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<document id="244903B03DAAE1216041B9036EB657EF" ID-DOI="10.11646/zootaxa.5351.3.5" ID-GBIF-Dataset="abba17ad-72fa-494c-8c63-d0458e0dc7ee" ID-ISSN="1175-5326" ID-Zenodo-Dep="8391764" ID-ZooBank="48A49241-4DEB-443C-834F-D8D0579B5E23" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1695990834078" checkinUser="plazi" docAuthor="Praveenraj, Jayasimhan" docDate="2023" docId="7E1D580BFFEB9009FF7AEDF36ABA2CB3" docLanguage="en" docName="zootaxa.5351.3.5.pdf" docOrigin="Zootaxa 5351 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5351.3.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Badis limaakumi Praveenraj 2023, new species" docType="treatment" docVersion="2" lastPageNumber="376" masterDocId="82242073FFEA900CFFEDE93D6B6F2B44" masterDocTitle="Badis limaakumi, a new species of badid fish from Nagaland, Northeast India (Teleostei: Percomorpha: Badidae)" masterLastPageNumber="379" masterPageNumber="371" pageNumber="372" updateTime="1696254706693" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D113233ECC12792DA975C95127E0D904">Badis limaakumi, a new species of badid fish from Nagaland, Northeast India (Teleostei: Percomorpha: Badidae)</mods:title>
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<mods:namePart id="2E58B42045872266426555FDEA439879">Praveenraj, Jayasimhan</mods:namePart>
<mods:nameIdentifier id="72687898BE280B8F6EB943557978874C" type="ORCID">0000-0002-5471-6931</mods:nameIdentifier>
<mods:affiliation id="DBCC9D873D2D323F7D246F0A343B38F1">Indian Council of Agricultural Research-Central Island Agricultural Research Institute, Port Blair, Andaman and Nicobar Islands, India, 744 101.</mods:affiliation>
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<mods:date id="426E18E6831DADA0A4427BD561EE6DF8">2023</mods:date>
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<treatment id="7E1D580BFFEB9009FF7AEDF36ABA2CB3" ID-DOI="http://doi.org/10.5281/zenodo.8391782" ID-GBIF-Taxon="213625462" ID-Zenodo-Dep="8391782" LSID="urn:lsid:plazi:treatment:7E1D580BFFEB9009FF7AEDF36ABA2CB3" httpUri="http://treatment.plazi.org/id/7E1D580BFFEB9009FF7AEDF36ABA2CB3" lastPageId="5" lastPageNumber="376" pageId="1" pageNumber="372">
<subSubSection id="BEAEBA96FFEB900DFF7AEDF3696B2FAC" box="[151,516,1230,1257]" pageId="1" pageNumber="372" type="nomenclature">
<paragraph id="F60BE91DFFEB900DFF7AEDF3696B2FAC" blockId="1.[151,516,1230,1293]" box="[151,516,1230,1257]" pageId="1" pageNumber="372">
<heading id="AD435E71FFEB900DFF7AEDF3696B2FAC" bold="true" box="[151,516,1230,1257]" fontSize="11" level="1" pageId="1" pageNumber="372" reason="1">
<emphasis id="C4C0350FFFEB900DFF7AEDF3696B2FAC" bold="true" box="[151,516,1230,1257]" pageId="1" pageNumber="372">
<emphasis id="C4C0350FFFEB900DFF7AEDF36A022FAD" bold="true" box="[151,365,1230,1257]" italics="true" pageId="1" pageNumber="372">
<taxonomicName id="31B4929EFFEB900DFF7AEDF36A082FAD" authority="Praveenraj, 2023" authorityName="Praveenraj" authorityYear="2023" box="[151,359,1230,1257]" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="372" phylum="Chordata" rank="species" species="limaakumi" status="new species">Badis limaakumi</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="DFF38874FFEB900DFE99EDF2696B2FAC" box="[372,516,1230,1257]" pageId="1" pageNumber="372" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BEAEBA96FFEB900DFF7AEDCE6A142E49" box="[151,379,1267,1293]" pageId="1" pageNumber="372" type="description">
<paragraph id="F60BE91DFFEB900DFF7AEDCE6A142E49" blockId="1.[151,516,1230,1293]" box="[151,379,1267,1293]" pageId="1" pageNumber="372">
(
<figureCitation id="6E8FF598FFEB900DFF72EDCE6A652E49" box="[159,266,1267,1293]" captionStart="FIGURE 1" captionStartId="2.[152,255,1415,1439]" captionTargetBox="[169,1418,181,1391]" captionTargetId="figure-34@2.[169,1418,181,1391]" captionTargetPageId="2" captionText="FIGURE 1. Badis limaakumi (a) Holotype, APRC/ZSI/V/P-2083, male, 66.8 mm SL (b) Paratype, APRC/ZSI/V/P-2084, male, 42.1 mm SL (c) paratype, APRC/ZSI/V/P-2084, female, 45.3 mm SL." figureDoi="http://doi.org/10.5281/zenodo.8391766" httpUri="https://zenodo.org/record/8391766/files/figure.png" pageId="1" pageNumber="372">Figures 1</figureCitation>
,
<figureCitation id="6E8FF598FFEB900DFEF5EDCE6A472E49" box="[280,296,1267,1293]" captionStart="FIGURE 2" captionStartId="3.[152,255,1767,1791]" captionTargetBox="[151,1436,1158,1743]" captionTargetId="figure-130@3.[151,1436,1158,1743]" captionTargetPageId="3" captionText="FIGURE 2. Badis limaakumi, in life, holotype, APRC/ZSI/V/P-2083, 66.8 mm SL." figureDoi="http://doi.org/10.5281/zenodo.8391768" httpUri="https://zenodo.org/record/8391768/files/figure.png" pageId="1" pageNumber="372">2</figureCitation>
,
<figureCitation id="6E8FF598FFEB900DFED9EDCE6A2C2E49" box="[308,323,1267,1293]" captionStart="FIGURE 3" captionStartId="4.[152,255,1993,2017]" captionTargetBox="[303,1284,1245,1970]" captionTargetId="figure-579@4.[303,1284,1245,1970]" captionTargetPageId="4" captionText="FIGURE 3. (A) Badis assamensis, CIARI/FF-93, 49.5 mm SL (B) Badis blosyrus, CIARI/FF-95, 34.6 mm SL." figureDoi="http://doi.org/10.5281/zenodo.8391770" httpUri="https://zenodo.org/record/8391770/files/figure.png" pageId="1" pageNumber="372">3</figureCitation>
,
<figureCitation id="6E8FF598FFEB900DFEBDEDCE6A1C2E49" box="[336,371,1267,1293]" captionStart="FIGURE 4" captionStartId="5.[152,255,1732,1756]" captionTargetBox="[453,1135,181,1709]" captionTargetId="figure-15@5.[453,1135,181,1709]" captionTargetPageId="5" captionText="FIGURE 4. Details of head of (A) Badis limaakumi, holotype, APRC/ZSI/V/P-2083, 66.8 mm SL (B) Badis assamensis, CIARI/FF-93, 49.5 mm SL, and (C) Badis blosyrus, CIARI/FF-95, 34.6 mm SL." figureDoi="http://doi.org/10.5281/zenodo.8391772" httpUri="https://zenodo.org/record/8391772/files/figure.png" pageId="1" pageNumber="372">4A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="BEAEBA96FFEB900DFF7AEC06686E2E85" pageId="1" pageNumber="372" type="materials_examined">
<paragraph id="F60BE91DFFEB900DFF7AEC0668292E3D" blockId="1.[151,1437,1339,1905]" pageId="1" pageNumber="372">
<materialsCitation id="46DCE340FFEB900DFF7AEC0668292E3D" ID-GBIF-Occurrence="4417459303" collectingDate="2022-02-13" collectionCode="APRC, V" collectorName="Limaakum" country="India" county="Mokochung District" latitude="26.443783" location="Milak River" longLatPrecision="1" longitude="94.4848" municipality="Milak River" pageId="1" pageNumber="372" specimenCode="APRC/ZSI/V/P-2083" specimenCount="1" stateProvince="Nagaland" typeStatus="holotype">
<emphasis id="C4C0350FFFEB900DFF7AEC066A642E11" bold="true" box="[151,267,1339,1365]" pageId="1" pageNumber="372">
<typeStatus id="290F57BFFFEB900DFF7AEC066A692E11" box="[151,262,1339,1365]" pageId="1" pageNumber="372">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="A6124166FFEB900DFEF6EC0669622E11" box="[283,525,1339,1365]" pageId="1" pageNumber="372">APRC/ZSI/V/P-2083</specimenCode>
,
<specimenCount id="E0B22294FFEB900DFDCFEC0669342E11" box="[546,603,1339,1365]" pageId="1" pageNumber="372" type="male">male</specimenCount>
, 67.0 mm SL;
<collectingCountry id="8EA3A98DFFEB900DFCC4EC06680A2E11" box="[809,869,1339,1365]" name="India" pageId="1" pageNumber="372">India</collectingCountry>
,
<collectingRegion id="347027FFFFEB900DFC97EC0668872E11" box="[890,1000,1339,1365]" country="India" name="Nagaland" pageId="1" pageNumber="372">Nagaland</collectingRegion>
,
<collectingCounty id="1F6A9191FFEB900DFC13EC066F822E11" box="[1022,1261,1339,1365]" pageId="1" pageNumber="372">Mokochung District</collectingCounty>
,
<collectingMunicipality id="166F7367FFEB900DFAEFEC066EF72E11" box="[1282,1432,1339,1365]" pageId="1" pageNumber="372">Milak River</collectingMunicipality>
;
<geoCoordinate id="93808FDAFFEB900DFF7AEC626A562E3D" box="[151,313,1375,1401]" degrees="26" direction="north" minutes="26.6270" orientation="latitude" pageId="1" pageNumber="372" precision="1" value="26.443783">26°26.6270N</geoCoordinate>
,
<geoCoordinate id="93808FDAFFEB900DFEA8EC626A8A2E3D" box="[325,485,1375,1401]" degrees="94" direction="east" minutes="29.0880" orientation="longitude" pageId="1" pageNumber="372" precision="1" value="94.4848">94°29.0880E</geoCoordinate>
;
<collectorName id="5B418CCBFFEB900DFE1DEC6269032E3D" box="[496,620,1375,1401]" pageId="1" pageNumber="372">Limaakum</collectorName>
,
<date id="820ACFDDFFEB900DFD95EC62682C2E3D" box="[632,835,1375,1401]" pageId="1" pageNumber="372" value="2022-02-13">
<collectingDate id="924E3635FFEB900DFD95EC62682C2E3D" box="[632,835,1375,1401]" pageId="1" pageNumber="372" value="2022-02-13">13 February 2022</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="F60BE91DFFEB900DFF2AECBE686E2E85" blockId="1.[151,1437,1339,1905]" pageId="1" pageNumber="372">
<materialsCitation id="46DCE340FFEB900DFF2AECBE68852ED9" ID-GBIF-Occurrence="4417459302" box="[199,1002,1410,1437]" collectingDate="2022-02-13" collectionCode="APRC" collectorName="Limaakum" country="India" county="Mokochung District" latitude="26.443783" location="Limaakum" longLatPrecision="1" longitude="94.4848" municipality="Milak River" pageId="1" pageNumber="372" specimenCode="APRC/ZSI/V/P-2084, CIARI/FF-92" specimenCount="9" specimenCount-female="4" specimenCount-male="4" stateProvince="Nagaland" typeStatus="paratype">
<emphasis id="C4C0350FFFEB900DFF2AECBE6A572ED9" bold="true" box="[199,312,1411,1437]" pageId="1" pageNumber="372">
<typeStatus id="290F57BFFFEB900DFF2AECBE6A5C2ED9" box="[199,307,1411,1437]" pageId="1" pageNumber="372">Paratype</typeStatus>
.
</emphasis>
<specimenCode id="A6124166FFEB900DFED0ECBE69472ED9" box="[317,552,1411,1437]" pageId="1" pageNumber="372">APRC/ZSI/V/P-2084</specimenCode>
,
<quantity id="314C44F8FFEB900DFDDFECBF69A12ED9" box="[562,718,1410,1437]" metricMagnitude="-2" metricUnit="m" metricValue="4.91" metricValueMax="5.61" metricValueMin="4.21" pageId="1" pageNumber="372" unit="mm" value="49.1" valueMax="56.1" valueMin="42.1">42.156.1 mm</quantity>
SL (
<specimenCount id="E0B22294FFEB900DFCECECBE68252ED9" box="[769,842,1411,1437]" pageId="1" pageNumber="372" type="male">4 male</specimenCount>
and
<specimenCount id="E0B22294FFEB900DFC91ECBE68B12ED9" box="[892,990,1411,1437]" pageId="1" pageNumber="372" type="female">4 female</specimenCount>
);
</materialsCitation>
<materialsCitation id="46DCE340FFEB900DFC02ECBE69922E85" ID-GBIF-Occurrence="4417459302" collectingDate="2022-02-13" collectionCode="APRC" collectorName="Limaakum" country="India" county="Mokochung District" latitude="26.443783" location="Limaakum" longLatPrecision="1" longitude="94.4848" municipality="Milak River" pageId="1" pageNumber="372" specimenCode="APRC/ZSI/V/P-2084, CIARI/FF-92" specimenCount="9" specimenCount-female="4" specimenCount-male="4" stateProvince="Nagaland" typeStatus="paratype">
<specimenCode id="A6124166FFEB900DFC02ECBE6FE92ED9" box="[1007,1158,1411,1437]" pageId="1" pageNumber="372">CIARI/FF-92</specimenCode>
,
<specimenCount id="E0B22294FFEB900DFB7DECBE6FAA2ED9" box="[1168,1221,1411,1437]" pageId="1" pageNumber="372" type="generic">1 ex.</specimenCount>
,
<specimenCount id="E0B22294FFEB900DFB3DECBE6E682ED9" box="[1232,1287,1411,1437]" pageId="1" pageNumber="372">male</specimenCount>
,
<quantity id="314C44F8FFEB900DFAFCECBF6E1F2ED9" box="[1297,1392,1410,1437]" metricMagnitude="-2" metricUnit="m" metricValue="5.18" pageId="1" pageNumber="372" unit="mm" value="51.8">51.8 mm</quantity>
SL, cleared and stained for osteology, same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BEAEBA96FFEB900DFF2AECF76F522C35" pageId="1" pageNumber="372" type="diagnosis">
<paragraph id="F60BE91DFFEB900DFF2AECF76E762C41" blockId="1.[151,1437,1339,1905]" pageId="1" pageNumber="372">
<emphasis id="C4C0350FFFEB900DFF2AECF76A512EA0" bold="true" box="[199,318,1482,1508]" pageId="1" pageNumber="372">Diagnosis.</emphasis>
<taxonomicName id="31B4929EFFEB900DFEAAECF6696B2EA1" authorityName="Praveenraj" authorityYear="2023" box="[327,516,1483,1509]" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="372" phylum="Chordata" rank="species" species="limaakumi">
<emphasis id="C4C0350FFFEB900DFEAAECF6696B2EA1" box="[327,516,1483,1509]" italics="true" pageId="1" pageNumber="372">Badis limaakumi</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="31B4929EFFEB900DFD5AECF668C72EA1" authority="SG" authorityName="SG" box="[695,936,1483,1509]" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="372" phylum="Chordata" rank="species" species="assamensis">
<emphasis id="C4C0350FFFEB900DFD5AECF668142EA0" box="[695,891,1483,1509]" italics="true" pageId="1" pageNumber="372">Badis assamensis</emphasis>
SG
</taxonomicName>
and is thus distinguished from all members of the other species groups by the presence of the opercular blotch, absence of the cleithral botch and a relatively large size.
<taxonomicName id="31B4929EFFEB900DFEE3EF2E6AA62D69" authorityName="Praveenraj" authorityYear="2023" box="[270,457,1555,1581]" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="372" phylum="Chordata" rank="species" species="limaakumi">
<emphasis id="C4C0350FFFEB900DFEE3EF2E6AA62D69" box="[270,457,1555,1581]" italics="true" pageId="1" pageNumber="372">Badis limaakumi</emphasis>
</taxonomicName>
is distinguished from other members of the
<taxonomicName id="31B4929EFFEB900DFC58EF2E6F112D69" authority="SG" authorityName="SG" box="[949,1150,1555,1581]" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="372" phylum="Chordata" rank="species" species="assamensis">
<emphasis id="C4C0350FFFEB900DFC58EF2E6F3F2D68" box="[949,1104,1555,1580]" italics="true" pageId="1" pageNumber="372">B. assamensis</emphasis>
SG
</taxonomicName>
,
<emphasis id="C4C0350FFFEB900DFB67EF2E6E222D68" box="[1162,1357,1555,1580]" italics="true" pageId="1" pageNumber="372">
i.e.
<taxonomicName id="31B4929EFFEB900DFB5EEF2E6E222D68" authority="SG" authorityName="SG" box="[1203,1357,1555,1580]" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="372" phylum="Chordata" rank="species" species="assamensis">B. assamensis</taxonomicName>
</emphasis>
and
<emphasis id="C4C0350FFFEB900DFA68EF2E6B942D15" italics="true" pageId="1" pageNumber="372">
<taxonomicName id="31B4929EFFEB900DFA68EF2E6B982D15" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="372" phylum="Chordata" rank="species" species="blosyrus">B. blosyrus</taxonomicName>
,
</emphasis>
by the absence of irregular blotches along sides of the body (vs. presence of two rows of irregular blackish blotches on sides) (
<figureCitation id="6E8FF598FFEB900DFE80EF666ADF2D31" box="[365,432,1627,1653]" captionStart="FIGURE 3" captionStartId="4.[152,255,1993,2017]" captionTargetBox="[303,1284,1245,1970]" captionTargetId="figure-579@4.[303,1284,1245,1970]" captionTargetPageId="4" captionText="FIGURE 3. (A) Badis assamensis, CIARI/FF-93, 49.5 mm SL (B) Badis blosyrus, CIARI/FF-95, 34.6 mm SL." figureDoi="http://doi.org/10.5281/zenodo.8391770" httpUri="https://zenodo.org/record/8391770/files/figure.png" pageId="1" pageNumber="372">Fig. 3</figureCitation>
), presence of 10 pairs of distinct vertical bars on body (vs. absence of vertical bars), snout rounded in lateral view (vs. pointed) (
<figureCitation id="6E8FF598FFEB900DFDD8EF4269182DDD" box="[565,631,1663,1689]" captionStart="FIGURE 4" captionStartId="5.[152,255,1732,1756]" captionTargetBox="[453,1135,181,1709]" captionTargetId="figure-15@5.[453,1135,181,1709]" captionTargetPageId="5" captionText="FIGURE 4. Details of head of (A) Badis limaakumi, holotype, APRC/ZSI/V/P-2083, 66.8 mm SL (B) Badis assamensis, CIARI/FF-93, 49.5 mm SL, and (C) Badis blosyrus, CIARI/FF-95, 34.6 mm SL." figureDoi="http://doi.org/10.5281/zenodo.8391772" httpUri="https://zenodo.org/record/8391772/files/figure.png" pageId="1" pageNumber="372">Fig. 4</figureCitation>
) and a shallower body (body depth 23.228.0 vs. 29.934.9% SL).
<taxonomicName id="31B4929EFFEB900DFAB3EF426A642DF9" authorityName="Praveenraj" authorityYear="2023" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="372" phylum="Chordata" rank="species" species="limaakumi">
<emphasis id="C4C0350FFFEB900DFAB3EF426A642DF9" italics="true" pageId="1" pageNumber="372">Badis limaakumi</emphasis>
</taxonomicName>
is additionally distinguished from
<taxonomicName id="31B4929EFFEB900DFD7EEF9E687F2DF9" box="[659,784,1699,1725]" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="372" phylum="Chordata" rank="species" species="blosyrus">
<emphasis id="C4C0350FFFEB900DFD7EEF9E687F2DF9" box="[659,784,1699,1725]" italics="true" pageId="1" pageNumber="372">B. blosyrus</emphasis>
</taxonomicName>
by its larger body size (&gt;
<quantity id="314C44F8FFEB900DFBDCEF9E6FEE2DF9" box="[1073,1153,1699,1725]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="372" unit="mm" value="60.0">60 mm</quantity>
vs. &lt;
<quantity id="314C44F8FFEB900DFB2DEF9E6E602DF9" box="[1216,1295,1699,1725]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="372" unit="mm" value="40.0">40 mm</quantity>
SL), shorter upper (8.310.2 vs. 12.013.6% SL) and lower jaw, (11.915.0 vs. 16.318.5% SL), more lateral-line scales (2931 vs. 2728), and absence of a prominent angle at lower jaw articulation (vs. presence of prominent angle).
</paragraph>
<paragraph id="F60BE91DFFEB900DFF2AEE326F522C35" blockId="1.[151,1437,1339,1905]" pageId="1" pageNumber="372">
Additional characters distinguishing
<taxonomicName id="31B4929EFFEB900DFD84EE3268472C6D" authorityName="Praveenraj" authorityYear="2023" box="[617,808,1807,1833]" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="372" phylum="Chordata" rank="species" species="limaakumi">
<emphasis id="C4C0350FFFEB900DFD84EE3268472C6D" box="[617,808,1807,1833]" italics="true" pageId="1" pageNumber="372">Badis limaakumi</emphasis>
</taxonomicName>
from members of the other SGs including
<taxonomicName id="31B4929EFFEB900DFAFAEE326BD22C09" authority="SG" authorityName="SG" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="372" phylum="Chordata" rank="species" species="badis">
<emphasis id="C4C0350FFFEB900DFAFAEE326EF22C6D" box="[1303,1437,1807,1833]" italics="true" pageId="1" pageNumber="372">Badis badis</emphasis>
SG
</taxonomicName>
,
<taxonomicName id="31B4929EFFEB900DFF21EE0E6AD92C09" authority="SG" authorityName="SG" box="[204,438,1843,1869]" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="372" phylum="Chordata" rank="species" species="corycaeus">
<emphasis id="C4C0350FFFEB900DFF21EE0E6AE82C08" box="[204,391,1843,1869]" italics="true" pageId="1" pageNumber="372">Badis corycaeus</emphasis>
SG
</taxonomicName>
,
<taxonomicName id="31B4929EFFEB900DFE28EE0E69132C09" authority="SG" authorityName="SG" box="[453,636,1843,1869]" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="372" phylum="Chordata" rank="species" species="ruber">
<emphasis id="C4C0350FFFEB900DFE28EE0E69232C09" box="[453,588,1843,1869]" italics="true" pageId="1" pageNumber="372">Badis ruber</emphasis>
SG
</taxonomicName>
,
<taxonomicName id="31B4929EFFEB900DFD66EE0E68152C09" authority="SG" authorityName="SG" box="[651,890,1843,1869]" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="372" phylum="Chordata" rank="species" species="autumnum">
<emphasis id="C4C0350FFFEB900DFD66EE0E68252C08" box="[651,842,1843,1869]" italics="true" pageId="1" pageNumber="372">Badis autumnum</emphasis>
SG
</taxonomicName>
, and other congeners from Northeast
<collectingCountry id="8EA3A98DFFEB900DFADDEE0E6E032C09" box="[1328,1388,1843,1869]" name="India" pageId="1" pageNumber="372">India</collectingCountry>
, the Western Ghats,
<collectingCountry id="8EA3A98DFFEB900DFEA4EE6A6AA02C35" box="[329,463,1879,1905]" name="Bangladesh" pageId="1" pageNumber="372">Bangladesh</collectingCountry>
,
<collectingCountry id="8EA3A98DFFEB900DFE34EE6A69292C35" box="[473,582,1879,1905]" name="Myanmar" pageId="1" pageNumber="372">Myanmar</collectingCountry>
and
<collectingCountry id="8EA3A98DFFEB900DFD91EE6A698F2C35" box="[636,736,1879,1905]" name="Thailand" pageId="1" pageNumber="372">Thailand</collectingCountry>
are provided in the discussion.
</paragraph>
</subSubSection>
<caption id="A2CBB995FFE8900EFF75ECBA68332E87" ID-DOI="http://doi.org/10.5281/zenodo.8391766" ID-Zenodo-Dep="8391766" httpUri="https://zenodo.org/record/8391766/files/figure.png" pageId="2" pageNumber="373" startId="2.[152,255,1415,1439]" targetBox="[169,1418,181,1391]" targetPageId="2" targetType="figure">
<paragraph id="F60BE91DFFE8900EFF75ECBA68332E87" blockId="2.[151,1436,1415,1475]" pageId="2" pageNumber="373">
<emphasis id="C4C0350FFFE8900EFF75ECBA6A7D2EDB" bold="true" box="[152,274,1415,1439]" pageId="2" pageNumber="373">FIGURE 1</emphasis>
.
<taxonomicName id="31B4929EFFE8900EFEF0ECBA6AA82EDB" authorityName="Praveenraj" authorityYear="2023" box="[285,455,1415,1439]" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="373" phylum="Chordata" rank="species" species="limaakumi">
<emphasis id="C4C0350FFFE8900EFEF0ECBA6AA82EDB" box="[285,455,1415,1439]" italics="true" pageId="2" pageNumber="373">Badis limaakumi</emphasis>
</taxonomicName>
(a) Holotype, APRC/ZSI/V/P-2083, male, 66.8 mm SL (b) Paratype, APRC/ZSI/V/P-2084, male, 42.1 mm SL (c) paratype, APRC/ZSI/V/P-2084, female, 45.3 mm SL.
</paragraph>
</caption>
<subSubSection id="BEAEBA96FFE89008FF7AEF376A682F81" lastPageId="4" lastPageNumber="375" pageId="2" pageNumber="373" type="description">
<paragraph id="F60BE91DFFE8900EFF7AEF376A472D60" blockId="2.[151,296,1546,1572]" box="[151,296,1546,1572]" pageId="2" pageNumber="373">
<heading id="AD435E71FFE8900EFF7AEF376A472D60" bold="true" box="[151,296,1546,1572]" fontSize="11" level="1" pageId="2" pageNumber="373" reason="1">
<emphasis id="C4C0350FFFE8900EFF7AEF376A472D60" bold="true" box="[151,296,1546,1572]" pageId="2" pageNumber="373">Description</emphasis>
</heading>
</paragraph>
<paragraph id="F60BE91DFFE8900EFF7AEF706F732CC3" blockId="2.[151,1437,1613,2036]" pageId="2" pageNumber="373">
General appearance as in
<figureCitation id="6E8FF598FFE8900EFE2DEF7069612D23" box="[448,526,1613,1639]" captionStart="FIGURE 1" captionStartId="2.[152,255,1415,1439]" captionTargetBox="[169,1418,181,1391]" captionTargetId="figure-34@2.[169,1418,181,1391]" captionTargetPageId="2" captionText="FIGURE 1. Badis limaakumi (a) Holotype, APRC/ZSI/V/P-2083, male, 66.8 mm SL (b) Paratype, APRC/ZSI/V/P-2084, male, 42.1 mm SL (c) paratype, APRC/ZSI/V/P-2084, female, 45.3 mm SL." figureDoi="http://doi.org/10.5281/zenodo.8391766" httpUri="https://zenodo.org/record/8391766/files/figure.png" pageId="2" pageNumber="373">Figs. 1</figureCitation>
and
<figureCitation id="6E8FF598FFE8900EFDA4EF7069362D23" box="[585,601,1613,1639]" captionStart="FIGURE 2" captionStartId="3.[152,255,1767,1791]" captionTargetBox="[151,1436,1158,1743]" captionTargetId="figure-130@3.[151,1436,1158,1743]" captionTargetPageId="3" captionText="FIGURE 2. Badis limaakumi, in life, holotype, APRC/ZSI/V/P-2083, 66.8 mm SL." figureDoi="http://doi.org/10.5281/zenodo.8391768" httpUri="https://zenodo.org/record/8391768/files/figure.png" pageId="2" pageNumber="373">2</figureCitation>
, and morphometric data provided in
<tableCitation id="BB36DCA6FFE8900EFBE9EF706F352D23" box="[1028,1114,1613,1639]" captionStart="TABLE 1" captionStartId="3.[152,249,150,176]" captionTargetPageId="3" captionText="TABLE 1. Morphometric data of Badis limaakumi (n=10) type series" httpUri="http://table.plazi.org/id/A2CBB995FFE9900FFF75E9AB68C82BF5" pageId="2" pageNumber="373" tableUuid="A2CBB995FFE9900FFF75E9AB68C82BF5">Table 1</tableCitation>
. Body elongate, moderately compressed laterally. In males, predorsal profile ascending in a straight line from tip of snout up to vertical through posterior region of eye, gently convex more posteriorly, and almost straight to base of caudal fin. Ventral contour descending gently until below preopercle, almost straight more posteriorly to anal-fin origin, and gently ascending to base of caudal fin. Eye situated midlaterally in anterior half of head. Snout shorter than eye diameter, rounded in lateral view. Jaws almost equal anteriorly, with lower jaw slightly projecting; maxilla reaching vertical through anterior margin of orbit, lower jaw articulation below middle of orbit. Opercular spine slender, with sharp tip. In females, predorsal and ventral profile slightly convex, and maximum size less than
<quantity id="314C44F8FFE8900EFBC2EE746F122C20" box="[1071,1149,1865,1892]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="373" unit="mm" value="50.0">50 mm</quantity>
SL. Adult males are more elongate than females. Gravid females appear plumper with distended abdomen.
</paragraph>
<paragraph id="F60BE91DFFE8900EFF2AEEAF6FFC2CB7" blockId="2.[151,1437,1613,2036]" pageId="2" pageNumber="373">Dentary pores 4 (d1d2), anguloarticular pores 2 (aa1aa2), preopercular pores 6 (p1p6), nasal pores 2 (n1 n2), supraorbital pores 4 (f1f4), extrascapular pores 3 (ex1ex3), posttemporal pores 2 (po1po2), coronalis pore 1 (cor), lachrymal pores 3 (l1l3), infraorbital pores 4 (io1io4), pterotic pores 3 (pt1pt3).</paragraph>
<caption id="A2CBB995FFE9900FFF75E9AB68C82BF5" ID-Table-UUID="A2CBB995FFE9900FFF75E9AB68C82BF5" box="[152,935,150,177]" httpUri="http://table.plazi.org/id/A2CBB995FFE9900FFF75E9AB68C82BF5" pageId="3" pageNumber="374" startId="3.[152,249,150,176]" targetBox="[163,1391,192,1113]" targetIsTable="true" targetPageId="3" targetType="table">
<paragraph id="F60BE91DFFE9900FFF75E9AB68C82BF5" blockId="3.[152,935,150,177]" box="[152,935,150,177]" pageId="3" pageNumber="374">
<emphasis id="C4C0350FFFE9900FFF75E9AB6A7A2BF4" bold="true" box="[152,277,150,176]" pageId="3" pageNumber="374">TABLE 1.</emphasis>
Morphometric data of
<taxonomicName id="31B4929EFFE9900FFDF4E9AA69BA2BF5" authorityName="Praveenraj" authorityYear="2023" box="[537,725,151,177]" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="374" phylum="Chordata" rank="species" species="limaakumi">
<emphasis id="C4C0350FFFE9900FFDF4E9AA69BA2BF5" box="[537,725,151,177]" italics="true" pageId="3" pageNumber="374">Badis limaakumi</emphasis>
</taxonomicName>
(n=10) type series
</paragraph>
</caption>
<paragraph id="F60BE91DFFE9900FFF0FE9F26E062F1D" pageId="3" pageNumber="374">
<table id="84B41BBDFFE96FF3FF4EE9FD6E002F1D" box="[163,1391,192,1113]" gridcols="4" gridrows="14" pageId="3" pageNumber="374">
<tr id="4884EB5FFFE96FF3FF4EE9FD6E002BB1" box="[163,1391,192,245]" gridrow="0" pageId="3" pageNumber="374">
<th id="0B558223FFE96FF3FF4EE9FD6A952BB1" box="[163,506,192,245]" gridcol="0" gridrow="0" pageId="3" pageNumber="374">
<emphasis id="C4C0350FFFE9900FFF0FE9F26A952BA3" bold="true" box="[226,506,207,231]" pageId="3" pageNumber="374">Morphometric characters</emphasis>
</th>
<th id="0B558223FFE96FF3FDBDE9FD69DC2BB1" box="[592,691,192,245]" gridcol="1" gridrow="0" pageId="3" pageNumber="374">
<emphasis id="C4C0350FFFE9900FFDBDE9FD69DC2B9C" bold="true" box="[592,691,192,216]" pageId="3" pageNumber="374">Holotype</emphasis>
</th>
<th id="0B558223FFE96FF3FCE9E9FD6FD32BB1" box="[772,1212,192,245]" gridcol="2" gridrow="0" pageId="3" pageNumber="374">
<emphasis id="C4C0350FFFE9900FFCE9E9FD6F6C2BB1" bold="true" pageId="3" pageNumber="374">Range (includes holotype and paratypes) (n=10)</emphasis>
</th>
<th id="0B558223FFE96FF3FB12E9FD6E002BB1" box="[1279,1391,192,245]" gridcol="3" gridrow="0" pageId="3" pageNumber="374">
<emphasis id="C4C0350FFFE9900FFB12E9FD6E002B9C" bold="true" box="[1279,1391,192,216]" pageId="3" pageNumber="374">mean± SD</emphasis>
</th>
</tr>
<tr id="4884EB5FFFE96FF3FF4EE83B6E002A5A" box="[163,1391,262,286]" gridrow="1" pageId="3" pageNumber="374">
<th id="0B558223FFE96FF3FF4EE83B6A952A5A" box="[163,506,262,286]" gridcol="0" gridrow="1" pageId="3" pageNumber="374">Standard length (mm)</th>
<td id="0B558223FFE96FF3FDBDE83B69DC2A5A" box="[592,691,262,286]" gridcol="1" gridrow="1" pageId="3" pageNumber="374">67.0</td>
<td id="0B558223FFE96FF3FCE9E83B6FD32A5A" box="[772,1212,262,286]" gridcol="2" gridrow="1" pageId="3" pageNumber="374">42.167.0</td>
<td id="0B558223FFE96FF3FB12E83B6E002A5A" box="[1279,1391,262,286]" gridcol="3" gridrow="1" pageId="3" pageNumber="374">49.5</td>
</tr>
<tr id="4884EB5FFFE96FF3FF4EE87F6E002A1E" box="[163,1391,322,346]" gridrow="2" pageId="3" pageNumber="374">
<th id="0B558223FFE96FF3FF4EE87F6E002A1E" box="[163,1391,322,346]" colspan="4" colspanRight="3" gridcol="0" gridrow="2" pageId="3" pageNumber="374">
<emphasis id="C4C0350FFFE9900FFF4EE87F6AAB2A1E" bold="true" box="[163,452,322,346]" pageId="3" pageNumber="374">In percent standard length</emphasis>
</th>
</tr>
<tr id="4884EB5FFFE96FF3FF4EE8406E002AD1" box="[163,1391,381,405]" gridrow="3" pageId="3" pageNumber="374">
<th id="0B558223FFE96FF3FF4EE8406A952AD1" box="[163,506,381,405]" gridcol="0" gridrow="3" pageId="3" pageNumber="374">Head length</th>
<td id="0B558223FFE96FF3FDBDE84069DC2AD1" box="[592,691,381,405]" gridcol="1" gridrow="3" pageId="3" pageNumber="374">27.6</td>
<td id="0B558223FFE96FF3FCE9E8406FD32AD1" box="[772,1212,381,405]" gridcol="2" gridrow="3" pageId="3" pageNumber="374">27.031.7</td>
<td id="0B558223FFE96FF3FB12E8406E002AD1" box="[1279,1391,381,405]" gridcol="3" gridrow="3" pageId="3" pageNumber="374">26.2±6.7</td>
</tr>
<tr id="4884EB5FFFE96FF3FF4EE8FF6E002A9F" box="[163,1391,450,475]" gridrow="4" pageId="3" pageNumber="374">
<th id="0B558223FFE96FF3FF4EE8FF6A952A9F" box="[163,506,450,475]" gridcol="0" gridrow="4" pageId="3" pageNumber="374">Snout length</th>
<td id="0B558223FFE96FF3FDBDE8FF69DC2A9F" box="[592,691,450,475]" gridcol="1" gridrow="4" pageId="3" pageNumber="374">7.1</td>
<td id="0B558223FFE96FF3FCE9E8FF6FD32A9F" box="[772,1212,450,475]" gridcol="2" gridrow="4" pageId="3" pageNumber="374">5.9 7.4</td>
<td id="0B558223FFE96FF3FB12E8FF6E002A9F" box="[1279,1391,450,475]" gridcol="3" gridrow="4" pageId="3" pageNumber="374">6.5±0.5</td>
</tr>
<tr id="4884EB5FFFE96FF3FF4EEB3C6E00295E" box="[163,1391,513,538]" gridrow="5" pageId="3" pageNumber="374">
<th id="0B558223FFE96FF3FF4EEB3C6A95295E" box="[163,506,513,538]" gridcol="0" gridrow="5" pageId="3" pageNumber="374">Eye diameter</th>
<td id="0B558223FFE96FF3FDBDEB3C69DC295E" box="[592,691,513,538]" gridcol="1" gridrow="5" pageId="3" pageNumber="374">6.8</td>
<td id="0B558223FFE96FF3FCE9EB3C6FD3295E" box="[772,1212,513,538]" gridcol="2" gridrow="5" pageId="3" pageNumber="374">6.89.3</td>
<td id="0B558223FFE96FF3FB12EB3C6E00295E" box="[1279,1391,513,538]" gridcol="3" gridrow="5" pageId="3" pageNumber="374">8.3±0.8</td>
</tr>
<tr id="4884EB5FFFE96FF3FF4EEB036E002912" box="[163,1391,574,598]" gridrow="6" pageId="3" pageNumber="374">
<th id="0B558223FFE96FF3FF4EEB036A952912" box="[163,506,574,598]" gridcol="0" gridrow="6" pageId="3" pageNumber="374">Interorbital width</th>
<td id="0B558223FFE96FF3FDBDEB0369DC2912" box="[592,691,574,598]" gridcol="1" gridrow="6" pageId="3" pageNumber="374">5.4</td>
<td id="0B558223FFE96FF3FCE9EB036FD32912" box="[772,1212,574,598]" gridcol="2" gridrow="6" pageId="3" pageNumber="374">5.17.2</td>
<td id="0B558223FFE96FF3FB12EB036E002912" box="[1279,1391,574,598]" gridcol="3" gridrow="6" pageId="3" pageNumber="374">5.8±0.7</td>
</tr>
<tr id="4884EB5FFFE96FF3FF4EEB446E002982" box="[163,1391,633,710]" gridrow="7" pageId="3" pageNumber="374">
<th id="0B558223FFE96FF3FF4EEB446A952982" box="[163,506,633,710]" gridcol="0" gridrow="7" pageId="3" pageNumber="374">Upper-jaw length Lower-jaw length</th>
<td id="0B558223FFE96FF3FDBDEB4469DC2982" box="[592,691,633,710]" gridcol="1" gridrow="7" pageId="3" pageNumber="374">8.8 11.9</td>
<td id="0B558223FFE96FF3FCE9EB446FD32982" box="[772,1212,633,710]" gridcol="2" gridrow="7" pageId="3" pageNumber="374">8.310.2 11.915.0</td>
<td id="0B558223FFE96FF3FB12EB446E002982" box="[1279,1391,633,710]" gridcol="3" gridrow="7" pageId="3" pageNumber="374">9.4±0.5 13.2±1.0</td>
</tr>
<tr id="4884EB5FFFE96FF3FF4EEBD96E002863" box="[163,1391,740,807]" gridrow="8" pageId="3" pageNumber="374">
<th id="0B558223FFE96FF3FF4EEBD96A952863" box="[163,506,740,807]" gridcol="0" gridrow="8" pageId="3" pageNumber="374">Body depth Pelvic-fin length</th>
<td id="0B558223FFE96FF3FDBDEBD969DC2863" box="[592,691,740,807]" gridcol="1" gridrow="8" pageId="3" pageNumber="374">26.4 21.1</td>
<td id="0B558223FFE96FF3FCE9EBD96FD32863" box="[772,1212,740,807]" gridcol="2" gridrow="8" pageId="3" pageNumber="374">23.228.0 21.125.7</td>
<td id="0B558223FFE96FF3FB12EBD96E002863" box="[1279,1391,740,807]" gridcol="3" gridrow="8" pageId="3" pageNumber="374">25.8±1.6 23.2±1.4</td>
</tr>
<tr id="4884EB5FFFE96FF3FF4EEA6E6E002828" box="[163,1391,851,876]" gridrow="9" pageId="3" pageNumber="374">
<th id="0B558223FFE96FF3FF4EEA6E6A952828" box="[163,506,851,876]" gridcol="0" gridrow="9" pageId="3" pageNumber="374">Pelvic to anal-fin distance</th>
<td id="0B558223FFE96FF3FDBDEA6E69DC2828" box="[592,691,851,876]" gridcol="1" gridrow="9" pageId="3" pageNumber="374">35.1</td>
<td id="0B558223FFE96FF3FCE9EA6E6FD32828" box="[772,1212,851,876]" gridcol="2" gridrow="9" pageId="3" pageNumber="374">31.537.7</td>
<td id="0B558223FFE96FF3FB12EA6E6E002828" box="[1279,1391,851,876]" gridcol="3" gridrow="9" pageId="3" pageNumber="374">34.0±1.9</td>
</tr>
<tr id="4884EB5FFFE96FF3FF4EEAAD6E0028EC" box="[163,1391,912,936]" gridrow="10" pageId="3" pageNumber="374">
<th id="0B558223FFE96FF3FF4EEAAD6E0028EC" box="[163,1391,912,936]" colspan="4" colspanRight="3" gridcol="0" gridrow="10" pageId="3" pageNumber="374">
<emphasis id="C4C0350FFFE9900FFF4EEAAD6ADD28EC" bold="true" box="[163,434,912,936]" pageId="3" pageNumber="374">In percent of head length</emphasis>
</th>
</tr>
<tr id="4884EB5FFFE96FF3FF4EEAF66E0028A0" box="[163,1391,971,996]" gridrow="11" pageId="3" pageNumber="374">
<th id="0B558223FFE96FF3FF4EEAF66A9528A0" box="[163,506,971,996]" gridcol="0" gridrow="11" pageId="3" pageNumber="374">Snout length</th>
<td id="0B558223FFE96FF3FDBDEAF669DC28A0" box="[592,691,971,996]" gridcol="1" gridrow="11" pageId="3" pageNumber="374">25.5</td>
<td id="0B558223FFE96FF3FCE9EAF66FD328A0" box="[772,1212,971,996]" gridcol="2" gridrow="11" pageId="3" pageNumber="374">20.825.5</td>
<td id="0B558223FFE96FF3FB12EAF66E0028A0" box="[1279,1391,971,996]" gridcol="3" gridrow="11" pageId="3" pageNumber="374">23.0±1.2</td>
</tr>
<tr id="4884EB5FFFE96FF3FF4EED3B6E002F5B" box="[163,1391,1030,1055]" gridrow="12" pageId="3" pageNumber="374">
<th id="0B558223FFE96FF3FF4EED3B6A952F5B" box="[163,506,1030,1055]" gridcol="0" gridrow="12" pageId="3" pageNumber="374">Eye diameter</th>
<td id="0B558223FFE96FF3FDBDED3B69DC2F5B" box="[592,691,1030,1055]" gridcol="1" gridrow="12" pageId="3" pageNumber="374">24.8</td>
<td id="0B558223FFE96FF3FCE9ED3B6FD32F5B" box="[772,1212,1030,1055]" gridcol="2" gridrow="12" pageId="3" pageNumber="374">24.833.4</td>
<td id="0B558223FFE96FF3FB12ED3B6E002F5B" box="[1279,1391,1030,1055]" gridcol="3" gridrow="12" pageId="3" pageNumber="374">29.2± 2.9</td>
</tr>
<tr id="4884EB5FFFE96FF3FF4EED7C6E002F1D" box="[163,1391,1089,1113]" gridrow="13" pageId="3" pageNumber="374">
<th id="0B558223FFE96FF3FF4EED7C6A952F1D" box="[163,506,1089,1113]" gridcol="0" gridrow="13" pageId="3" pageNumber="374">Interorbital width</th>
<td id="0B558223FFE96FF3FDBDED7C69DC2F1D" box="[592,691,1089,1113]" gridcol="1" gridrow="13" pageId="3" pageNumber="374">19.7</td>
<td id="0B558223FFE96FF3FCE9ED7C6FD32F1D" box="[772,1212,1089,1113]" gridcol="2" gridrow="13" pageId="3" pageNumber="374">17.225.3</td>
<td id="0B558223FFE96FF3FB12ED7C6E002F1D" box="[1279,1391,1089,1113]" gridcol="3" gridrow="13" pageId="3" pageNumber="374">20.3± 2.4</td>
</tr>
</table>
</paragraph>
<caption id="A2CBB995FFE9900FFF75EFDA68822DBB" ID-DOI="http://doi.org/10.5281/zenodo.8391768" ID-Zenodo-Dep="8391768" box="[152,1005,1767,1791]" httpUri="https://zenodo.org/record/8391768/files/figure.png" pageId="3" pageNumber="374" startId="3.[152,255,1767,1791]" targetBox="[151,1436,1158,1743]" targetPageId="3" targetType="figure">
<paragraph id="F60BE91DFFE9900FFF75EFDA68822DBB" blockId="3.[152,1005,1767,1791]" box="[152,1005,1767,1791]" pageId="3" pageNumber="374">
<emphasis id="C4C0350FFFE9900FFF75EFDA6A7D2DBB" bold="true" box="[152,274,1767,1791]" pageId="3" pageNumber="374">FIGURE 2</emphasis>
.
<emphasis id="C4C0350FFFE9900FFEF2EFDA6ABF2DBB" box="[287,464,1767,1791]" italics="true" pageId="3" pageNumber="374">
<taxonomicName id="31B4929EFFE9900FFEF2EFDA6AA22DBB" authorityName="Praveenraj" authorityYear="2023" box="[287,461,1767,1791]" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="374" phylum="Chordata" rank="species" species="limaakumi">Badis limaakumi</taxonomicName>
,
</emphasis>
in life, holotype, APRC/ZSI/V/P-2083, 66.8 mm SL.
</paragraph>
</caption>
<paragraph id="F60BE91DFFE99008FF2AEE106F502925" blockId="3.[151,1436,1837,2044]" lastBlockId="4.[151,1437,150,1221]" lastPageId="4" lastPageNumber="375" pageId="3" pageNumber="374">
Scales ctenoid on sides, cycloid on top of head. Predorsal scales anterior to coronalis pore 3 (7) or 4* (3), posteriorly 7* (7) or 8 (3). Cheek and opercular scales ctenoid, cheek scales extending posteriorly from termination of gape. Scales rows on cheek 4 (1), 5 (6) or 6* (3), opercle with 3 rows of scales, 1 row each on preopercle, subopercle and interopercle. Scales in lateral row 30 (6) or 31* (4). Lateral-line canal on body in pored-scales, arranged into two segments. Upper lateral-line canal starting above operculum with 24 (1), 25 (3) and 26* (6) pored scales, lower lateral-line canal starting at one scale anterior to vertical through posterior base of anal fin with 4 (1), 5* (8) and 6 (1) pored scales. Circumpeduncular scales 20, with 9 (10) scales above, 9 (10) below lateral line. Dorsal-fin scale cover 24 scales wide; anal-fin scales cover 3 scales wide. Vertical scale rows 2 (2), 2½ (2), 3 (4), 3½* (2) above, and 7 (3), 7½ (4), 8(2), 8½*(1) below lateral line. Dorsal fin with 17* (8) or 18 (2) spines and 9 (10) soft rays, its origin vertically above pectoral-fin origin, and ending at two scales beyond vertical through base of last anal-fin ray. Dorsal- and anal-fin lappets with rounded tips. Posterior tips of dorsal fin reaching to anterior third of caudal fin when adpressed. Anal fin with 3* (10) spines and 6* (6) or 7 (4) soft rays, reaching base of caudal fin when adpressed. Caudal fin rounded, with 14 principal rays. Pectoral fin rounded, with 13 (5), 14 (1) or 15
<superScript id="01C14455FFEE9008FADFE8536E552A38" attach="left" box="[1330,1338,366,380]" fontSize="6" pageId="4" pageNumber="375">*</superScript>
(4) rays, extending to about 1/4 of distance to anal-fin origin when adpressed. Pelvic fin pointed with 1 spine and 5 (10) rays. Gill rakers on first gill arch 7 (3). Branchiostegal rays 6 (10). Vertebrae
<quantity id="314C44F8FFEE9008FC46E88A688C2A95" box="[939,995,439,465]" metricMagnitude="-1" metricUnit="m" metricValue="7.62" pageId="4" pageNumber="375" unit="in" value="30.0">30 in</quantity>
total, with 17 abdominal and 13 caudal. Dorsal fin with 1
<superScript id="01C14455FFEE9008FEBAE8E66A0D2AAD" attach="left" box="[343,354,475,489]" fontSize="6" pageId="4" pageNumber="375">st</superScript>
pterygiophore inserted between neural spines of 2
<superScript id="01C14455FFEE9008FC76E8E768C42AAC" attach="left" box="[923,939,474,488]" fontSize="6" pageId="4" pageNumber="375">nd</superScript>
and 3
<superScript id="01C14455FFEE9008FC1DE8E768912AAC" attach="left" box="[1008,1022,474,488]" fontSize="6" pageId="4" pageNumber="375">rd</superScript>
vertebrae. Anal fin 1
<superScript id="01C14455FFEE9008FB00E8E66F972AAD" attach="left" box="[1261,1272,475,489]" fontSize="6" pageId="4" pageNumber="375">st</superScript>
pterygiophore inserted between haemal spines of 17
<superScript id="01C14455FFEE9008FDD9E8C3692E2948" attach="left" box="[564,577,510,524]" fontSize="6" pageId="4" pageNumber="375">th</superScript>
and 18
<superScript id="01C14455FFEE9008FD7CE8C369F12948" attach="left" box="[657,670,510,524]" fontSize="6" pageId="4" pageNumber="375">th</superScript>
vertebrae. Both jaws with tiny unicuspid teeth arranged in wide band of about four rows anteriorly in each jaw and two rows posteriorly. Anterior-most teeth larger than others. Vomer, adjacent palatine, parasphenoid, and hypobranchial 3 toothed; basihyal not toothed.
</paragraph>
<paragraph id="F60BE91DFFEE9008FF2AEB576E6429B5" blockId="4.[151,1437,150,1221]" pageId="4" pageNumber="375">
<emphasis id="C4C0350FFFEE9008FF2AEB576A8F29C1" bold="true" box="[199,480,618,645]" pageId="4" pageNumber="375">Colouration in ethanol.</emphasis>
Body pale grey, becoming darker towards dorsal-fin base and scaled portion of finrays, interradial fin membranes grey with tinges of black. Bars on body faint. Anal fin dark-grey. Caudal fin greyish, scaled portion dark grey. Caudal blotch black. Pectoral and pelvic fin pale grey. Preorbital and post orbital stripe greyish. Cheek greyish, preopercle and opercle dark grey. Opercular blotch black. Lower jaw greyish.
</paragraph>
<paragraph id="F60BE91DFFEE9008FF2AEBC76A682F81" blockId="4.[151,1437,150,1221]" pageId="4" pageNumber="375">
<emphasis id="C4C0350FFFEE9008FF2AEBC76AC82850" bold="true" box="[199,423,762,788]" pageId="4" pageNumber="375">Colouration in life.</emphasis>
Ground color of body brownish-red, head olive-green or greyish, with black scales on nape. Preorbital stripe dark-grey, continuing to posterior region of lower jaw. Postorbital stripe with black or bluish scales. No supraorbital or suborbital stripe. Opercle with bluish iridescent blotch at base of opercular spine, two scales high and three scales wide. Cleithral blotch absent. Body with 10 pairs of distinct black vertical bars: 6 pairs between head and anal-fin origin, 2 pairs between anal-fin origin and posterior insertion of it, and remaining 2 between posterior of anal-fin origin and caudal fin. Bars 15, 9 and 10, broad and straight; bars 68 slightly curved. Bar 10 on caudal peduncle composed of a medial bluish caudal spot, which usually elongate and extending across last scale of side and base of caudal fin. Bars becoming darker towards dorsal-fin base, and continuing onto scaled portion of dorsal fin. Dorsal fin anterior portion begins with broad, brownish submargin, and narrow, orange and pale-blue dorsal margin continued up to spinous portion only; posterior soft end of dorsal fin hyaline, its dorsal margin bluish. Anal fin hyaline, spines greyish with white margins. Pectoral fin hyaline. Pelvic fin hyaline, interradial membranes and basal margin greyish. Caudal fin hyaline, yellowish at scaled portion, posterodorsal and posteroventral margins pale-blue.
</paragraph>
</subSubSection>
<caption id="A2CBB995FFEE9008FF75EEF46E682CA6" ID-DOI="http://doi.org/10.5281/zenodo.8391770" ID-Zenodo-Dep="8391770" box="[152,1287,1993,2018]" httpUri="https://zenodo.org/record/8391770/files/figure.png" pageId="4" pageNumber="375" startId="4.[152,255,1993,2017]" targetBox="[303,1284,1245,1970]" targetPageId="4" targetType="figure">
<paragraph id="F60BE91DFFEE9008FF75EEF46E682CA6" blockId="4.[152,1287,1993,2018]" box="[152,1287,1993,2018]" pageId="4" pageNumber="375">
<emphasis id="C4C0350FFFEE9008FF75EEF46A772CA5" bold="true" box="[152,280,1993,2017]" pageId="4" pageNumber="375">FIGURE 3.</emphasis>
(A)
<taxonomicName id="31B4929EFFEE9008FEA5EEF769212CA6" authority=", CIARI" authorityName="CIARI" box="[328,590,1994,2018]" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="375" phylum="Chordata" rank="species" species="assamensis">
<emphasis id="C4C0350FFFEE9008FEA5EEF76A952CA5" box="[328,506,1994,2018]" italics="true" pageId="4" pageNumber="375">Badis assamensis</emphasis>
, CIARI
</taxonomicName>
/FF-93, 49.5 mm SL (B)
<taxonomicName id="31B4929EFFEE9008FCA4EEF76F5C2CA5" authority=", CIARI" authorityName="CIARI" box="[841,1075,1993,2018]" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="375" phylum="Chordata" rank="species" species="blosyrus">
<emphasis id="C4C0350FFFEE9008FCA4EEF768B02CA6" box="[841,991,1994,2018]" italics="true" pageId="4" pageNumber="375">Badis blosyrus</emphasis>
, CIARI
</taxonomicName>
/FF-95, 34.6 mm SL.
</paragraph>
</caption>
<caption id="A2CBB995FFEF9009FF75EFF968A52C45" ID-DOI="http://doi.org/10.5281/zenodo.8391772" ID-Zenodo-Dep="8391772" httpUri="https://zenodo.org/record/8391772/files/figure.png" pageId="5" pageNumber="376" startId="5.[152,255,1732,1756]" targetBox="[453,1135,181,1709]" targetPageId="5" targetType="figure">
<paragraph id="F60BE91DFFEF9009FF75EFF968A52C45" blockId="5.[151,1436,1732,1793]" pageId="5" pageNumber="376">
<emphasis id="C4C0350FFFEF9009FF75EFF96A742D98" bold="true" box="[152,283,1732,1756]" pageId="5" pageNumber="376">FIGURE 4.</emphasis>
Details of head of (A)
<emphasis id="C4C0350FFFEF9009FDF7EFF869A02D99" box="[538,719,1733,1757]" italics="true" pageId="5" pageNumber="376">
<taxonomicName id="31B4929EFFEF9009FDF7EFF869A42D99" authorityName="Praveenraj" authorityYear="2023" box="[538,715,1733,1757]" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="limaakumi">Badis limaakumi</taxonomicName>
,
</emphasis>
holotype, APRC/ZSI/V/P-2083, 66.8 mm SL (B)
<taxonomicName id="31B4929EFFEF9009FB0CEFF86BB12C45" authority=", CIARI" authorityName="CIARI" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="assamensis">
<emphasis id="C4C0350FFFEF9009FB0CEFF86EF92D98" box="[1249,1430,1733,1757]" italics="true" pageId="5" pageNumber="376">Badis assamensis</emphasis>
, CIARI
</taxonomicName>
/FF-93, 49.5 mm SL, and (C)
<taxonomicName id="31B4929EFFEF9009FDE6EFD4699A2C44" authority=", CIARI" authorityName="CIARI" box="[523,757,1768,1793]" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="blosyrus">
<emphasis id="C4C0350FFFEF9009FDE6EFD469CE2C45" box="[523,673,1769,1793]" italics="true" pageId="5" pageNumber="376">Badis blosyrus</emphasis>
, CIARI
</taxonomicName>
/FF-95, 34.6 mm SL.
</paragraph>
</caption>
<subSubSection id="BEAEBA96FFEF9009FF2AEE14697D2CF4" pageId="5" pageNumber="376" type="distribution">
<paragraph id="F60BE91DFFEF9009FF2AEE14697D2CF4" blockId="5.[151,1437,1833,2040]" pageId="5" pageNumber="376">
<emphasis id="C4C0350FFFEF9009FF2AEE146A982C07" bold="true" box="[199,503,1833,1860]" pageId="5" pageNumber="376">Distribution and habitat.</emphasis>
<taxonomicName id="31B4929EFFEF9009FDEDEE1469D02C07" authorityName="Praveenraj" authorityYear="2023" box="[512,703,1833,1859]" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="limaakumi">
<emphasis id="C4C0350FFFEF9009FDEDEE1469D02C07" box="[512,703,1833,1859]" italics="true" pageId="5" pageNumber="376">Badis limaakumi</emphasis>
</taxonomicName>
is presently known only from the Milak River in Mokokchung district,
<collectingRegion id="347027FFFFEF9009FF1BEE706A0B2C23" box="[246,356,1869,1895]" country="India" name="Nagaland" pageId="5" pageNumber="376">Nagaland</collectingRegion>
, Northeast
<collectingCountry id="8EA3A98DFFEF9009FE04EE7069492C23" box="[489,550,1869,1895]" name="India" pageId="5" pageNumber="376">India</collectingCountry>
. The
<typeStatus id="290F57BFFFEF9009FD84EE7369F62C2C" box="[617,665,1870,1896]" pageId="5" pageNumber="376">type</typeStatus>
locality during
<date id="820ACFDDFFEF9009FCB9EE7368822C2C" box="[852,1005,1870,1896]" pageId="5" pageNumber="376" value="2022-01">January 2022</date>
had a depth of
<quantity id="314C44F8FFEF9009FB44EE706F812C2C" box="[1193,1262,1869,1896]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="376" unit="m" value="0.5">0.5 m</quantity>
, with substrate comprising numerous boulders and rocks (
<figureCitation id="6E8FF598FFEF9009FD86EE4C69C12CC8" box="[619,686,1905,1932]" captionStart="FIGURE 5" captionStartId="6.[152,255,840,864]" captionTargetBox="[370,1217,181,815]" captionTargetId="figure-34@6.[370,1217,181,815]" captionTargetPageId="6" captionText="FIGURE 5. Type locality of Badis limaakumi, Milak River at Mokochung District, Nagaland, India." figureDoi="http://doi.org/10.5281/zenodo.8391774" httpUri="https://zenodo.org/record/8391774/files/figure.png" pageId="5" pageNumber="376">Fig. 5</figureCitation>
). Co-occurring species include
<taxonomicName id="31B4929EFFEF9009FBE4EE4F6FD22CCF" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[1033,1213,1905,1931]" family="Cyprinidae" genus="Esomus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="danrica">
<emphasis id="C4C0350FFFEF9009FBE4EE4F6FD22CCF" box="[1033,1213,1905,1931]" italics="true" pageId="5" pageNumber="376">Esomus danrica</emphasis>
</taxonomicName>
,
<taxonomicName id="31B4929EFFEF9009FB24EE4C6EFA2CCF" authorityName="Nath &amp; Dey" authorityYear="1989" box="[1225,1429,1905,1931]" family="Amblycipitidae" genus="Amblyceps" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="apangi">
<emphasis id="C4C0350FFFEF9009FB24EE4C6EFA2CCF" box="[1225,1429,1905,1931]" italics="true" pageId="5" pageNumber="376">Amblyceps apangi</emphasis>
</taxonomicName>
,
<taxonomicName id="31B4929EFFEF9009FF7AEEAB6A6A2CF4" box="[151,261,1942,1968]" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="C4C0350FFFEF9009FF7AEEAB6BB22CF4" box="[151,221,1942,1968]" italics="true" pageId="5" pageNumber="376">Garra</emphasis>
<emphasis id="C4C0350FFFEF9009FF09EEAB6A6A2CF4" bold="true" box="[228,261,1942,1968]" pageId="5" pageNumber="376">sp.</emphasis>
</taxonomicName>
, and
<taxonomicName id="31B4929EFFEF9009FEAEEEA869642CF4" box="[323,523,1941,1968]" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="C4C0350FFFEF9009FEAEEEA86A8C2CEB" box="[323,483,1941,1967]" italics="true" pageId="5" pageNumber="376">Psilorhynchus</emphasis>
<emphasis id="C4C0350FFFEF9009FE07EEAB69642CF4" bold="true" box="[490,523,1942,1968]" pageId="5" pageNumber="376">sp.</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="BEAEBA96FFEF9009FF2AEE876ABA2CB3" pageId="5" pageNumber="376" type="etymology">
<paragraph id="F60BE91DFFEF9009FF2AEE876ABA2CB3" blockId="5.[151,1437,1833,2040]" pageId="5" pageNumber="376">
<emphasis id="C4C0350FFFEF9009FF2AEE876A212C90" bold="true" box="[199,334,1978,2004]" pageId="5" pageNumber="376">Etymology.</emphasis>
The species name honours Limaakum, Assistant Professor at the Fazl Ali College,
<collectingRegion id="347027FFFFEF9009FB1EEE846E0E2C97" box="[1267,1377,1977,2003]" country="India" name="Nagaland" pageId="5" pageNumber="376">Nagaland</collectingRegion>
, who discovered this new species.
</paragraph>
</subSubSection>
</treatment>
</document>