142 lines
16 KiB
XML
142 lines
16 KiB
XML
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<mods:title id="E2058AB75907FDBF619A08C03F3B2BFC">One the genus Tocama Reitter (Coleoptera, Scarabaeidae, Melolonthinae), with descriptions of two new species from Indochina</mods:title>
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<mods:namePart id="BCA29570FBC4618537FC4EC634A23709">Li, Chun-Lin</mods:namePart>
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<mods:namePart id="4DC35183AB0FA10A137BED6C6F567815">Wang, Chuan-Chan</mods:namePart>
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<mods:namePart id="C7B53C40E79390F5399E889750683683">Keith, Denis</mods:namePart>
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<mods:namePart id="D50826B5A638358D92A2032F77B75BB2">Yang, Ping-Shih</mods:namePart>
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<mods:title id="EF83A6B76D6B83D43D87B6D23347EBBA">ZooKeys</mods:title>
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<mods:date id="E99DE00CBABFFACE48F76D29EF57A772">2012</mods:date>
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<mods:detail id="9472621F1F9D234D195C01E2F1929AD2" type="volume">
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<mods:number id="1812D4E73B58B79D9527C87547AF6320">177</mods:number>
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<mods:url id="C27433121CC946E672CFF9FBB5350C55">http://dx.doi.org/10.3897/zookeys.177.2482</mods:url>
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<mods:classification id="3A89B78A42CD2BC3D57E5A643782B210">journal article</mods:classification>
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<mods:identifier id="A90FC157CD0619911048540288A7CAC0" type="DOI">http://dx.doi.org/10.3897/zookeys.177.2482</mods:identifier>
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<mods:identifier id="3192BBAF1F7338BF1698595BFF60C9B0" type="Pensoft-Pub">1313-2970-177-37</mods:identifier>
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<treatment id="7D5D8FFA8FAF58EC67053EDCFCA504D1" ID-GBIF-Taxon="152033630" LSID="urn:lsid:plazi:treatment:7D5D8FFA8FAF58EC67053EDCFCA504D1" httpUri="http://treatment.plazi.org/id/7D5D8FFA8FAF58EC67053EDCFCA504D1" lastPageId="6" lastPageNumber="43" pageId="5" pageNumber="42" scope_family="Scarabaeidae" scope_order="Coleoptera" scope_subFamily="Melolonthinae">
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<subSubSection id="42872932B4BEC3E8AE0503AE9BD5AF88" pageId="5" pageNumber="42" type="nomenclature">
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<paragraph id="24532595080E7997AE96511EFE5D3A77" pageId="5" pageNumber="42">
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<taxonomicName id="3564E48422FF22C18A01335CB5D3032F" ID-CoL="7CFZD" authority="Moser, 1913" authorityName="Moser" authorityYear="1913" class="Insecta" family="Scarabaeidae" genus="Tocama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tocama tonkinensis" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="tonkinensis">Tocama tonkinensis (Moser, 1913)</taxonomicName>
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<taxonomicNameLabel id="D47D1F728DE85CBF18B5B20B0582BD2D" pageId="5" pageNumber="42">comb. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="7F8172C5B16A4179C988C97831B3CBD7" pageId="5" pageNumber="42" type="reference_group">
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<paragraph id="C908783FEB281247AEF7781A546B62CF" pageId="5" pageNumber="42">
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<taxonomicName id="59541B7F174FBFF94C8B4BC724F84028" class="Insecta" family="Scarabaeidae" genus="Hoplosternus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplosternus tonkinensis" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="tonkinensis">Hoplosternus tonkinensis</taxonomicName>
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<bibRefCitation id="95219BC2628493A38A0436DAC3557667" author="Moser, J" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="8" pageNumber="45" pagination="271 - 297" title="Beitrag zur kenntnis der Melolonthiden (Col.)." volume="1913" year="1913">Moser 1913</bibRefCitation>
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: 290. Original combination
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</paragraph>
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<paragraph id="4B3869957D389C27E3C19D5681F9560C" pageId="5" pageNumber="42">
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<taxonomicName id="28F526AD9134CB0DCCA432A1045B1CFF" class="Insecta" family="Scarabaeidae" genus="Hoplosternus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplosternus pygidialis" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="pygidialis">Hoplosternus pygidialis</taxonomicName>
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Moser, 1915: 589, syn. n.
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</paragraph>
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<paragraph id="9ABD74BEEE543DB3865DF4CC472688C2" pageId="5" pageNumber="42">
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<taxonomicName id="FD6C807BA37F6FD1F53AD276672EA882" class="Insecta" family="Scarabaeidae" genus="Tocama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tocama atra subsp. atra" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="subSpecies" species="atra" subSpecies="atra">Tocama atra atra</taxonomicName>
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Keith, 2006: 225, syn. n.
