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<mods:title id="AA1FF9EDB0FE076300E82CB0C4822714">Molecular and morphological data suggest a new species of big-eared bat (Vespertilionidae: Corynorhinus) endemic to northeastern Mexico</mods:title>
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<mods:namePart id="A4FA452827C5F5CF85EC83A206641703">López-Cuamatzi, Issachar L.</mods:namePart>
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<mods:namePart id="69FC1E7B84ACB8B4D11D8EEF2005C487">Ortega, Jorge</mods:namePart>
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<mods:namePart id="3A1DD0C0BEFEDE2929C186BFE6C1BAFB">Ospina-Garcés, Sandra M.</mods:namePart>
<mods:affiliation id="71D9BE4B011FC3E7908FB531FD8F42BA">Centro de Investigaciones Tropicales, Universidad Veracruzana, Xalapa de Enríquez, Veracruz, Mexico, &amp; Centro de Investigación en Biodiversidad y Conservación, Universidad Autónoma del Estado de Morelos, Cuernavaca, Morelos, Mexic</mods:affiliation>
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<mods:date id="BBC3AB0056FA8448F4171F9D7664025E">2024</mods:date>
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<emphasis id="C4F96355F8108D23DA5FFA15FC10739D" bold="true" box="[533,915,1442,1471]" italics="true" pageId="22" pageNumber="23">Corynorhinus leonpaniaguae</emphasis>
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urn:lsid:zoobank.org:act: 
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Vernacular name: León Paniagua’s Big-eared Bat, Murciélago-mula de León-Paniagua 
<emphasis id="C4F96355F8108D23DA5FF9AEFD9D7013" bold="true" box="[533,542,1561,1585]" pageId="22" pageNumber="23">(</emphasis>
Spanish
<emphasis id="C4F96355F8108D23DA38F9AEFDF87013" bold="true" box="[626,635,1561,1585]" pageId="22" pageNumber="23">)</emphasis>
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<emphasis id="C4F96355F8108D23DA7FF98BFD2D7076" bold="true" box="[565,686,1596,1620]" pageId="22" pageNumber="23">Synonyms.</emphasis>
<taxonomicName id="318DC4C4F8108D23DA8EF98CFB2B7076" authority="Miller, 1897" authorityName="Miller" authorityYear="1897" box="[708,1192,1595,1620]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="22" pageNumber="23" phylum="Chordata" rank="subSpecies" species="macrotis" subSpecies="pallescens">
<emphasis id="C4F96355F8108D23DA8EF98CFBA37071" box="[708,1056,1595,1619]" italics="true" pageId="22" pageNumber="23">Corynorhinus macrotis pallescens</emphasis>
Miller, 1897
</taxonomicName>
, (Part)
</paragraph>
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<taxonomicName id="318DC4C4F8108D23DA7FF9E9FBA27054" authority="Allen, 1916" authorityName="Allen" authorityYear="1916" box="[565,1057,1630,1654]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="22" pageNumber="23" phylum="Chordata" rank="subSpecies" species="megalotis" subSpecies="mexicanus">
<emphasis id="C4F96355F8108D23DA7FF9E9FC207054" box="[565,931,1630,1654]" italics="true" pageId="22" pageNumber="23">Corynorhinus megalotis mexicanus</emphasis>
Allen, 1916
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<taxonomicName id="318DC4C4F8108D23DA7FF937FBBB70BB" authority="Miller, 1924" authorityName="Miller" authorityYear="1924" box="[565,1080,1664,1689]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="22" pageNumber="23" phylum="Chordata" rank="subSpecies" species="rafinesquii" subSpecies="mexicanus">
<emphasis id="C4F96355F8108D23DA7FF937FC3370BA" box="[565,944,1664,1689]" italics="true" pageId="22" pageNumber="23">Corynorhinus rafinesquii mexicanus</emphasis>
Miller, 1924
</taxonomicName>
, (Part)
</paragraph>
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<taxonomicName id="318DC4C4F8108D23DA7FF914FB997099" authority="Dalquest, 1953" authorityName="Dalquest" authorityYear="1953" box="[565,1050,1699,1723]" class="Mammalia" family="Vespertilionidae" genus="Plecotus" kingdom="Animalia" order="Chiroptera" pageId="22" pageNumber="23" phylum="Chordata" rank="subSpecies" species="rafinesquii" subSpecies="mexicanus">
<emphasis id="C4F96355F8108D23DA7FF914FCF67099" box="[565,885,1699,1723]" italics="true" pageId="22" pageNumber="23">Plecotus rafinesquii mexicanus</emphasis>
Dalquest, 1953
</taxonomicName>
, (Part)
</paragraph>
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<taxonomicName id="318DC4C4F8108D23DA7FF971FC1C70FC" authority="Handley, 1959" authorityName="Handley" authorityYear="1959" box="[565,927,1734,1758]" class="Mammalia" family="Vespertilionidae" genus="Plecotus" kingdom="Animalia" order="Chiroptera" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="mexicanus">
<emphasis id="C4F96355F8108D23DA7FF971FC8370FC" box="[565,768,1734,1758]" italics="true" pageId="22" pageNumber="23">Plecotus mexicanus</emphasis>
Handley, 1959
</taxonomicName>
</paragraph>
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<taxonomicName id="318DC4C4F8108D23DA7FF95FFC7D7123" authority="G. M. Allen, 1916" authorityName="G. M. Allen" authorityYear="1916" box="[565,1022,1768,1793]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="mexicanus">
<emphasis id="C4F96355F8108D23DA7FF95FFCB87122" box="[565,827,1768,1792]" italics="true" pageId="22" pageNumber="23">Corynorhinus mexicanus</emphasis>
G. M. Allen, 1916
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, (Part)
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<materialsCitation id="46E5B51AF8108D23DA7FF8BCFA4A714B" ID-GBIF-Occurrence="4988226301" collectingDate="2022-04-18" collectionCode="MZFC-M" collectorName="Juan Cruzado &amp; Silvino Hernandez &amp; Issachar L. Lopez-Cuamatzi" country="Mexico" latitude="25.16096" location="Cave El Hundido" longLatPrecision="1" longitude="-100.622986" municipality="Nuevo León" pageId="22" pageNumber="23" specimenCode="MZFC-M16326" specimenCount="1" specimenCount-adult="1" specimenCount-female="1" typeStatus="holotype">
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<typeStatus id="293601E5F8108D23DA7FF8BCFD1D7101" box="[565,670,1803,1827]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
.
</emphasis>
Colección de Mamíferos, 
<collectingMunicipality id="1656253DF8108D23DB83F8BCFAE17101" box="[969,1378,1803,1827]" pageId="22" pageNumber="23">Museo de Zoología Alfonso L. Herrera</collectingMunicipality>
(
<collectionCode id="909C2782F8108D23DD3AF8BCFA5D7101" box="[1392,1502,1803,1827]" pageId="22" pageNumber="23">MZFC-M</collectionCode>
), No. 
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is an 
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<specimenCount id="E08B74CEF8108D23DB29F899FC2A7164" box="[867,937,1838,1862]" count="1" pageId="22" pageNumber="23" type="female">female</specimenCount>
with skin and skull dry preserved and collected on 
<date id="82339987F8108D23DD82F899FD03714B" pageId="22" pageNumber="23" value="2022-04-18">
<collectingDate id="9277606FF8108D23DD82F899FD03714B" pageId="22" pageNumber="23" value="2022-04-18">
April 18 
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, 2022
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</date>
, by 
<collectorName id="5B78DA91F8108D23DAE3F8E6FCBD714B" box="[681,830,1873,1897]" pageId="22" pageNumber="23">Juan Cruzado</collectorName>
, 
<collectorName id="5B78DA91F8108D23DB02F8E6FB8D714A" box="[840,1038,1872,1897]" pageId="22" pageNumber="23">Silvino Hernández</collectorName>
and 
<collectorName id="5B78DA91F8108D23DC08F8E6FAEA714A" box="[1090,1385,1872,1897]" pageId="22" pageNumber="23">Issachar L. López-Cuamatzi</collectorName>
(
<figureCitation id="6EB6A3C2F8108D23DD32F8E6FA3A714B" box="[1400,1465,1873,1897]" captionStart="Fig 12" captionStartId="23.[533,561,1443,1462]" captionTargetBox="[533,1493,208,1421]" captionTargetId="figure-132@23.[533,1493,208,1421]" captionTargetPageId="23" captionText="Fig 12. Holotype pictures. Coloration of the dorsal and ventral fur of the holotype (MZFC-M16326) of Corynorhinus leonpaniaguae sp. nov. (A, B). Cranium, mandible (C, D, E) and lateral view of the first upper double-cuspid incisor (F) of the holotype. https://doi.org/10.1371/journal.pone.0296275.g012" figureDoi="http://doi.org/10.5281/zenodo.14290654" httpUri="https://zenodo.org/record/14290654/files/figure.png" pageId="22" pageNumber="23">
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12
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).
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<emphasis id="C4F96355F8118D22DA5FFA14FD7F7394" bold="true" box="[533,764,1443,1462]" pageId="23" pageNumber="24">Fig 12. Holotype pictures.</emphasis>
Coloration of the dorsal and ventral fur of the holotype (MZFC-M16326) of 
<taxonomicName id="318DC4C4F8118D22DDCBFA14FD0C73ED" authorityName="López-Cuamatzi &amp; Ortega &amp; Ospina-Garcés &amp; Zúñiga &amp; G." authorityYear="2024" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="leonpaniaguae" status="sp. nov.">
<emphasis id="C4F96355F8118D22DDCBFA14FD0C73ED" italics="true" pageId="23" pageNumber="24">Corynorhinus leonpaniaguae</emphasis>
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(A, B). Cranium, mandible (C, D, E) and lateral view of the first upper double-cuspid incisor (F) of the holotype.
</paragraph>
<paragraph id="F632BF47F8118D22DA5FFA48FC2E7030" blockId="23.[533,1524,1443,1554]" box="[533,941,1535,1554]" pageId="23" pageNumber="24">https://doi.org/10.1371/journal.pone.0296275.g012</paragraph>
</caption>
<paragraph id="F632BF47F8118D22DA7FF983FB85704D" blockId="23.[533,1536,1588,1890]" pageId="23" pageNumber="24">
<emphasis id="C4F96355F8118D22DA7FF983FD48706E" bold="true" box="[565,715,1588,1612]" pageId="23" pageNumber="24">Type locality.</emphasis>
<location id="F352E99CF8118D22DAAAF983FC1B706E" LSID="urn:lsid:plazi:treatment:7E240E51F8108D2FDA5FFA15FBCD70E7:F352E99CF8118D22DAAAF983FC1B706E" box="[736,920,1588,1612]" country="Mexico" latitude="25.16096" longLatPrecision="1" longitude="-100.622986" name="Cave El Hundido" pageId="23" pageNumber="24" stateProvince="Nuevo Leon">Cave El Hundido</location>
(
<geoCoordinate id="93B9D980F8118D22DBEFF983FB8B706E" box="[933,1032,1588,1612]" degrees="25.16096" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="25.16096">25.16096</geoCoordinate>
, 
<geoCoordinate id="93B9D980F8118D22DC58F983FB15706E" box="[1042,1174,1588,1612]" degrees="100.622985" direction="west" orientation="longitude" pageId="23" pageNumber="24" precision="1" value="-100.622986">-100.622985</geoCoordinate>
; 2072 msnm), 2.5 
<collectorName id="5B78DA91F8118D22DD1CF983FA24706E" box="[1366,1447,1588,1612]" pageId="23" pageNumber="24">Km NE</collectorName>
from 
<collectorName id="5B78DA91F8118D22DA5FF9E0FD35704D" box="[533,694,1623,1647]" pageId="23" pageNumber="24">Puerto Grande</collectorName>
, 
<collectorName id="5B78DA91F8118D22DAF5F9E0FC94704D" box="[703,791,1623,1647]" pageId="23" pageNumber="24">Galeana</collectorName>
, 
<collectingRegion id="344971A5F8118D22DB6BF9E0FC25704D" box="[801,934,1623,1647]" country="Mexico" name="Nuevo Leon" pageId="23" pageNumber="24">Nuevo León</collectingRegion>
, 
<collectingCountry id="8E9AFFD7F8118D22DBFAF9E0FB81704D" box="[944,1026,1623,1647]" name="Mexico" pageId="23" pageNumber="24">
<collectingRegion id="344971A5F8118D22DBFAF9E0FB81704D" box="[944,1026,1623,1647]" country="Mexico" name="Mexico" pageId="23" pageNumber="24">Mexico</collectingRegion>
</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="BE97ECCCF8118D22DA7FF9CDFA31713E" pageId="23" pageNumber="24" type="etymology">
<paragraph id="F632BF47F8118D22DA7FF9CDFA31713E" blockId="23.[533,1536,1588,1890]" pageId="23" pageNumber="24">
<emphasis id="C4F96355F8118D22DA7FF9CDFD3770B0" bold="true" box="[565,692,1658,1682]" pageId="23" pageNumber="24">Etymology.</emphasis>
<taxonomicName id="318DC4C4F8118D22DA83F9CEFC7470B3" box="[713,1015,1657,1681]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="leonpaniaguae">
<emphasis id="C4F96355F8118D22DA83F9CEFC7470B3" box="[713,1015,1657,1681]" italics="true" pageId="23" pageNumber="24">Corynorhinus leonpaniaguae</emphasis>
</taxonomicName>
is named in honor of Dr. Livia S. León Paniagua, in recognition of her outstanding contribution to the knowledge of the systematics and natural history of Mexican mammals. In addition to being a pioneering woman in Mexican mammalogy, Livia has been a great mentor, dedicated to the training of new scientists. This species name is a noun in the genitive case formed by adding -ae to the stem of the name [
<bibRefCitation id="921CC2B6F8118D22DD37F8B3FA20713E" author="ICZN" box="[1405,1443,1796,1820]" pageId="23" pageNumber="24" refId="ref20249" refString="110. ICZN. International Code of Zoological Nomenclature. 4 th ed. London: International Commission on Zoological Nomenclature; 1999." type="book" year="1999">110</bibRefCitation>
].
</paragraph>
</subSubSection>
<subSubSection id="BE97ECCCF8118D2DDA7FF890FD27732F" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="biology_ecology">
<paragraph id="F632BF47F8118D2DDA7FF890FD27732F" blockId="23.[533,1536,1588,1890]" lastBlockId="24.[533,1531,957,1883]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
<emphasis id="C4F96355F8118D22DA7FF890FCCE711D" bold="true" box="[565,845,1831,1855]" pageId="23" pageNumber="24">Habitat and distribution.</emphasis>
<taxonomicName id="318DC4C4F8118D22DB28F890FB12711D" box="[866,1169,1831,1855]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="leonpaniaguae">
<emphasis id="C4F96355F8118D22DB28F890FB12711D" box="[866,1169,1831,1855]" italics="true" pageId="23" pageNumber="24">Corynorhinus leonpaniaguae</emphasis>
</taxonomicName>
lives in open pine forests located on slopes and canyons of the northern Sierra Madre Oriental and mountains of the state of 
<collectingRegion id="344971A5F81E8D2DDA5FFC0AFDFB75F7" box="[533,632,957,981]" country="Mexico" name="Coahuila de Zaragoza" pageId="24" pageNumber="25">Coahuila</collectingRegion>
. Records of the presence of this species occur mainly between 300 and 2000 meters above sea level (masl) in the states of 
<collectingRegion id="344971A5F81E8D2DDBD7FC57FC7C75DA" box="[925,1023,992,1016]" country="Mexico" name="Coahuila de Zaragoza" pageId="24" pageNumber="25">Coahuila</collectingRegion>
, 
<collectingRegion id="344971A5F81E8D2DDC43FC57FB0E75DA" box="[1033,1165,992,1016]" country="Mexico" name="Nuevo Leon" pageId="24" pageNumber="25">Nuevo León</collectingRegion>
, and 
<collectingRegion id="344971A5F81E8D2DDC8FFC57FAC775DA" box="[1221,1348,992,1016]" country="Mexico" name="Tamaulipas" pageId="24" pageNumber="25">Tamaulipas</collectingRegion>
. It is an endemic species of 
<collectingCountry id="8E9AFFD7F81E8D2DDACAFBB4FD4C7239" box="[640,719,1027,1051]" name="Mexico" pageId="24" pageNumber="25">Mexico</collectingCountry>
with distribution restricted to the northeast of the country (
<figureCitation id="6EB6A3C2F81E8D2DDD01FBB4FA0F7239" box="[1355,1420,1027,1051]" captionStart="Fig 13" captionStartId="24.[533,561,812,831]" captionTargetBox="[533,1493,208,790]" captionTargetId="figure-423@24.[533,1493,208,790]" captionTargetPageId="24" captionText="Fig 13. Historical records of the presence of Corynorhinus mexicanus and Corynorhinus leonpaniaguae sp. nov. The Mexican administrative boundaries layer was downloaded from the GADM (https://gadm.org/ downloadcountryv3.html)." figureDoi="http://doi.org/10.5281/zenodo.14290656" httpUri="https://zenodo.org/record/14290656/files/figure.png" pageId="24" pageNumber="25">Fig 13</figureCitation>
). In the localities of Galeana, 
<collectingRegion id="344971A5F81E8D2DDABFFB92FCF9721F" box="[757,890,1061,1085]" country="Mexico" name="Nuevo Leon" pageId="24" pageNumber="25">Nuevo León</collectingRegion>
, and Sierra de Zapalinamé in Arteaga, 
<collectingRegion id="344971A5F81E8D2DDD59FB92FAF6721F" box="[1299,1397,1061,1085]" country="Mexico" name="Coahuila de Zaragoza" pageId="24" pageNumber="25">Coahuila</collectingRegion>
, specimens were captured in sites close to forests where globose and cylindrical cacti are present. Other plant species present are 
<taxonomicName id="318DC4C4F81E8D2DDB57FBDCFC5172A0" box="[797,978,1130,1154]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="species" species="cembroides">
<emphasis id="C4F96355F81E8D2DDB57FBDCFC5172A0" box="[797,978,1130,1154]" italics="true" pageId="24" pageNumber="25">Pinus cembroides</emphasis>
</taxonomicName>
, 
<taxonomicName id="318DC4C4F81E8D2DDB94FBDCFBBB72A1" box="[990,1080,1131,1155]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="C4F96355F81E8D2DDB94FBDCFB9A72A0" box="[990,1049,1131,1154]" italics="true" pageId="24" pageNumber="25">Pinus</emphasis>
sp
</taxonomicName>
, 
<taxonomicName id="318DC4C4F81E8D2DDC08FBDCFB4672A1" box="[1090,1221,1131,1155]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="C4F96355F81E8D2DDC08FBDCFB2572A0" box="[1090,1190,1131,1154]" italics="true" pageId="24" pageNumber="25">Juniperus</emphasis>
sp
</taxonomicName>
, 
<taxonomicName id="318DC4C4F81E8D2DDC85FBDCFADB72A0" box="[1231,1368,1130,1154]" class="Liliopsida" family="Asparagaceae" genus="Yucca" kingdom="Plantae" order="Asparagales" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="species" species="filifera">
<emphasis id="C4F96355F81E8D2DDC85FBDCFADB72A0" box="[1231,1368,1130,1154]" italics="true" pageId="24" pageNumber="25">Yucca filifera</emphasis>
</taxonomicName>
, 
<taxonomicName id="318DC4C4F81E8D2DDD2EFBDCFA7172A0" box="[1380,1522,1130,1155]" class="Liliopsida" family="Asparagaceae" genus="Yucca" kingdom="Plantae" order="Asparagales" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="species" species="linearifolia">
<emphasis id="C4F96355F81E8D2DDD2EFBDCFAF772A0" box="[1380,1396,1131,1154]" italics="true" pageId="24" pageNumber="25">Y</emphasis>
. 
<emphasis id="C4F96355F81E8D2DDDCAFBDDFA7172A0" box="[1408,1522,1130,1154]" italics="true" pageId="24" pageNumber="25">linearifolia</emphasis>
</taxonomicName>
, and 
<taxonomicName id="318DC4C4F81E8D2DDA09FB39FC8A7287" box="[579,777,1166,1189]" class="Liliopsida" family="Asparagaceae" genus="Yucca" kingdom="Plantae" order="Asparagales" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="species" species="carnerosana">
<emphasis id="C4F96355F81E8D2DDA09FB39FC8A7287" box="[579,777,1166,1189]" italics="true" pageId="24" pageNumber="25">Yucca carnerosana</emphasis>
</taxonomicName>
. The characteristics of the roosts used by this species are unknown; however, in 
<collectingRegion id="344971A5F81E8D2DDADDFB07FD7A72EA" box="[663,761,1200,1224]" country="Mexico" name="Coahuila de Zaragoza" pageId="24" pageNumber="25">Coahuila</collectingRegion>
, 
<specimenCount id="E08B74CEF81E8D2DDB49FB07FC1572EA" box="[771,918,1200,1224]" count="1" pageId="24" pageNumber="25" type="generic">one specimen</specimenCount>
was captured during the day inside an abandoned mine, and, in 
<collectingRegion id="344971A5F81E8D2DDA2FFB64FD6972C9" box="[613,746,1235,1259]" country="Mexico" name="Nuevo Leon" pageId="24" pageNumber="25">Nuevo León</collectingRegion>
, a colony with pregnant females was found in a limestone, dolomite, and gypsum cave.
