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<document ID-DOI="10.3897/zookeys.36.306" ID-GBIF-Dataset="4db6b327-7482-432e-a5f6-36f91c79fef3" ID-ISSN="13132970" ID-Zenodo-Dep="576620" ID-ZooBank="ADCACC88-6C78-4386-8E33-3F98234ECE92" approvalRequired="105" approvalRequired_for_textStreams="105" checkinTime="1587967053581" checkinUser="plazi" docAuthor="Rix, Michael &amp; Harvey, Mark" docDate="2010" docId="7E13878EFFEF1B21FF321D7EFE2E8FF1" docLanguage="en" docName="ZK_article_2154.pdf" docOrigin="ZooKeys 36 (36)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@6ed9075d" docStyleName="zookeys.2008.journal_article" docTitle="Olgania Hickman 1979" docType="treatment" docVersion="5" lastPageNumber="91" masterDocId="822AFFF6FFB61B7BFFBE1F57FFE98D05" masterDocTitle="The spider family Micropholcommatidae (Arachnida: Araneae: Araneoidea): a relimitation and revision at the generic level" masterLastPageNumber="321" masterPageNumber="1" pageNumber="90" updateTime="1669157185164" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:title>The spider family Micropholcommatidae (Arachnida: Araneae: Araneoidea): a relimitation and revision at the generic level</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rix, Michael</mods:namePart>
<mods:affiliation>Western Australian Museum, Welshpool DC, Perth, Australia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Harvey, Mark</mods:namePart>
<mods:affiliation>Western Australian Museum, Perth, Welshpool, Australia</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2010</mods:date>
<mods:detail type="pubDate">
<mods:number>2010-02-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>36</mods:number>
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<mods:detail type="issue">
<mods:number>36</mods:number>
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<mods:start>1</mods:start>
<mods:end>321</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.36.306</mods:identifier>
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<mods:identifier type="ZooBank">ADCACC88-6C78-4386-8E33-3F98234ECE92</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3789468" ID-GBIF-Taxon="163748995" ID-Zenodo-Dep="3789468" LSID="urn:lsid:plazi:treatment:7E13878EFFEF1B21FF321D7EFE2E8FF1" httpUri="http://treatment.plazi.org/id/7E13878EFFEF1B21FF321D7EFE2E8FF1" lastPageId="90" lastPageNumber="91" pageId="89" pageNumber="90">
<subSubSection box="[140,515,553,580]" pageId="89" pageNumber="90" type="nomenclature">
<paragraph blockId="89.[140,515,553,580]" box="[140,515,553,580]" pageId="89" pageNumber="90">
<heading box="[140,515,553,580]" fontSize="11" level="2" pageId="89" pageNumber="90" reason="3">
<emphasis bold="true" box="[140,515,553,580]" pageId="89" pageNumber="90">
Genus
<taxonomicName authority="Hickman, 1979" authorityName="Hickman" authorityYear="1979" box="[223,515,553,580]" class="Arachnida" family="Micropholcommatidae" genus="Olgania" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[223,320,553,580]" italics="true" pageId="89" pageNumber="90">Olgania</emphasis>
<bibRefCitation author="Hickman VV" box="[327,515,553,579]" pageId="89" pageNumber="90" pagination="53 - 79" refId="ref70870" refString="Hickman VV (1979) Some Tasmanian spiders of the families Oonopidae, Anapidae and Mysmenidae. Papers and Proceedings of the Royal Society of Tasmania 113: 53 - 79." type="journal article" year="1979">Hickman, 1979</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="89" pageNumber="90" type="reference_group">
<paragraph blockId="89.[140,1107,624,721]" pageId="89" pageNumber="90">
<treatmentCitationGroup pageId="89" pageNumber="90">
<treatmentCitation author="Hickman VV" box="[140,474,624,650]" page="71" pageId="89" pageNumber="90" year="1979">
<taxonomicName authority="Hickman, 1979: 71" authorityName="Hickman" authorityPageNumber="71" authorityYear="1979" box="[140,474,624,650]" class="Arachnida" family="Micropholcommatidae" genus="Olgania" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis box="[140,228,624,650]" italics="true" pageId="89" pageNumber="90">Olgania</emphasis>
<bibRefCitation author="Hickman VV" box="[239,474,624,650]" pageId="89" pageNumber="90" pagination="53 - 79" refId="ref70870" refString="Hickman VV (1979) Some Tasmanian spiders of the families Oonopidae, Anapidae and Mysmenidae. Papers and Proceedings of the Royal Society of Tasmania 113: 53 - 79." type="journal article" year="1979">Hickman, 1979: 71</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus box="[487,542,625,651]" pageId="89" pageNumber="90">Type</typeStatus>
species by original designation
<taxonomicName authority="Hickman, 1979" authorityName="Hickman P" authorityYear="1979" class="Arachnida" family="Micropholcommatidae" genus="Olgania" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="excavata">
<emphasis box="[916,1107,624,650]" italics="true" pageId="89" pageNumber="90">Olgania excavata</emphasis>
<bibRefCitation author="Hickman VV" box="[185,372,659,686]" pageId="89" pageNumber="90" pagination="53 - 79" refId="ref70870" refString="Hickman VV (1979) Some Tasmanian spiders of the families Oonopidae, Anapidae and Mysmenidae. Papers and Proceedings of the Royal Society of Tasmania 113: 53 - 79." type="journal article" year="1979">Hickman, 1979</bibRefCitation>
</taxonomicName>
.
<treatmentCitation author="Brignoli PM" box="[386,620,659,686]" page="372" pageId="89" pageNumber="90" year="1983">
<bibRefCitation author="Brignoli PM" box="[386,620,659,686]" pageId="89" pageNumber="90" refId="ref69575" refString="Brignoli PM (1983) A Catalogue of the Araneae Described Between 1940 and 1981. Manchester University Press, Manchester, 755 pp." type="book" year="1983">Brignoli, 1983: 372</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation author="Davies VT" box="[633,836,659,686]" page="55" pageId="89" pageNumber="90" year="1985">
<bibRefCitation author="Davies VT" box="[633,836,659,686]" pageId="89" pageNumber="90" pagination="48 - 125" refId="ref69989" refString="Davies VT (1985) Araneomorphae (in part). In: Walton DW (Ed) Zoological Catalogue of Australia, Vol. 3. Mygalomorphae, Araneomorphae (in part), Pseudoscorpionida, Amblypygi and Palpigradi. Australian Government Publishing Service, Canberra, 48 - 125." type="book chapter" year="1985">Davies, 1985: 55</bibRefCitation>
</treatmentCitation>
. Transferred from
<taxonomicName authorityName="Simon" authorityYear="1895" class="Arachnida" family="Anapidae" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="90" phylum="Arthropoda" rank="family">Anapidae</taxonomicName>
to
<taxonomicName box="[284,544,695,721]" class="Arachnida" family="Micropholcommatidae" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="90" phylum="Arthropoda" rank="family">Micropholcommatidae</taxonomicName>
by
<treatmentCitation author="Platnick NI &amp; Forster RR" box="[586,915,695,721]" page="76" pageId="89" pageNumber="90" year="1989">
<bibRefCitation author="Platnick NI &amp; Forster RR" box="[586,915,695,721]" pageId="89" pageNumber="90" pagination="1 - 139" refId="ref71971" refString="Platnick NI, Forster RR (1989) A revision of the temperate South American and Australasian spiders of the family Anapidae (Araneae, Araneoidea). Bulletin of the American Museum of Natural History 190: 1 - 139." type="journal article" year="1989">Platnick &amp; Forster, 1989: 76</bibRefCitation>
</treatmentCitation>
.
<bibRefCitation author="Platnick NI" box="[926,1096,695,721]" pageId="89" pageNumber="90" refId="ref71855" refString="Platnick NI (2009) The World Spider Catalog, Version 10.0. American Museum of Natural History, New York. http: // research. amnh. org / entomology / spiders / catalog / [accessed 20. X. 2009]" type="url" year="2009">Platnick, 2009</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="89.[140,1108,765,1671]" pageId="89" pageNumber="90">
<emphasis bold="true" box="[140,256,765,791]" pageId="89" pageNumber="90">
<emphasis box="[140,251,765,791]" inLineHeading="true" pageId="89" pageNumber="90" reason="1">Affinities</emphasis>
.
</emphasis>
The genus
<taxonomicName box="[390,478,765,791]" class="Arachnida" family="Micropholcommatidae" genus="Olgania" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis box="[390,478,765,791]" italics="true" pageId="89" pageNumber="90">Olgania</emphasis>
</taxonomicName>
appears to be the sister-lineage to
<taxonomicName authorityName="Simon" authorityYear="1880" box="[878,982,765,791]" class="Arachnida" family="Micropholcommatidae" genus="Taphiassa" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis box="[878,982,765,791]" italics="true" pageId="89" pageNumber="90">Taphiassa</emphasis>
</taxonomicName>
from Australasia (
<figureCitation box="[234,302,800,826]" captionStart="Figure 4" captionStartId="137.[140,212,1307,1329]" captionTargetBox="[161,1087,180,1278]" captionTargetId="figure@137.[140,1107,165,1288]" captionTargetPageId="137" captionText="Figure 4. Optimal cladogram for the Micropholcommatidae under equal and implied weights, and one of six equally parsimonious trees recovered from an equal weights analysis of the new matrix (length = 135; see Appendix II). With the exception of the highlighted (*) node Patelliella adusta + Micropholcommatini, all clades shown were recovered in all iterations of an implied weights analysis (K=16); the phylogenetic position of P. adusta changed to P. adusta + Textricellini under K-values 15. Unambiguous character optimisations are shown for each node, and bold species are newly described, with generic type species highlighted (*)." figureDoi="http://doi.org/10.5281/zenodo.3767693" httpUri="https://zenodo.org/record/3767693/files/figure.png" pageId="89" pageNumber="90">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="89" pageNumber="90" type="diagnosis">
<paragraph blockId="89.[140,1108,765,1671]" pageId="89" pageNumber="90">
<emphasis bold="true" box="[185,312,835,861]" pageId="89" pageNumber="90">
<emphasis box="[185,307,835,861]" inLineHeading="true" pageId="89" pageNumber="90" reason="1">Diagnosis</emphasis>
.
</emphasis>
Species of
<taxonomicName box="[443,531,835,861]" class="Arachnida" family="Micropholcommatidae" genus="Olgania" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis box="[443,531,835,861]" italics="true" pageId="89" pageNumber="90">Olgania</emphasis>
</taxonomicName>
can be distinguished from species of
<taxonomicName authorityName="Simon" authorityYear="1880" box="[966,1070,835,861]" class="Arachnida" family="Micropholcommatidae" genus="Taphiassa" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis box="[966,1070,835,861]" italics="true" pageId="89" pageNumber="90">Taphiassa</emphasis>
</taxonomicName>
by the presence of six or fewer eyes (Figs 177C, 182C), and by the presence of a very large, wedge-shaped anterior sclerite which covers the entire anterior portion of the abdomen (Figs 176D, 177A). Other diagnostic characters include the presence of bulging anterior projections on the male chelicerae (Figs 188A, 190F), the presence of a ring of prominent sclerotic sigillae on the posterior abdomen (Fig. 177B), and the troglomorphic morphology of most species (Fig. 179).
</paragraph>
</subSubSection>
<subSubSection lastPageId="90" lastPageNumber="91" pageId="89" pageNumber="90" type="description">
<paragraph blockId="89.[140,1108,765,1671]" pageId="89" pageNumber="90">
<emphasis bold="true" box="[185,335,1081,1107]" pageId="89" pageNumber="90">
<emphasis box="[185,330,1081,1107]" inLineHeading="true" pageId="89" pageNumber="90" reason="1">Description</emphasis>
.
</emphasis>
Very small, entelegyne
<taxonomicName box="[615,746,1082,1108]" order="Null" pageId="89" pageNumber="132" rank="superFamily" superFamily="Araneoidea">Araneoidea</taxonomicName>
: total length 0.90 to 1.20.
<emphasis italics="true" pageId="89" pageNumber="90">Cephalothorax</emphasis>
: Carapace with glandular depressions above maxillae; cuticle of carapace and sternum heavily punctate, covered with glandular pits (Fig. 188); margins fused to sternum via pleural sclerites. Six (Fig. 177C) or no (Fig. 182C) eyes present on anterior margin of pars cephalica; AME absent. Chelicerae with or (rarely) without bulging anterior projections in males; promargin with two true teeth (Fig. 190B); fused setal sockets, peg teeth and ectal stridulatory ridges absent.
</paragraph>
<paragraph blockId="89.[140,1108,765,1671]" pageId="89" pageNumber="90">
<emphasis box="[185,459,1328,1354]" italics="true" pageId="89" pageNumber="90">Legs and female pedipalp</emphasis>
: Legs three-clawed (Fig. 191D), covered with smooth or serrate hair-like setae. Trichobothria present on legs; tibiae each with three (legs IIII) or four (legs IIIIV) trichobothria; metatarsi (legs IIII) each with single trichobothrium (Fig. 191C). Female pedipalp entire, five-segmented; claw absent (Fig. 190CD).
</paragraph>
<paragraph blockId="89.[140,1108,765,1671]" lastBlockId="90.[140,1108,167,756]" lastPageId="90" lastPageNumber="91" pageId="89" pageNumber="90">
<emphasis box="[185,287,1469,1495]" italics="true" pageId="89" pageNumber="90">Abdomen</emphasis>
: Abdomen subtriangular in lateral view, with very large, wedge-shaped anterior sclerite present around epigastric region and petiole, extending anteriorly and laterally to cover entire anterior portion of abdomen (Figs 176D, 177A); dorsal scute absent on males and females (Fig. 176AB); posterior abdomen thickened around dorsal and lateral margins of anterior sclerite, with ring of sclerotic sigillae (Fig. 177B) and sclerotic ring surrounding spinnerets and colulus. Six spinnerets situated posterior to fleshy colulus; ALS largest; PMS smallest; spigot morphology unknown. Anterior tracheal system well-developed, with multiple anteriorly-directed tracheae (Fig. 178A); posterior tracheal spiracle vestigial (Figs 189CD).
</paragraph>
<paragraph blockId="90.[140,1108,167,756]" pageId="90" pageNumber="91">
<emphasis box="[185,286,272,298]" italics="true" pageId="90" pageNumber="91">Genitalia</emphasis>
: Male pedipalp (Fig. 192) small to large; patella with retrolaterally-directed, hooked ligulate retrolateral apophysis and strongly recurved distal apophysis; tibia with or without pair of large prolateral macrosetae; tegulum smooth, with curved, marginal evaginated tegular ridge; embolus exposed, long (length&gt; 5× width), sometimes coiled. Female genitalia (Fig. 178) with pair of separate, globular anterior receptacula each with internal ducts; insemination ducts looped or coiled; fertilisation ducts simple, curved.
</paragraph>
</subSubSection>
<subSubSection pageId="90" pageNumber="91" type="distribution">
<paragraph blockId="90.[140,1108,167,756]" box="[185,796,483,510]" pageId="90" pageNumber="91">
<emphasis bold="true" box="[185,344,483,509]" pageId="90" pageNumber="91">
<emphasis box="[185,339,483,509]" inLineHeading="true" pageId="90" pageNumber="91" reason="1">Distribution</emphasis>
.
</emphasis>
Known only from
<collectingRegion box="[561,668,483,509]" country="Australia" name="Tasmania" pageId="90" pageNumber="91">Tasmania</collectingRegion>
(Fig. 213).
</paragraph>
<paragraph blockId="90.[140,1108,167,756]" pageId="90" pageNumber="91">
<emphasis bold="true" box="[185,351,518,544]" pageId="90" pageNumber="91">
<emphasis box="[185,346,518,544]" inLineHeading="true" pageId="90" pageNumber="91" reason="1">Composition</emphasis>
.
</emphasis>
<taxonomicName authority="Hickman, 1979" authorityName="Hickman P" authorityYear="1979" box="[358,732,519,545]" class="Arachnida" family="Micropholcommatidae" genus="Olgania" kingdom="Animalia" order="Araneae" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="excavata">
<emphasis box="[358,545,519,545]" italics="true" pageId="90" pageNumber="91">Olgania excavata</emphasis>
<bibRefCitation author="Hickman VV" box="[552,732,519,545]" pageId="90" pageNumber="91" pagination="53 - 79" refId="ref70870" refString="Hickman VV (1979) Some Tasmanian spiders of the families Oonopidae, Anapidae and Mysmenidae. Papers and Proceedings of the Royal Society of Tasmania 113: 53 - 79." type="journal article" year="1979">Hickman, 1979</bibRefCitation>
</taxonomicName>
and the four new species
<taxonomicName authorityName="Rix &amp; Harvey" authorityYear="2010" class="Arachnida" family="Micropholcommatidae" genus="Olgania" kingdom="Animalia" order="Araneae" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="cracroft">
<emphasis italics="true" pageId="90" pageNumber="91">O. cracroft</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Rix &amp; Harvey" authorityYear="2010" box="[192,330,554,580]" class="Arachnida" family="Micropholcommatidae" genus="Olgania" kingdom="Animalia" order="Araneae" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="eberhardi">
<emphasis box="[192,330,554,580]" italics="true" pageId="90" pageNumber="91">O. eberhardi</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Rix &amp; Harvey" authorityYear="2010" box="[345,492,554,580]" class="Arachnida" family="Micropholcommatidae" genus="Olgania" kingdom="Animalia" order="Araneae" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="troglodytes">
<emphasis box="[345,492,554,580]" italics="true" pageId="90" pageNumber="91">O. troglodytes</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Rix &amp; Harvey" authorityYear="2010" box="[549,635,554,580]" class="Arachnida" family="Micropholcommatidae" genus="Olgania" kingdom="Animalia" order="Araneae" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="weld">
<emphasis box="[549,635,554,580]" italics="true" pageId="90" pageNumber="91">O. weld</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph blockId="90.[140,1108,167,756]" pageId="90" pageNumber="91">
<emphasis bold="true" box="[185,430,589,615]" pageId="90" pageNumber="91">
<emphasis box="[185,318,589,615]" inLineHeading="true" pageId="90" pageNumber="91" reason="1">Taxonomic</emphasis>
remarks.
</emphasis>
The genus
<taxonomicName box="[564,652,589,615]" class="Arachnida" family="Micropholcommatidae" genus="Olgania" kingdom="Animalia" order="Araneae" pageId="90" pageNumber="91" phylum="Arthropoda" rank="genus">
<emphasis box="[564,652,589,615]" italics="true" pageId="90" pageNumber="91">Olgania</emphasis>
</taxonomicName>
was first described by
<bibRefCitation author="Hickman VV" box="[914,1107,589,615]" pageId="90" pageNumber="91" pagination="53 - 79" refId="ref70870" refString="Hickman VV (1979) Some Tasmanian spiders of the families Oonopidae, Anapidae and Mysmenidae. Papers and Proceedings of the Royal Society of Tasmania 113: 53 - 79." type="journal article" year="1979">Hickman (1979)</bibRefCitation>
in the family
<taxonomicName authorityName="Simon" authorityYear="1895" box="[289,399,624,650]" class="Arachnida" family="Anapidae" kingdom="Animalia" order="Araneae" pageId="90" pageNumber="91" phylum="Arthropoda" rank="family">Anapidae</taxonomicName>
, before being transferred to the
<taxonomicName box="[754,1014,624,650]" class="Arachnida" family="Micropholcommatidae" kingdom="Animalia" order="Araneae" pageId="90" pageNumber="91" phylum="Arthropoda" rank="family">Micropholcommatidae</taxonomicName>
by
<bibRefCitation author="Platnick NI &amp; Forster RR" pageId="90" pageNumber="91" pagination="1 - 139" refId="ref71971" refString="Platnick NI, Forster RR (1989) A revision of the temperate South American and Australasian spiders of the family Anapidae (Araneae, Araneoidea). Bulletin of the American Museum of Natural History 190: 1 - 139." type="journal article" year="1989">Platnick and Forster (1989: 76)</bibRefCitation>
. The close phylogenetic relationship between
<taxonomicName box="[970,1058,659,685]" class="Arachnida" family="Micropholcommatidae" genus="Olgania" kingdom="Animalia" order="Araneae" pageId="90" pageNumber="91" phylum="Arthropoda" rank="genus">
<emphasis box="[970,1058,659,685]" italics="true" pageId="90" pageNumber="91">Olgania</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Simon" authorityYear="1880" box="[140,244,695,721]" class="Arachnida" family="Micropholcommatidae" genus="Taphiassa" kingdom="Animalia" order="Araneae" pageId="90" pageNumber="91" phylum="Arthropoda" rank="genus">
<emphasis box="[140,244,695,721]" italics="true" pageId="90" pageNumber="91">Taphiassa</emphasis>
</taxonomicName>
was first noted by
<bibRefCitation author="Rix MG &amp; Harvey MS &amp; Roberts JD" box="[451,635,695,721]" pageId="90" pageNumber="91" pagination="1031 - 1048" refId="ref72243" refString="Rix MG, Harvey MS, Roberts JD (2008) Molecular phylogenetics of the spider family Micropholcommatidae (Arachnida: Araneae) using nuclear rRNA genes (18 S and 28 S). Molecular Phylogenetics and Evolution 46: 1031 - 1048." type="journal article" year="2008">Rix et al. (2008)</bibRefCitation>
, and the two genera now form the distinctive subfamily
<taxonomicName authorityName="Rix &amp; Harvey" authorityYear="2010" box="[304,451,730,756]" family="Null" pageId="90" pageNumber="91" rank="subFamily" subFamily="Taphiassinae">Taphiassinae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>