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<document id="1297DDE31F444208A6C839B7BCF25647" ID-DOI="10.5281/zenodo.158526" ID-GBIF-Dataset="c60c307a-0287-42a2-9b9b-db8bec485488" ID-ISSN="1175­5326" ID-Zenodo-Dep="158526" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461024504380" checkinUser="plazi" docAuthor="Gill, Anthony C. &amp; Allen, Gerald R." docDate="2004" docId="7D10D8198D53FF98902AFBA8FC79FB28" docLanguage="en" docName="zt00604.pdf" docOrigin="Zootaxa 604" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Pseudochromis tonozukai Gill &amp; Allen, 2004, new species" docType="treatment" docVersion="10" lastPageNumber="11" masterDocId="8129A0618D55FF929122FFB8FFECFFE6" masterDocTitle="Pseudochromis lugubris and P. tonozukai, two new dottyback fish species from the Indo­Australian Archipelago (Perciformes: Pseudochromidae: Pseudochrominae)" masterLastPageNumber="12" masterPageNumber="1" pageNumber="7" updateTime="1698571520642" updateUser="plazi">
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<mods:title id="B851A52B39DC272FF3AB59881677BF9E">Pseudochromis lugubris and P. tonozukai, two new dottyback fish species from the Indo­Australian Archipelago (Perciformes: Pseudochromidae: Pseudochrominae)</mods:title>
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<mods:namePart id="BA87FDF0EAFF44D8C4D9F9B8267A4070">Gill, Anthony C.</mods:namePart>
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<mods:namePart id="D577A5E1A99BDD0B1D7E7EE9C249FB73">Allen, Gerald R.</mods:namePart>
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<mods:title id="D3C0C6648EB705E1645D4A1A6DEA00CC">Zootaxa</mods:title>
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<mods:date id="7E08CFD9055CEE5F4D7185A1616AAD3B">2004</mods:date>
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<mods:identifier id="85F1D943533BD313D1BE85B18866827A" type="DOI">10.5281/zenodo.158526</mods:identifier>
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<treatment id="7D10D8198D53FF98902AFBA8FC79FB28" ID-DOI="http://doi.org/10.5281/zenodo.5667623" ID-GBIF-Taxon="119605929" ID-Zenodo-Dep="5667623" LSID="urn:lsid:plazi:treatment:7D10D8198D53FF98902AFBA8FC79FB28" httpUri="http://treatment.plazi.org/id/7D10D8198D53FF98902AFBA8FC79FB28" lastPageId="10" lastPageNumber="11" pageId="6" pageNumber="7">
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<paragraph id="F506690F8D53FF94902AFBA8FD3BFBCC" blockId="6.[264,727,1040,1141]" box="[264,727,1040,1066]" pageId="6" pageNumber="7">
<heading id="AE4EDE638D53FF94902AFBA8FD3BFBCC" bold="true" box="[264,727,1040,1066]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
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<taxonomicName id="32B9128C8D53FF94902AFBA8FDACFBCC" ID-CoL="4NTNQ" box="[264,576,1040,1066]" class="Actinopterygii" family="Pseudochromidae" genus="Pseudochromis" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="tonozukai" status="sp. nov.">
<emphasis id="C7CDB51D8D53FF94902AFBA8FDACFBCC" bold="true" box="[264,576,1040,1066]" italics="true" pageId="6" pageNumber="7">Pseudochromis tonozukai</emphasis>
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<taxonomicNameLabel id="DCFE08668D53FF949365FBA8FD3BFBCC" box="[583,727,1040,1066]" pageId="6" pageNumber="7" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="F506690F8D53FF94902AFB8BFDE6FBAB" blockId="6.[264,727,1040,1141]" box="[264,522,1075,1101]" pageId="6" pageNumber="7">Spot­stripe Dottyback</paragraph>
</subSubSection>
<subSubSection id="BDA33A848D53FF94902AFBE3FAE4FB24" pageId="6" pageNumber="7" type="description">
<paragraph id="F506690F8D53FF94902AFBE3FDC7FB93" blockId="6.[264,727,1040,1141]" box="[264,555,1115,1141]" pageId="6" pageNumber="7">
(
<figureCitation id="6D82758A8D53FF949032FBE3FE6EFB93" box="[272,386,1115,1141]" captionStart="FIGURE 2" captionStartId="6.[264,368,891,915]" captionTargetBox="[390,1198,324,874]" captionTargetId="figure@6.[380,1217,302,900]" captionTargetPageId="6" captionText="FIGURE 2. Distributional records for Pseudochromis bitaeniatus (closed circles; after Gill 2004: fig. 32), P. lugubris (triangles) and P. t o n o z u k a i (square)." httpUri="https://zenodo.org/record/158528/files/figure.png" pageId="6" pageNumber="7">Figures 2</figureCitation>
,
<figureCitation id="6D82758A8D53FF9490ACFBE3FE51FB93" box="[398,445,1115,1141]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="9.[264,367,1276,1300]" captionStartId-1="9.[264,368,1847,1871]" captionTargetBox-0="[378,1208,906,1268]" captionTargetBox-1="[353,1233,1407,1838]" captionTargetId-0="figure@9.[333,1271,903,1270]" captionTargetId-1="figure@9.[316,1286,1405,1840]" captionTargetPageId-0="9" captionTargetPageId-1="9" captionText-0="FIGURE 4. Pseudochromis tonozukai, female, underwater photo, Weh Island, off northern Sumatra, Indonesia. (Photo by T. Tonozuka)." captionText-1="FIGURE 5. Pseudochromis tonozukai, male, underwater photo, Weh Island, off northern Sumatra, Indonesia. (Photo by T. Tonozuka)" httpUri-0="https://zenodo.org/record/169300/files/figure.png" httpUri-1="https://zenodo.org/record/169301/files/figure.png" pageId="6" pageNumber="7">45</figureCitation>
;
<tableCitation id="B83B5CB48D53FF9490E8FBE3FDF3FB93" box="[458,543,1115,1141]" captionStart="TABLE 1" captionStartId="5.[264,352,283,307]" captionTargetBox="[267,1314,529,742]" captionTargetPageId="5" captionText="TABLE 1. Frequency distributions for selected meristic characters of Pseudochromis bitaeniatus (after Gill 2004), P. lugubris and P. t o n o z u k a i. * indicates characters for which bilateral counts are included." httpUri="http://table.plazi.org/id/A1C639878D50FF97902AFEA3FE32FE9E" pageId="6" pageNumber="7" tableUuid="A1C639878D50FF97902AFEA3FE32FE9E">Table 1</tableCitation>
)
</paragraph>
<paragraph id="F506690F8D53FF94902AFB13FAE4FB24" blockId="6.[264,1288,1195,1218]" box="[264,1288,1195,1218]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="D5A94E218D53FF94902AFB13FAE4FB24" box="[264,1288,1195,1218]" pageId="6" pageNumber="7">
<taxonomicName id="32B9128C8D53FF94902AFB13FE47FB24" ID-CoL="6ZLM" box="[264,427,1195,1218]" class="Actinopterygii" family="Pseudochromidae" genus="Pseudochromis" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="C7CDB51D8D53FF94902AFB13FE47FB24" box="[264,427,1195,1218]" italics="true" pageId="6" pageNumber="7">Pseudochromis</emphasis>
</taxonomicName>
sp.;
<treatmentCitation id="74184F1E8D53FF9490FDFB13FD42FB24" author="Michael" box="[479,686,1195,1218]" page="139" pageId="6" pageNumber="7" year="2004">
<bibRefCitation id="912814FE8D53FF9490FDFB13FD90FB24" author="Michael" box="[479,636,1195,1218]" pageId="6" pageNumber="12" refString="Michael, S. W. (2004) Reef Fish Series Book 2. Basslets, Dottybacks and Hawkfishes. T. F. H. Publications, Neptune City, 296 pp." type="book" year="2004">Michael, 2004</bibRefCitation>
: 139
</treatmentCitation>
140 (colour photos; sexual dimorphism; natural history
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BDA33A848D53FF94902AFB4BFB4BFA6B" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="F506690F8D53FF94902AFB4BFE9BFA83" blockId="6.[264,1324,1267,1901]" pageId="6" pageNumber="7">
<emphasis id="C7CDB51D8D53FF94902AFB4BFE9FFAEA" box="[264,371,1267,1292]" italics="true" pageId="6" pageNumber="7">
<typeStatus id="2A02D7AD8D53FF94902AFB4BFE9FFAEA" box="[264,371,1267,1292]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectionCode id="93A8F1CA8D53FF9490A4FB4BFE25FAEB" box="[390,457,1267,1293]" httpUri="http://grbio.org/cool/h4qb-ekf6" name="Pusat Penelitian dan Pengembangan Oseanologi" pageId="6" pageNumber="7">NCIP</collectionCode>
6187,
<quantity id="3241C4EA8D53FF949302FB4BFD67FAEB" box="[544,651,1267,1293]" metricMagnitude="-2" metricUnit="m" metricValue="6.59" pageId="6" pageNumber="7" unit="mm" value="65.9">65.9 mm</quantity>
SL, gravid female,
<collectingCountry id="8DAE299F8D53FF9492A3FB4BFC1AFAEB" box="[897,1014,1267,1293]" name="Indonesia" pageId="6" pageNumber="7">Indonesia</collectingCountry>
, northern Sumatra, Aceh, Weh Island, near Point Seukundo (
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<geoCoordinate id="908D0FC88D53FF9493DDFA9BFC65FADB" box="[767,905,1307,1341]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="15" value="95.2975">95o17'51&quot;E</geoCoordinate>
),
<quantity id="3241C4EA8D53FF949282FA9BFC32FADB" box="[928,990,1315,1341]" metricMagnitude="1" metricUnit="m" metricValue="3.6" pageId="6" pageNumber="7" unit="m" value="36.0">36 m</quantity>
, spear, G.R. Allen,
<date id="81074FCF8D53FF9495E4FA9BFE9FFA83" pageId="6" pageNumber="7" value="1999-01-27">27 January 1999</date>
.
</paragraph>
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<emphasis id="C7CDB51D8D53FF94901AFACBFE4FFA6A" box="[312,419,1395,1420]" italics="true" pageId="6" pageNumber="7">
<typeStatus id="2A02D7AD8D53FF94901AFACBFE4FFA6A" box="[312,419,1395,1420]" pageId="6" pageNumber="7" type="paratype">Paratype</typeStatus>
</emphasis>
:
<collectionCode id="93A8F1CA8D53FF949090FACBFE16FA6B" LSID="urn:lsid:biocol.org:col:34981" box="[434,506,1395,1421]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="6" pageNumber="7">WAM</collectionCode>
P.31526­001,
<quantity id="3241C4EA8D53FF949381FACBFCE0FA6B" box="[675,780,1395,1421]" metricMagnitude="-2" metricUnit="m" metricValue="6.26" pageId="6" pageNumber="7" unit="mm" value="62.6">62.6 mm</quantity>
SL, male, collected with
<typeStatus id="2A02D7AD8D53FF94951AFACBFB4FFA6B" box="[1080,1187,1395,1421]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="BDA33A848D53FF94901AFA23FE2FF9CB" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="F506690F8D53FF94901AFA23FE2FF9CB" blockId="6.[264,1324,1267,1901]" pageId="6" pageNumber="7">
<emphasis id="C7CDB51D8D53FF94901AFA23FE43FA52" box="[312,431,1435,1460]" italics="true" pageId="6" pageNumber="7">Diagnosis</emphasis>
. The following combination of characters distinguishes
<taxonomicName id="32B9128C8D53FF94957AFA23FB06FA52" box="[1112,1258,1435,1460]" class="Actinopterygii" family="Pseudochromidae" genus="Pseudochromis" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="tonozukai">
<emphasis id="C7CDB51D8D53FF94957AFA23FB06FA52" box="[1112,1258,1435,1460]" italics="true" pageId="6" pageNumber="7">P. tonozukai</emphasis>
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from all other pseudochromines: dorsal­fin rays III,25; anal­fin rays III,1415; scales in lateral series 3436; caudal fin rhomboid to trifurcate; and upper part of pectoral fin without small dark spot.
</paragraph>
</subSubSection>
<subSubSection id="BDA33A848D53FF98901AF983FE62FDA8" lastPageId="10" lastPageNumber="11" pageId="6" pageNumber="7" type="description">
<paragraph id="F506690F8D53FF95901AF983FBF3FEB8" blockId="6.[264,1324,1267,1901]" lastBlockId="7.[264,1325,284,1870]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="C7CDB51D8D53FF94901AF983FE2EF9B2" box="[312,450,1595,1620]" italics="true" pageId="6" pageNumber="7">Description</emphasis>
. Dorsal­fin rays III,25, all segmented rays branched; anal­fin rays III,14 15 (III,15), all segmented rays branched; pectoral­fin rays 1718 (18/18); upper procurrent caudal­fin rays 6; lower procurrent caudal­fin rays 56 (5); total caudal­fin rays 2829 (28); scales in lateral series 3436 (35/36); anterior lateral­line scales 2527 (26/27); anterior lateral line terminating beneath segmented dorsal­fin ray 1819 (19/18); posterior lateral­line scales 713 + 01 (7 + 0/7 + 0); scales between lateral lines 3; horizontal scale rows above anal­fin origin 1114 + 1 + 3 (11 + 1 + 3/12 + 1 + 3); circumpeduncular scales 16; predorsal scales 21; scales behind eye 3; scales to preopercular angle 5; gill rakers 6 + 12; pseudobranch filaments 1112 (11); circumorbital pores 2431 (24/25); preopercular pores 1224 (13/12); dentary pores 4; posterior interorbital pores 1.
</paragraph>
<paragraph id="F506690F8D52FF95901AFED4FCAFFD08" blockId="7.[264,1325,284,1870]" pageId="7" pageNumber="8">Lower lip incomplete; dorsal and anal fins without distinct scale sheaths, though with scales intermittently overlapping fin bases; predorsal scales extending anteriorly to posterior nasal pores; opercle with 5 moderately distinct serrations; teeth of outer ceratobranchial­1 gill rakers well developed on raker tips only; anterior dorsal­fin pterygiophore formula S/S/S + 3/1 + 1/1/1/1/1/1 + 1/1; dorsal­fin spines moderately stout and pungent; anterior anal­fin pterygiophore formula 3/1 + 1/1/1 + 1/1/1 + 1; anal­fin spines moderately stout and pungent, the second spine stouter than the third; pelvic­fin spine moderately stout and pungent; second segmented pelvic­fin ray longest; caudal fin angular with upper, lower and mid rays weakly to strongly produced to give rhomboid to trifurcate shape; vertebrae 10 + 16; epineurals 1315 (15); epurals 3.</paragraph>
<paragraph id="F506690F8D52FF95901AFD44FACBFBC8" blockId="7.[264,1325,284,1870]" pageId="7" pageNumber="8">Upper jaw with 2 pairs of curved, enlarged caniniform teeth anteriorly, medial pair smallest, and 6 or 7 (at symphysis) to 2 or 3 (on sides of jaw) inner rows of small conical teeth; lower jaw with 1 or 2 pairs of curved, enlarged caniniform teeth anteriorly, medial pair smallest, and 4 or 5 (at symphysis) to 1 (on sides of jaw) inner rows of small, conical teeth, teeth on middle of jaw larger and curved; vomer with 2 rows of small conical teeth, forming chevron; palatine with 13 rows of small conical teeth arranged in elongate, suboval patch, anterior part of the tooth patch more­or­less contiguous with posterolateral arm of vomerine tooth patch; ectopterygoid edentate; tongue moderately pointed and edentate.</paragraph>
<paragraph id="F506690F8D52FF95901AFB84FB16F8A8" blockId="7.[264,1325,284,1870]" pageId="7" pageNumber="8">As percentage of SL: head length 23.624.4 (24.4); orbit diameter 8.18.3 (8.3); snout length 6.26.4 (6.4); fleshy interorbital width 5.55.9 (5.9); bony interorbital width 3.8 4.6 (4.6); body width 11.511.7 (11.5); snout tip to posterior tip of retroarticular bone 13.814.1 (13.8); predorsal length 31.632.6 (32.6); prepelvic length 30.730.8 (30.8); posterior tip of retroarticular bone to pelvic­fin origin 17.9; dorsal­fin origin to pelvic­fin origin 27.628.2 (28.2); dorsal­fin origin to middle dorsal­fin ray 32.334.6 (34.6); dorsal­fin origin to anal­fin origin 41.141.4 (41.4); pelvic­fin origin to anal­fin origin 28.8 30.2 (28.8); middle dorsal­fin ray to dorsal­fin termination 26.727.0 (27.0); middle dorsal­fin ray to anal­fin origin 24.425.0 (25.0); anal­fin origin to dorsal­fin termination 33.834.8 (33.8); anal­fin base length 27.528.4 (27.5); dorsal­fin termination to anal­fin termination 14.514.7 (14.7); dorsal­fin termination to caudal peduncle dorsal edge 12.3 12.7 (12.7); dorsal­fin termination to caudal peduncle ventral edge 18.719.0 (19.0); anal­fin termination to caudal peduncle dorsal edge 19.920.0 (19.9); anal­fin termination to caudal peduncle ventral edge 13.514.1 (13.5); first dorsal­fin spine 1.82.4 (2.4); second dorsal­fin spine 4.55.6 (5.6); third dorsal­fin spine 7.38.0 (8.0); first segmented dorsal­fin ray 12.612.8 (12.6); fourth last segmented dorsal­fin ray 16.818.5 (16.8); first anal­fin spine 1.62.6 (2.6); second anal­fin spine 4.25.0 (5.0); third anal­fin spine 6.7 7.3 (7.3); first segmented anal­fin ray 9.111.4 (11.4); fourth last segmented anal­fin ray 15.817.4 (15.8); third pectoral­fin ray 14.314.5 (14.3); pelvic­fin spine 10.510.9 (10.9); second segmented pelvic­fin ray 29.330.7 (29.3); caudal­fin length 29.234.7 (34.7).</paragraph>
<paragraph id="F506690F8D5DFF9A901AFEA4FD61F9B8" blockId="8.[264,1324,284,1870]" pageId="8" pageNumber="9">
Live coloration (based on underwater photographs of individuals taken at the
<typeStatus id="2A02D7AD8D5DFF9A95DAFEA4FAC7FED0" box="[1272,1323,284,310]" pageId="8" pageNumber="9">type</typeStatus>
locality by T. Tonozuka). Females (
<figureCitation id="6D82758A8D5DFF9A9384FEFCFD02FEB8" box="[678,750,324,350]" captionStart="FIGURE 4" captionStartId="9.[264,367,1276,1300]" captionTargetBox="[378,1208,906,1268]" captionTargetId="figure@9.[333,1271,903,1270]" captionTargetPageId="9" captionText="FIGURE 4. Pseudochromis tonozukai, female, underwater photo, Weh Island, off northern Sumatra, Indonesia. (Photo by T. Tonozuka)." httpUri="https://zenodo.org/record/169300/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
): head grey, becoming pale pink ventrally; indistinct reddish to maroon stripe extending from posterior edge of eye to opercle, with scattered blue to mauve spots within stripe; lower part of stripe on operculum with fine gold vermiculations; blue to mauve curved streak extending from mid­posterior edge of orbit around outer edge of infraorbital stripe; iris yellowish grey, with bright blue suboval ring around pupil; dorsal contour of body grey, becoming yellowish grey posteriorly; edges of scales within grey to yellowish grey area dark grey, forming chequered to reticulate pattern; remainder of body white with dark reddish grey to dark grey stripe extending from maroon stripe on head to mid­lower part of caudal fin, becoming brown to black on fin; scales within stripe each with dark blue to mauve basal spot; dorsal fin yellowish grey on basal half, remainder of fin yellowish hyaline with indistinct mauve spots and mauve distal margin; anal fin hyaline, narrowly pale blue on base, with distal margin pale blue; brown to black stripe on caudal fin broadly bordered with white on dorsal edge and pale grey on ventral edge; white area bordered dorsally with dark yellowish grey stripe, this converging posteriorly with lower brown to black stripe; remainder of caudal fin yellowish hyaline dorsally and mauvish hyaline ventrally; pelvic fin pale blue to white basally, remainder of fin hyaline; pectoral fin pinkish hyaline. Males (
<figureCitation id="6D82758A8D5DFF9A9271FC7CFC72FC38" box="[851,926,964,990]" captionStart="FIGURE 5" captionStartId="9.[264,368,1847,1871]" captionTargetBox="[353,1233,1407,1838]" captionTargetId="figure@9.[316,1286,1405,1840]" captionTargetPageId="9" captionText="FIGURE 5. Pseudochromis tonozukai, male, underwater photo, Weh Island, off northern Sumatra, Indonesia. (Photo by T. Tonozuka)" httpUri="https://zenodo.org/record/169301/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
): head grey, becoming pale pink ventrally; indistinct reddish to maroon stripe extending from posterior edge of eye to opercle, with scattered blue to mauve spots within stripe; lower part of stripe on operculum with fine gold vermiculations; blue to mauve curved streak extending from mid­posterior edge of orbit around outer edge of infraorbital stripe; iris yellowish grey, with turquoise suboval ring around pupil; dorsal contour of body grey, edges of scales within grey area dark grey, forming chequered to reticulate pattern; remainder of body white with broad maroon (anteriorly) to orange stripe extending from maroon stripe on head to mid­lower part of caudal fin, becoming brown to black on posterior part of caudal fin; scales within stripe each with dark blue to mauve basal spot; dorsal fin greyish hyaline on basal half, remainder of fin hyaline with indistinct mauve spots and mauve distal margin; anal fin hyaline, narrowly pale blue on base, with distal margin indistinctly mauve; orange to black stripe on caudal fin bordered with white, broadly on dorsal edge and narrowly on ventral edge; upper white area bordered dorsally with yellowish grey stripe, this converging posteriorly with lower orange to black stripe; remainder of caudal fin yellowish hyaline dorsally and mauvish hyaline ventrally; pelvic fin pale blue to white basally, remainder of fin hyaline; pectoral fin pinkish hyaline.
</paragraph>
<paragraph id="F506690F8D5DFF9B901AF9D4FEA3FEB8" blockId="8.[264,1324,284,1870]" lastBlockId="9.[264,1322,284,350]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">Preserved coloration. Pattern similar to live coloration: pale areas on head, body and fins become pale yellow to pale brown; blue to mauve spots on head and body become brown to dark grey; dorsal part of head and body become greyish brown, with dark reticulations on dorsal part of body becoming dark grey­brown; dark reddish grey to dark grey stripe on female becoming dark greyish brown to dark grey; maroon to orange stripe on male becoming dark greyish brown and distinct anteriorly, and dusky brown and indistinct posteriorly; dark stripes on caudal fin remain, becoming greyish brown to dark grey or black.</paragraph>
<caption id="A1C639878D5CFF9B902AFCB0FE79FCA4" httpUri="https://zenodo.org/record/169299/files/figure.png" pageId="9" pageNumber="10" targetBox="[370,1216,433,765]" targetPageId="9">
<paragraph id="F506690F8D5CFF9B902AFCB0FE79FCA4" blockId="9.[264,1323,776,834]" pageId="9" pageNumber="10">
<emphasis id="C7CDB51D8D5CFF9B902AFCB0FE62FCC6" bold="true" box="[264,398,776,800]" pageId="9" pageNumber="10">FIGURE 3.</emphasis>
<taxonomicName id="32B9128C8D5CFF9B90B8FCB0FD52FCF9" box="[410,702,776,799]" class="Actinopterygii" family="Pseudochromidae" genus="Pseudochromis" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="bitaeniatus">
<emphasis id="C7CDB51D8D5CFF9B90B8FCB0FD52FCF9" box="[410,702,776,799]" italics="true" pageId="9" pageNumber="10">Pseudochromis bitaeniatus</emphasis>
</taxonomicName>
, underwater photo, Mabul, Sabah, Malaysia. (Photo by R.H. Kuiter).
</paragraph>
</caption>
<caption id="A1C639878D5CFF9B902AFB44FD0BFAD0" httpUri="https://zenodo.org/record/169300/files/figure.png" pageId="9" pageNumber="10" targetBox="[378,1208,906,1268]" targetPageId="9">
<paragraph id="F506690F8D5CFF9B902AFB44FD0BFAD0" blockId="9.[264,1322,1276,1334]" pageId="9" pageNumber="10">
<emphasis id="C7CDB51D8D5CFF9B902AFB44FE7EFAF2" bold="true" box="[264,402,1276,1300]" pageId="9" pageNumber="10">FIGURE 4.</emphasis>
<taxonomicName id="32B9128C8D5CFF9B9081FB45FD50FAF2" box="[419,700,1277,1300]" class="Actinopterygii" family="Pseudochromidae" genus="Pseudochromis" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="tonozukai">
<emphasis id="C7CDB51D8D5CFF9B9081FB45FD50FAF2" box="[419,700,1277,1300]" italics="true" pageId="9" pageNumber="10">Pseudochromis tonozukai</emphasis>
</taxonomicName>
, female, underwater photo, Weh Island, off northern Sumatra, Indonesia. (Photo by T. Tonozuka).
</paragraph>
</caption>
<caption id="A1C639878D5CFF9B902AF88FFD95F897" httpUri="https://zenodo.org/record/169301/files/figure.png" pageId="9" pageNumber="10" targetBox="[353,1233,1407,1838]" targetPageId="9">
<paragraph id="F506690F8D5CFF9B902AF88FFD95F897" blockId="9.[264,1322,1847,1905]" pageId="9" pageNumber="10">
<emphasis id="C7CDB51D8D5CFF9B902AF88FFE67F8A9" bold="true" box="[264,395,1847,1871]" pageId="9" pageNumber="10">FIGURE 5.</emphasis>
<taxonomicName id="32B9128C8D5CFF9B90B0F88FFD48F8A8" box="[402,676,1847,1870]" class="Actinopterygii" family="Pseudochromidae" genus="Pseudochromis" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="tonozukai">
<emphasis id="C7CDB51D8D5CFF9B90B0F88FFD48F8A8" box="[402,676,1847,1870]" italics="true" pageId="9" pageNumber="10">Pseudochromis tonozukai</emphasis>
</taxonomicName>
, male, underwater photo, Weh Island, off northern Sumatra, Indonesia. (Photo by T. Tonozuka)
</paragraph>
</caption>
<paragraph id="F506690F8D5FFF98901AFEA4FC0EFE30" blockId="10.[264,1323,284,1230]" pageId="10" pageNumber="11">
<emphasis id="C7CDB51D8D5FFF98901AFEA4FDB6FED3" box="[312,602,284,309]" italics="true" pageId="10" pageNumber="11">Habitat and distribution</emphasis>
.
<taxonomicName id="32B9128C8D5FFF989349FEA4FC76FED3" box="[619,922,284,309]" class="Actinopterygii" family="Pseudochromidae" genus="Pseudochromis" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="tonozukai">
<emphasis id="C7CDB51D8D5FFF989349FEA4FC76FED3" box="[619,922,284,309]" italics="true" pageId="10" pageNumber="11">Pseudochromis tonozukai</emphasis>
</taxonomicName>
is known only from Weh Island, off northern Sumatra,
<collectingCountry id="8DAE299F8D5FFF989330FEFCFD68FEB8" box="[530,644,324,350]" name="Indonesia" pageId="10" pageNumber="11">Indonesia</collectingCountry>
(
<figureCitation id="6D82758A8D5FFF9893B4FEFCFD33FEB8" box="[662,735,324,350]" captionStart="FIGURE 2" captionStartId="6.[264,368,891,915]" captionTargetBox="[390,1198,324,874]" captionTargetId="figure@6.[380,1217,302,900]" captionTargetPageId="6" captionText="FIGURE 2. Distributional records for Pseudochromis bitaeniatus (closed circles; after Gill 2004: fig. 32), P. lugubris (triangles) and P. t o n o z u k a i (square)." httpUri="https://zenodo.org/record/158528/files/figure.png" pageId="10" pageNumber="11">Fig. 2</figureCitation>
). It was observed and collected in
<quantity id="3241C4EA8D5FFF9895A1FEFCFB1CFEB8" box="[1155,1264,324,350]" metricMagnitude="1" metricUnit="m" metricValue="3.8" metricValueMax="4.0" metricValueMin="3.6" pageId="10" pageNumber="11" unit="m" value="38.0" valueMax="40.0" valueMin="36.0">
36
<specimenCount id="E3BFA2868D5FFF98958DFEFCFB1CFEB8" box="[1199,1264,324,350]" pageId="10" pageNumber="11" type="generic">40 m</specimenCount>
</quantity>
. The habitat consisted of a 45° slope with mixed sand­rubble substrate. Approximately 10 individuals were observed in the proximity of dead coral outcrops. Several of these were solitary individuals, but at least four male­female pairs were seen.
</paragraph>
<paragraph id="F506690F8D5FFF98901AFE5CFE62FDA8" blockId="10.[264,1323,284,1230]" pageId="10" pageNumber="11">
<emphasis id="C7CDB51D8D5FFF98901AFE5CFE38FE1B" box="[312,468,484,509]" italics="true" pageId="10" pageNumber="11">Comparisons</emphasis>
.
<taxonomicName id="32B9128C8D5FFF9890C6FE5CFCFFFE1B" box="[484,787,484,509]" class="Actinopterygii" family="Pseudochromidae" genus="Pseudochromis" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="tonozukai">
<emphasis id="C7CDB51D8D5FFF9890C6FE5CFCFFFE1B" box="[484,787,484,509]" italics="true" pageId="10" pageNumber="11">Pseudochromis tonozukai</emphasis>
</taxonomicName>
appears to be related to
<taxonomicName id="32B9128C8D5FFF98951BFE5CFB35FE1B" box="[1081,1241,484,509]" class="Actinopterygii" family="Pseudochromidae" genus="Pseudochromis" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="bitaeniatus">
<emphasis id="C7CDB51D8D5FFF98951BFE5CFB35FE1B" box="[1081,1241,484,509]" italics="true" pageId="10" pageNumber="11">P. bitaeniatus</emphasis>
</taxonomicName>
and
<taxonomicName id="32B9128C8D5FFF989437FE5CFE85FDC3" class="Actinopterygii" family="Pseudochromidae" genus="Pseudochromis" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="lugubris">
<emphasis id="C7CDB51D8D5FFF989437FE5CFE85FDC3" italics="true" pageId="10" pageNumber="11">P. lugubris</emphasis>
</taxonomicName>
. Characters distinguishing the three species are discussed under Comparisons for
<taxonomicName id="32B9128C8D5FFF98902AFD8CFE6AFDAB" box="[264,390,564,589]" class="Actinopterygii" family="Pseudochromidae" genus="Pseudochromis" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="lugubris">
<emphasis id="C7CDB51D8D5FFF98902AFD8CFE6AFDAB" box="[264,390,564,589]" italics="true" pageId="10" pageNumber="11">P. lugubris</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="BDA33A848D5FFF98901AFDE4FDCCFB98" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="F506690F8D5FFF98901AFDE4FBCCFC80" blockId="10.[264,1323,284,1230]" pageId="10" pageNumber="11">
<taxonomicName id="32B9128C8D5FFF98901AFDE4FD8BFD93" ID-CoL="4NTNQ" box="[312,615,604,629]" class="Actinopterygii" family="Pseudochromidae" genus="Pseudochromis" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="tonozukai">
<emphasis id="C7CDB51D8D5FFF98901AFDE4FD8BFD93" box="[312,615,604,629]" italics="true" pageId="10" pageNumber="11">Pseudochromis tonozukai</emphasis>
</taxonomicName>
might also be confused with
<taxonomicName id="32B9128C8D5FFF9892E9FDE4FE7CFD78" ID-CoL="4NTN3" authority="Gill and Woodland (1992)" authorityName="Gill and Woodland" authorityYear="1992" class="Actinopterygii" family="Pseudochromidae" genus="Pseudochromis" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="reticulatus">
<emphasis id="C7CDB51D8D5FFF9892E9FDE4FB8BFD93" box="[971,1127,604,629]" italics="true" pageId="10" pageNumber="11">P. reticulatus</emphasis>
<bibRefCitation id="912814FE8D5FFF989552FDE4FE7CFD78" author="Gill" pageId="10" pageNumber="12" refString="Gill, A. C. &amp; Woodland, D. J. (1992) Description of a new dottyback of the genus Pseudochromis (Pisces: Pseudochromidae) from Western Australia. Records of the Australian Museum, 44 (3), 247 - 251." type="journal article" year="1992">Gill and Woodland (1992)</bibRefCitation>
</taxonomicName>
from northwestern
<collectingCountry id="8DAE299F8D5FFF98935EFD3CFD0BFD78" box="[636,743,644,670]" name="Australia" pageId="10" pageNumber="11">Australia</collectingCountry>
and
<taxonomicName id="32B9128C8D5FFF989206FD3CFB47FD78" ID-CoL="4NTMP" authority="Gill and Randall (1998)" authorityName="Gill and Randall" authorityYear="1998" box="[804,1195,644,670]" class="Actinopterygii" family="Pseudochromidae" genus="Pseudochromis" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="pictus">
<emphasis id="C7CDB51D8D5FFF989206FD3CFC64FD7B" box="[804,904,644,669]" italics="true" pageId="10" pageNumber="11">P. pictus</emphasis>
<bibRefCitation id="912814FE8D5FFF9892ADFD3CFB47FD78" author="Gill" box="[911,1195,644,670]" pageId="10" pageNumber="12" refString="Gill, A. C. &amp; Randall, J. E. (1998) Five new species of the dottyback genus Pseudochromis from Indonesia (Teleostei: Pseudochromidae). Revue francaise d'Aquariologie Herpetologie, 25 (1 - 2), 17 - 26." type="journal article" year="1998">Gill and Randall (1998)</bibRefCitation>
</taxonomicName>
from Alor Island,
<collectingCountry id="8DAE299F8D5FFF98907DFD14FE39FD20" box="[351,469,684,710]" name="Indonesia" pageId="10" pageNumber="11">Indonesia</collectingCountry>
. Both of these species have a pointed caudal fin (rounded with the middle rays produced) and are pale with dark reticulations on the upper part of the body. However, neither species has a stripe on the lower part of the body, and both have more segmented dorsal­fin rays than
<taxonomicName id="32B9128C8D5FFF989358FC9CFCE6FCDB" ID-CoL="4NTNQ" box="[634,778,804,829]" class="Actinopterygii" family="Pseudochromidae" genus="Pseudochromis" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="tonozukai">
<emphasis id="C7CDB51D8D5FFF989358FC9CFCE6FCDB" box="[634,778,804,829]" italics="true" pageId="10" pageNumber="11">P. tonozukai</emphasis>
</taxonomicName>
(2627 versus 25).
<taxonomicName id="32B9128C8D5FFF9892D7FC9CFB1AFCDB" ID-CoL="4NTMP" box="[1013,1270,804,829]" class="Actinopterygii" family="Pseudochromidae" genus="Pseudochromis" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="pictus">
<emphasis id="C7CDB51D8D5FFF9892D7FC9CFB1AFCDB" box="[1013,1270,804,829]" italics="true" pageId="10" pageNumber="11">Pseudochromis pictus</emphasis>
</taxonomicName>
further differs in having more circumpeduncular scales (20 versus 16).
</paragraph>
<paragraph id="F506690F8D5FFF98901AFCCCFDCCFB98" blockId="10.[264,1323,284,1230]" pageId="10" pageNumber="11">
Due largely to the unusual caudal­fin shape, specimens of
<taxonomicName id="32B9128C8D5FFF9892DFFCCCFB63FC6B" ID-CoL="4NTNQ" box="[1021,1167,884,909]" class="Actinopterygii" family="Pseudochromidae" genus="Pseudochromis" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="tonozukai">
<emphasis id="C7CDB51D8D5FFF9892DFFCCCFB63FC6B" box="[1021,1167,884,909]" italics="true" pageId="10" pageNumber="11">P. tonozukai</emphasis>
</taxonomicName>
will not key unambiguously beyond couplet 52 of Gills (2004) key to
<taxonomicName id="32B9128C8D5FFF9892F6FC24FB64FC53" ID-CoL="6ZLM" box="[980,1160,924,949]" class="Actinopterygii" family="Pseudochromidae" genus="Pseudochromis" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="C7CDB51D8D5FFF9892F6FC24FB64FC53" box="[980,1160,924,949]" italics="true" pageId="10" pageNumber="11">Pseudochromis</emphasis>
</taxonomicName>
species. It is readily distinguished from species at or following that couplet in having the following character combination: caudal fin rhomboid to trifurcate; segmented dorsal­fin rays 25; segmented anal­fin rays 1415; and upper part of pectoral fin without small dark spot. The live and preserved colorations also readily distinguish
<taxonomicName id="32B9128C8D5FFF9892A0FB84FBFDFBB3" ID-CoL="4NTNQ" box="[898,1041,1084,1109]" class="Actinopterygii" family="Pseudochromidae" genus="Pseudochromis" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="tonozukai">
<emphasis id="C7CDB51D8D5FFF9892A0FB84FBFDFBB3" box="[898,1041,1084,1109]" italics="true" pageId="10" pageNumber="11">P. tonozukai</emphasis>
</taxonomicName>
from these and all other pseudochromid species.
</paragraph>
</subSubSection>
<subSubSection id="BDA33A848D5FFF98901AFB34FC79FB28" pageId="10" pageNumber="11" type="discussion">
<paragraph id="F506690F8D5FFF98901AFB34FC79FB28" blockId="10.[264,1323,284,1230]" pageId="10" pageNumber="11">
<emphasis id="C7CDB51D8D5FFF98901AFB34FE72FB43" box="[312,414,1164,1189]" italics="true" pageId="10" pageNumber="11">Remarks</emphasis>
. The specific epithet is for Takamasa Tonozuka, who first discovered the species and brought it to the attention of the second author.
</paragraph>
</subSubSection>
</treatment>
</document>