</paragraph>
</subSubSection>
<caption id="A2F2EFCFF81E8D2DDA5FFC9BFD7A7550" ID-DOI="http://doi.org/10.5281/zenodo.14290656" ID-Zenodo-Dep="14290656" httpUri="https://zenodo.org/record/14290656/files/figure.png" pageId="24" pageNumber="25" startId="24.[533,561,812,831]" targetBox="[533,1493,208,790]" targetPageId="24" targetType="figure">
<paragraph id="F632BF47F81E8D2DDA5FFC9BFD7A7550" blockId="24.[533,1508,811,882]" pageId="24" pageNumber="25">
<emphasis id="C4F96355F81E8D2DDA5FFC9BFA67751D" bold="true" box="[533,1508,811,831]" pageId="24" pageNumber="25">
Fig 13. Historical records of the presence of 
<taxonomicName id="318DC4C4F81E8D2DDBD0FC9CFBF7751D" authorityName="G. M. Allen" authorityYear="1916" box="[922,1140,811,831]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="mexicanus">
<emphasis id="C4F96355F81E8D2DDBD0FC9CFBF7751D" bold="true" box="[922,1140,811,831]" italics="true" pageId="24" pageNumber="25">Corynorhinus mexicanus</emphasis>
</taxonomicName>
and 
<taxonomicName id="318DC4C4F81E8D2DDCD5FC9CFA1F751C" authorityName="López-Cuamatzi &amp; Ortega &amp; Ospina-Garcés &amp; Zúñiga &amp; G." authorityYear="2024" box="[1183,1436,811,830]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="leonpaniaguae" status="sp. nov.">
<emphasis id="C4F96355F81E8D2DDCD5FC9CFA1F751C" bold="true" box="[1183,1436,811,830]" italics="true" pageId="24" pageNumber="25">Corynorhinus leonpaniaguae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DFCADE2EF81E8D2DDDEAFC9BFA67751D" box="[1440,1508,812,831]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
The Mexican administrative boundaries layer was downloaded from the GADM (https://gadm.org/ downloadcountryv3.html).
</paragraph>
</caption>
<paragraph id="F632BF47F81E8D2DDA5FFC3FFC2E75B9" blockId="24.[533,941,904,923]" box="[533,941,904,923]" pageId="24" pageNumber="25">https://doi.org/10.1371/journal.pone.0296275.g013</paragraph>
<subSubSection id="BE97ECCCF81E8D2CDA7FFAAFFB6C7141" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="description">
<paragraph id="F632BF47F81E8D2DDA7FFAAFFB767022" blockId="24.[533,1531,957,1883]" pageId="24" pageNumber="25">
<emphasis id="C4F96355F81E8D2DDA7FFAAFFCFA7312" bold="true" box="[565,889,1304,1328]" pageId="24" pageNumber="25">Description and comparison.</emphasis>
The specimens from 
<collectingRegion id="344971A5F81E8D2DDC27FAAFFB737312" box="[1133,1264,1304,1328]" country="Mexico" name="Nuevo Leon" pageId="24" pageNumber="25">Nuevo León</collectingRegion>
present a brownish to grayish colored dorsal fur with dark bases and slightly lighter tips that appear to contrast between bands. However, this contrast is not equivalent to that observed in 
<taxonomicName id="318DC4C4F81E8D2DDD7FFAEAFD0473BA" baseAuthorityName="Cooper" baseAuthorityYear="1838" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="townsendii">
<emphasis id="C4F96355F81E8D2DDD7FFAEAFD0473BA" italics="true" pageId="24" pageNumber="25">Corynorhinus townsendii</emphasis>
</taxonomicName>
(
<figureCitation id="6EB6A3C2F81E8D2DDADCFA37FD5573BA" box="[662,726,1408,1432]" captionStart="Fig 14" captionStartId="25.[533,561,1542,1561]" captionTargetBox="[533,1493,208,1520]" captionTargetId="figure-45@25.[533,1493,208,1520]" captionTargetPageId="25" captionText="Fig 14. Comparison between C. mexicanus, C. leonpaniaguae, and C. townsendii. Specimens shown correspond to C. mexicanus from (A) Puebla (ENCB27986), (B) Tlaxcala (ENCB4405), and (C) Durango (CRD11777); C. leonpaniaguae from the cave of San Josecito, in Gral. Zaragoza, Nuevo Leon, Mexico (D- CRD11774; E- CRD11778; and F- CRD11777). Specimen of C. townsendii from Durango (G- CRD4831). Some differences in color bands on the dorsal fur are shown for C. leonpaniaguae (H), C. mexicanus (I), and C. townsendii (J). Double-cuspid on upper incisor tooth (white arrow) observed in C. leonpaniaguae (K), C. mexicanus (L), and C. townsendii (M). https://doi.org/10.1371/journal.pone.0296275.g014" figureDoi="http://doi.org/10.5281/zenodo.14290658" httpUri="https://zenodo.org/record/14290658/files/figure.png" pageId="24" pageNumber="25">Fig 14</figureCitation>
). Like 
<taxonomicName id="318DC4C4F81E8D2DDB6AFA37FC2873B5" authorityName="G. M. Allen" authorityYear="1916" box="[800,939,1408,1432]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="mexicanus">
<emphasis id="C4F96355F81E8D2DDB6AFA37FCB273B5" box="[800,817,1408,1431]" italics="true" pageId="24" pageNumber="25">C</emphasis>
. 
<emphasis id="C4F96355F81E8D2DDB76FA37FC2873B5" box="[828,939,1408,1431]" italics="true" pageId="24" pageNumber="25">mexicanus</emphasis>
</taxonomicName>
and 
<taxonomicName id="318DC4C4F81E8D2DDB95FA37FBEF73BA" authorityName="Lesson" authorityYear="1827" box="[991,1132,1408,1432]" class="Mammalia" family="Vespertilionidae" genus="Plecotus" kingdom="Animalia" order="Chiroptera" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="rafinesquii">
<emphasis id="C4F96355F81E8D2DDB95FA37FC7373B5" box="[991,1008,1408,1431]" italics="true" pageId="24" pageNumber="25">C</emphasis>
. 
<emphasis id="C4F96355F81E8D2DDBB6FA37FBEF73BA" box="[1020,1132,1408,1432]" italics="true" pageId="24" pageNumber="25">rafinesquii</emphasis>
</taxonomicName>
, 
<taxonomicName id="318DC4C4F81E8D2DDC32FA37FAAF73BA" box="[1144,1324,1408,1432]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="leonpaniaguae">
<emphasis id="C4F96355F81E8D2DDC32FA37FB0A73B5" box="[1144,1161,1408,1431]" italics="true" pageId="24" pageNumber="25">C</emphasis>
. 
<emphasis id="C4F96355F81E8D2DDCDEFA37FAAF73BA" box="[1172,1324,1408,1432]" italics="true" pageId="24" pageNumber="25">leonpaniaguae</emphasis>
</taxonomicName>
presents a double cuspid in the first upper incisor (
<figureCitation id="6EB6A3C2F81E8D2DDB25FA14FC387399" box="[879,955,1443,1467]" captionStart="Fig 12" captionStartId="23.[533,561,1443,1462]" captionTargetBox="[533,1493,208,1421]" captionTargetId="figure-132@23.[533,1493,208,1421]" captionTargetPageId="23" captionText="Fig 12. Holotype pictures. Coloration of the dorsal and ventral fur of the holotype (MZFC-M16326) of Corynorhinus leonpaniaguae sp. nov. (A, B). Cranium, mandible (C, D, E) and lateral view of the first upper double-cuspid incisor (F) of the holotype. https://doi.org/10.1371/journal.pone.0296275.g012" figureDoi="http://doi.org/10.5281/zenodo.14290654" httpUri="https://zenodo.org/record/14290654/files/figure.png" pageId="24" pageNumber="25">Figs 12</figureCitation>
and 
<figureCitation id="6EB6A3C2F81E8D2DDBA5FA14FB8B7399" box="[1007,1032,1443,1467]" captionStart="Fig 14" captionStartId="25.[533,561,1542,1561]" captionTargetBox="[533,1493,208,1520]" captionTargetId="figure-45@25.[533,1493,208,1520]" captionTargetPageId="25" captionText="Fig 14. Comparison between C. mexicanus, C. leonpaniaguae, and C. townsendii. Specimens shown correspond to C. mexicanus from (A) Puebla (ENCB27986), (B) Tlaxcala (ENCB4405), and (C) Durango (CRD11777); C. leonpaniaguae from the cave of San Josecito, in Gral. Zaragoza, Nuevo Leon, Mexico (D- CRD11774; E- CRD11778; and F- CRD11777). Specimen of C. townsendii from Durango (G- CRD4831). Some differences in color bands on the dorsal fur are shown for C. leonpaniaguae (H), C. mexicanus (I), and C. townsendii (J). Double-cuspid on upper incisor tooth (white arrow) observed in C. leonpaniaguae (K), C. mexicanus (L), and C. townsendii (M). https://doi.org/10.1371/journal.pone.0296275.g014" figureDoi="http://doi.org/10.5281/zenodo.14290658" httpUri="https://zenodo.org/record/14290658/files/figure.png" pageId="24" pageNumber="25">14</figureCitation>
). Its dental formula is i 2/3, c 1/1, p 2/3, and m 3/3, with a total of 36 teeth in its adult stage. The average values of cranial measurements and external measurements for females and males are reported in 
<tableCitation id="BB0F8AFCF81E8D2DDCEBFA5FFB6C7022" box="[1185,1263,1512,1536]" captionStart="Table 4" captionStartId="26.[96,146,208,227]" captionTargetId="graphics-394@26.[96,1536,235,670]" captionTargetPageId="26" captionText="Table 4. Morphological measurements of external structures obtained in Corynorhinus leonpaniaguae." httpUri="http://table.plazi.org/id/A2F2EFCFF81C8D2FD82AFF67FC6B76C1" pageId="24" pageNumber="25" tableUuid="A2F2EFCFF81C8D2FD82AFF67FC6B76C1">Table 4</tableCitation>
.
</paragraph>
<paragraph id="F632BF47F81E8D2DDA7FF9BCFD007179" blockId="24.[533,1531,957,1883]" pageId="24" pageNumber="25">
In external appearance, it is similar to and is almost indistinguishable from 
<taxonomicName id="318DC4C4F81E8D2DDD18F9BCFA5D7000" authorityName="G. M. Allen" authorityYear="1916" box="[1362,1502,1547,1571]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="mexicanus">
<emphasis id="C4F96355F81E8D2DDD18F9BCFAE07000" box="[1362,1379,1547,1570]" italics="true" pageId="24" pageNumber="25">C</emphasis>
. 
<emphasis id="C4F96355F81E8D2DDD25F9BCFA5D7000" box="[1391,1502,1547,1570]" italics="true" pageId="24" pageNumber="25">mexicanus</emphasis>
</taxonomicName>
specimens from the Sierra Madre Occidental (SMOC). 
<taxonomicName id="318DC4C4F81E8D2DDC14F99AFA0E7067" box="[1118,1421,1581,1605]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="leonpaniaguae">
<emphasis id="C4F96355F81E8D2DDC14F99AFA0E7067" box="[1118,1421,1581,1605]" italics="true" pageId="24" pageNumber="25">Corynorhinus leonpaniaguae</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="318DC4C4F81E8D2DDAE3F9E7FCB4704A" baseAuthorityName="Cooper" baseAuthorityYear="1838" box="[681,823,1616,1640]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="townsendii">
<emphasis id="C4F96355F81E8D2DDAE3F9E7FD397045" box="[681,698,1616,1639]" italics="true" pageId="24" pageNumber="25">C</emphasis>
. 
<emphasis id="C4F96355F81E8D2DDA8FF9E7FCB4704A" box="[709,823,1616,1640]" italics="true" pageId="24" pageNumber="25">townsendii</emphasis>
</taxonomicName>
because the latter presents a tragus of&gt; 
<quantity id="317512A2F81E8D2DDCACF9E7FAB2704A" box="[1254,1329,1616,1640]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="24" pageNumber="25" unit="mm" value="13.0">13 mm</quantity>
and a naked uropatagium with ten or more interfemoral grooves, while the former presents a tragus of &lt;
<quantity id="317512A2F81E8D2DDD8EF9C4FDC2708F" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="24" pageNumber="25" unit="mm" value="13.0">13 mm</quantity>
and a hairy uropatagium with nine or fewer interfemoral grooves. Moreover, 
<taxonomicName id="318DC4C4F81E8D2DDD30F921FDB270F2" baseAuthorityName="Cooper" baseAuthorityYear="1838" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="townsendii">
<emphasis id="C4F96355F81E8D2DDD30F921FA08708F" box="[1402,1419,1686,1709]" italics="true" pageId="24" pageNumber="25">C</emphasis>
. 
<emphasis id="C4F96355F81E8D2DDDDDF921FDB270F2" italics="true" pageId="24" pageNumber="25">townsendii</emphasis>
</taxonomicName>
usually has a larger forearm (
<quantity id="317512A2F81E8D2DDB23F90FFC7270F2" box="[873,1009,1720,1744]" metricMagnitude="-2" metricUnit="m" metricValue="4.3" metricValueMax="4.7" metricValueMin="3.9" pageId="24" pageNumber="25" unit="mm" value="43.0" valueMax="47.0" valueMin="39.0">39 to 47 mm</quantity>
) and cranium maximum length (
<quantity id="317512A2F81E8D2DDD18F90FFDC170D1" metricMagnitude="-2" metricUnit="m" metricValue="1.625" metricValueMax="1.73" metricValueMin="1.52" pageId="24" pageNumber="25" unit="mm" value="16.25" valueMax="17.3" valueMin="15.2">15.2 to 17.3 mm</quantity>
) than 
<taxonomicName id="318DC4C4F81E8D2DDACCF96CFCB870D0" box="[646,827,1754,1779]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="leonpaniaguae">
<emphasis id="C4F96355F81E8D2DDACCF96CFD1470D0" box="[646,663,1755,1778]" italics="true" pageId="24" pageNumber="25">C</emphasis>
. 
<emphasis id="C4F96355F81E8D2DDAE9F96DFCB870D0" box="[675,827,1754,1778]" italics="true" pageId="24" pageNumber="25">leonpaniaguae</emphasis>
</taxonomicName>
. In external appearance, 
<taxonomicName id="318DC4C4F81E8D2DDC0FF96CFB7A70D0" box="[1093,1273,1754,1779]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="leonpaniaguae">
<emphasis id="C4F96355F81E8D2DDC0FF96CFBD570D0" box="[1093,1110,1755,1778]" italics="true" pageId="24" pageNumber="25">C</emphasis>
. 
<emphasis id="C4F96355F81E8D2DDC2BF96DFB7A70D0" box="[1121,1273,1754,1778]" italics="true" pageId="24" pageNumber="25">leonpaniaguae</emphasis>
</taxonomicName>
is also similar to 
<taxonomicName id="318DC4C4F81E8D2DDDFAF96CFDE37137" authorityName="Lesson" authorityYear="1827" class="Mammalia" family="Vespertilionidae" genus="Plecotus" kingdom="Animalia" order="Chiroptera" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="rafinesquii">
<emphasis id="C4F96355F81E8D2DDDFAF96CFA4270D0" box="[1456,1473,1755,1778]" italics="true" pageId="24" pageNumber="25">C</emphasis>
. 
<emphasis id="C4F96355F81E8D2DDD87F96CFDE37137" italics="true" pageId="24" pageNumber="25">rafinesquii</emphasis>
</taxonomicName>
; however, 
<taxonomicName id="318DC4C4F81E8D2DDA98F949FC047137" box="[722,903,1789,1813]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="leonpaniaguae">
<emphasis id="C4F96355F81E8D2DDA98F949FD607137" box="[722,739,1790,1813]" italics="true" pageId="24" pageNumber="25">C</emphasis>
. 
<emphasis id="C4F96355F81E8D2DDAA5F94AFC047137" box="[751,903,1789,1813]" italics="true" pageId="24" pageNumber="25">leonpaniaguae</emphasis>
</taxonomicName>
has bicolored ventral fur with brown bases and light tips, while 
<taxonomicName id="318DC4C4F81E8D2DDA1FF897FD62711A" authorityName="Lesson" authorityYear="1827" box="[597,737,1824,1848]" class="Mammalia" family="Vespertilionidae" genus="Plecotus" kingdom="Animalia" order="Chiroptera" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="rafinesquii">
<emphasis id="C4F96355F81E8D2DDA1FF897FDE57115" box="[597,614,1824,1847]" italics="true" pageId="24" pageNumber="25">C</emphasis>
. 
<emphasis id="C4F96355F81E8D2DDA3BF897FD62711A" box="[625,737,1824,1848]" italics="true" pageId="24" pageNumber="25">rafinesquii</emphasis>
</taxonomicName>
has more contrasting fur due to the presence of hairs with black bases and white tips.
</paragraph>
<caption id="A2F2EFCFF81F8D2CDA5FF9B1FC2E70E3" ID-DOI="http://doi.org/10.5281/zenodo.14290658" ID-Zenodo-Dep="14290658" httpUri="https://zenodo.org/record/14290658/files/figure.png" pageId="25" pageNumber="26" startId="25.[533,561,1542,1561]" targetBox="[533,1493,208,1520]" targetPageId="25" targetType="figure">
<paragraph id="F632BF47F81F8D2CDA5FF9B1FABF70B5" blockId="25.[533,1533,1541,1729]" pageId="25" pageNumber="26">
<emphasis id="C4F96355F81F8D2CDA5FF9B1FB5B703B" bold="true" box="[533,1240,1541,1561]" pageId="25" pageNumber="26">
Fig 14. Comparison between 
<taxonomicName id="318DC4C4F81F8D2CDB5DF9B2FC08703A" authorityName="G. M. Allen" authorityYear="1916" box="[791,907,1541,1561]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="mexicanus">
<emphasis id="C4F96355F81F8D2CDB5DF9B2FCA6703A" bold="true" box="[791,805,1541,1560]" italics="true" pageId="25" pageNumber="26">C</emphasis>
. 
<emphasis id="C4F96355F81F8D2CDB65F9B2FC08703A" bold="true" box="[815,907,1541,1560]" italics="true" pageId="25" pageNumber="26">mexicanus</emphasis>
</taxonomicName>
, 
<taxonomicName id="318DC4C4F81F8D2CDBDFF9B2FBAE703A" box="[917,1069,1541,1561]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="leonpaniaguae">
<emphasis id="C4F96355F81F8D2CDBDFF9B2FC20703A" bold="true" box="[917,931,1541,1560]" italics="true" pageId="25" pageNumber="26">C</emphasis>
. 
<emphasis id="C4F96355F81F8D2CDBE7F9B2FBAE703A" bold="true" box="[941,1069,1541,1560]" italics="true" pageId="25" pageNumber="26">leonpaniaguae</emphasis>
</taxonomicName>
, and 
<taxonomicName id="318DC4C4F81F8D2CDC17F9B2FB50703A" baseAuthorityName="Cooper" baseAuthorityYear="1838" box="[1117,1235,1541,1561]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="townsendii">
<emphasis id="C4F96355F81F8D2CDC17F9B2FBE8703A" bold="true" box="[1117,1131,1541,1560]" italics="true" pageId="25" pageNumber="26">C</emphasis>
. 
<emphasis id="C4F96355F81F8D2CDC3EF9B2FB50703A" bold="true" box="[1140,1235,1541,1560]" italics="true" pageId="25" pageNumber="26">townsendii</emphasis>
</taxonomicName>
.
</emphasis>
Specimens shown correspond to 
<taxonomicName id="318DC4C4F81F8D2CDA5FF9A8FD067013" authorityName="G. M. Allen" authorityYear="1916" box="[533,645,1567,1586]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="mexicanus">
<emphasis id="C4F96355F81F8D2CDA5FF9A8FDA17010" box="[533,546,1567,1586]" italics="true" pageId="25" pageNumber="26">C</emphasis>
. 
<emphasis id="C4F96355F81F8D2CDA66F9A8FD067013" box="[556,645,1567,1585]" italics="true" pageId="25" pageNumber="26">mexicanus</emphasis>
</taxonomicName>
from (A) Puebla (ENCB27986), (B) Tlaxcala (ENCB4405), and (C) Durango (CRD11777); 
<taxonomicName id="318DC4C4F81F8D2CDDCEF9A8FD0C7069" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="25" pageNumber="29" phylum="Chordata" rank="species" species="leonpaniaguae">
C. 
<emphasis id="C4F96355F81F8D2CDA5FF98FFD0C7069" box="[533,655,1592,1611]" italics="true" pageId="25" pageNumber="26">leonpaniaguae</emphasis>
</taxonomicName>
from the cave of San Josecito, in Gral. Zaragoza, Nuevo Leon, Mexico (D- CRD11774; E- CRD11778; and F- CRD11777). Specimen of 
<taxonomicName id="318DC4C4F81F8D2CDB66F9E5FC1D7046" baseAuthorityName="Cooper" baseAuthorityYear="1838" box="[812,926,1617,1637]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="townsendii">
<emphasis id="C4F96355F81F8D2CDB66F9E5FCBA7047" box="[812,825,1618,1637]" italics="true" pageId="25" pageNumber="26">C</emphasis>
. 
<emphasis id="C4F96355F81F8D2CDB09F9E6FC1D7046" box="[835,926,1617,1636]" italics="true" pageId="25" pageNumber="26">townsendii</emphasis>
</taxonomicName>
from Durango (G- CRD4831). Some differences in color bands on the dorsal fur are shown for 
<taxonomicName id="318DC4C4F81F8D2CDAAEF9DCFCF6705C" box="[740,885,1643,1662]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="leonpaniaguae">
<emphasis id="C4F96355F81F8D2CDAAEF9DCFD72705C" box="[740,753,1643,1662]" italics="true" pageId="25" pageNumber="26">C</emphasis>
. 
<emphasis id="C4F96355F81F8D2CDAB1F9DCFCF6705C" box="[763,885,1643,1662]" italics="true" pageId="25" pageNumber="26">leonpaniaguae</emphasis>
</taxonomicName>
(H), 
<taxonomicName id="318DC4C4F81F8D2CDBE8F9DCFB91705F" authorityName="G. M. Allen" authorityYear="1916" box="[930,1042,1643,1662]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="mexicanus">
<emphasis id="C4F96355F81F8D2CDBE8F9DCFC2C705C" box="[930,943,1643,1662]" italics="true" pageId="25" pageNumber="26">C</emphasis>
. 
<emphasis id="C4F96355F81F8D2CDBF3F9DCFB91705F" box="[953,1042,1643,1661]" italics="true" pageId="25" pageNumber="26">mexicanus</emphasis>
</taxonomicName>
(I), and 
<taxonomicName id="318DC4C4F81F8D2CDC10F9DCFB4F705C" baseAuthorityName="Cooper" baseAuthorityYear="1838" box="[1114,1228,1643,1662]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="townsendii">
<emphasis id="C4F96355F81F8D2CDC10F9DCFBE4705C" box="[1114,1127,1643,1662]" italics="true" pageId="25" pageNumber="26">C</emphasis>
. 
<emphasis id="C4F96355F81F8D2CDC3BF9DCFB4F705C" box="[1137,1228,1643,1662]" italics="true" pageId="25" pageNumber="26">townsendii</emphasis>
</taxonomicName>
(J). Double-cuspid on upper incisor tooth (white arrow) observed in 
<taxonomicName id="318DC4C4F81F8D2CDB6CF933FC3470B5" box="[806,951,1668,1687]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="leonpaniaguae">
<emphasis id="C4F96355F81F8D2CDB6CF933FCB070B5" box="[806,819,1668,1687]" italics="true" pageId="25" pageNumber="26">C</emphasis>
. 
<emphasis id="C4F96355F81F8D2CDB77F933FC3470B5" box="[829,951,1668,1687]" italics="true" pageId="25" pageNumber="26">leonpaniaguae</emphasis>
</taxonomicName>
(K), 
<taxonomicName id="318DC4C4F81F8D2CDBA8F933FBD170B5" authorityName="G. M. Allen" authorityYear="1916" box="[994,1106,1668,1687]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="mexicanus">
<emphasis id="C4F96355F81F8D2CDBA8F933FC6C70B5" box="[994,1007,1668,1687]" italics="true" pageId="25" pageNumber="26">C</emphasis>
. 
<emphasis id="C4F96355F81F8D2CDBB3F932FBD170B5" box="[1017,1106,1669,1687]" italics="true" pageId="25" pageNumber="26">mexicanus</emphasis>
</taxonomicName>
(L), and 
<taxonomicName id="318DC4C4F81F8D2CDCD4F933FA9370B5" baseAuthorityName="Cooper" baseAuthorityYear="1838" box="[1182,1296,1668,1687]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="townsendii">
<emphasis id="C4F96355F81F8D2CDCD4F933FB2870B5" box="[1182,1195,1668,1687]" italics="true" pageId="25" pageNumber="26">C</emphasis>
. 
<emphasis id="C4F96355F81F8D2CDCFFF933FA9370B5" box="[1205,1296,1668,1687]" italics="true" pageId="25" pageNumber="26">townsendii</emphasis>
</taxonomicName>
(M).
</paragraph>
<paragraph id="F632BF47F81F8D2CDA5FF919FC2E70E3" blockId="25.[533,1533,1541,1729]" box="[533,941,1710,1729]" pageId="25" pageNumber="26">https://doi.org/10.1371/journal.pone.0296275.g014</paragraph>
</caption>
<paragraph id="F632BF47F81F8D2CDA7FF954FA957162" blockId="25.[533,1530,1763,1891]" pageId="25" pageNumber="26">
<emphasis id="C4F96355F81F8D2CDA7FF954FCA470D9" bold="true" box="[565,807,1763,1787]" pageId="25" pageNumber="26">Geographic variation.</emphasis>
With the restricted distribution and limited sample size, it was not possible to detect morphological variation associated with geography. Only two haplotypes of Cyt- 
<emphasis id="C4F96355F81F8D2CDA0EF89FFDD27162" box="[580,593,1832,1856]" italics="true" pageId="25" pageNumber="26">b</emphasis>
sequences have been detected within 
<taxonomicName id="318DC4C4F81F8D2CDBABF89FFA937162" box="[993,1296,1832,1856]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="leonpaniaguae">
<emphasis id="C4F96355F81F8D2CDBABF89FFA937162" box="[993,1296,1832,1856]" italics="true" pageId="25" pageNumber="26">Corynorhinus leonpaniaguae</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="F632BF47F81F8D2CDA7FF8FCFB6C7141" blockId="25.[533,1530,1763,1891]" box="[565,1263,1867,1891]" pageId="25" pageNumber="26">
<emphasis id="C4F96355F81F8D2CDA7FF8FCFD317141" bold="true" box="[565,690,1867,1891]" pageId="25" pageNumber="26">Subspecies.</emphasis>
<taxonomicName id="318DC4C4F81F8D2CDA82F8FCFC747141" box="[712,1015,1867,1891]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="leonpaniaguae">
<emphasis id="C4F96355F81F8D2CDA82F8FCFC747141" box="[712,1015,1867,1891]" italics="true" pageId="25" pageNumber="26">Corynorhinus leonpaniaguae</emphasis>
</taxonomicName>
is a monotypic species.
</paragraph>
</subSubSection>
<caption id="A2F2EFCFF81C8D2FD82AFF67FC6B76C1" ID-Table-UUID="A2F2EFCFF81C8D2FD82AFF67FC6B76C1" box="[96,1000,207,227]" httpUri="http://table.plazi.org/id/A2F2EFCFF81C8D2FD82AFF67FC6B76C1" pageId="26" pageNumber="27" startId="26.[96,146,208,227]" targetBox="[96,1497,247,716]" targetIsTable="true" targetPageId="26" targetType="table">
<paragraph id="F632BF47F81C8D2FD82AFF67FC6B76C1" blockId="26.[96,1000,207,227]" box="[96,1000,207,227]" pageId="26" pageNumber="27">
Table 4. Morphological measurements of external structures obtained in 
<taxonomicName id="318DC4C4F81C8D2FDAACFF78FC6076C0" box="[742,995,207,226]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="leonpaniaguae">
<emphasis id="C4F96355F81C8D2FDAACFF78FC6076C0" bold="true" box="[742,995,207,226]" italics="true" pageId="26" pageNumber="27">Corynorhinus leonpaniaguae</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="F632BF47F81C8D2FDAD2FF40FA8B74EE" pageId="26" pageNumber="27">
<table id="848D4DE7F81C72CAD82AFF40FA5A74EE" box="[96,1497,247,716]" gridcols="9" gridrows="14" pageId="26" pageNumber="27">
<tr id="48BDBD05F81C72CAD82AFF40FA5A7728" box="[96,1497,247,266]" gridrow="0" pageId="26" pageNumber="27" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-8="1">
<th id="0B6CD479F81C72CADAD2FF40FCB27728" box="[664,817,247,266]" gridcol="3" gridrow="0" pageId="26" pageNumber="27">
<emphasis id="C4F96355F81C8D2FDAD2FF40FD627728" bold="true" box="[664,737,247,266]" pageId="26" pageNumber="27">Females</emphasis>
</th>
<th id="0B6CD479F81C72CADC9BFF40FAE67728" box="[1233,1381,247,266]" gridcol="7" gridrow="0" pageId="26" pageNumber="27">
<emphasis id="C4F96355F81C8D2FDC9BFF40FA867728" bold="true" box="[1233,1285,247,266]" pageId="26" pageNumber="27">Males</emphasis>
</th>
</tr>
<tr id="48BDBD05F81C72CAD82AFEAEFA5A770E" box="[96,1497,281,300]" gridrow="1" pageId="26" pageNumber="27" rowspan-0="1">
<td id="0B6CD479F81C72CAD980FEAEFE7F770E" box="[458,508,281,300]" gridcol="1" gridrow="1" pageId="26" pageNumber="27">
<emphasis id="C4F96355F81C8D2FD980FEAEFE7F770E" bold="true" box="[458,508,281,300]" pageId="26" pageNumber="27">Mean</emphasis>
</td>
<td id="0B6CD479F81C72CADA06FEAEFDF3770E" box="[588,624,281,300]" gridcol="2" gridrow="1" pageId="26" pageNumber="27">
<emphasis id="C4F96355F81C8D2FDA1AFEAEFDE8770E" bold="true" box="[592,619,281,300]" pageId="26" pageNumber="27">SD</emphasis>
</td>
<td id="0B6CD479F81C72CADAD2FEAEFCB2770E" box="[664,817,281,300]" gridcol="3" gridrow="1" pageId="26" pageNumber="27">
<emphasis id="C4F96355F81C8D2FDA96FEAEFC97770E" bold="true" box="[732,788,281,300]" pageId="26" pageNumber="27">Range</emphasis>
</td>
<td id="0B6CD479F81C72CADBCAFEAEFC2D770E" box="[896,942,281,300]" gridcol="4" gridrow="1" pageId="26" pageNumber="27">
<emphasis id="C4F96355F81C8D2FDBC3FEAEFC24770E" bold="true" box="[905,935,281,300]" pageId="26" pageNumber="27">CV</emphasis>
</td>
<td id="0B6CD479F81C72CADBB4FEAEFBB3770E" box="[1022,1072,281,300]" gridcol="5" gridrow="1" pageId="26" pageNumber="27">
<emphasis id="C4F96355F81C8D2FDBB4FEAEFBB3770E" bold="true" box="[1022,1072,281,300]" pageId="26" pageNumber="27">Mean</emphasis>
</td>
<td id="0B6CD479F81C72CADCCBFEAEFB26770E" box="[1153,1189,281,300]" gridcol="6" gridrow="1" pageId="26" pageNumber="27">
<emphasis id="C4F96355F81C8D2FDCCFFEAEFB23770E" bold="true" box="[1157,1184,281,300]" pageId="26" pageNumber="27">SD</emphasis>
</td>
<td id="0B6CD479F81C72CADC9BFEAEFAE6770E" box="[1233,1381,281,300]" gridcol="7" gridrow="1" pageId="26" pageNumber="27">
<emphasis id="C4F96355F81C8D2FDD5BFEAEFACA770E" bold="true" box="[1297,1353,281,300]" pageId="26" pageNumber="27">Range</emphasis>
</td>
<td id="0B6CD479F81C72CADDFFFEAEFA5A770E" box="[1461,1497,281,300]" gridcol="8" gridrow="1" pageId="26" pageNumber="27">
<emphasis id="C4F96355F81C8D2FDDF2FEAEFA55770E" bold="true" box="[1464,1494,281,300]" pageId="26" pageNumber="27">CV</emphasis>
</td>
</tr>
<tr id="48BDBD05F81C72CAD82AFE8DFA5A776F" box="[96,1497,314,333]" gridrow="2" pageId="26" pageNumber="27">
<th id="0B6CD479F81C72CAD82AFE8DFEF9776F" box="[96,378,314,333]" gridcol="0" gridrow="2" pageId="26" pageNumber="27">Maximum length</th>
<td id="0B6CD479F81C72CAD980FE8DFE7F776F" box="[458,508,314,333]" gridcol="1" gridrow="2" pageId="26" pageNumber="27">14.87</td>
<td id="0B6CD479F81C72CADA06FE8DFDF3776F" box="[588,624,314,333]" gridcol="2" gridrow="2" pageId="26" pageNumber="27">0.34</td>
<td id="0B6CD479F81C72CADAD2FE8DFCB2776F" box="[664,817,314,333]" gridcol="3" gridrow="2" pageId="26" pageNumber="27">14.19–15.20</td>
<td id="0B6CD479F81C72CADBCAFE8DFC2D776F" box="[896,942,314,333]" gridcol="4" gridrow="2" pageId="26" pageNumber="27">2.28</td>
<td id="0B6CD479F81C72CADBB4FE8DFBB3776F" box="[1022,1072,314,333]" gridcol="5" gridrow="2" pageId="26" pageNumber="27">15.10</td>
<td id="0B6CD479F81C72CADCCBFE8DFB26776F" box="[1153,1189,314,333]" gridcol="6" gridrow="2" pageId="26" pageNumber="27">0.31</td>
<td id="0B6CD479F81C72CADC9BFE8DFAE6776F" box="[1233,1381,314,333]" gridcol="7" gridrow="2" pageId="26" pageNumber="27">14.8–15.68</td>
<td id="0B6CD479F81C72CADDFFFE8DFA5A776F" box="[1461,1497,314,333]" gridcol="8" gridrow="2" pageId="26" pageNumber="27">2.05</td>
</tr>
<tr id="48BDBD05F81C72CAD82AFEECFA5A774C" box="[96,1497,347,366]" gridrow="3" pageId="26" pageNumber="27">
<th id="0B6CD479F81C72CAD82AFEECFEF9774C" box="[96,378,347,366]" gridcol="0" gridrow="3" pageId="26" pageNumber="27">Zygomatic width</th>
<td id="0B6CD479F81C72CAD980FEECFE7F774C" box="[458,508,347,366]" gridcol="1" gridrow="3" pageId="26" pageNumber="27">7.94</td>
<td id="0B6CD479F81C72CADA06FEECFDF3774C" box="[588,624,347,366]" gridcol="2" gridrow="3" pageId="26" pageNumber="27">0.36</td>
<td id="0B6CD479F81C72CADAD2FEECFCB2774C" box="[664,817,347,366]" gridcol="3" gridrow="3" pageId="26" pageNumber="27">7.05–8.37</td>
<td id="0B6CD479F81C72CADBCAFEECFC2D774C" box="[896,942,347,366]" gridcol="4" gridrow="3" pageId="26" pageNumber="27">4.60</td>
<td id="0B6CD479F81C72CADBB4FEECFBB3774C" box="[1022,1072,347,366]" gridcol="5" gridrow="3" pageId="26" pageNumber="27">8.08</td>
<td id="0B6CD479F81C72CADCCBFEECFB26774C" box="[1153,1189,347,366]" gridcol="6" gridrow="3" pageId="26" pageNumber="27">0.24</td>
<td id="0B6CD479F81C72CADC9BFEECFAE6774C" box="[1233,1381,347,366]" gridcol="7" gridrow="3" pageId="26" pageNumber="27">7.79–8.38</td>
<td id="0B6CD479F81C72CADDFFFEECFA5A774C" box="[1461,1497,347,366]" gridcol="8" gridrow="3" pageId="26" pageNumber="27">3.01</td>
</tr>
<tr id="48BDBD05F81C72CAD82AFECAFA5A77B2" box="[96,1497,381,400]" gridrow="4" pageId="26" pageNumber="27">
<th id="0B6CD479F81C72CAD82AFECAFEF977B2" box="[96,378,381,400]" gridcol="0" gridrow="4" pageId="26" pageNumber="27">Postorbital width</th>
<td id="0B6CD479F81C72CAD980FECAFE7F77B2" box="[458,508,381,400]" gridcol="1" gridrow="4" pageId="26" pageNumber="27">3.36</td>
<td id="0B6CD479F81C72CADA06FECAFDF377B2" box="[588,624,381,400]" gridcol="2" gridrow="4" pageId="26" pageNumber="27">0.08</td>
<td id="0B6CD479F81C72CADAD2FECAFCB277B2" box="[664,817,381,400]" gridcol="3" gridrow="4" pageId="26" pageNumber="27">3.24–3.49</td>
<td id="0B6CD479F81C72CADBCAFECAFC2D77B2" box="[896,942,381,400]" gridcol="4" gridrow="4" pageId="26" pageNumber="27">2.43</td>
<td id="0B6CD479F81C72CADBB4FECAFBB377B2" box="[1022,1072,381,400]" gridcol="5" gridrow="4" pageId="26" pageNumber="27">3.36</td>
<td id="0B6CD479F81C72CADCCBFECAFB2677B2" box="[1153,1189,381,400]" gridcol="6" gridrow="4" pageId="26" pageNumber="27">0.12</td>
<td id="0B6CD479F81C72CADC9BFECAFAE677B2" box="[1233,1381,381,400]" gridcol="7" gridrow="4" pageId="26" pageNumber="27">3.17–3.54</td>
<td id="0B6CD479F81C72CADDFFFECAFA5A77B2" box="[1461,1497,381,400]" gridcol="8" gridrow="4" pageId="26" pageNumber="27">3.64</td>
</tr>
<tr id="48BDBD05F81C72CAD82AFE29FA5A7793" box="[96,1497,414,433]" gridrow="5" pageId="26" pageNumber="27">
<th id="0B6CD479F81C72CAD82AFE29FEF97793" box="[96,378,414,433]" gridcol="0" gridrow="5" pageId="26" pageNumber="27">Width of the braincase</th>
<td id="0B6CD479F81C72CAD980FE29FE7F7793" box="[458,508,414,433]" gridcol="1" gridrow="5" pageId="26" pageNumber="27">7.73</td>
<td id="0B6CD479F81C72CADA06FE29FDF37793" box="[588,624,414,433]" gridcol="2" gridrow="5" pageId="26" pageNumber="27">0.27</td>
<td id="0B6CD479F81C72CADAD2FE29FCB27793" box="[664,817,414,433]" gridcol="3" gridrow="5" pageId="26" pageNumber="27">7.33–8.07</td>
<td id="0B6CD479F81C72CADBCAFE29FC2D7793" box="[896,942,414,433]" gridcol="4" gridrow="5" pageId="26" pageNumber="27">3.43</td>
<td id="0B6CD479F81C72CADBB4FE29FBB37793" box="[1022,1072,414,433]" gridcol="5" gridrow="5" pageId="26" pageNumber="27">7.72</td>
<td id="0B6CD479F81C72CADCCBFE29FB267793" box="[1153,1189,414,433]" gridcol="6" gridrow="5" pageId="26" pageNumber="27">0.34</td>
<td id="0B6CD479F81C72CADC9BFE29FAE67793" box="[1233,1381,414,433]" gridcol="7" gridrow="5" pageId="26" pageNumber="27">7.35–8.38</td>
<td id="0B6CD479F81C72CADDFFFE29FA5A7793" box="[1461,1497,414,433]" gridcol="8" gridrow="5" pageId="26" pageNumber="27">4.44</td>
</tr>
<tr id="48BDBD05F81C72CAD82AFE08FA5A77F0" box="[96,1497,447,466]" gridrow="6" pageId="26" pageNumber="27">
<th id="0B6CD479F81C72CAD82AFE08FEF977F0" box="[96,378,447,466]" gridcol="0" gridrow="6" pageId="26" pageNumber="27">Maxillary toothrow length</th>
<td id="0B6CD479F81C72CAD980FE08FE7F77F0" box="[458,508,447,466]" gridcol="1" gridrow="6" pageId="26" pageNumber="27">4.74</td>
<td id="0B6CD479F81C72CADA06FE08FDF377F0" box="[588,624,447,466]" gridcol="2" gridrow="6" pageId="26" pageNumber="27">0.08</td>
<td id="0B6CD479F81C72CADAD2FE08FCB277F0" box="[664,817,447,466]" gridcol="3" gridrow="6" pageId="26" pageNumber="27">4.67–4.94</td>
<td id="0B6CD479F81C72CADBCAFE08FC2D77F0" box="[896,942,447,466]" gridcol="4" gridrow="6" pageId="26" pageNumber="27">1.78</td>
<td id="0B6CD479F81C72CADBB4FE08FBB377F0" box="[1022,1072,447,466]" gridcol="5" gridrow="6" pageId="26" pageNumber="27">4.69</td>
<td id="0B6CD479F81C72CADCCBFE08FB2677F0" box="[1153,1189,447,466]" gridcol="6" gridrow="6" pageId="26" pageNumber="27">0.07</td>
<td id="0B6CD479F81C72CADC9BFE08FAE677F0" box="[1233,1381,447,466]" gridcol="7" gridrow="6" pageId="26" pageNumber="27">4.60–4.80</td>
<td id="0B6CD479F81C72CADDFFFE08FA5A77F0" box="[1461,1497,447,466]" gridcol="8" gridrow="6" pageId="26" pageNumber="27">1.47</td>
</tr>
<tr id="48BDBD05F81C72CAD82AFE56FA5A77D6" box="[96,1497,481,500]" gridrow="7" pageId="26" pageNumber="27">
<th id="0B6CD479F81C72CAD82AFE56FEF977D6" box="[96,378,481,500]" gridcol="0" gridrow="7" pageId="26" pageNumber="27">Width between upper molars</th>
<td id="0B6CD479F81C72CAD980FE56FE7F77D6" box="[458,508,481,500]" gridcol="1" gridrow="7" pageId="26" pageNumber="27">5.82</td>
<td id="0B6CD479F81C72CADA06FE56FDF377D6" box="[588,624,481,500]" gridcol="2" gridrow="7" pageId="26" pageNumber="27">0.11</td>
<td id="0B6CD479F81C72CADAD2FE56FCB277D6" box="[664,817,481,500]" gridcol="3" gridrow="7" pageId="26" pageNumber="27">5.66–5.96</td>
<td id="0B6CD479F81C72CADBCAFE56FC2D77D6" box="[896,942,481,500]" gridcol="4" gridrow="7" pageId="26" pageNumber="27">1.88</td>
<td id="0B6CD479F81C72CADBB4FE56FBB377D6" box="[1022,1072,481,500]" gridcol="5" gridrow="7" pageId="26" pageNumber="27">5.74</td>
<td id="0B6CD479F81C72CADCCBFE56FB2677D6" box="[1153,1189,481,500]" gridcol="6" gridrow="7" pageId="26" pageNumber="27">0.16</td>
<td id="0B6CD479F81C72CADC9BFE56FAE677D6" box="[1233,1381,481,500]" gridcol="7" gridrow="7" pageId="26" pageNumber="27">5.57–5.97</td>
<td id="0B6CD479F81C72CADDFFFE56FA5A77D6" box="[1461,1497,481,500]" gridcol="8" gridrow="7" pageId="26" pageNumber="27">2.78</td>
</tr>
<tr id="48BDBD05F81C72CAD82AFDB5FA5A7437" box="[96,1497,514,533]" gridrow="8" pageId="26" pageNumber="27">
<th id="0B6CD479F81C72CAD82AFDB5FEF97437" box="[96,378,514,533]" gridcol="0" gridrow="8" pageId="26" pageNumber="27">Total length</th>
<td id="0B6CD479F81C72CAD980FDB5FE7F7437" box="[458,508,514,533]" gridcol="1" gridrow="8" pageId="26" pageNumber="27">91.59</td>
<td id="0B6CD479F81C72CADA06FDB5FDF37437" box="[588,624,514,533]" gridcol="2" gridrow="8" pageId="26" pageNumber="27">5.73</td>
<td id="0B6CD479F81C72CADAD2FDB5FCB27437" box="[664,817,514,533]" gridcol="3" gridrow="8" pageId="26" pageNumber="27">84.37–102.79</td>
<td id="0B6CD479F81C72CADBCAFDB5FC2D7437" box="[896,942,514,533]" gridcol="4" gridrow="8" pageId="26" pageNumber="27">6.25</td>
<td id="0B6CD479F81C72CADBB4FDB5FBB37437" box="[1022,1072,514,533]" gridcol="5" gridrow="8" pageId="26" pageNumber="27">90.56</td>
<td id="0B6CD479F81C72CADCCBFDB5FB267437" box="[1153,1189,514,533]" gridcol="6" gridrow="8" pageId="26" pageNumber="27">6.00</td>
<td id="0B6CD479F81C72CADC9BFDB5FAE67437" box="[1233,1381,514,533]" gridcol="7" gridrow="8" pageId="26" pageNumber="27">82.08–101.14</td>
<td id="0B6CD479F81C72CADDFFFDB5FA5A7437" box="[1461,1497,514,533]" gridcol="8" gridrow="8" pageId="26" pageNumber="27">6.62</td>
</tr>
<tr id="48BDBD05F81C72CAD82AFD94FA5A7414" box="[96,1497,547,566]" gridrow="9" pageId="26" pageNumber="27">
<th id="0B6CD479F81C72CAD82AFD94FEF97414" box="[96,378,547,566]" gridcol="0" gridrow="9" pageId="26" pageNumber="27">Tail length</th>
<td id="0B6CD479F81C72CAD980FD94FE7F7414" box="[458,508,547,566]" gridcol="1" gridrow="9" pageId="26" pageNumber="27">43.37</td>
<td id="0B6CD479F81C72CADA06FD94FDF37414" box="[588,624,547,566]" gridcol="2" gridrow="9" pageId="26" pageNumber="27">6.15</td>
<td id="0B6CD479F81C72CADAD2FD94FCB27414" box="[664,817,547,566]" gridcol="3" gridrow="9" pageId="26" pageNumber="27">34.95–55.75</td>
<td id="0B6CD479F81C72CADBCAFD94FC2D7414" box="[896,942,547,566]" gridcol="4" gridrow="9" pageId="26" pageNumber="27">14.19</td>
<td id="0B6CD479F81C72CADBB4FD94FBB37414" box="[1022,1072,547,566]" gridcol="5" gridrow="9" pageId="26" pageNumber="27">40.12</td>
<td id="0B6CD479F81C72CADCCBFD94FB267414" box="[1153,1189,547,566]" gridcol="6" gridrow="9" pageId="26" pageNumber="27">3.53</td>
<td id="0B6CD479F81C72CADC9BFD94FAE67414" box="[1233,1381,547,566]" gridcol="7" gridrow="9" pageId="26" pageNumber="27">35.20–44.91</td>
<td id="0B6CD479F81C72CADDFFFD94FA5A7414" box="[1461,1497,547,566]" gridcol="8" gridrow="9" pageId="26" pageNumber="27">8.79</td>
</tr>
<tr id="48BDBD05F81C72CAD82AFDF2FA5A747A" box="[96,1497,581,600]" gridrow="10" pageId="26" pageNumber="27">
<th id="0B6CD479F81C72CAD82AFDF2FEF9747A" box="[96,378,581,600]" gridcol="0" gridrow="10" pageId="26" pageNumber="27">Ear length</th>
<td id="0B6CD479F81C72CAD980FDF2FE7F747A" box="[458,508,581,600]" gridcol="1" gridrow="10" pageId="26" pageNumber="27">28.66</td>
<td id="0B6CD479F81C72CADA06FDF2FDF3747A" box="[588,624,581,600]" gridcol="2" gridrow="10" pageId="26" pageNumber="27">1.24</td>
<td id="0B6CD479F81C72CADAD2FDF2FCB2747A" box="[664,817,581,600]" gridcol="3" gridrow="10" pageId="26" pageNumber="27">27.37–30.43</td>
<td id="0B6CD479F81C72CADBCAFDF2FC2D747A" box="[896,942,581,600]" gridcol="4" gridrow="10" pageId="26" pageNumber="27">4.33</td>
<td id="0B6CD479F81C72CADBB4FDF2FBB3747A" box="[1022,1072,581,600]" gridcol="5" gridrow="10" pageId="26" pageNumber="27">28.43</td>
<td id="0B6CD479F81C72CADCCBFDF2FB26747A" box="[1153,1189,581,600]" gridcol="6" gridrow="10" pageId="26" pageNumber="27">1.39</td>
<td id="0B6CD479F81C72CADC9BFDF2FAE6747A" box="[1233,1381,581,600]" gridcol="7" gridrow="10" pageId="26" pageNumber="27">27.13–31.30</td>
<td id="0B6CD479F81C72CADDFFFDF2FA5A747A" box="[1461,1497,581,600]" gridcol="8" gridrow="10" pageId="26" pageNumber="27">4.89</td>
</tr>
<tr id="48BDBD05F81C72CAD82AFDD1FA5A745B" box="[96,1497,614,633]" gridrow="11" pageId="26" pageNumber="27">
<th id="0B6CD479F81C72CAD82AFDD1FEF9745B" box="[96,378,614,633]" gridcol="0" gridrow="11" pageId="26" pageNumber="27">Tragus length</th>
<td id="0B6CD479F81C72CAD980FDD1FE7F745B" box="[458,508,614,633]" gridcol="1" gridrow="11" pageId="26" pageNumber="27">10.89</td>
<td id="0B6CD479F81C72CADA06FDD1FDF3745B" box="[588,624,614,633]" gridcol="2" gridrow="11" pageId="26" pageNumber="27">0.93</td>
<td id="0B6CD479F81C72CADAD2FDD1FCB2745B" box="[664,817,614,633]" gridcol="3" gridrow="11" pageId="26" pageNumber="27">9.54–12</td>
<td id="0B6CD479F81C72CADBCAFDD1FC2D745B" box="[896,942,614,633]" gridcol="4" gridrow="11" pageId="26" pageNumber="27">8.54</td>
<td id="0B6CD479F81C72CADBB4FDD1FBB3745B" box="[1022,1072,614,633]" gridcol="5" gridrow="11" pageId="26" pageNumber="27">10.23</td>
<td id="0B6CD479F81C72CADCCBFDD1FB26745B" box="[1153,1189,614,633]" gridcol="6" gridrow="11" pageId="26" pageNumber="27">0.80</td>
<td id="0B6CD479F81C72CADC9BFDD1FAE6745B" box="[1233,1381,614,633]" gridcol="7" gridrow="11" pageId="26" pageNumber="27">8.69–10.79</td>
<td id="0B6CD479F81C72CADDFFFDD1FA5A745B" box="[1461,1497,614,633]" gridcol="8" gridrow="11" pageId="26" pageNumber="27">7.80</td>
</tr>
<tr id="48BDBD05F81C72CAD82AFD30FA5A74B8" box="[96,1497,647,666]" gridrow="12" pageId="26" pageNumber="27">
<th id="0B6CD479F81C72CAD82AFD30FEF974B8" box="[96,378,647,666]" gridcol="0" gridrow="12" pageId="26" pageNumber="27">Forearm length</th>
<td id="0B6CD479F81C72CAD980FD30FE7F74B8" box="[458,508,647,666]" gridcol="1" gridrow="12" pageId="26" pageNumber="27">41.91</td>
<td id="0B6CD479F81C72CADA06FD30FDF374B8" box="[588,624,647,666]" gridcol="2" gridrow="12" pageId="26" pageNumber="27">1.48</td>
<td id="0B6CD479F81C72CADAD2FD30FCB274B8" box="[664,817,647,666]" gridcol="3" gridrow="12" pageId="26" pageNumber="27">38.71–43.94</td>
<td id="0B6CD479F81C72CADBCAFD30FC2D74B8" box="[896,942,647,666]" gridcol="4" gridrow="12" pageId="26" pageNumber="27">3.54</td>
<td id="0B6CD479F81C72CADBB4FD30FBB374B8" box="[1022,1072,647,666]" gridcol="5" gridrow="12" pageId="26" pageNumber="27">40.52</td>
<td id="0B6CD479F81C72CADCCBFD30FB2674B8" box="[1153,1189,647,666]" gridcol="6" gridrow="12" pageId="26" pageNumber="27">0.99</td>
<td id="0B6CD479F81C72CADC9BFD30FAE674B8" box="[1233,1381,647,666]" gridcol="7" gridrow="12" pageId="26" pageNumber="27">39.54–42.08</td>
<td id="0B6CD479F81C72CADDFFFD30FA5A74B8" box="[1461,1497,647,666]" gridcol="8" gridrow="12" pageId="26" pageNumber="27">2.44</td>
</tr>
<tr id="48BDBD05F81C72CAD82AFD0EFA5A74EE" box="[96,1497,697,716]" gridrow="13" pageId="26" pageNumber="27">
<th id="0B6CD479F81C72CAD82AFD0EFA5A74EE" box="[96,1497,697,716]" colspan="9" colspanRight="8" gridcol="0" gridrow="13" pageId="26" pageNumber="27">Mean, standard deviation (SD), range, and coefficient of variation (CV) values are shown. Measurements are reported in millimeters (mm).</th>
</tr>
</table>
</paragraph>
<paragraph id="F632BF47F81C8D2FD82AFD5EFE7074DE" blockId="26.[96,499,745,764]" box="[96,499,745,764]" pageId="26" pageNumber="27">
<tableNote id="0B6BBEC9F81C8D2FD82AFD5EFE7074DE" box="[96,499,745,764]" pageId="26" pageNumber="27" targetBox="[96,1497,247,716]" targetPageId="26">https://doi.org/10.1371/journal.pone.0296275.t004</tableNote>
</paragraph>
<subSubSection id="BE97ECCCF81C8D2FDA7FFC90FBCD70E7" pageId="26" pageNumber="27" type="biology_ecology">
<paragraph id="F632BF47F81C8D2FDA7FFC90FB1D7255" blockId="26.[533,1536,807,1733]" pageId="26" pageNumber="27">
<emphasis id="C4F96355F81C8D2FDA7FFC90FD64751D" bold="true" box="[565,743,807,831]" pageId="26" pageNumber="27">Natural history.</emphasis>
There is no published information documenting the natural history of this species. Some observations made during the fieldwork of this study suggest certain aspects of the biology of 
<taxonomicName id="318DC4C4F81C8D2FDA8DFCDBFC7575A6" box="[711,1014,876,900]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="leonpaniaguae">
<emphasis id="C4F96355F81C8D2FDA8DFCDBFC7575A6" box="[711,1014,876,900]" italics="true" pageId="26" pageNumber="27">Corynorhinus leonpaniaguae</emphasis>
</taxonomicName>
. In two expeditions conducted in 
<date id="82339987F81C8D2FDD15FCDAFA6675A7" box="[1375,1509,877,901]" pageId="26" pageNumber="27" value="2021-08">August 2021</date>
and 
<date id="82339987F81C8D2FDA09FC38FD377585" box="[579,692,911,935]" pageId="26" pageNumber="27" value="2022-04">April 2022</date>
at the 
<typeStatus id="293601E5F81C8D2FDAB3FC38FCA57585" box="[761,806,911,935]" pageId="26" pageNumber="27">type</typeStatus>
locality of this species, several specimens of 
<taxonomicName id="318DC4C4F81C8D2FDCB3FC27FA3A7585" authorityName="Miller" authorityYear="1897" box="[1273,1465,911,935]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="thysanodes">
<emphasis id="C4F96355F81C8D2FDCB3FC27FA3A7585" box="[1273,1465,911,935]" italics="true" pageId="26" pageNumber="27">Myotis thysanodes</emphasis>
</taxonomicName>
, 
<taxonomicName id="318DC4C4F81C8D2FDD8FFC38FD6875E8" baseAuthorityName="Cooper" baseAuthorityYear="1838" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="townsendii">
<emphasis id="C4F96355F81C8D2FDD8FFC38FD6875E8" italics="true" pageId="26" pageNumber="27">Corynorhinus townsendii</emphasis>
</taxonomicName>
, 
<taxonomicName id="318DC4C4F81C8D2FDABDFC05FC4F75E8" baseAuthorityName="Saussure" baseAuthorityYear="1860" box="[759,972,946,970]" class="Mammalia" family="Phyllostomidae" genus="Leptonycteris" kingdom="Animalia" order="Chiroptera" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="nivalis">
<emphasis id="C4F96355F81C8D2FDABDFC05FC4F75E8" box="[759,972,946,970]" italics="true" pageId="26" pageNumber="27">Leptonycteris nivalis</emphasis>
</taxonomicName>
, 
<taxonomicName id="318DC4C4F81C8D2FDB9DFC05FB2C75E8" baseAuthorityName="G. M. Allen" baseAuthorityYear="1916" box="[983,1199,946,970]" class="Mammalia" family="Vespertilionidae" genus="Idionycteris" kingdom="Animalia" order="Chiroptera" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="phyllotis">
<emphasis id="C4F96355F81C8D2FDB9DFC05FB2C75E8" box="[983,1199,946,970]" italics="true" pageId="26" pageNumber="27">Idionycteris phyllotis</emphasis>
</taxonomicName>
, and 
<taxonomicName id="318DC4C4F81C8D2FDCA3FC05FA2D75E8" baseAuthorityName="LeConte" baseAuthorityYear="1856" box="[1257,1454,946,970]" class="Mammalia" family="Vespertilionidae" genus="Antrozous" kingdom="Animalia" order="Chiroptera" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="pallidus">
<emphasis id="C4F96355F81C8D2FDCA3FC05FA2D75E8" box="[1257,1454,946,970]" italics="true" pageId="26" pageNumber="27">Antrozous pallidus</emphasis>
</taxonomicName>
were captured. These specimens were captured at the entrance of the El Hundido Cave both entering and exiting the cave. This suggests that 
<taxonomicName id="318DC4C4F81C8D2FDB97FC40FA88722D" box="[989,1291,1015,1039]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="leonpaniaguae">
<emphasis id="C4F96355F81C8D2FDB97FC40FA88722D" box="[989,1291,1015,1039]" italics="true" pageId="26" pageNumber="27">Corynorhinus leonpaniaguae</emphasis>
</taxonomicName>
may share its roosts with these species. Of those mentioned above, 
<taxonomicName id="318DC4C4F81C8D2FDC4BFBACFB147210" authorityName="Miller" authorityYear="1897" box="[1025,1175,1050,1074]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="thysanodes">
<emphasis id="C4F96355F81C8D2FDC4BFBACFB9B7210" box="[1025,1048,1051,1074]" italics="true" pageId="26" pageNumber="27">M</emphasis>
. 
<emphasis id="C4F96355F81C8D2FDC6EFBADFB147210" box="[1060,1175,1050,1074]" italics="true" pageId="26" pageNumber="27">thysanodes</emphasis>
</taxonomicName>
and 
<taxonomicName id="318DC4C4F81C8D2FDC80FBADFADA7210" baseAuthorityName="Cooper" baseAuthorityYear="1838" box="[1226,1369,1050,1074]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="townsendii">
<emphasis id="C4F96355F81C8D2FDC80FBADFB587213" box="[1226,1243,1050,1073]" italics="true" pageId="26" pageNumber="27">C</emphasis>
. 
<emphasis id="C4F96355F81C8D2FDCADFBADFADA7210" box="[1255,1369,1050,1074]" italics="true" pageId="26" pageNumber="27">townsendii</emphasis>
</taxonomicName>
were the only species, apart from 
<taxonomicName id="318DC4C4F81C8D2FDAA9FB8BFB927276" box="[739,1041,1084,1108]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="leonpaniaguae">
<emphasis id="C4F96355F81C8D2FDAA9FB8BFB927276" box="[739,1041,1084,1108]" italics="true" pageId="26" pageNumber="27">Corynorhinus leonpaniaguae</emphasis>
</taxonomicName>
, that were captured on both field trips, indicating that all three species are probably residents of the cave.
</paragraph>
<paragraph id="F632BF47F81C8D2FDA7FFB35FC1D7365" blockId="26.[533,1536,807,1733]" pageId="26" pageNumber="27">
In 
<date id="82339987F81C8D2FDA19FB35FD5972B8" box="[595,730,1154,1178]" pageId="26" pageNumber="27" value="2021-08">August 2021</date>
, 
<specimenCount id="E08B74CEF81C8D2FDAAEFB35FCE872B8" box="[740,875,1154,1178]" count="4" pageId="26" pageNumber="27" type="juvenile">four juvenile</specimenCount>
male individuals were captured, indicating that weaning of these specimens had occurred in June-July, which is similar to that reported for 
<taxonomicName id="318DC4C4F81C8D2FDD28FB12FA6D729E" authorityName="G. M. Allen" authorityYear="1916" box="[1378,1518,1189,1213]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="mexicanus">
<emphasis id="C4F96355F81C8D2FDD28FB12FAF0729E" box="[1378,1395,1189,1212]" italics="true" pageId="26" pageNumber="27">C</emphasis>
. 
<emphasis id="C4F96355F81C8D2FDD35FB12FA6D729E" box="[1407,1518,1189,1212]" italics="true" pageId="26" pageNumber="27">mexicanus</emphasis>
</taxonomicName>
[
<bibRefCitation id="921CC2B6F81C8D2FDA54FB70FDB472FD" author="Lacki MJ &amp; Dodd LE" box="[542,567,1223,1247]" pageId="26" pageNumber="27" pagination="39 - 52" refId="ref16724" refString="39. Lacki MJ, Dodd LE. Diet and foraging behavior of Corynorhinus in eastern North America. In: Loeb SC, Lacki MJ, Miller DA, editors. Conservation and management of eastern big-eared bats: a symposium. Asheville: Department of Agriculture Forest Service, Southern Research Station; 2011. pp. 39 - 52." type="book chapter" year="2011">39</bibRefCitation>
]. In 
<date id="82339987F81C8D2FDA20FB70FD5E72FD" box="[618,733,1223,1247]" pageId="26" pageNumber="27" value="2022-04">April 2022</date>
, 
<specimenCount id="E08B74CEF81C8D2FDAACFB70FCE572FD" box="[742,870,1223,1247]" count="12" pageId="26" pageNumber="27" type="adult">twelve adult</specimenCount>
females in an advanced state of pregnancy were captured, suggesting that births probably occur between the end of April and mid-May. All specimens captured in May and August presented a considerable number of ectoparasitic flies, presumably of the 
<taxonomicName id="318DC4C4F81C8D2FDA12FA98FCC47365" authorityName="Cockerell" authorityYear="1910" box="[600,839,1327,1351]" class="Insecta" family="Hippoboscidae" genus="Trichobius" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="corynorhini">
<emphasis id="C4F96355F81C8D2FDA12FA98FCC47365" box="[600,839,1327,1351]" italics="true" pageId="26" pageNumber="27">Trichobius corynorhini</emphasis>
</taxonomicName>
species.
</paragraph>
<paragraph id="F632BF47F81C8D2FDA7FFAE5FBCD70E7" blockId="26.[533,1536,807,1733]" pageId="26" pageNumber="27">
The acoustic characteristics of 
<taxonomicName id="318DC4C4F81C8D2FDB30FAE5FB2B7348" box="[890,1192,1362,1386]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="leonpaniaguae">
<emphasis id="C4F96355F81C8D2FDB30FAE5FB2B7348" box="[890,1192,1362,1386]" italics="true" pageId="26" pageNumber="27">Corynorhinus leonpaniaguae</emphasis>
</taxonomicName>
were obtained from ten specimens recorded using the hand-release technique and an Echo Meter Touch 2 Pro bat detector. Recordings were analyzed with the Batsound 
<emphasis id="C4F96355F81C8D2FDBBAFA20FB88738D" bold="true" box="[1008,1035,1431,1455]" pageId="26" pageNumber="27">1</emphasis>
software using Hanning window and 2048 of FFT size. Two sonotypes composed of modulated and harmonic pulses were observed from the recordings (
<figureCitation id="6EB6A3C2F81C8D2FDAF7FA6AFD7D73D7" box="[701,766,1501,1525]" captionStart="Fig 15" captionStartId="27.[533,561,1320,1339]" captionTargetBox="[544,1486,209,1252]" captionTargetId="figure-3@27.[533,1493,208,1298]" captionTargetPageId="27" captionText="Fig 15. Sonotypes depicting echolocation pulses of Corynorhinus leonpaniaguae. Sonotype I (upper panel) comprising combinations of two pulses, and sonotype II (lower panel) comprising single pulses of longer duration. https://doi.org/10.1371/journal.pone.0296275.g015" figureDoi="http://doi.org/10.5281/zenodo.14290660" httpUri="https://zenodo.org/record/14290660/files/figure.png" pageId="26" pageNumber="27">Fig 15</figureCitation>
). The first sonotype consisted of combinations of two pulses repeated serially with a longer time interval between combinations than between the pulses in the combination. Although two-pulse combinations were predominant, three-pulse combinations were observed in some specimens. The second sonotype was characterized by the absence of pulse combinations and the presence of modulated pulses of longer duration. Details of frequency, duration, and interval are presented in 
<tableCitation id="BB0F8AFCF81C8D2FDC46F93DFBD97080" box="[1036,1114,1674,1698]" captionStart="Table 5" captionStartId="27.[96,146,1481,1500]" captionTargetId="graphics-43@27.[96,1536,1509,1769]" captionTargetPageId="27" captionText="Table 5. Spectro-temporal characteristics of the echolocation pulses of Corynorhinus leonpaniaguae." httpUri="http://table.plazi.org/id/A2F2EFCFF81D8D2ED82AFA7EFC5773FE" pageId="26" pageNumber="27" tableUuid="A2F2EFCFF81D8D2ED82AFA7EFC5773FE">Table 5</tableCitation>
. Differences in acoustic characteristics among the species of 
<taxonomicName id="318DC4C4F81C8D2FDABDF91BFC0B70E6" authorityName="H. Allen" authorityYear="1865" box="[759,904,1708,1732]" class="Mammalia" family="Vespertilionidae" genus="Corynorhinus" kingdom="Animalia" order="Chiroptera" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="C4F96355F81C8D2FDABDF91BFC0B70E6" box="[759,904,1708,1732]" italics="true" pageId="26" pageNumber="27">Corynorhinus</emphasis>
</taxonomicName>
are still unknown.
</paragraph>
</subSubSection>
</treatment>
</document>