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</paragraph>
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<paragraph id="45BB759978C8544D345DC83E3C0A8BF7" pageId="5" pageNumber="42">
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<taxonomicName id="828F48EDFFC99168EF6F82CB1B68AC16" class="Insecta" family="Scarabaeidae" genus="Tocama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tocama atra subsp. reichenbachi" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="subSpecies" species="atra" subSpecies="reichenbachi">Tocama atra reichenbachi</taxonomicName>
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Keith, 2007: 338, syn. n.
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</paragraph>
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</subSubSection>
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<subSubSection id="DF4685A3AF345D3EC6BED2685A1A2AA0" pageId="5" pageNumber="42" type="distribution">
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<paragraph id="0AF30A7F3E95B79342DA7D355FCF4578" pageId="5" pageNumber="42">Distribution.</paragraph>
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<paragraph id="2992E438B746FE031C9E3CE962819AD8" pageId="5" pageNumber="42">
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China; Vietnam; Laos (new country record with a male collected in Mt. Phu Pan, Houaphan province, Laos, in the first
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<normalizedToken id="F489CCB20993B23B68BC397CCE948D3F" originalValue="author’s">author's</normalizedToken>
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collections); Myanmar (new country record with two females collected nearby Putao, Kachin state, Myanmar, in the first
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<normalizedToken id="726D717277FA422A581E565F613B630A" originalValue="author’s">author's</normalizedToken>
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collections)
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</paragraph>
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</subSubSection>
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<subSubSection id="B426B1E6EF991773DEADF5291EA11B54" lastPageId="6" lastPageNumber="43" pageId="5" pageNumber="42" type="remarks">
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<paragraph id="ECDE4085653BB3E8256A6F7820E5C6B3" pageId="5" pageNumber="42">Remarks.</paragraph>
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<paragraph id="CC154A89ABC05DA4025C29B276DF909E" lastPageId="6" lastPageNumber="43" pageId="5" pageNumber="42">
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<bibRefCitation id="8B608889E46F58A048080424DB340A1D" author="Moser, J" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="8" pageNumber="45" pagination="271 - 297" title="Beitrag zur kenntnis der Melolonthiden (Col.)." volume="1913" year="1913">Moser (1913)</bibRefCitation>
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described
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<taxonomicName id="33787FFBF4378D57C347DC30B298ACEF" class="Insecta" family="Scarabaeidae" genus="Hoplosternus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplosternus tonkinensis" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="tonkinensis">Hoplosternus tonkinensis</taxonomicName>
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based from a female (labeled as "Tonkin [presently northern Vietnam] Montes Manson April. Mai 2
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<normalizedToken id="42FD39E5B93A9C57EB00DF7156D8441F" originalValue="–3000’">-3000'</normalizedToken>
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H Fruhstorfer (printed) // Hoplosternus tonkinensis Type Mos (handwritten)", currently deposited at Museum
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<normalizedToken id="C0988BFFFB7C2441D13980F4FBE9DA68" originalValue="für">fuer</normalizedToken>
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Naturkunde der Humboldt Universitat (ZMHB)),
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<pageBreakToken id="D22DBD096CE4FBE3A701608C6E6FE5B2" pageId="6" pageNumber="43" start="start">Berlin</pageBreakToken>
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, Germany). Subsequently, he described
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<taxonomicName id="3D3C64755E79746EAF68AC0E2A2F39DE" class="Insecta" family="Scarabaeidae" genus="Hoplosternus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplosternus pygidialis" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="pygidialis">Hoplosternus pygidialis</taxonomicName>
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Moser, 1915, from a male (labeled as "Kiautschou [Guizhou Province] China (printed)// Hoplosternus pygidialis Type Mos (handwritten), deposited at ZMHB). We examined both type specimens and found that they share series of characters of pronotum shape, punctuation, setation, and male genitalia shape; these characters clearly indicate that the specimens are members of
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<taxonomicName id="87244A9085CD9200C1A67415986F2699" class="Insecta" family="Scarabaeidae" genus="Tocama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tocama" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="genus">Tocama</taxonomicName>
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. Moreover, having examined large number of
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<taxonomicName id="8AECFC4E8D5434920CF9D77C94A99559" class="Insecta" family="Scarabaeidae" genus="Tocama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tocama tonkinensis" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="tonkinensis">Tocama tonkinensis</taxonomicName>
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specimens (45 males and 43 females) from Indochina and China, we found that the sexual dimorphism of the species is particularly distinct in the shape of the pygidium, which might be the reason of separation for the previous two species by Julius Moser. We conclude that these two type specimens are conspecific.
