113 lines
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113 lines
12 KiB
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<mods:title id="F41185A09C42EF2CBCCE8BF1D2DD0158">Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae)</mods:title>
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<mods:namePart id="E1EACC1A7AA3612F49FAE1211572CD7E">Kohlmann, Bert</mods:namePart>
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<mods:namePart id="C361C7BF4BF3033A6D3DF39E7A0EA4A8">Arriaga-Jimenez, Alfonsina</mods:namePart>
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<mods:namePart id="71F90094E7D23F0459CF4189641FF340">Roes, Matthias</mods:namePart>
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<mods:number id="D3E43245AEDD73F8A2EEC663CA984690">743</mods:number>
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<treatment id="7E2DEA7201CB96E41326FCCC778FC035" ID-GBIF-Taxon="142283987" LSID="urn:lsid:plazi:treatment:7E2DEA7201CB96E41326FCCC778FC035" httpUri="http://treatment.plazi.org/id/7E2DEA7201CB96E41326FCCC778FC035" lastPageId="10" lastPageNumber="77" pageId="9" pageNumber="76">
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<subSubSection id="04C75D22445A14118A180C07FFEAF603" pageId="9" pageNumber="76" type="nomenclature">
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<paragraph id="15A49BD8E4554776CBEDA40C2689F04B" pageId="9" pageNumber="76">
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<taxonomicName id="505112D69A3D5B8270EDF93CA35099EA" ID-CoL="4FSJM" authority="Edmonds, 1979" authorityName="Edmonds" authorityYear="1979" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus (Notiophanaeus) halffterorum" order="Coleoptera" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="halffterorum" subGenus="Notiophanaeus">Phanaeus (Notiophanaeus) halffterorum Edmonds, 1979</taxonomicName>
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</paragraph>
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<subSubSection id="8C8EA1135CCC68F5059677702FF927AC" lastPageId="10" lastPageNumber="77" pageId="9" pageNumber="76" type="reference_group">
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<paragraph id="2D07F5454457D9436A930B81696626BA" lastPageId="10" lastPageNumber="77" pageId="9" pageNumber="76">
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<taxonomicName id="D6F4A7F0CC54698CFA895B5E5235231E" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus halffterorum" order="Coleoptera" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="halffterorum">Phanaeus halffterorum</taxonomicName>
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:
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<bibRefCitation id="DB3C765E0C438F95BF9268D578999066" author="Edmonds, WD" journalOrPublisher="Pan-Pacific Entomologist" pageId="20" pageNumber="87" pagination="99 - 105" title="A new species of Phanaeus from Mexico (Coleoptera: Scarabaeidae)." volume="55" year="1979">Edmonds (1979</bibRefCitation>
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: 99; partim),
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<bibRefCitation id="F3297D266C823C7A36442701AA0B57DC" author="Halffter, G" journalOrPublisher="Instituto de Ecologia, Mexico City" pageId="20" pageNumber="87" title="The nesting behavior of dung beetles (Scarabaeinae). An ecological and evolutive approach." year="1982">Halffter and Edmonds (1982</bibRefCitation>
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: 88-89),
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<bibRefCitation id="1CC1C8821A3CB3C02F1C5904EDE7065C" pageId="9" pageNumber="76">Anduaga and Halffter (1991</bibRefCitation>
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: 157),
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<bibRefCitation id="E7C0D7769EBA42ED4A1B0972C2199416" author="Delgado-Castillo, L" journalOrPublisher="The Coleopterists Bulletin" pageId="19" pageNumber="86" pagination="121 - 126" title="A new Mexican species of Onthophagus with mycophagous habits (Coleoptera: Scarabaeidae: Scarabaeinae)." volume="47" year="1993">Delgado-Castillo et al. (1993</bibRefCitation>
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: 125),
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<bibRefCitation id="5B043C7B8D70534E546A3B1F62175814" author="Edmonds, WD" journalOrPublisher="Natural History Museum of Los Angeles County, Contributions in Science" pageId="20" pageNumber="87" pagination="1 - 105" title="Revision of Phanaeus Macleay, a New World Genus of Scarabaeinae Dung Beetles (Coleoptera: Scarabaeidae, Scarabaeinae)." volume="442" year="1994">Edmonds (1994</bibRefCitation>
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: 39-43, 101),
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<bibRefCitation id="08556BEEB4FEC20F89B51C6E2C97F15A" author="Anduaga, S" journalOrPublisher="Acta Zoologica Mexicana NS" pageId="19" pageNumber="86" pagination="119 - 130" title="Escarabajos coprofagos (Coleoptera: Scarabaeoidea) asociados a hongos en la Sierra Madre Occidental, Durango, Mexico: con una compilacion de las especies micetofagas." volume="80" year="2000">Anduaga (2000</bibRefCitation>
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: 125, 130),
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<bibRefCitation id="DB8B4E8BDDE0B89F1960FB2C2769F4C8" author="Arnaud, P" journalOrPublisher="Hillside Books, Canterbury, Kent" pageId="19" pageNumber="86" title="Phanaeini. Les Coleopteres du Monde, vol. 28." year="2002">Arnaud (2002</bibRefCitation>
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: 95-
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<pageBreakToken id="461CA0E5C7B89E55B0AB21F0727818BF" pageId="10" pageNumber="77" start="start">96</pageBreakToken>
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),
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<bibRefCitation id="A80D68EA5641FC550AB307FABAF1D78E" pageId="10" pageNumber="77">Edmonds (2003</bibRefCitation>
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: 61, 65),
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<bibRefCitation id="17490A621EE56CC0C7587C9BFF86C1A1" author="Edmonds, WD" journalOrPublisher="Zootaxa" pageId="20" pageNumber="87" pagination="31 - 37" title="A new species of Phanaeus MacLeay (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini) from Oaxaca, Mexico." volume="1171" year="2006">Edmonds (2006</bibRefCitation>
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: 31, 34, 36),
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<bibRefCitation id="E656C37CC7F8EE220A0CB71B36E0F870" author="Ceballos, G" journalOrPublisher="Gobierno del Estado de Mexico, Toluca de Lerdo" pageId="19" pageNumber="86" title="La diversidad biologica del Estado de Mexico. Estudio de estado." url="http://www.biodiversidad.gob.mx/" year="2009">Ceballos et al. (2009</bibRefCitation>
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: 397),
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<bibRefCitation id="8B779A54EA94342BA17E742BA9288B37" author="Edmonds, WD" journalOrPublisher="Insecta Mundi" pageId="20" pageNumber="87" pagination="1 - 108" title="Taxonomy of Phanaeus revisited: Revised keys to and comments on species of the New World dung beetle genus Phanaeus MacLeay, 1819 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini)." volume="0274" year="2012">
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Edmonds and
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<normalizedToken id="22B04FBDF2D3A85F3C6DDE68B965C335" originalValue="Zídek">Zidek</normalizedToken>
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(2012
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</bibRefCitation>
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: 3, 5, 12, 52, 54),
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<bibRefCitation id="C9280484681FCEA0B92D89D10DC3AE74" pageId="10" pageNumber="77">Deloya et al. (2014</bibRefCitation>
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: 77),
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<bibRefCitation id="3907D1BB646B3FA055E49AE4BE230722" author="Moctezuma, V" journalOrPublisher="Coleopterists Bulletin" pageId="21" pageNumber="88" pagination="47 - 56" title="A new species of Phanaeus MacLeay (Coleoptera: Scarabaeidae: Scarabaeinae) from Los Chimalapas, Oaxaca, Mexico." url="https://doi.org/10.1649/0010-065X-71.1.47" volume="71" year="2017">Moctezuma and Halffter (2017</bibRefCitation>
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: 52, 54-55),
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<bibRefCitation id="9A1C225055875A1F1E0FD3018EC882B9" author="Moctezuma, V" journalOrPublisher="ZooKeys" pageId="21" pageNumber="88" pagination="113 - 135" title="Two new species of the Phanaeusendymion group (Coleopter; Scarabaeidae, Scarabaeinae)." url="https://doi.org/10.3897/zookeys.702.14728" volume="702" year="2017">Moctezuma et al. (2017</bibRefCitation>
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: 115),
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<bibRefCitation id="838572E1DA2681AB2EFDD4091F5C74FD" author="Lizardo, V" journalOrPublisher="Evolution" pageId="20" pageNumber="87" url="https://doi.org/10.11646/zootaxa.4358.2.3" year="in press">Lizardo et al. (in press)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="5C8705F147B9A47600053E26B2FBF480" pageId="10" pageNumber="77" type="remarks">
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<paragraph id="BB05348B211C08A594887B4AE199AEF5" pageId="10" pageNumber="77">Remarks.</paragraph>
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<paragraph id="89596EEAE6F845DB7DE0B740B38DAE42" pageId="10" pageNumber="77">This study identifies new localities of this species present in the Blackaller private collection, which seems to have a rather restricted distribution in the State of Mexico.</paragraph>
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<paragraph id="948CF627F2BD04116E2F2D35F844C756" pageId="10" pageNumber="77">
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Estado de
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<normalizedToken id="266F687EDD6AF85095FAA45BC91DDF70" originalValue="México">Mexico</normalizedToken>
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(8 specimens): Sierra de Nanchititla, 6 km al Este de Nanchititla, Bosque de Pino-Encino, CD en Hongos, 1840 m, 15-VIII-2015,
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<geoCoordinate id="A14EC7CEC3174704C76679C10514E57B" direction="north" orientation="latitude" precision="925" value="18.866667">18°52'N</geoCoordinate>
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,
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<geoCoordinate id="1F283B9E4AB30769445864B97C84C322" direction="west" orientation="longitude" precision="925" value="-100.4">100°24'W</geoCoordinate>
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, J.Blackaller y L.
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<normalizedToken id="71DF5200AA8578FFE947ABD1DE4A8AB0" originalValue="Zacarías">Zacarias</normalizedToken>
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cols.; 06-VIII-2011, Trampa de Luz, Blackaller y Robacker cols.; 21-VIII-2011, J. Blackaller col.
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</paragraph>
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<paragraph id="7B6BF502B218DAE37F2EB0AF692ED71D" pageId="10" pageNumber="77">Comparing this species with its closest relative, Ph. bravoensis, one can add as a further difference between the two species the fact that Ph. halffterorum has its pygidium covered with coarse punctures while Ph. bravoensis has faintly impressed small punctures. Although it is not specified in the original description of this species (Edmonds 1979), specimens were collected directly by hand feeding on fungi pertaining to the genus Boletus.</paragraph>
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</subSubSection>
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</treatment>
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