365 lines
40 KiB
XML
365 lines
40 KiB
XML
<document ID-DOI="10.3897/zookeys.36.306" ID-GBIF-Dataset="4db6b327-7482-432e-a5f6-36f91c79fef3" ID-ISSN="1313–2970" ID-Zenodo-Dep="576620" ID-ZooBank="ADCACC88-6C78-4386-8E33-3F98234ECE92" approvalRequired="105" approvalRequired_for_textStreams="105" checkinTime="1587967053581" checkinUser="plazi" docAuthor="Rix, Michael & Harvey, Mark" docDate="2010" docId="7E13878EFFA01B6CFF321E2EFDE28972" docLanguage="en" docName="ZK_article_2154.pdf" docOrigin="ZooKeys 36 (36)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@6ed9075d" docStyleName="zookeys.2008.journal_article" docTitle="Micropholcomma Crosby & Bishop 1927" docType="treatment" docVersion="5" lastPageNumber="24" masterDocId="822AFFF6FFB61B7BFFBE1F57FFE98D05" masterDocTitle="The spider family Micropholcommatidae (Arachnida: Araneae: Araneoidea): a relimitation and revision at the generic level" masterLastPageNumber="321" masterPageNumber="1" pageNumber="23" updateTime="1669157185164" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The spider family Micropholcommatidae (Arachnida: Araneae: Araneoidea): a relimitation and revision at the generic level</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Rix, Michael</mods:namePart>
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<mods:affiliation>Western Australian Museum, Welshpool DC, Perth, Australia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Harvey, Mark</mods:namePart>
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<mods:affiliation>Western Australian Museum, Perth, Welshpool, Australia</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2010</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2010-02-22</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>36</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>36</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>321</mods:end>
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</mods:extent>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.36.306</mods:identifier>
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<mods:identifier type="GBIF-Dataset">4db6b327-7482-432e-a5f6-36f91c79fef3</mods:identifier>
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<mods:identifier type="ISSN">1313–2970</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576620</mods:identifier>
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<mods:identifier type="ZooBank">ADCACC88-6C78-4386-8E33-3F98234ECE92</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3789411" ID-GBIF-Taxon="163749020" ID-Zenodo-Dep="3789411" LSID="urn:lsid:plazi:treatment:7E13878EFFA01B6CFF321E2EFDE28972" httpUri="http://treatment.plazi.org/id/7E13878EFFA01B6CFF321E2EFDE28972" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
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<subSubSection box="[140,722,377,404]" pageId="22" pageNumber="23" type="nomenclature">
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<paragraph blockId="22.[140,722,377,404]" box="[140,722,377,404]" pageId="22" pageNumber="23">
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<heading box="[140,722,377,404]" fontSize="11" level="2" pageId="22" pageNumber="23" reason="3">
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||
<emphasis bold="true" box="[140,722,377,404]" pageId="22" pageNumber="23">
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Genus
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<taxonomicName authority="Crosby & Bishop, 1927" authorityName="Crosby & Bishop" authorityYear="1927" box="[223,722,377,404]" class="Arachnida" family="Micropholcommatidae" genus="Micropholcomma" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
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<emphasis bold="true" box="[223,431,377,404]" italics="true" pageId="22" pageNumber="23">Micropholcomma</emphasis>
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<bibRefCitation author="Crosby CR & Bishop SC" box="[438,722,377,403]" pageId="22" pageNumber="23" pagination="147 - 154" refId="ref69920" refString="Crosby CR, Bishop SC (1927) New species of Erigoneae and Theridiidae. Journal of the New York Entomological Society 35: 147 - 154." type="journal article" year="1927">Crosby & Bishop, 1927</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection pageId="22" pageNumber="23" type="reference_group">
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<paragraph blockId="22.[140,1108,448,792]" pageId="22" pageNumber="23">
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<treatmentCitationGroup pageId="22" pageNumber="23">
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<treatmentCitation author="Crosby CR & Bishop SC" box="[140,680,448,475]" page="152" pageId="22" pageNumber="23" year="1927">
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<taxonomicName authority="Crosby & Bishop, 1927: 152" authorityName="Crosby & Bishop" authorityPageNumber="152" authorityYear="1927" box="[140,680,448,475]" class="Arachnida" family="Micropholcommatidae" genus="Micropholcomma" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
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||
<emphasis box="[140,329,448,474]" italics="true" pageId="22" pageNumber="23">Micropholcomma</emphasis>
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||
<bibRefCitation author="Crosby CR & Bishop SC" box="[338,680,448,475]" pageId="22" pageNumber="23" pagination="147 - 154" refId="ref69920" refString="Crosby CR, Bishop SC (1927) New species of Erigoneae and Theridiidae. Journal of the New York Entomological Society 35: 147 - 154." type="journal article" year="1927">Crosby & Bishop, 1927: 152</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
.
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||
<typeStatus box="[693,748,449,475]" pageId="22" pageNumber="23">Type</typeStatus>
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species by original designation
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<taxonomicName authority="Crosby & Bishop, 1927" authorityName="Crosby & Bishop" authorityYear="1927" box="[185,787,483,510]" class="Arachnida" family="Micropholcommatidae" genus="Micropholcomma" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="caeligenus">
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<emphasis box="[185,492,483,509]" italics="true" pageId="22" pageNumber="23">Micropholcomma caeligenus</emphasis>
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<bibRefCitation author="Crosby CR & Bishop SC" box="[502,787,483,510]" pageId="22" pageNumber="23" pagination="147 - 154" refId="ref69920" refString="Crosby CR, Bishop SC (1927) New species of Erigoneae and Theridiidae. Journal of the New York Entomological Society 35: 147 - 154." type="journal article" year="1927">Crosby & Bishop, 1927</bibRefCitation>
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</taxonomicName>
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||
.