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</paragraph>
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<paragraph id="33D86762551AF781F2441B2AEEB6035E" pageId="6" pageNumber="43">
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Additionally, the intraspecific variation of
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<taxonomicName id="661C0E7976F86150371DE12D44F5B8FF" class="Insecta" family="Scarabaeidae" genus="Tocama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tocama tonkinensis" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="tonkinensis">Tocama tonkinensis</taxonomicName>
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is significant in body color. There are two main forms of body color, black and castaneous, that led
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<bibRefCitation id="AD79F67E7BFCC9BE4943A7730233ADCA" author="Keith, D" journalOrPublisher="Lambillionea" pageId="7" pageNumber="44" pagination="223 - 229" title="Sur le genre Tocama Reitter, 1902 (Col. Scarabaeoidea Melolonthidae Melolonthinae)." volume="106" year="2006">Keith (2006</bibRefCitation>
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, 2007) to separation of
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<taxonomicName id="16D4B9FC77FF333197909B0DD437F67B" class="Insecta" family="Scarabaeidae" genus="Tocama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tocama atra subsp. atra" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="atra" subSpecies="atra">Tocama atra atra</taxonomicName>
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(black form) and
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<taxonomicName id="D9BF6F9000779478455A0D0F3011D9C2" class="Insecta" family="Scarabaeidae" genus="Tocama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tocama atra subsp. reichenbachi" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="atra" subSpecies="reichenbachi">Tocama atra reichenbachi</taxonomicName>
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(castaneous form). We consider that they are all within the variability of
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<taxonomicName id="B6F7506BB3764FD7BD4B15FCB911E8DF" class="Insecta" family="Scarabaeidae" genus="Tocama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tocama tonkinensis" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="tonkinensis">Tocama tonkinensis</taxonomicName>
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based on the examination of the type series of
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<taxonomicName id="30D44BF0C3E5D475E8AF34CE9F7BB919" class="Insecta" family="Scarabaeidae" genus="Tocama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tocama atra subsp. atra" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="atra" subSpecies="atra">Tocama atra atra</taxonomicName>
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and
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<taxonomicName id="02AFF5613FE8A7131030F6894BEC1AFA" class="Insecta" family="Scarabaeidae" genus="Tocama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tocama atra subsp. reichenbachi" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="atra" subSpecies="reichenbachi">Tocama atra reichenbachi</taxonomicName>
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(see also Figs 5-6 in
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<bibRefCitation id="7C3D789D06844F1BAB0AF36BEC6EEC71" author="Keith, D" journalOrPublisher="Lambillionea" pageId="7" pageNumber="44" pagination="223 - 229" title="Sur le genre Tocama Reitter, 1902 (Col. Scarabaeoidea Melolonthidae Melolonthinae)." volume="106" year="2006">Keith (2006)</bibRefCitation>
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for the shape of male genitalia). Accordingly, we herein transfer
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<taxonomicName id="A5D882FCA169E503A1B5AEAAB846E2B7" class="Insecta" family="Scarabaeidae" genus="Hoplosternus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplosternus tonkinensis" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="tonkinensis">Hoplosternus tonkinensis</taxonomicName>
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to
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<taxonomicName id="52C54D3378003F05C8FB25C8FF79E914" class="Insecta" family="Scarabaeidae" genus="Tocama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tocama" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="genus">Tocama</taxonomicName>
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and synonymize the names
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<taxonomicName id="86673355109EF9289A8D1C7A5AF9F173" class="Insecta" family="Scarabaeidae" genus="Hoplosternus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplosternus pygidialis" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="pygidialis">Hoplosternus pygidialis</taxonomicName>
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,
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<taxonomicName id="D16887E654FAD5A270037225929153CA" class="Insecta" family="Scarabaeidae" genus="Tocama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tocama atra subsp. atra" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="atra" subSpecies="atra">Tocama atra atra</taxonomicName>
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and
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<taxonomicName id="8BD78EFC0EC93285C5520DED0D831958" class="Insecta" family="Scarabaeidae" genus="Tocama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tocama atra subsp. reichenbachi" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="atra" subSpecies="reichenbachi">Tocama atra reichenbachi</taxonomicName>
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with
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<taxonomicName id="1139CE7E119691CC47BE2E5BE2D5D0B7" class="Insecta" family="Scarabaeidae" genus="Tocama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tocama tonkinensis" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="tonkinensis">Tocama tonkinensis</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |