39" pageNumber="40">sp. n.</taxonomicNameLabel>
Fig. 9BMap 2
</paragraph>
</subSubSection>
<subSubSection pageId="39" pageNumber="40" type="type locality">
<paragraph pageId="39" pageNumber="40">Type locality.</paragraph>
<paragraph pageId="39" pageNumber="40">
COSTA RICA: Guanacaste: 9 km S Santa Cecilia, Pitilla Research Station [
<geoCoordinate direction="north" orientation="latitude" precision="92" value="10.988334">10°59.3'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-85.425">85°25.5'W</geoCoordinate>
].
</paragraph>
<paragraph pageId="39" pageNumber="40">T</paragraph>
</subSubSection>
<subSubSection pageId="39" pageNumber="40" type="ype material">
<paragraph pageId="39" pageNumber="40">ype material.</paragraph>
<paragraph pageId="39" pageNumber="40">Holotype male: &quot;Est. Pitilla, 9km S. Sta. Cecilia, P. N. Guanacaste, Prov. Guana, COSTA RICA. 700 m Ene [January] 1994, C. Moraga, L N 330200_380200 #2563&quot; (INBIO). Paratypes (18): 17: same data as holotype (INBIO, FMNH, AKTC, MSCC); 1: same data as holotype, but iii.1994 (INBIO).</paragraph>
</subSubSection>
<subSubSection pageId="39" pageNumber="40" type="description">
<paragraph pageId="39" pageNumber="40">Diagnostic description.</paragraph>
<paragraph pageId="39" pageNumber="40">
This species is very similar in most characters to
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus mirabilis" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mirabilis">Operclipygus mirabilis</taxonomicName>
and is only described to the extent that it differs from it. Length: 1.72-1.75 mm, width: 1.40-1.44 mm; male with lateral submarginal pronotal stria diverging from marginal in anterior two-thirds, ending behind anterior corner, median pair of pronotal gland openings borne on very weak tubercles; pronotal disk not depressed anteriorly around tubercles; lateral submarginal pronotal stria of female situated slightly further from margin, particularly anteriorly, than that of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus mirabilis" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mirabilis">Operclipygus mirabilis</taxonomicName>
. Male genitalia with apical guides of S8 slightly narrower apically than in
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus mirabilis" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mirabilis">Operclipygus mirabilis</taxonomicName>
, S9 also slightly narrower and more elongate; aedeagus essentially indistinguishable.
</paragraph>
</subSubSection>
<subSubSection pageId="39" pageNumber="40" type="remarks">
<paragraph pageId="39" pageNumber="40">Remarks.</paragraph>
<paragraph pageId="39" pageNumber="40">
This species and
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus mirabilis" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mirabilis">Operclipygus mirabilis</taxonomicName>
are very similar, differing mainly in the exaggeration of the tubercle of the male pronotum, which in
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus pustulifer" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="pustulifer">Operclipygus pustulifer</taxonomicName>
lacks any surrounding depression (Fig. 9B).
</paragraph>
</subSubSection>
<subSubSection pageId="39" pageNumber="40" type="etymology">
<paragraph pageId="39" pageNumber="40">Etymology.</paragraph>
<paragraph pageId="39" pageNumber="40">
This
<normalizedToken originalValue="species">species'</normalizedToken>
name refers to the raised gland opening of the male pronotum.
</paragraph>
</subSubSection>
</treatment>
</document>