226 lines
20 KiB
XML
226 lines
20 KiB
XML
<document ID-DOI="10.3897/zookeys.36.306" ID-GBIF-Dataset="4db6b327-7482-432e-a5f6-36f91c79fef3" ID-ISSN="1313–2970" ID-Zenodo-Dep="576620" ID-ZooBank="ADCACC88-6C78-4386-8E33-3F98234ECE92" approvalRequired="105" approvalRequired_for_textStreams="105" checkinTime="1587967053581" checkinUser="plazi" docAuthor="Rix, Michael & Harvey, Mark" docDate="2010" docId="7E13878EFF9E1B52FF321CDEFCF0897D" docLanguage="en" docName="ZK_article_2154.pdf" docOrigin="ZooKeys 36 (36)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@6ed9075d" docStyleName="zookeys.2008.journal_article" docTitle="Eterosonycha ocellata Rix & Harvey 2010, sp. n." docType="treatment" docUuid="46E2827A-B315-4C35-855E-705A20E6B220" docUuidSource="ZooBank" docVersion="5" lastPageNumber="42" masterDocId="822AFFF6FFB61B7BFFBE1F57FFE98D05" masterDocTitle="The spider family Micropholcommatidae (Arachnida: Araneae: Araneoidea): a relimitation and revision at the generic level" masterLastPageNumber="321" masterPageNumber="1" pageNumber="41" updateTime="1669157185164" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The spider family Micropholcommatidae (Arachnida: Araneae: Araneoidea): a relimitation and revision at the generic level</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rix, Michael</mods:namePart>
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<mods:affiliation>Western Australian Museum, Welshpool DC, Perth, Australia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Harvey, Mark</mods:namePart>
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<mods:affiliation>Western Australian Museum, Perth, Welshpool, Australia</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2010</mods:date>
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||
<mods:detail type="pubDate">
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<mods:number>2010-02-22</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>36</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>36</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>321</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.36.306</mods:identifier>
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<mods:identifier type="GBIF-Dataset">4db6b327-7482-432e-a5f6-36f91c79fef3</mods:identifier>
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||
<mods:identifier type="ISSN">1313–2970</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576620</mods:identifier>
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||
<mods:identifier type="ZooBank">ADCACC88-6C78-4386-8E33-3F98234ECE92</mods:identifier>
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||
</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3789405" ID-GBIF-Taxon="163749043" ID-Zenodo-Dep="3789405" LSID="urn:lsid:zoobank.org:act:46E2827A-B315-4C35-855E-705A20E6B220" httpUri="http://treatment.plazi.org/id/7E13878EFF9E1B52FF321CDEFCF0897D" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
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||
<subSubSection pageId="40" pageNumber="41" type="nomenclature">
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||
<paragraph blockId="40.[140,955,905,1003]" box="[140,644,905,932]" pageId="40" pageNumber="41">
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||
<heading box="[140,644,905,932]" fontSize="11" level="2" pageId="40" pageNumber="41" reason="3">
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<emphasis bold="true" box="[140,644,905,932]" pageId="40" pageNumber="41">
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<taxonomicName authority="Rix & Harvey" authorityName="Rix & Harvey" authorityYear="2010" box="[140,567,905,932]" class="Arachnida" family="Micropholcommatidae" genus="Eterosonycha" kingdom="Animalia" order="Araneae" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="ocellata" status="sp. n.">
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<emphasis bold="true" box="[140,395,905,932]" italics="true" pageId="40" pageNumber="41">
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<emphasis box="[140,296,905,932]" inLineHeading="true" pageId="40" pageNumber="41" reason="1">Eterosonycha</emphasis>
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ocellata
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</emphasis>
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Rix & Harvey
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</taxonomicName>
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,
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<taxonomicNameLabel box="[579,644,906,932]" pageId="40" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="40.[140,955,905,1003]" box="[140,955,941,968]" pageId="40" pageNumber="41">
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||
<uri box="[140,955,941,968]" pageId="40" pageNumber="41">
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||
urn:lsid:zoobank.org:act:
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||
<uuid box="[418,955,941,968]" pageId="40" pageNumber="41">46E2827A-B315-4C35-855E-705A20E6B220</uuid>
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||
</uri>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection box="[140,332,976,1003]" pageId="40" pageNumber="41" type="description">
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<paragraph blockId="40.[140,955,905,1003]" box="[140,332,976,1003]" pageId="40" pageNumber="41">Figs 51D, 54–55</paragraph>
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</subSubSection>
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<subSubSection pageId="40" pageNumber="41" type="materials_examined">
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<paragraph blockId="40.[140,1108,1046,1672]" pageId="40" pageNumber="41">
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<emphasis bold="true" box="[140,312,1046,1073]" pageId="40" pageNumber="41">
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<emphasis box="[140,198,1047,1073]" inLineHeading="true" pageId="40" pageNumber="41" reason="1">Type</emphasis>
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material.
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</emphasis>
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<materialsCitation ID-GBIF-Occurrence="2608825887" collectingDate="2006-04-23" collectionCode="NMV" collectorName="M. Rix" country="Australia" county="Apollo Bay" latitude="-38.75528" location="Great Otway National Park" longLatPrecision="19" longitude="143.55501" municipality="Maits Rest" pageId="40" pageNumber="41" specimenCount="1" stateProvince="Victoria" typeStatus="holotype">
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<typeStatus box="[319,426,1047,1073]" pageId="40" pageNumber="41">Holotype</typeStatus>
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male:
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<location LSID="urn:lsid:plazi:treatment:7E13878EFF9E1B52FF321CDEFCF0897D:F3656043FF9E1B53FE481B40FCD88934" box="[502,817,1047,1073]" country="Australia" county="Apollo Bay" latitude="-38.75528" longLatPrecision="19" longitude="143.55501" municipality="Maits Rest" name="Great Otway National Park" pageId="40" pageNumber="41" stateProvince="Victoria">Great Otway National Park</location>
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,
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<collectingMunicipality box="[829,953,1047,1073]" pageId="40" pageNumber="41">Maits Rest</collectingMunicipality>
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,
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<quantity box="[964,1038,1047,1073]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="40" pageNumber="41" unit="km" value="10.0">10 km</quantity>
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W. of
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<collectingCounty box="[140,268,1082,1108]" pageId="40" pageNumber="41">Apollo Bay</collectingCounty>
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,
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<collectingRegion box="[280,374,1082,1108]" country="Australia" name="Victoria" pageId="40" pageNumber="41">Victoria</collectingRegion>
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,
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<collectingCountry box="[388,491,1082,1108]" name="Australia" pageId="40" pageNumber="41">Australia</collectingCountry>
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, sifting moss from trunk of
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<taxonomicName box="[817,1101,1082,1108]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="40" pageNumber="41" phylum="Tracheophyta" rank="species" species="cunninghamii">
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<emphasis box="[817,1101,1082,1108]" italics="true" pageId="40" pageNumber="41">Nothofagus cunninghamii</emphasis>
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</taxonomicName>
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,
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<geoCoordinate box="[140,272,1117,1144]" degrees="38" direction="south" minutes="45" orientation="latitude" pageId="40" pageNumber="41" precision="15" seconds="19" value="-38.75528">38°45'19"S</geoCoordinate>
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,
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<geoCoordinate box="[284,435,1117,1143]" degrees="143" direction="east" minutes="33" orientation="longitude" pageId="40" pageNumber="41" precision="15" seconds="18" value="143.55501">143°33'18"E</geoCoordinate>
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,
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<date box="[447,578,1117,1143]" pageId="40" pageNumber="41" value="2006-04-23">
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<collectingDate box="[447,578,1117,1143]" pageId="40" pageNumber="41" value="2006-04-23">23.IV.2006</collectingDate>
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</date>
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,
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<collectorName box="[590,671,1117,1144]" pageId="40" pageNumber="41">M. Rix</collectorName>
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(
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<collectionCode box="[686,757,1117,1143]" country="Australia" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="40" pageNumber="41" type="Museum">NMV</collectionCode>
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<accessionNumber box="[764,863,1117,1144]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/K10768" pageId="40" pageNumber="41">K10768</accessionNumber>
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).
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</materialsCitation>
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</paragraph>
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<paragraph blockId="40.[140,1108,1046,1672]" pageId="40" pageNumber="41">
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<typeStatus box="[185,295,1153,1179]" pageId="40" pageNumber="41">Paratypes</typeStatus>
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:
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<materialsCitation ID-GBIF-Occurrence="2608825881" collectingDate="2006-04-23" collectionCode="NMV, WAM" collectorName="M. Rix" country="Australia" county="Apollo Bay" latitude="-38.75528" location="Great Otway National Park" longLatPrecision="19" longitude="143.55501" municipality="Maits Rest" pageId="40" pageNumber="41" specimenCount="1" specimenCount-male="1" stateProvince="Victoria" typeStatus="allotype">
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<typeStatus box="[307,402,1152,1178]" pageId="40" pageNumber="41">Allotype</typeStatus>
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female, same data as holotype (
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<collectionCode box="[763,834,1152,1178]" country="Australia" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="40" pageNumber="41" type="Museum">NMV</collectionCode>
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<accessionNumber box="[842,939,1152,1179]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/K10769" pageId="40" pageNumber="41">K10769</accessionNumber>
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);
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<specimenCount box="[960,1039,1152,1178]" pageId="40" pageNumber="41" type="male">1 male</specimenCount>
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, same data as
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<typeStatus box="[223,322,1187,1213]" pageId="40" pageNumber="41">holotype</typeStatus>
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(
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<collectionCode box="[337,410,1188,1214]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="40" pageNumber="41" type="Museum">WAM</collectionCode>
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<accessionNumber box="[417,515,1188,1214]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/T94443" pageId="40" pageNumber="41">T94443</accessionNumber>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="40" pageNumber="41" type="etymology">
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<paragraph blockId="40.[140,1108,1046,1672]" pageId="40" pageNumber="41">
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<emphasis bold="true" box="[185,320,1223,1249]" pageId="40" pageNumber="41">
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<emphasis box="[185,315,1223,1249]" inLineHeading="true" pageId="40" pageNumber="41" reason="1">Etymology</emphasis>
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.
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</emphasis>
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The specific epithet is derived from the Latin ‘ocellatus’, meaning ‘having little eyes’ (
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<bibRefCitation author="Brown RW" box="[355,501,1258,1285]" pageId="40" pageNumber="41" refId="ref69600" refString="Brown RW (1956) Composition of Scientific Words: A Manual of Methods and a Lexicon of Materials for the Practice of Logotechnics. Smithsonian Books, Washington, DC, 882 pp." type="book" year="1956">Brown 1956</bibRefCitation>
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), and refers to the small eyes of this species.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="40" pageNumber="41" type="diagnosis">
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<paragraph blockId="40.[140,1108,1046,1672]" pageId="40" pageNumber="41">
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||
<emphasis bold="true" box="[185,312,1293,1319]" pageId="40" pageNumber="41">
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<emphasis box="[185,307,1293,1319]" inLineHeading="true" pageId="40" pageNumber="41" reason="1">Diagnosis</emphasis>
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.
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</emphasis>
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Males of
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<taxonomicName authorityName="Rix & Harvey" authorityYear="2010" box="[427,654,1293,1319]" class="Arachnida" family="Micropholcommatidae" genus="Eterosonycha" kingdom="Animalia" order="Araneae" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="ocellata">
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<emphasis box="[427,654,1293,1319]" italics="true" pageId="40" pageNumber="41">Eterosonycha ocellata</emphasis>
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</taxonomicName>
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can be distinguished from all other described congeners by the shape of the distal conductor, which has a tubular prolateral process and bifurcate retrolateral process (
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<figureCitation box="[672,757,1363,1390]" captionStart="Figure 55" captionStartId="178.[140,212,1622,1644]" captionTargetBox="[163,1070,943,1592]" captionTargetId="figure@178.[159,1088,929,1602]" captionTargetPageId="178" captionText="Figure 55. Eterosonycha ocellata sp. n., paratype male from the Otway Ranges, Victoria (WAM T94443): A left pedipalp, ventral view B left pedipalp, retrolateral view. Scale bar = 0.065 mm (65 µm)." figureDoi="http://doi.org/10.5281/zenodo.3767769" httpUri="https://zenodo.org/record/3767769/files/figure.png" pageId="40" pageNumber="41">Fig. 55</figureCitation>
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). Females can be distinguished from all other described congeners by the distinctive shape of the external epigyne (
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<figureCitation box="[148,255,1434,1461]" captionStart="Figure 54" captionStartId="178.[140,212,826,848]" captionTargetBox="[140,1108,165,807]" captionTargetId="figure@178.[140,1109,165,810]" captionTargetPageId="178" captionText="Figure 54. Eterosonycha ocellata sp. n. from the Otway Ranges, Victoria. A, C holotype male (NMV K10768): A habitus, dorsal view C cephalothorax, antero-dorsal view. B, D allotype female (NMV K10769): B habitus, dorsal view D abdomen, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3767767" httpUri="https://zenodo.org/record/3767767/files/figure.png" pageId="40" pageNumber="41">Fig. 54D</figureCitation>
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). Both sexes can also be recognised by the relatively small eyes (
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<figureCitation box="[970,1075,1434,1461]" captionStart="Figure 54" captionStartId="178.[140,212,826,848]" captionTargetBox="[140,1108,165,807]" captionTargetId="figure@178.[140,1109,165,810]" captionTargetPageId="178" captionText="Figure 54. Eterosonycha ocellata sp. n. from the Otway Ranges, Victoria. A, C holotype male (NMV K10768): A habitus, dorsal view C cephalothorax, antero-dorsal view. B, D allotype female (NMV K10769): B habitus, dorsal view D abdomen, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3767767" httpUri="https://zenodo.org/record/3767767/files/figure.png" pageId="40" pageNumber="41">Fig. 54C</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="description">
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<paragraph blockId="40.[140,1108,1046,1672]" lastBlockId="41.[140,1108,167,1144]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
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<emphasis bold="true" box="[185,340,1469,1495]" pageId="40" pageNumber="41">
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<emphasis box="[185,335,1469,1495]" inLineHeading="true" pageId="40" pageNumber="41" reason="1">Description</emphasis>
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.
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</emphasis>
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<emphasis box="[348,507,1469,1495]" italics="true" pageId="40" pageNumber="41">
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<typeStatus box="[348,446,1469,1495]" pageId="40" pageNumber="41">Holotype</typeStatus>
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male
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</emphasis>
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: Total length 0.83. Carapace 0.42 long, 0.35 wide. Abdomen 0.57 long, 0.41 wide. Leg I femur 0.33. Body colour pale tan-yellow, carapace slightly darker. Carapace raised anteriorly, not fused to sternum except around petiole; dorsal surface of pars cephalica slightly convex in lateral view. Eight eyes present on anterior margin of pars cephalica; AME 0.5x diameter of ALE; PME separated by slightly more than their own diameter. Chelicerae each with large, bulging anterior projection; promargin with three peg teeth. Legs relatively short (leg I femur-carapace ratio 0.79); distal, prolateral macroseta present on tibia I. Abdomen oval, covered with hair-like setae; dorsal scute large, covering most of dorsal surface of abdomen; lateral sclerotic strips absent. Pedipalpal patella with retrolaterally-directed, hooked lRPA and expanded distal region with ornate, ridged cuticular microstructure; tibia enlarged, with pronounced, curved posterior process; tegulum large, bulging posteriorly, with excavate ETR and large, divergent, fleshy distal conductor bearing tubular prolateral process and bifurcate retrolateral process; embolus short (length <5× width), spur-like, situated adjacent to base of conductor (
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||
<figureCitation box="[275,363,483,510]" captionStart="Figure 55" captionStartId="178.[140,212,1622,1644]" captionTargetBox="[163,1070,943,1592]" captionTargetId="figure@178.[159,1088,929,1602]" captionTargetPageId="178" captionText="Figure 55. Eterosonycha ocellata sp. n., paratype male from the Otway Ranges, Victoria (WAM T94443): A left pedipalp, ventral view B left pedipalp, retrolateral view. Scale bar = 0.065 mm (65 µm)." figureDoi="http://doi.org/10.5281/zenodo.3767769" httpUri="https://zenodo.org/record/3767769/files/figure.png" pageId="41" pageNumber="42">Fig. 55</figureCitation>
|
||
).
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</paragraph>
|
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<paragraph blockId="41.[140,1108,167,1144]" pageId="41" pageNumber="42">
|
||
<emphasis box="[185,357,519,545]" italics="true" pageId="41" pageNumber="42">
|
||
<typeStatus box="[185,274,519,545]" pageId="41" pageNumber="42">Allotype</typeStatus>
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female
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</emphasis>
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: Total length 1.00. Carapace 0.47 long, 0.33 wide. Abdomen 0.62 long, 0.46 wide. Leg I femur 0.35. Cephalothorax, abdominal sclerites, legs tan-yellow; abdomen dark olive brown dorsally, with chequered dark-olive and tanyellow pattern laterally and ventrally. Carapace raised anteriorly, not fused to sternum except around petiole; dorsal surface of pars cephalica slightly convex in lateral view. Eight eyes present on anterior margin of pars cephalica; AME 0.5x diameter of ALE; PME separated by their own diameter. Chelicerae without bulging anterior projections; promargin without peg teeth. Legs relatively short (leg I femurcarapace ratio 0.74); macrosetae absent. Abdomen oval, covered with hair-like setae; dorsal scute and lateral sclerotic strips absent. Pedipalp entire, five-segmented. Epigyne heavily sclerotised externally, with distinctive morphology (
|
||
<figureCitation box="[913,1027,870,897]" captionStart="Figure 54" captionStartId="178.[140,212,826,848]" captionTargetBox="[140,1108,165,807]" captionTargetId="figure@178.[140,1109,165,810]" captionTargetPageId="178" captionText="Figure 54. Eterosonycha ocellata sp. n. from the Otway Ranges, Victoria. A, C holotype male (NMV K10768): A habitus, dorsal view C cephalothorax, antero-dorsal view. B, D allotype female (NMV K10769): B habitus, dorsal view D abdomen, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3767767" httpUri="https://zenodo.org/record/3767767/files/figure.png" pageId="41" pageNumber="42">Fig. 54D</figureCitation>
|
||
); spermathecae looped and twisted; insemination ducts broad, curved; fertilisation ducts filiform (Fig. 51D).
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||
</paragraph>
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</subSubSection>
|
||
<subSubSection box="[185,906,976,1003]" pageId="41" pageNumber="42" type="distribution">
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||
<paragraph blockId="41.[140,1108,167,1144]" box="[185,906,976,1003]" pageId="41" pageNumber="42">
|
||
<emphasis bold="true" box="[185,344,976,1002]" pageId="41" pageNumber="42">
|
||
<emphasis box="[185,339,976,1002]" inLineHeading="true" pageId="41" pageNumber="42" reason="1">Distribution</emphasis>
|
||
.
|
||
</emphasis>
|
||
Known only from the Otway Ranges of
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<collectingRegion box="[806,901,976,1002]" country="Australia" name="Victoria" pageId="41" pageNumber="42">Victoria</collectingRegion>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="41" pageNumber="42" type="discussion">
|
||
<paragraph blockId="41.[140,1108,167,1144]" pageId="41" pageNumber="42">
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||
<emphasis bold="true" box="[185,298,1011,1037]" pageId="41" pageNumber="42">
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||
<emphasis box="[185,293,1011,1037]" inLineHeading="true" pageId="41" pageNumber="42" reason="1">Remarks</emphasis>
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.
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</emphasis>
|
||
<taxonomicName authorityName="Rix & Harvey" authorityYear="2010" box="[310,541,1011,1037]" class="Arachnida" family="Micropholcommatidae" genus="Eterosonycha" kingdom="Animalia" order="Araneae" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="ocellata">
|
||
<emphasis box="[310,541,1011,1037]" italics="true" pageId="41" pageNumber="42">Eterosonycha ocellata</emphasis>
|
||
</taxonomicName>
|
||
is an unusual species of
|
||
<taxonomicName authorityName="Butler" authorityYear="1932" box="[848,986,1011,1037]" class="Arachnida" family="Micropholcommatidae" genus="Eterosonycha" kingdom="Animalia" order="Araneae" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[848,986,1011,1037]" italics="true" pageId="41" pageNumber="42">Eterosonycha</emphasis>
|
||
</taxonomicName>
|
||
, currently known only from the Otway Ranges of
|
||
<collectingRegion box="[582,677,1047,1073]" country="Australia" name="Victoria" pageId="41" pageNumber="42">Victoria</collectingRegion>
|
||
. The
|
||
<typeStatus box="[736,784,1047,1073]" pageId="41" pageNumber="42">type</typeStatus>
|
||
specimens were found living together in moss on the trunk of a large Southern Beech (
|
||
<taxonomicName box="[815,1098,1082,1108]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="41" pageNumber="42" phylum="Tracheophyta" rank="species" species="cunninghamii">
|
||
<emphasis box="[815,1098,1082,1108]" italics="true" pageId="41" pageNumber="42">Nothofagus cunninghamii</emphasis>
|
||
</taxonomicName>
|
||
) tree, in dark, wet, cool-temperate rainforest at Maits Rest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |