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<document id="40A2140F88AEFAB80CB0BF68BFE2EDE3" ID-DOI="10.5252/geodiversitas2020v42a3" ID-GBIF-Dataset="80d102d3-ab38-4035-9e03-f5f04022aa5a" ID-ISSN="1638-9395" ID-Zenodo-Dep="3696928" ID-ZooBank="urn:lsid:zoobank.org:pub:344FC2F5-A395-449E-915A-EAC53F366764" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583333301723" checkinUser="valdenar" docAuthor="Wang, Xiaoming, Tseng, Z. Jack, Wu, Wen-yu, Ye, Jie, Meng, Jin &amp; Bi, Shundong" docDate="2020" docId="7C6787D2FFF4FFFBC635EB5DFA573A1B" docLanguage="en" docName="GEODIVERSITAS.42.3.29-45.pdf.imf" docOrigin="Geodiversitas 42 (3)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Tungurictis small sp. Colbert 1939" docType="treatment" docVersion="8" lastPageNumber="40" masterDocId="805EFFAAFFF8FFF6C235ED08FFC23D15" masterDocTitle="A new species of Tungurictis Colbert, 1939 (Carnivora, Hyaenidae) from the middle Miocene of Junggar Basin, northwestern China and the early divergence of basal hyaenids in East Asia" masterLastPageNumber="45" masterPageNumber="29" pageNumber="39" updateTime="1698724711154" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="90495EFA603AF2D51666DAE6EEDA045B">A new species of Tungurictis Colbert, 1939 (Carnivora, Hyaenidae) from the middle Miocene of Junggar Basin, northwestern China and the early divergence of basal hyaenids in East Asia</mods:title>
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<mods:namePart id="C4BCD355B140804B6E25BBFF470D7A1D">Wang, Xiaoming</mods:namePart>
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<mods:namePart id="9EFBE4AF2128BA7CC988FD7FFB530BF7">Tseng, Z. Jack</mods:namePart>
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<mods:namePart id="B1501714A5624BE26317795E7C3D0A86">Wu, Wen-yu</mods:namePart>
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<mods:namePart id="951C55DEAD8AC0FFF673B611B6BCD6EB">Ye, Jie</mods:namePart>
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<mods:namePart id="BC86F515C629ED97C2A55C0AE87D291D">Meng, Jin</mods:namePart>
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<mods:namePart id="61260BD5206CD2DB79619438A944E73B">Bi, Shundong</mods:namePart>
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<treatment id="7C6787D2FFF4FFFBC635EB5DFA573A1B" ID-DOI="http://doi.org/10.5281/zenodo.3703736" ID-GBIF-Taxon="162385034" ID-Zenodo-Dep="3703736" LSID="urn:lsid:plazi:treatment:7C6787D2FFF4FFFBC635EB5DFA573A1B" httpUri="http://treatment.plazi.org/id/7C6787D2FFF4FFFBC635EB5DFA573A1B" lastPageId="13" lastPageNumber="40" pageId="12" pageNumber="39">
<subSubSection id="BCD4654FFFF4FFFAC635EB5DFB1E3B7A" box="[1024,1244,1620,1647]" pageId="12" pageNumber="39" type="nomenclature">
<paragraph id="F47136C4FFF4FFFAC635EB5DFB1E3B7A" blockId="12.[1024,1244,1620,1679]" box="[1024,1244,1620,1647]" pageId="12" pageNumber="39">
<heading id="AF3981A8FFF4FFFAC635EB5DFB1E3B7A" box="[1024,1244,1620,1647]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="39" reason="2">
<taxonomicName id="33CE4D47FFF4FFFAC635EB5DFB1E3B7A" authorityName="Colbert" authorityYear="1939" box="[1024,1244,1620,1647]" class="Mammalia" family="Hyaenidae" genus="Tungurictis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="12" pageNumber="32" phylum="Chordata" rank="species" species="small sp.">
<emphasis id="C6BAEAD6FFF4FFFAC635EB5DFBBA3B7B" box="[1024,1144,1621,1646]" italics="true" pageId="12" pageNumber="39">Tungurictis</emphasis>
small sp.
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="BCD4654FFFF4FFFBC673EB7CFA573A1B" lastPageId="13" lastPageNumber="40" pageId="12" pageNumber="39" type="description">
<paragraph id="F47136C4FFF4FFFAC673EB7CFB543B9A" blockId="12.[1024,1244,1620,1679]" box="[1094,1174,1652,1679]" pageId="12" pageNumber="39">
(
<figureCitation id="6CF52A41FFF4FFFAC67BEB7CFB4F3B9A" box="[1102,1165,1652,1679]" captionStart="FIG" captionStartId="12.[132,143,1351,1369]" captionTargetBox="[450,1143,224,1307]" captionTargetId="figure@12.[435,1153,214,1312]" captionTargetPageId="12" captionText="FIG. 7. — Tungurictis small sp., IVPP V 11497, left dentary fragment with m1 and m2 alveolus. A, stereo photos of occlusal view; B, lingual view; C, buccal view. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3696942" httpUri="https://zenodo.org/record/3696942/files/figure.png" pageId="12" pageNumber="39">Fig. 7</figureCitation>
)
</paragraph>
<paragraph id="F47136C4FFF4FFFAC118EBA5FA813BD0" blockId="12.[813,1347,1709,1734]" box="[813,1347,1709,1734]" pageId="12" pageNumber="39">
<emphasis id="C6BAEAD6FFF4FFFAC118EBA5FC7F3BD0" box="[813,957,1709,1733]" italics="true" pageId="12" pageNumber="39">Protictitherium</emphasis>
small sp.
<bibRefCitation id="905F4B35FFF4FFFAC605EBA6FAC93BD0" author="WANG X. &amp; YE J. &amp; MENG J. &amp; WU W. - Y. &amp; LIU L. - P. &amp; BI S. - D." box="[1072,1291,1709,1734]" pageId="12" pageNumber="39" pagination="218 - 243" refId="ref12118" refString="WANG X., YE J., MENG J., WU W. - Y., LIU L. - P. &amp; BI S. - D. 1998. - Carnivore from middle Miocene of Northern Junggar Basin, Xinjiang Autonomous Region, China. Vertebrata PalAsiatica 36: 218 - 243." type="journal article" year="1998">
Wang
<emphasis id="C6BAEAD6FFF4FFFAC65AEBA6FB5F3BD0" box="[1135,1181,1709,1733]" italics="true" pageId="12" pageNumber="39">et al.</emphasis>
1998: 221
</bibRefCitation>
, 236.
</paragraph>
<paragraph id="F47136C4FFF4FFFAC118EBD4FAE03A4B" blockId="12.[813,1455,1756,1886]" pageId="12" pageNumber="39">
REFERRED SPECIMENS. — IVPP
<accessionNumber id="EB9DAB27FFF4FFFAC644EBD5FB0F3BE0" box="[1137,1229,1757,1781]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/V11497" pageId="12" pageNumber="39" type="EnaNcbi">V 11497</accessionNumber>
, left dentary fragment with m1 and m2 alveolus from Botamoyin, lower part of Halamagai Formation (
<bibRefCitation id="905F4B35FFF4FFFAC197EA1AFB8A3A3F" author="WANG X. &amp; YE J. &amp; MENG J. &amp; WU W. - Y. &amp; LIU L. - P. &amp; BI S. - D." box="[930,1096,1809,1834]" pageId="12" pageNumber="39" pagination="218 - 243" refId="ref12118" refString="WANG X., YE J., MENG J., WU W. - Y., LIU L. - P. &amp; BI S. - D. 1998. - Carnivore from middle Miocene of Northern Junggar Basin, Xinjiang Autonomous Region, China. Vertebrata PalAsiatica 36: 218 - 243." type="journal article" year="1998">
Wang
<emphasis id="C6BAEAD6FFF4FFFAC1D4EA1AFBD23A3C" box="[993,1040,1809,1833]" italics="true" pageId="12" pageNumber="39">et al.</emphasis>
1998
</bibRefCitation>
: fig. 3C) (
<figureCitation id="6CF52A41FFF4FFFAC698EA19FB263A3C" box="[1197,1252,1809,1833]" captionStart="FIG" captionStartId="12.[132,143,1351,1369]" captionTargetBox="[450,1143,224,1307]" captionTargetId="figure@12.[435,1153,214,1312]" captionTargetPageId="12" captionText="FIG. 7. — Tungurictis small sp., IVPP V 11497, left dentary fragment with m1 and m2 alveolus. A, stereo photos of occlusal view; B, lingual view; C, buccal view. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3696942" httpUri="https://zenodo.org/record/3696942/files/figure.png" pageId="12" pageNumber="39">Fig. 7</figureCitation>
). — IVPP
<accessionNumber id="EB9DAB27FFF4FFFAC767EA1AFA693A3F" box="[1362,1451,1810,1834]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/V11498" pageId="12" pageNumber="39" type="EnaNcbi">V 11498</accessionNumber>
, left dentary with c-p1 alveoli, p2 broken, p3-m2 alveoli, from Tieersihabahe, lower part of Halamagai Formation.
</paragraph>
<paragraph id="F47136C4FFF4FFFAC118EA9BFA5F3AB8" blockId="12.[813,1456,1939,2030]" box="[813,1437,1939,1965]" pageId="12" pageNumber="39">
COMPARISON WITH JUNGGAR
<taxonomicName id="33CE4D47FFF4FFFAC65AEA9BFA963ABE" authorityName="Wang &amp; Tseng &amp; Ye &amp; Meng &amp; Bi" authorityYear="2020" box="[1135,1364,1939,1964]" class="Mammalia" family="Hyaenidae" genus="Tungurictis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="12" pageNumber="42" phylum="Chordata" rank="species" species="peignei">
<emphasis id="C6BAEAD6FFF4FFFAC65AEA9BFA963ABE" box="[1135,1364,1939,1964]" italics="true" pageId="12" pageNumber="39">TUNGURICTIS PEIGNEI</emphasis>
</taxonomicName>
, N. SP.
</paragraph>
<paragraph id="F47136C4FFF4FFFAC118EABBFA6D3AF8" blockId="12.[813,1456,1939,2030]" pageId="12" pageNumber="39">
Only two dentary fragments are available for this taxon. The length of m1 is only 80% of the average for Junggar
<taxonomicName id="33CE4D47FFF4FFFAC70BEADCFA6D3AF8" authorityName="Colbert" authorityYear="1939" box="[1342,1455,2004,2029]" class="Mammalia" family="Hyaenidae" genus="Tungurictis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="12" pageNumber="32" phylum="Chordata" rank="genus">
<emphasis id="C6BAEAD6FFF4FFFAC70BEADCFA6D3AF8" box="[1342,1455,2004,2029]" italics="true" pageId="12" pageNumber="39">Tungurictis</emphasis>
</taxonomicName>
</paragraph>
<caption id="A0B1664CFFF5FFFBC2B1EB4BFDDD3B79" ID-DOI="http://doi.org/10.5281/zenodo.3696944" ID-Zenodo-Dep="3696944" httpUri="https://zenodo.org/record/3696944/files/figure.png" pageId="13" pageNumber="40" startId="13.[132,143,1603,1621]" targetBox="[479,1107,215,1558]" targetPageId="13">
<paragraph id="F47136C4FFF5FFFBC2B1EB4BFDDD3B79" blockId="13.[132,1457,1603,1644]" pageId="13" pageNumber="40">
FIG. 8. —
<emphasis id="C6BAEAD6FFF5FFFBC2E7EB4BFE7A3B40" box="[210,440,1603,1621]" italics="true" pageId="13" pageNumber="40">Protictitherium intermedium</emphasis>
Schmidt-Kittler,1976,SNSB-BSPG 1968 VI 736,holotype,left dentary with p2 and m1-2.
<emphasis id="C6BAEAD6FFF5FFFBC6AEEB4CFB6A3B43" bold="true" box="[1179,1192,1604,1622]" pageId="13" pageNumber="40">A</emphasis>
, stereo photos of occlusal view;
<emphasis id="C6BAEAD6FFF5FFFBC2B1EB52FF533B79" bold="true" box="[132,145,1626,1644]" pageId="13" pageNumber="40">B</emphasis>
, lingual; and
<emphasis id="C6BAEAD6FFF5FFFBC334EB52FECD3B79" bold="true" box="[257,271,1626,1644]" pageId="13" pageNumber="40">C</emphasis>
, buccal view. Scale bar: 10 mm.
</paragraph>
</caption>
<paragraph id="F47136C4FFF5FFFBC2B1EBBDFCC53ABB" blockId="13.[132,775,1716,2030]" pageId="13" pageNumber="40">
<emphasis id="C6BAEAD6FFF5FFFBC2B1EBBDFF0B3BDB" box="[132,201,1717,1742]" italics="true" pageId="13" pageNumber="40">peignei</emphasis>
,
<taxonomicNameLabel id="DD8957ADFFF5FFFBC2E3EBBDFECC3BDA" box="[214,270,1717,1743]" pageId="13" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
specimens (
<tableCitation id="B94C037FFFF5FFFBC3B8EBBCFE183BDB" box="[397,474,1716,1742]" captionStart="TABLE" captionStartId="7.[132,143,520,538]" captionText="TABLE 2. — Comparative measurements of lower teeth of Tungurictis Colbert, 1939 from Junggar Basin. Those for Protictitherium Kretzoi, 1938 from Turkey are from Schmidt-Kittler (1976) and those for Tungurictis follow Wang (2004: table 1)." httpUri="http://table.plazi.org/id/A0B1664CFFFFFFF1C2B1EF00FCE73F24" pageId="13" pageNumber="40" tableUuid="A0B1664CFFFFFFF1C2B1EF00FCE73F24">Table 2</tableCitation>
), much too small to be comfortably included in the latter. Although overall consistent with the latter in dental morphology, this small
<taxonomicName id="33CE4D47FFF5FFFBC00BEBFCFD7A3A1B" box="[574,696,1780,1806]" form="stands" pageId="13" pageNumber="40" rank="form">form stands</taxonomicName>
out for its relatively more cuspidate and lingually slanted entoconid and much smaller hypoconulid, which is consistent with assigning this taxon to
<taxonomicName id="33CE4D47FFF5FFFBC33EEA5CFEB93A78" authorityName="Colbert" authorityYear="1939" box="[267,379,1876,1901]" class="Mammalia" family="Hyaenidae" genus="Tungurictis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="13" pageNumber="32" phylum="Chordata" rank="genus">
<emphasis id="C6BAEAD6FFF5FFFBC33EEA5CFEB93A78" box="[267,379,1876,1901]" italics="true" pageId="13" pageNumber="40">Tungurictis</emphasis>
</taxonomicName>
. Furthermore, the ramus height at the position of m1 appears to be visibly shallower in this unnamed taxon relative to m1 crown height, compared to
<taxonomicName id="33CE4D47FFF5FFFBC05DEA9CFD073AB8" authority="Wang &amp; Tseng &amp; Ye &amp; Meng &amp; Bi, 2020" authorityName="Wang &amp; Tseng &amp; Ye &amp; Meng &amp; Bi" authorityYear="2020" box="[616,709,1940,1965]" class="Mammalia" family="Hyaenidae" genus="Tungurictis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="13" pageNumber="40" phylum="Chordata" rank="species" species="peignei" status="sp. nov.">
<emphasis id="C6BAEAD6FFF5FFFBC05DEA9CFD073AB8" box="[616,709,1940,1965]" italics="true" pageId="13" pageNumber="40">T. peignei</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="DD8957ADFFF5FFFBC0E4EA9CFCC53ABB" box="[721,775,1940,1966]" pageId="13" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="F47136C4FFF5FFFBC2AEEABBFA573A1B" blockId="13.[132,775,1716,2030]" lastBlockId="13.[813,1456,1716,1806]" pageId="13" pageNumber="40">Intraspecific variations and sexual dimorphism in extinct basal hyaenids are difficult to evaluate when most species from any single locality amount to no more than a handful of specimens. For the present, the small size and dental morphology seems to suggest a different species, pending additional material.</paragraph>
</subSubSection>
</treatment>
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