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<treatmentCitation author="Roewer CF" box="[801,1027,484,510]" page="392" pageId="22" pageNumber="23" year="1942">
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<bibRefCitation author="Roewer CF" box="[801,1027,484,510]" pageId="22" pageNumber="23" refId="ref72288" refString="Roewer CF (1942) Katalog der Araneae von 1758 bis 1940. 1. Band (Mesothelae, Orthognatha, Lapidognatha: Dysderaeformia, Scytodiformia, Pholciformia, Zodariiformia, Hersiliaeformia, Argyopiformia). Paul Budy, Berlin, 1040 pp." type="book" year="1942">Roewer, 1942: 392</bibRefCitation>
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</treatmentCitation>
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.
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<treatmentCitation author="Hickman VV" page="183" pageId="22" pageNumber="23" year="1944">
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<bibRefCitation author="Hickman VV" pageId="22" pageNumber="23" pagination="179 - 195" refId="ref70811" refString="Hickman VV (1944) On some new Australian Apneumonomorphae with notes on their respiratory system. Papers and Proceedings of the Royal Society of Tasmania 1944: 179 - 195." type="journal article" year="1944">Hickman, 1944: 183</bibRefCitation>
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</treatmentCitation>
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.
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<treatmentCitation author="Bonnet P" box="[389,627,518,545]" page="2905" pageId="22" pageNumber="23" year="1957">
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<bibRefCitation author="Bonnet P" box="[389,627,518,545]" pageId="22" pageNumber="23" pagination="1927 - 3026" refId="ref69400" refString="Bonnet P (1957) Bibliographia Araneorum, Tome II (3 me partie: G- M). Douladoure, Toulouse, 1927 - 3026." type="book chapter" year="1957">Bonnet, 1957: 2905</bibRefCitation>
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</treatmentCitation>
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||
.
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||
<treatmentCitation author="Forster RR" box="[641,858,518,545]" page="297" pageId="22" pageNumber="23" year="1959">
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<bibRefCitation author="Forster RR" box="[641,858,518,545]" pageId="22" pageNumber="23" pagination="269 - 329" refId="ref70149" refString="Forster RR (1959) The spiders of the family Symphytognathidae. Transactions of the Royal Society of New Zealand 86: 269 - 329." type="journal article" year="1959">Forster, 1959: 297</bibRefCitation>
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</treatmentCitation>
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.
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<treatmentCitation author="Hickman VV" box="[871,1104,518,545]" page="65" pageId="22" pageNumber="23" year="1981">
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<bibRefCitation author="Hickman VV" box="[871,1104,518,545]" pageId="22" pageNumber="23" pagination="47 - 68" refId="ref70902" refString="Hickman VV (1981) New Tasmanian spiders of the families Archaeidae, Cycloctenidae, Amaurobiidae and Micropholcommatidae. Papers and Proceedings of the Royal Society of Tasmania 115: 47 - 68." type="journal article" year="1981">Hickman, 1981: 65</bibRefCitation>
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</treatmentCitation>
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.
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<treatmentCitation author="Brignoli PM" box="[185,413,554,580]" page="374" pageId="22" pageNumber="23" year="1983">
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<bibRefCitation author="Brignoli PM" box="[185,413,554,580]" pageId="22" pageNumber="23" refId="ref69575" refString="Brignoli PM (1983) A Catalogue of the Araneae Described Between 1940 and 1981. Manchester University Press, Manchester, 755 pp." type="book" year="1983">Brignoli, 1983: 374</bibRefCitation>
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</treatmentCitation>
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. Rix, 2008: 343.
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<bibRefCitation author="Platnick NI" box="[610,779,554,580]" pageId="22" pageNumber="23" refId="ref71855" refString="Platnick NI (2009) The World Spider Catalog, Version 10.0. American Museum of Natural History, New York. http: // research. amnh. org / entomology / spiders / catalog / [accessed 20. X. 2009]" type="url" year="2009">Platnick, 2009</bibRefCitation>
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.
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||
</treatmentCitationGroup>
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</paragraph>
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<paragraph blockId="22.[140,1108,448,792]" pageId="22" pageNumber="23">
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<treatmentCitationGroup pageId="22" pageNumber="23">
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||
<treatmentCitation author="Butler LSG" box="[140,513,589,615]" page="103" pageId="22" pageNumber="23" year="1932">
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||
<taxonomicName authority="Butler, 1932: 103" authorityName="Butler" authorityPageNumber="103" authorityYear="1932" box="[140,513,589,615]" class="Arachnida" family="Micropholcommatidae" genus="Microlinypheus" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
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<emphasis box="[140,305,589,615]" italics="true" pageId="22" pageNumber="23">Microlinypheus</emphasis>
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<bibRefCitation author="Butler LSG" box="[312,513,589,615]" pageId="22" pageNumber="23" pagination="103 - 117" refId="ref69719" refString="Butler LSG (1932) Studies in Australian spiders, No. 2. Proceedings of the Royal Society of Victoria 44: 103 - 117." type="journal article" year="1932">Butler, 1932: 103</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
.
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||
<typeStatus box="[523,578,590,616]" pageId="22" pageNumber="23">Type</typeStatus>
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||
species by monotypy
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||
<taxonomicName authority="Butler, 1932" authorityName="Butler" authorityYear="1932" class="Arachnida" family="Micropholcommatidae" genus="Microlinypheus" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="bryophilus">
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<emphasis box="[824,1107,589,615]" italics="true" pageId="22" pageNumber="23">Microlinypheus bryophilus</emphasis>
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||
<bibRefCitation author="Butler LSG" box="[185,330,624,650]" pageId="22" pageNumber="23" pagination="103 - 117" refId="ref69719" refString="Butler LSG (1932) Studies in Australian spiders, No. 2. Proceedings of the Royal Society of Victoria 44: 103 - 117." type="journal article" year="1932">Butler, 1932</bibRefCitation>
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||
</taxonomicName>
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||
.
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||
<treatmentCitation author="Roewer CF" box="[341,560,624,651]" page="617" pageId="22" pageNumber="23" year="1942">
|
||
<bibRefCitation author="Roewer CF" box="[341,560,624,651]" pageId="22" pageNumber="23" refId="ref72288" refString="Roewer CF (1942) Katalog der Araneae von 1758 bis 1940. 1. Band (Mesothelae, Orthognatha, Lapidognatha: Dysderaeformia, Scytodiformia, Pholciformia, Zodariiformia, Hersiliaeformia, Argyopiformia). Paul Budy, Berlin, 1040 pp." type="book" year="1942">Roewer, 1942: 617</bibRefCitation>
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||
</treatmentCitation>
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||
. Synonymised by
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||
<treatmentCitation author="Hickman VV" box="[764,1005,624,650]" page="186" pageId="22" pageNumber="23" year="1944">
|
||
<bibRefCitation author="Hickman VV" box="[764,1005,624,650]" pageId="22" pageNumber="23" pagination="179 - 195" refId="ref70811" refString="Hickman VV (1944) On some new Australian Apneumonomorphae with notes on their respiratory system. Papers and Proceedings of the Royal Society of Tasmania 1944: 179 - 195." type="journal article" year="1944">Hickman, 1944: 186</bibRefCitation>
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</treatmentCitation>
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||
.
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||
<treatmentCitation author="Bonnet P" page="2885" pageId="22" pageNumber="23" year="1957">
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||
<bibRefCitation author="Bonnet P" pageId="22" pageNumber="23" pagination="1927 - 3026" refId="ref69400" refString="Bonnet P (1957) Bibliographia Araneorum, Tome II (3 me partie: G- M). Douladoure, Toulouse, 1927 - 3026." type="book chapter" year="1957">Bonnet, 1957: 2885</bibRefCitation>
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||
</treatmentCitation>
|
||
.
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||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph blockId="22.[140,1108,448,792]" pageId="22" pageNumber="23">
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||
<treatmentCitationGroup pageId="22" pageNumber="23">
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||
<treatmentCitation author="Butler LSG" box="[140,477,695,721]" page="106" pageId="22" pageNumber="23" year="1932">
|
||
<taxonomicName authority="Butler, 1932: 106" authorityName="Butler" authorityPageNumber="106" authorityYear="1932" box="[140,477,695,721]" class="Arachnida" family="Micropholcommatidae" genus="Plectochetos" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
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||
<emphasis box="[140,265,695,721]" italics="true" pageId="22" pageNumber="23">Plectochetos</emphasis>
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||
<bibRefCitation author="Butler LSG" box="[273,477,695,721]" pageId="22" pageNumber="23" pagination="103 - 117" refId="ref69719" refString="Butler LSG (1932) Studies in Australian spiders, No. 2. Proceedings of the Royal Society of Victoria 44: 103 - 117." type="journal article" year="1932">Butler, 1932: 106</bibRefCitation>
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||
</taxonomicName>
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||
</treatmentCitation>
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||
.
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<typeStatus box="[488,543,695,721]" pageId="22" pageNumber="23">Type</typeStatus>
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species by monotypy
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||
<taxonomicName authority="Butler, 1932" authorityName="Butler" authorityYear="1932" class="Arachnida" family="Micropholcommatidae" genus="Plectochetos" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="longissimus">
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<emphasis box="[795,1048,695,721]" italics="true" pageId="22" pageNumber="23">Plectochetos longissimus</emphasis>
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||
<bibRefCitation author="Butler LSG" pageId="22" pageNumber="23" pagination="103 - 117" refId="ref69719" refString="Butler LSG (1932) Studies in Australian spiders, No. 2. Proceedings of the Royal Society of Victoria 44: 103 - 117." type="journal article" year="1932">Butler, 1932</bibRefCitation>
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||
</taxonomicName>
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||
.
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||
<treatmentCitation author="Roewer CF" box="[311,541,730,756]" page="617" pageId="22" pageNumber="23" year="1942">
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||
<bibRefCitation author="Roewer CF" box="[311,541,730,756]" pageId="22" pageNumber="23" refId="ref72288" refString="Roewer CF (1942) Katalog der Araneae von 1758 bis 1940. 1. Band (Mesothelae, Orthognatha, Lapidognatha: Dysderaeformia, Scytodiformia, Pholciformia, Zodariiformia, Hersiliaeformia, Argyopiformia). Paul Budy, Berlin, 1040 pp." type="book" year="1942">Roewer, 1942: 617</bibRefCitation>
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||
</treatmentCitation>
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||
.
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||
<treatmentCitation author="Bonnet P" box="[558,802,730,757]" page="3707" pageId="22" pageNumber="23" year="1958">
|
||
<bibRefCitation author="Bonnet P" box="[558,802,730,757]" pageId="22" pageNumber="23" pagination="3027 - 4230" refId="ref69427" refString="Bonnet P (1958) Bibliographia Araneorum, Tome II (4 me partie: N- S). Douladoure, Toulouse, 3027 - 4230." type="book chapter" year="1958">Bonnet, 1958: 3707</bibRefCitation>
|
||
</treatmentCitation>
|
||
. Synonymised by
|
||
<treatmentCitation author="Forster RR" page="297" pageId="22" pageNumber="23" year="1959">
|
||
<bibRefCitation author="Forster RR" pageId="22" pageNumber="23" pagination="269 - 329" refId="ref70149" refString="Forster RR (1959) The spiders of the family Symphytognathidae. Transactions of the Royal Society of New Zealand 86: 269 - 329." type="journal article" year="1959">Forster, 1959: 297</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph blockId="22.[140,1108,835,1672]" pageId="22" pageNumber="23">
|
||
<emphasis bold="true" box="[140,256,835,861]" pageId="22" pageNumber="23">
|
||
<emphasis box="[140,251,835,861]" inLineHeading="true" pageId="22" pageNumber="23" reason="1">Affinities</emphasis>
|
||
.
|
||
</emphasis>
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||
The genus
|
||
<taxonomicName authorityName="Crosby & Bishop" authorityYear="1927" box="[382,571,835,861]" class="Arachnida" family="Micropholcommatidae" genus="Micropholcomma" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
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||
<emphasis box="[382,571,835,861]" italics="true" pageId="22" pageNumber="23">Micropholcomma</emphasis>
|
||
</taxonomicName>
|
||
appears to be the sister-lineage to
|
||
<taxonomicName authorityName="Forster C" authorityYear="1959" class="Arachnida" family="Micropholcommatidae" genus="Pua" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="novaezealandiae">
|
||
<emphasis italics="true" pageId="22" pageNumber="23">Pua novaezealandiae</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry box="[290,443,871,897]" name="New Zealand" pageId="22" pageNumber="23">New Zealand</collectingCountry>
|
||
(
|
||
<figureCitation box="[458,526,871,897]" captionStart="Figure 4" captionStartId="137.[140,212,1307,1329]" captionTargetBox="[161,1087,180,1278]" captionTargetId="figure@137.[140,1107,165,1288]" captionTargetPageId="137" captionText="Figure 4. Optimal cladogram for the Micropholcommatidae under equal and implied weights, and one of six equally parsimonious trees recovered from an equal weights analysis of the ‘new’ matrix (length = 135; see Appendix II). With the exception of the highlighted (*) node ‘Patelliella adusta + Micropholcommatini’, all clades shown were recovered in all iterations of an implied weights analysis (K=1–6); the phylogenetic position of P. adusta changed to ‘P. adusta + Textricellini’ under K-values 1–5. Unambiguous character optimisations are shown for each node, and bold species are newly described, with generic type species highlighted (*)." figureDoi="http://doi.org/10.5281/zenodo.3767693" httpUri="https://zenodo.org/record/3767693/files/figure.png" pageId="22" pageNumber="23">Fig. 4</figureCitation>
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||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="22" pageNumber="23" type="diagnosis">
|
||
<paragraph blockId="22.[140,1108,835,1672]" pageId="22" pageNumber="23">
|
||
<emphasis bold="true" box="[185,312,905,931]" pageId="22" pageNumber="23">
|
||
<emphasis box="[185,307,905,931]" inLineHeading="true" pageId="22" pageNumber="23" reason="1">Diagnosis</emphasis>
|
||
.
|
||
</emphasis>
|
||
Species of
|
||
<taxonomicName authorityName="Crosby & Bishop" authorityYear="1927" box="[435,624,906,932]" class="Arachnida" family="Micropholcommatidae" genus="Micropholcomma" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[435,624,906,932]" italics="true" pageId="22" pageNumber="23">Micropholcomma</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from all other
|
||
<taxonomicName family="Null" pageId="22" pageNumber="146" rank="tribe" tribe="Micropholcommatini">Micropholcommatini</taxonomicName>
|
||
except
|
||
<taxonomicName authorityName="Forster C" authorityYear="1959" box="[399,626,941,967]" class="Arachnida" family="Micropholcommatidae" genus="Pua" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="novaezealandiae">
|
||
<emphasis box="[399,626,941,967]" italics="true" pageId="22" pageNumber="23">Pua novaezealandiae</emphasis>
|
||
</taxonomicName>
|
||
by the presence of a normal, non-bifurcate embolus (
|
||
<figureCitation box="[250,337,976,1002]" captionStart="Figure 9" captionStartId="142.[140,212,687,709]" captionTargetBox="[151,1072,196,640]" captionTargetId="figure@142.[140,1108,184,645]" captionTargetPageId="142" captionText="Figure 9. Left pedipalps of male Micropholcommatini, Textricellini and Taphiassinae, showing trajectory of the sperm ducts: A Micropholcomma caeligenum Crosby & Bishop B Tinytrella pusilla (Forster) C Taphiassa punctata (Forster). Scale bars = 0.065 mm (65 µm). Numbered arrows denote the trajectory of the ducts, from the subtegular fundibule (1) to the base of the embolus." figureDoi="http://doi.org/10.5281/zenodo.3767705" httpUri="https://zenodo.org/record/3767705/files/figure.png" pageId="22" pageNumber="23">Fig. 9A</figureCitation>
|
||
); and can be distinguished from
|
||
<taxonomicName authorityName="Forster" authorityYear="1959" box="[702,744,977,1002]" class="Arachnida" family="Micropholcommatidae" genus="Pua" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[702,744,977,1002]" italics="true" pageId="22" pageNumber="23">Pua</emphasis>
|
||
</taxonomicName>
|
||
by the presence of a long embolus (length> 5× width) which is usually looped or coiled (
|
||
<figureCitation box="[801,891,1011,1038]" captionStart="Figure 9" captionStartId="142.[140,212,687,709]" captionTargetBox="[151,1072,196,640]" captionTargetId="figure@142.[140,1108,184,645]" captionTargetPageId="142" captionText="Figure 9. Left pedipalps of male Micropholcommatini, Textricellini and Taphiassinae, showing trajectory of the sperm ducts: A Micropholcomma caeligenum Crosby & Bishop B Tinytrella pusilla (Forster) C Taphiassa punctata (Forster). Scale bars = 0.065 mm (65 µm). Numbered arrows denote the trajectory of the ducts, from the subtegular fundibule (1) to the base of the embolus." figureDoi="http://doi.org/10.5281/zenodo.3767705" httpUri="https://zenodo.org/record/3767705/files/figure.png" pageId="22" pageNumber="23">Figs 9A</figureCitation>
|
||
, 12A, 17B). Other diagnostic characters include the presence (and presumed retention) of posterior tracheae (shared with
|
||
<taxonomicName authorityName="Rix & Harvey" authorityYear="2010" box="[364,449,1082,1108]" class="Arachnida" family="Micropholcommatidae" genus="Gigiella" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[364,449,1082,1108]" italics="true" pageId="22" pageNumber="23">Gigiella</emphasis>
|
||
</taxonomicName>
|
||
) and the presence of a male leg I-pedipalpal stridulatory system (Fig. 21D).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="description">
|
||
<paragraph blockId="22.[140,1108,835,1672]" pageId="22" pageNumber="23">
|
||
<emphasis bold="true" box="[185,335,1152,1178]" pageId="22" pageNumber="23">
|
||
<emphasis box="[185,330,1152,1178]" inLineHeading="true" pageId="22" pageNumber="23" reason="1">Description</emphasis>
|
||
.
|
||
</emphasis>
|
||
Very small, entelegyne
|
||
<taxonomicName box="[599,729,1152,1178]" order="Null" pageId="22" pageNumber="132" rank="superFamily" superFamily="Araneoidea">Araneoidea</taxonomicName>
|
||
; total length 0.70 to 1.30.
|
||
<emphasis italics="true" pageId="22" pageNumber="23">Cephalothorax</emphasis>
|
||
: Carapace without glandular depressions above maxillae; cuticle without glandular pits (Fig. 18C); margins fused to sternum via pleural sclerites. Eight (Fig. 18C), rarely six or (in troglomorphic taxa) fewer eyes present on anterior margin of pars cephalica; AME, if present, smallest. Chelicerae with or without bulging anterior projections in males; promargin with true teeth, a pair of fused setal sockets adjacent to base of fang and single peg tooth in males (Fig. 19F).
|
||
</paragraph>
|
||
<paragraph blockId="22.[140,1108,835,1672]" pageId="22" pageNumber="23">
|
||
<emphasis box="[185,463,1399,1425]" italics="true" pageId="22" pageNumber="23">Legs and female pedipalp</emphasis>
|
||
: Legs three-clawed (
|
||
<figureCitation box="[700,809,1399,1425]" captionStart="Figure 20" captionStartId="151.[140,212,940,962]" captionTargetBox="[140,1108,292,921]" captionTargetId="figure@151.[140,1108,291,923]" captionTargetPageId="151" captionText="Figure 20. Scanning electron micrographs of female Micropholcomma bryophilum (Butler) from near Marysville, Victoria (WAM T94453): A leg I tarsal organ B leg I lyriform organ C leg I tibial trichobothrium D leg IV claws, showing elongate inferior claw." figureDoi="http://doi.org/10.5281/zenodo.3767707" httpUri="https://zenodo.org/record/3767707/files/figure.png" pageId="22" pageNumber="23">Fig. 20D</figureCitation>
|
||
), covered with smooth or serrate hair-like setae; femur I of males with prolateral stridulatory denticles (Fig. 18C). Trichobothria present on legs; tibiae each with two or three trichobothria (
|
||
<figureCitation box="[989,1094,1469,1495]" captionStart="Figure 20" captionStartId="151.[140,212,940,962]" captionTargetBox="[140,1108,292,921]" captionTargetId="figure@151.[140,1108,291,923]" captionTargetPageId="151" captionText="Figure 20. Scanning electron micrographs of female Micropholcomma bryophilum (Butler) from near Marysville, Victoria (WAM T94453): A leg I tarsal organ B leg I lyriform organ C leg I tibial trichobothrium D leg IV claws, showing elongate inferior claw." figureDoi="http://doi.org/10.5281/zenodo.3767707" httpUri="https://zenodo.org/record/3767707/files/figure.png" pageId="22" pageNumber="23">Fig. 20C</figureCitation>
|
||
); metatarsi without trichobothria. Female pedipalp four-segmented (with fused tibiatarsus) or reduced to a vestigial nubbin; claw absent (Figs 19C–D).
|
||
</paragraph>
|
||
<paragraph blockId="22.[140,1108,835,1672]" lastBlockId="23.[140,1108,167,1144]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
|
||
<emphasis box="[185,287,1575,1601]" italics="true" pageId="22" pageNumber="23">Abdomen</emphasis>
|
||
: Abdomen oval or globose; anterior sclerite present around epigastric region and petiole; large dorsal scute present on males (Fig. 14), absent (Fig. 16B) or (rarely) present (Fig. 13B) on females; lateral sclerotic strips present on males; posterior sclerotic ring surrounding spinnerets and colulus. Six spinnerets situated posterior to fleshy colulus (
|
||
<figureCitation box="[306,474,202,228]" captionStart="Figure 22" captionStartId="153.[140,212,817,839]" captionTargetBox="[140,1108,165,796]" captionTargetId="figure@153.[140,1108,165,798]" captionTargetPageId="153" captionText="Figure 22. Scanning electron micrographs of spinnerets of female Micropholcomma bryophilum (Butler) from near Marysville, Victoria (WAM T94453) (left) and paratype female Austropholcomma florentine sp. n. from the Florentine Valley, Tasmania (FMC) (right): A–B anterior lateral spinnerets C–D posterior lateral spinnerets. Arrows denote the anterior, mesal direction." figureDoi="http://doi.org/10.5281/zenodo.3767709" httpUri="https://zenodo.org/record/3767709/files/figure.png" pageId="23" pageNumber="24">Figs 22A, 22C</figureCitation>
|
||
,
|
||
<figureCitation box="[486,514,202,228]" captionStart="Figure 23" captionStartId="154.[140,212,1453,1475]" captionTargetBox="[140,1108,165,1434]" captionTargetId="figure@154.[140,1108,165,1436]" captionTargetPageId="154" captionText="Figure 23. Scanning electron micrographs of posterior median spinnerets of female (left) and male (right) Micropholcommatini: A–B Micropholcomma bryophilum (Butler) from near Marysville, Victoria (WAM T94453) C–D Pua novaezealandiae Forster from the Makarora Valley, New Zealand (AMNH) E–F Austropholcomma florentine sp. n. from the Florentine Valley, Tasmania (FMC) G–H Tricellina gertschi (Forster & Platnick) from near Chaitén, Chile (AMNH). Note the apparent loss of the anterior CY gland spigot in female Austropholcomma and Tricellina. Arrows denote the anterior, mesal direction." figureDoi="http://doi.org/10.5281/zenodo.3767711" httpUri="https://zenodo.org/record/3767711/files/figure.png" pageId="23" pageNumber="24">23</figureCitation>
|
||
A–B); PMS of
|
||
<taxonomicName baseAuthorityName="Butler" baseAuthorityYear="1932" box="[684,849,202,228]" class="Arachnida" family="Micropholcommatidae" genus="Micropholcomma" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="bryophilum">
|
||
<emphasis box="[684,849,202,228]" italics="true" pageId="23" pageNumber="24">M. bryophilum</emphasis>
|
||
</taxonomicName>
|
||
with single medial AC gland spigot; PLS of
|
||
<taxonomicName baseAuthorityName="Butler" baseAuthorityYear="1932" box="[372,536,237,263]" class="Arachnida" family="Micropholcommatidae" genus="Micropholcomma" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="bryophilum">
|
||
<emphasis box="[372,536,237,263]" italics="true" pageId="23" pageNumber="24">M. bryophilum</emphasis>
|
||
</taxonomicName>
|
||
with complete triad. Anterior tracheal system welldeveloped, with relatively large lateral atria and multiple radiating tracheae (Fig. 15B); second tracheal tubes on each side looping mesally and extending through petiole into cephalothorax (Fig. 12D); posterior tracheal spiracle present, tracheae quadritracheate.
|
||
</paragraph>
|
||
<paragraph blockId="23.[140,1108,167,1144]" pageId="23" pageNumber="24">
|
||
<emphasis box="[185,288,378,404]" italics="true" pageId="23" pageNumber="24">Genitalia</emphasis>
|
||
: Male pedipalp (Fig. 21) small to large, relatively simple; patella with distally-directed, flanged or hooked ligulate retrolateral apophysis and retrolateral stridulatory ridges; tegulum smooth, with curved evaginated tegular ridge; embolus exposed, long (length> 5× width), usually looped or coiled. Female genitalia (Figs 15A–E) with pair of separate, globular anterior spermathecae; insemination ducts coiled around fertilisation ducts; fertilisation ducts posteriorly-directed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="23" pageNumber="24" type="distribution">
|
||
<paragraph blockId="23.[140,1108,167,1144]" pageId="23" pageNumber="24">
|
||
<emphasis bold="true" box="[185,344,589,615]" pageId="23" pageNumber="24">
|
||
<emphasis box="[185,339,589,615]" inLineHeading="true" pageId="23" pageNumber="24" reason="1">Distribution</emphasis>
|
||
.
|
||
</emphasis>
|
||
Eastern and western mainland
|
||
<collectingCountry box="[710,809,589,615]" name="Australia" pageId="23" pageNumber="24">Australia</collectingCountry>
|
||
and
|
||
<collectingRegion box="[869,976,589,615]" country="Australia" name="Tasmania" pageId="23" pageNumber="24">Tasmania</collectingRegion>
|
||
(Fig. 217). Two undescribed female specimens from Rivière Bleue and Mont Mou,
|
||
<collectingCountry name="New Caledonia" pageId="23" pageNumber="24">New Caledonia</collectingCountry>
|
||
(QMB
|
||
<accessionNumber box="[268,359,659,686]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/S60522" pageId="23" pageNumber="24">S60522</accessionNumber>
|
||
, QMB
|
||
<accessionNumber box="[447,536,660,686]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/S72474" pageId="23" pageNumber="24">S72474</accessionNumber>
|
||
) probably also belong in this genus, but males are required to confirm the identification.
|
||
</paragraph>
|
||
<paragraph blockId="23.[140,1108,167,1144]" pageId="23" pageNumber="24">
|
||
<emphasis bold="true" box="[185,351,729,755]" pageId="23" pageNumber="24">
|
||
<emphasis box="[185,346,729,755]" inLineHeading="true" pageId="23" pageNumber="24" reason="1">Composition</emphasis>
|
||
.
|
||
</emphasis>
|
||
Seven described species,
|
||
<taxonomicName authority="(Butler, 1932)" baseAuthorityName="Butler" baseAuthorityYear="1932" class="Arachnida" family="Micropholcommatidae" genus="Micropholcomma" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="bryophilum">
|
||
<emphasis box="[674,1005,730,756]" italics="true" pageId="23" pageNumber="24">Micropholcomma bryophilum</emphasis>
|
||
(
|
||
<bibRefCitation author="Butler LSG" pageId="23" pageNumber="24" pagination="103 - 117" refId="ref69719" refString="Butler LSG (1932) Studies in Australian spiders, No. 2. Proceedings of the Royal Society of Victoria 44: 103 - 117." type="journal article" year="1932">Butler, 1932</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Crosby & Bishop, 1927" authorityName="Crosby & Bishop B" authorityYear="1927" box="[223,667,765,791]" class="Arachnida" family="Micropholcommatidae" genus="Micropholcomma" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="caeligenum">
|
||
<emphasis box="[223,384,765,791]" italics="true" pageId="23" pageNumber="24">M. caeligenum</emphasis>
|
||
<bibRefCitation author="Crosby CR & Bishop SC" box="[392,667,765,791]" pageId="23" pageNumber="24" pagination="147 - 154" refId="ref69920" refString="Crosby CR, Bishop SC (1927) New species of Erigoneae and Theridiidae. Journal of the New York Entomological Society 35: 147 - 154." type="journal article" year="1927">Crosby & Bishop, 1927</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Rix, 2008" authorityName="Rix" authorityYear="2008" box="[680,921,765,791]" class="Arachnida" family="Micropholcommatidae" genus="Micropholcomma" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="linnaei">
|
||
<emphasis box="[680,797,765,791]" italics="true" pageId="23" pageNumber="24">M. linnaei</emphasis>
|
||
Rix, 2008
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="(Butler, 1932)" baseAuthorityName="Butler" baseAuthorityYear="1932" class="Arachnida" family="Micropholcommatidae" genus="Micropholcomma" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="longissimum">
|
||
<emphasis box="[933,1107,765,791]" italics="true" pageId="23" pageNumber="24">M. longissimum</emphasis>
|
||
(
|
||
<bibRefCitation author="Butler LSG" box="[148,294,800,826]" pageId="23" pageNumber="24" pagination="103 - 117" refId="ref69719" refString="Butler LSG (1932) Studies in Australian spiders, No. 2. Proceedings of the Royal Society of Victoria 44: 103 - 117." type="journal article" year="1932">Butler, 1932</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Hickman, 1944" authorityName="Hickman" authorityYear="1944" box="[314,618,800,826]" class="Arachnida" family="Micropholcommatidae" genus="Micropholcomma" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="mirum">
|
||
<emphasis box="[314,430,801,826]" italics="true" pageId="23" pageNumber="24">M. mirum</emphasis>
|
||
<bibRefCitation author="Hickman VV" box="[436,618,800,826]" pageId="23" pageNumber="24" pagination="179 - 195" refId="ref70811" refString="Hickman VV (1944) On some new Australian Apneumonomorphae with notes on their respiratory system. Papers and Proceedings of the Royal Society of Tasmania 1944: 179 - 195." type="journal article" year="1944">Hickman, 1944</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Hickman, 1944" authorityName="Hickman" authorityYear="1944" box="[630,974,800,826]" class="Arachnida" family="Micropholcommatidae" genus="Micropholcomma" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="parmatum">
|
||
<emphasis box="[630,785,801,826]" italics="true" pageId="23" pageNumber="24">M. parmatum</emphasis>
|
||
<bibRefCitation author="Hickman VV" box="[792,974,800,826]" pageId="23" pageNumber="24" pagination="179 - 195" refId="ref70811" refString="Hickman VV (1944) On some new Australian Apneumonomorphae with notes on their respiratory system. Papers and Proceedings of the Royal Society of Tasmania 1944: 179 - 195." type="journal article" year="1944">Hickman, 1944</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Hickman, 1981" authorityName="Hickman" authorityYear="1981" class="Arachnida" family="Micropholcommatidae" genus="Micropholcomma" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="turbans">
|
||
<emphasis box="[986,1107,800,826]" italics="true" pageId="23" pageNumber="24">M. turbans</emphasis>
|
||
<bibRefCitation author="Hickman VV" box="[140,325,835,862]" pageId="23" pageNumber="24" pagination="47 - 68" refId="ref70902" refString="Hickman VV (1981) New Tasmanian spiders of the families Archaeidae, Cycloctenidae, Amaurobiidae and Micropholcommatidae. Papers and Proceedings of the Royal Society of Tasmania 115: 47 - 68." type="journal article" year="1981">Hickman, 1981</bibRefCitation>
|
||
</taxonomicName>
|
||
, and the new species
|
||
<taxonomicName authorityName="Rix & Harvey" authorityYear="2010" box="[579,679,836,861]" class="Arachnida" family="Micropholcommatidae" genus="Micropholcomma" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="junee">
|
||
<emphasis box="[579,679,836,861]" italics="true" pageId="23" pageNumber="24">M. junee</emphasis>
|
||
</taxonomicName>
|
||
. Undescribed species are also known from throughout the range of the genus.
|
||
</paragraph>
|
||
<paragraph blockId="23.[140,1108,167,1144]" pageId="23" pageNumber="24">
|
||
<emphasis bold="true" box="[185,477,906,932]" pageId="23" pageNumber="24">
|
||
<emphasis box="[185,364,906,932]" inLineHeading="true" pageId="23" pageNumber="24" reason="1">Nomenclatural</emphasis>
|
||
remarks.
|
||
</emphasis>
|
||
The gender of the genus name “
|
||
<taxonomicName authorityName="Crosby & Bishop" authorityYear="1927" box="[857,1046,906,932]" class="Arachnida" family="Micropholcommatidae" genus="Micropholcomma" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[857,1046,906,932]" italics="true" pageId="23" pageNumber="24">Micropholcomma</emphasis>
|
||
</taxonomicName>
|
||
” was not specified by
|
||
<bibRefCitation author="Crosby CR & Bishop SC" box="[330,640,941,967]" pageId="23" pageNumber="24" pagination="147 - 154" refId="ref69920" refString="Crosby CR, Bishop SC (1927) New species of Erigoneae and Theridiidae. Journal of the New York Entomological Society 35: 147 - 154." type="journal article" year="1927">Crosby and Bishop (1927)</bibRefCitation>
|
||
, although the
|
||
<typeStatus box="[808,856,942,968]" pageId="23" pageNumber="24">type</typeStatus>
|
||
species was described with a masculine specific epithet (as
|
||
<emphasis box="[554,580,977,1002]" italics="true" pageId="23" pageNumber="24">M</emphasis>
|
||
. “
|
||
<taxonomicName authorityName="Crosby & Bishop" authorityYear="1927" box="[607,714,976,1002]" class="Arachnida" family="Micropholcommatidae" genus="Micropholcomma" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="caeligenus">
|
||
<emphasis box="[607,714,976,1002]" italics="true" pageId="23" pageNumber="24">caeligenus</emphasis>
|
||
</taxonomicName>
|
||
”). The name
|
||
<taxonomicName authorityName="Crosby & Bishop" authorityYear="1927" box="[870,1059,976,1002]" class="Arachnida" family="Micropholcommatidae" genus="Micropholcomma" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[870,1059,976,1002]" italics="true" pageId="23" pageNumber="24">Micropholcomma</emphasis>
|
||
</taxonomicName>
|
||
was clearly derived from the theridiid taxon name “
|
||
<taxonomicName authority="Thorell" authorityName="Thorell" authorityYear="1869" box="[663,790,1011,1037]" class="Arachnida" family="Theridiidae" genus="Pholcomma" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[663,790,1011,1037]" italics="true" pageId="23" pageNumber="24">Pholcomma</emphasis>
|
||
</taxonomicName>
|
||
” Thorell, and as highlighted by
|
||
<bibRefCitation author="Levi HW" box="[177,315,1047,1074]" pageId="23" pageNumber="24" pagination="32 - 39" refId="ref71219" refString="Levi HW (1964) The American spiders of the genera Styposis and Pholcomma (Araneae, Theridiidae). Psyche 71: 32 - 39." type="journal article" year="1964">Levi (1964)</bibRefCitation>
|
||
, this name should be treated as neuter. To this end,
|
||
<bibRefCitation author="Brignoli PM" box="[928,1107,1047,1073]" pageId="23" pageNumber="24" refId="ref69575" refString="Brignoli PM (1983) A Catalogue of the Araneae Described Between 1940 and 1981. Manchester University Press, Manchester, 755 pp." type="book" year="1983">Brignoli (1983)</bibRefCitation>
|
||
and
|
||
<bibRefCitation author="Davies VT" box="[189,349,1082,1109]" pageId="23" pageNumber="24" pagination="48 - 125" refId="ref69989" refString="Davies VT (1985) Araneomorphae (in part). In: Walton DW (Ed) Zoological Catalogue of Australia, Vol. 3. Mygalomorphae, Araneomorphae (in part), Pseudoscorpionida, Amblypygi and Palpigradi. Australian Government Publishing Service, Canberra, 48 - 125." type="book chapter" year="1985">Davies (1985)</bibRefCitation>
|
||
amended the suffix endings of the species names, all of which were previously masculine or feminine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |