treatments-xml/data/7C/4E/8E/7C4E8E28A261FF959D196F98FC135549.xml

<document id="45E358B4D1309ACDE26FB264DF42D33E" ID-DOI="10.5852/ejt.2018.425" ID-GBIF-Dataset="845847ee-c68b-49a6-b66f-9bfe7b5e3fcd" ID-ISSN="2118-9773" ID-Zenodo-Dep="3806131" ID-ZooBank="AB5C3414-92EC-4CE3-8963-880E07648D11" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="admin" IM.tables_requiresApprovalFor="existingObjects,operationResults,plazi" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1588793040195" checkinUser="valdenar" docAuthor="Santos, Erika, Rubal, Marcos, Veiga, Puri, da Rocha, Clélia M. C. &amp; Fontoura, Paulo" docDate="2018" docId="7C4E8E28A261FF959D196F98FC135549" docLanguage="en" docName="ejt-425_santos_rubal_veiga.pdf.imf" docOrigin="European Journal of Taxonomy 425" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Batillipes lusitanus Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura 2018, sp. nov." docType="treatment" docUuid="EA994F4C-C1D9-4B43-871C-327A3CB644C4" docUuidSource="ZooBank" docVersion="12" lastPageNumber="17" masterDocId="8077F650A268FF859F6C6E0BFFD15401" masterDocTitle="Batillipes (Tardigrada, Arthrotardigrada) from the Portuguese coast with the description of two new species and a new dichotomous key for all species" masterLastPageNumber="32" masterPageNumber="1" pageNumber="10" updateTime="1740668371324" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="683500B52E5D219FDAB3D3C1F79FE39B">Batillipes (Tardigrada, Arthrotardigrada) from the Portuguese coast with the description of two new species and a new dichotomous key for all species</mods:title>
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<mods:namePart id="79A7C7ACB203F2BD9C36B53BC283F0A6">Santos, Erika</mods:namePart>
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<mods:affiliation id="82AB700A6588AC53B05C9384E92698BE">Department of Biology, Faculty of Sciences, University of Porto, Rua Campo Alegre, 4169 - 007, Porto, Portugal. &amp; MARE, Marine and Environmental Sciences Centre, ISPA - Instituto Universitário, Rua Jardim do Tabaco 34, 1149 - 041 Lisboa, Portugal.</mods:affiliation>
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<mods:namePart id="74297E0C24F7EA6A869B57BFCADED598">Rubal, Marcos</mods:namePart>
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<mods:affiliation id="BF04F849FE1FFF9A5524EF3FF63D67DA">Department of Biology, Faculty of Sciences, University of Porto, Rua Campo Alegre, 4169 - 007, Porto, Portugal. &amp; CIIMAR / CIMAR, Laboratory of Coastal Biodiversity, Centre of Marine and Environmental Research, Terminal de cruzeiros do Porto de Leixões, Avenida General Norton de Matos S / N, 4450 - 208 Matosinhos Portugal.</mods:affiliation>
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<mods:namePart id="9E8BA806AD383352DB822EC741BFB768">Veiga, Puri</mods:namePart>
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<mods:affiliation id="1A21DE25D2E5306900B1AA2058BD22D3">Department of Biology, Faculty of Sciences, University of Porto, Rua Campo Alegre, 4169 - 007, Porto, Portugal. &amp; CIIMAR / CIMAR, Laboratory of Coastal Biodiversity, Centre of Marine and Environmental Research, Terminal de cruzeiros do Porto de Leixões, Avenida General Norton de Matos S / N, 4450 - 208 Matosinhos Portugal.</mods:affiliation>
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<mods:namePart id="1627F4BFE49E0FDBCE20EE1288552939">da Rocha, Clélia M. C.</mods:namePart>
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<mods:affiliation id="3B53FC51D18374C33D5CF2DB98EEDB19">Departamento de Biologia, Universidade Federal Rural de Pernambuco, Av. Manoel de Medeiros, S / N, Dois Irmãos, Recife, PE, Brasil. 52171 - 090.</mods:affiliation>
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<mods:namePart id="EB60A5241DE301B05521B789423C8AFC">Fontoura, Paulo</mods:namePart>
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<mods:affiliation id="8B878B0BC981F04009D54303B56946AB">Department of Biology, Faculty of Sciences, University of Porto, Rua Campo Alegre, 4169 - 007, Porto, Portugal. &amp; MARE, Marine and Environmental Sciences Centre, ISPA - Instituto Universitário, Rua Jardim do Tabaco 34, 1149 - 041 Lisboa, Portugal.</mods:affiliation>
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<mods:title id="004C9B3CE54588E64B286C3819DA438A">European Journal of Taxonomy</mods:title>
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<mods:date id="FEE3FB07F624B992E0B294F031B25421">2018</mods:date>
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<treatment id="7C4E8E28A261FF959D196F98FC135549" ID-DOI="http://doi.org/10.5281/zenodo.3816371" ID-GBIF-Taxon="163971347" ID-Zenodo-Dep="3816371" LSID="urn:lsid:zoobank.org:act:EA994F4C-C1D9-4B43-871C-327A3CB644C4" httpUri="http://treatment.plazi.org/id/7C4E8E28A261FF959D196F98FC135549" lastPageId="16" lastPageNumber="17" pageId="9" pageNumber="10" scope_class="Heterotardigrada">
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<paragraph id="F4583F3EA261FF8C9D196F98FC6E55AF" blockId="9.[363,1224,403,500]" box="[629,959,403,430]" pageId="9" pageNumber="10">
<heading id="AF108852A261FF8C9D196F98FC6E55AF" box="[629,959,403,430]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="10" reason="2">
<taxonomicName id="33E744BDA261FF8C9D196F98FC8C55AF" ID-CoL="c684b3a7-83f6-4d22-aec2-c4b281154ea2" authority="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura, 2018" authorityName="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura" authorityYear="2018" box="[629,861,403,430]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="9" pageNumber="10" phylum="Tardigrada" rank="species" species="lusitanus" status="sp. nov.">
<emphasis id="C693E32CA261FF8C9D196F98FC8C55AF" bold="true" box="[629,861,403,430]" italics="true" pageId="9" pageNumber="10">Batillipes lusitanus</emphasis>
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<taxonomicNameLabel id="DDA05E57A261FF8C9C086F9FFC6E55AF" box="[868,959,404,430]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
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<figureCitation id="6CDC23BBA261FF8C9DD86FD1FCCF55F5" box="[692,798,474,500]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="9.[189,232,1960,1986]" captionStartId-1="10.[189,232,1846,1872]" captionStartId-2="13.[189,232,1847,1873]" captionStartId-3="14.[189,232,1886,1912]" captionTargetBox-0="[193,1397,1034,1917]" captionTargetBox-1="[191,1397,933,1805]" captionTargetBox-2="[190,1398,902,1805]" captionTargetBox-3="[193,1395,1122,1838]" captionTargetId-0="figure@9.[187,1397,1034,1917]" captionTargetId-1="figure@10.[191,1397,933,1805]" captionTargetId-2="figure@13.[189,1399,902,1805]" captionTargetId-3="figure@14.[192,1395,1122,1838]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="13" captionTargetPageId-3="14" captionText-0="Fig. 4. Batillipes lusitanus sp. nov. A. Schematic drawing, dorsal view. B. Habitus of the holotype, ♀, ventral view. Scale bars = 50 µm." captionText-1="Fig. 5. Batillipes lusitanus sp. nov. A–B, D. Holotype, ♀. A–B. Detail of the head. A. Dorsal view, showing internal cirrus (black arrowhead), lateral cirrus (asterisk), secondary clava (white arrowhead). B. Ventral view, showing primary clava (white arrowhead), external cirrus (black arrowhead). C. Detail of body lateral projections of a paratype (slide C.IX-46) (black arrowheads); D. Detail of leg sensory organ on leg IV. Scale bars = 10 µm." captionText-2="Fig. 6. Batillipes lusitanus sp. nov. A. Fourth leg of a paratype (slide C.IX-46) with details of toes (1–6), note the shape of the ventro-lateral projection between legs III–IV (black arrowhead). B. Fourth leg of a paratype (slide C.IX-1), showing the rigid process on the distal extreme (black arrowhead). C. Female gonopore (white arrowhead) and anus (black arrowhead). D. Male gonopore (white arrowhead) and anus (black arrowhead). Scale bars = 10 µm." captionText-3="Fig. 7. Batillipes lusitanus sp. nov. A. Posterior portion of the body, showing the dorsal sculpture. B. Caudal apparatus, showing details of cuticular pillars. C. SEM photo showing the dorsal (white arrowhead) and ventral (black arrowhead) aspect of the cuticle. Pillars are also visible (black arrow). Scale bars = 10 µm." figureDoi-0="http://doi.org/10.5281/zenodo.3806143" figureDoi-1="http://doi.org/10.5281/zenodo.3806145" figureDoi-2="http://doi.org/10.5281/zenodo.3806149" figureDoi-3="http://doi.org/10.5281/zenodo.3806153" httpUri-0="https://zenodo.org/record/3806143/files/figure.png" httpUri-1="https://zenodo.org/record/3806145/files/figure.png" httpUri-2="https://zenodo.org/record/3806149/files/figure.png" httpUri-3="https://zenodo.org/record/3806153/files/figure.png" pageId="9" pageNumber="10">Figs 4–7</figureCitation>
; 
<tableCitation id="B9650A85A261FF8C9C456FD1FC5155F5" box="[809,896,474,500]" captionStart="Table 2" captionStartId="11.[188,257,564,592]" captionTargetPageId="11" captionText="Table 2 (continued on next page). Measurements (µm) of selected morphological structures of specimens of Batillipes lusitanus sp. nov. (Holo = Holotype; Allo = Allotype; SD = Standard deviation; ‘Range’ refers to the smallest and largest measured specimen / structure; N = number of specimens/structures measured)." pageId="9" pageNumber="10">Table 2</tableCitation>
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<subSubSection id="BCFD6CB5A261FF8C9FD16C10FC00577F" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="F4583F3EA261FF8C9FD16C10FEE75634" blockId="9.[189,310,539,565]" box="[189,310,539,565]" pageId="9" pageNumber="10">
<heading id="AF108852A261FF8C9FD16C10FEE75634" bold="true" box="[189,310,539,565]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="C693E32CA261FF8C9FD16C10FEE75634" bold="true" box="[189,310,539,565]" pageId="9" pageNumber="10">Diagnosis</emphasis>
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<taxonomicName id="33E744BDA261FF8C9FD16C41FEFC5665" authorityName="Richters" authorityYear="1909" box="[189,301,586,612]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="9" pageNumber="10" phylum="Tardigrada" rank="genus">
<emphasis id="C693E32CA261FF8C9FD16C41FEFC5665" box="[189,301,586,612]" italics="true" pageId="9" pageNumber="10">Batillipes</emphasis>
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with tubular undivided primary clavae with a wrinkled surface. Conspicuous papillar secondary clavae. Cephalic cirri with lance-like distal tips. Sensory spines on all legs. A small rigid process is present in the distal extreme of legs IV. Toes 3 and 4 on feet of legs IV equal in length. A distinct head separated from the body by a neck constriction followed by well-developed lateral processes. Ventro-lateral body projections between all leg pairs present: small, often indistinct, between legs I– III and well developed, semicircular or slightly pointed, between legs III–IV. The caudal apparatus consists of a semicircular cuticular expansion. Dorsal cuticle constituted by large pillars (ca 
<quantity id="331F92DBA261FF8C9A4C6D16FAA65736" box="[1312,1399,797,823]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="9" pageNumber="10" unit="mm" value="3.2">3.2 µm</quantity>
high), appearing as large tubercles (diameter about 
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). Ventral cuticle finely punctated. Rosettelike female gonopore separated from the trilobed anus by a groove.
</paragraph>
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<paragraph id="F4583F3EA261FF8C9FD169A3FD9753E7" blockId="9.[189,1398,1960,2023]" pageId="9" pageNumber="10">
<emphasis id="C693E32CA261FF8C9FD169A3FEDA53C3" bold="true" box="[189,267,1960,1986]" pageId="9" pageNumber="10">Fig. 4.</emphasis>
<taxonomicName id="33E744BDA261FF8C9E7869A2FE2753C2" authority="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura, 2018" authorityName="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura" authorityYear="2018" box="[276,502,1961,1987]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="9" pageNumber="10" phylum="Tardigrada" rank="species" species="lusitanus" status="sp. nov.">
<emphasis id="C693E32CA261FF8C9E7869A2FE2753C2" box="[276,502,1961,1987]" italics="true" pageId="9" pageNumber="10">Batillipes lusitanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDA05E57A261FF8C9E9269A2FD8B53C2" box="[510,602,1961,1987]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="C693E32CA261FF8C9D0E69A3FDA653C3" bold="true" box="[610,631,1960,1986]" pageId="9" pageNumber="10">A</emphasis>
. Schematic drawing, dorsal view. 
<emphasis id="C693E32CA261FF8C9B6369A2FBF253C2" bold="true" box="[1039,1059,1961,1987]" pageId="9" pageNumber="10">B</emphasis>
. Habitus of the holotype, ♀, ventral view. Scale bars = 50 µm.
</paragraph>
</caption>
<subSubSection id="BCFD6CB5A262FF8F9FD16F00FDEB5576" pageId="10" pageNumber="11" type="etymology">
<paragraph id="F4583F3EA262FF8F9FD16F00FE955524" blockId="10.[189,324,267,293]" box="[189,324,267,293]" pageId="10" pageNumber="11">
<heading id="AF108852A262FF8F9FD16F00FE955524" bold="true" box="[189,324,267,293]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="C693E32CA262FF8F9FD16F00FE955524" bold="true" box="[189,324,267,293]" pageId="10" pageNumber="11">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="F4583F3EA262FF8F9FD16F32FDEB5576" blockId="10.[189,1398,313,375]" pageId="10" pageNumber="11">
The specific name ‘ 
<taxonomicName id="33E744BDA262FF8F9EDC6F32FDCB5552" authorityName="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura" authorityYear="2018" box="[432,538,313,339]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="10" pageNumber="11" phylum="Tardigrada" rank="species" species="lusitanus">
<emphasis id="C693E32CA262FF8F9EDC6F32FDCB5552" box="[432,538,313,339]" italics="true" pageId="10" pageNumber="11">lusitanus</emphasis>
</taxonomicName>
’ refers to the locus typicus, 
<collectingCountry id="8CF07FAEA262FF8F9C126F32FC355552" box="[894,996,313,339]" name="Portugal" pageId="10" pageNumber="11">Portugal</collectingCountry>
, primitively known as Lusitânia, 
<taxonomicName id="33E744BDA262FF8F9FD16F56FEF65576" authorityName="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura" authorityYear="2018" box="[189,295,349,375]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="10" pageNumber="11" phylum="Tardigrada" rank="species" species="lusitanus">
<emphasis id="C693E32CA262FF8F9FD16F56FEF65576" box="[189,295,349,375]" italics="true" pageId="10" pageNumber="11">lusitanus</emphasis>
</taxonomicName>
= inhabiting Lusitânia.
</paragraph>
</subSubSection>
<subSubSection id="BCFD6CB5A262FF889FD16F95FDEE5548" lastPageId="13" lastPageNumber="14" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="F4583F3EA262FF8F9FD16F95FE7B55B9" blockId="10.[189,426,414,440]" box="[189,426,414,440]" pageId="10" pageNumber="11">
<heading id="AF108852A262FF8F9FD16F95FE7B55B9" bold="true" box="[189,426,414,440]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="C693E32CA262FF8F9FD16F95FE7B55B9" bold="true" box="[189,426,414,440]" pageId="10" pageNumber="11">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="448F3563A262FF8F9F866FC6FE3F562C" ID-GBIF-Occurrence="2610639305" collectionCode="C" country="Portugal" latitude="37.116943" location="Meia-Praia Beach" longLatPrecision="19" longitude="-8.643611" municipality="Lagos" pageId="10" pageNumber="11" specimenCode="C.IX-1" specimenCount="1" specimenCount-female="1" stateProvince="Algarve" typeStatus="holotype">
<paragraph id="F4583F3EA262FF8F9F866FC6FE8D55E6" blockId="10.[189,1399,461,557]" box="[234,348,461,487]" pageId="10" pageNumber="11">
<emphasis id="C693E32CA262FF8F9F866FC6FE8D55E6" bold="true" box="[234,348,461,487]" pageId="10" pageNumber="11">
<typeStatus id="2B5C819CA262FF8F9F866FC6FE8D55E6" box="[234,348,461,487]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="F4583F3EA262FF8F9FD16FFBFE3F562C" blockId="10.[189,1399,461,557]" pageId="10" pageNumber="11">
<collectingCountry id="8CF07FAEA262FF8F9FD16FFBFE8C560B" box="[189,349,496,522]" name="Portugal" pageId="10" pageNumber="11">PORTUGAL</collectingCountry>
: 
<specimenCount id="E2E1F4B7A262FF8F9E0A6FFBFE10560B" box="[358,449,496,522]" pageId="10" pageNumber="11" type="female">♀ adult</specimenCount>
, collected at 
<location id="F13869E5A262FF8F9D346FFBFCFC560B" LSID="urn:lsid:plazi:treatment:7C4E8E28A261FF959D196F98FC135549:F13869E5A262FF8F9D346FFBFCFC560B" box="[600,813,496,522]" country="Portugal" latitude="37.116943" longLatPrecision="19" longitude="-8.643611" municipality="Lagos" name="Meia-Praia Beach" pageId="10" pageNumber="11" stateProvince="Algarve">Meia-Praia Beach</location>
, 
<collectingMunicipality id="143CA544A262FF8F9C546FFBFC52560B" box="[824,899,496,522]" pageId="10" pageNumber="11">Lagos</collectingMunicipality>
, 
<collectingRegion id="3623F1DCA262FF8F9CE06FFBFC3C560B" box="[908,1005,496,522]" pageId="10" pageNumber="11">Algarve</collectingRegion>
, 
<geoCoordinate id="91D359F9A262FF8F9C946FE5FBA1560B" box="[1016,1136,494,522]" degrees="37" direction="north" minutes="7" orientation="latitude" pageId="10" pageNumber="11" precision="15" seconds="1" value="37.116943">37°7′1″ N</geoCoordinate>
, 
<geoCoordinate id="91D359F9A262FF8F9B176FE5FAD4560B" box="[1147,1285,494,522]" degrees="8" direction="west" minutes="38" orientation="longitude" pageId="10" pageNumber="11" precision="15" seconds="37" value="-8.643611">8°38′37″ W</geoCoordinate>
, mounted in glycerol (slide 
<specimenCode id="A4419745A262FF8F9EE76C18FE30562C" box="[395,481,531,557]" pageId="10" pageNumber="11">C.IX-1</specimenCode>
).
</paragraph>
</materialsCitation>
<materialsCitation id="448F3563A262FF8F9F866C5FFE3E56B4" ID-GBIF-Occurrence="2610639302" collectionCode="C" country="Portugal" latitude="37.116943" location="Meia-Praia Beach" longLatPrecision="19" longitude="-8.643611" municipality="Lagos" pageId="10" pageNumber="11" specimenCode="C.IX-2" specimenCount="1" specimenCount-adult="1" stateProvince="Algarve" typeStatus="allotype">
<paragraph id="F4583F3EA262FF8F9F866C5FFE85566F" blockId="10.[189,1399,596,693]" box="[234,340,596,622]" pageId="10" pageNumber="11">
<emphasis id="C693E32CA262FF8F9F866C5FFE85566F" bold="true" box="[234,340,596,622]" pageId="10" pageNumber="11">
<typeStatus id="2B5C819CA262FF8F9F866C5FFE85566F" box="[234,340,596,622]" pageId="10" pageNumber="11" type="allotype">Allotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="F4583F3EA262FF8F9FD16C73FE3E56B4" blockId="10.[189,1399,596,693]" pageId="10" pageNumber="11">
<collectingCountry id="8CF07FAEA262FF8F9FD16C73FE8C5693" box="[189,349,632,658]" name="Portugal" pageId="10" pageNumber="11">PORTUGAL</collectingCountry>
: 
<specimenCount id="E2E1F4B7A262FF8F9E0A6C7DFE105693" box="[358,449,630,658]" count="1" pageId="10" pageNumber="11" type="adult">♂ adult</specimenCount>
, collected at 
<location id="F13869E5A262FF8F9D346C73FCFC5693" LSID="urn:lsid:plazi:treatment:7C4E8E28A261FF959D196F98FC135549:F13869E5A262FF8F9D346C73FCFC5693" box="[600,813,632,658]" country="Portugal" latitude="37.116943" longLatPrecision="19" longitude="-8.643611" municipality="Lagos" name="Meia-Praia Beach" pageId="10" pageNumber="11" stateProvince="Algarve">Meia-Praia Beach</location>
, 
<collectingMunicipality id="143CA544A262FF8F9C546C73FC525693" box="[824,899,632,658]" pageId="10" pageNumber="11">Lagos</collectingMunicipality>
, 
<collectingRegion id="3623F1DCA262FF8F9CE06C73FC3C5693" box="[908,1005,632,658]" pageId="10" pageNumber="11">Algarve</collectingRegion>
, 
<geoCoordinate id="91D359F9A262FF8F9C946C7DFBA15693" box="[1016,1136,630,658]" degrees="37" direction="north" minutes="7" orientation="latitude" pageId="10" pageNumber="11" precision="15" seconds="1" value="37.116943">37°7′1″ N</geoCoordinate>
, 
<geoCoordinate id="91D359F9A262FF8F9B176C7DFAD45693" box="[1147,1285,630,658]" degrees="8" direction="west" minutes="38" orientation="longitude" pageId="10" pageNumber="11" precision="15" seconds="37" value="-8.643611">8°38′37″ W</geoCoordinate>
, mounted in glycerol (slide 
<specimenCode id="A4419745A262FF8F9EE76C90FE3356B4" box="[395,482,667,693]" pageId="10" pageNumber="11">C.IX-2</specimenCode>
).
</paragraph>
</materialsCitation>
<materialsCitation id="448F3563A262FF8F9F866CD7FBB2573C" ID-GBIF-Occurrence="2610639307" collectionCode="C" country="Portugal" latitude="37.116943" location="Meia-Praia Beach" longLatPrecision="19" longitude="-8.643611" pageId="10" pageNumber="11" specimenCode="C.IX-35-C.IX-66" specimenCount="78" specimenCount-female="19" specimenCount-juvenile="37" specimenCount-male="21" typeStatus="paratype">
<paragraph id="F4583F3EA262FF8F9F866CD7FEB656F7" blockId="10.[189,1399,732,829]" box="[234,359,732,758]" pageId="10" pageNumber="11">
<emphasis id="C693E32CA262FF8F9F866CD7FEB656F7" bold="true" box="[234,359,732,758]" pageId="10" pageNumber="11">
<typeStatus id="2B5C819CA262FF8F9F866CD7FEB656F7" box="[234,359,732,758]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="F4583F3EA262FF889FD16D0BFDEE5548" blockId="10.[189,1399,732,829]" lastBlockId="13.[189,1399,265,329]" lastPageId="13" lastPageNumber="14" pageId="10" pageNumber="11">
<collectingCountry id="8CF07FAEA262FF8F9FD16D0BFE8C571B" box="[189,349,768,794]" name="Portugal" pageId="10" pageNumber="11">PORTUGAL</collectingCountry>
: 78 specimens (
<specimenCount id="E2E1F4B7A262FF8F9D7E6D0BFDB2571B" box="[530,611,768,794]" count="19" pageId="10" pageNumber="11" type="female">19 ♀♀</specimenCount>
, 
<specimenCount id="E2E1F4B7A262FF8F9D1C6D0BFD6D571B" box="[624,700,766,794]" count="21" pageId="10" pageNumber="11" type="male">21 ♂♂</specimenCount>
, 
<specimenCount id="E2E1F4B7A262FF8F9DA16D0BFC8B5718" box="[717,858,767,794]" count="37" pageId="10" pageNumber="11" type="juvenile">37 juveniles</specimenCount>
and 
<specimenCount id="E2E1F4B7A262FF8F9CFC6D0BFB87571B" box="[912,1110,767,794]" count="1" pageId="10" pageNumber="11" type="larva">1 four-toed larva</specimenCount>
), collected at 
<location id="F13869E5A262FF8F9B986CF4FED8573C" LSID="urn:lsid:plazi:treatment:7C4E8E28A261FF959D196F98FC135549:F13869E5A262FF8F9B986CF4FED8573C" country="Portugal" latitude="37.116943" longLatPrecision="19" longitude="-8.643611" name="Meia-Praia Beach" pageId="10" pageNumber="11">Meia-Praia Beach</location>
, 
<geoCoordinate id="91D359F9A262FF8F9E7A6D2AFE5E573C" box="[278,399,801,829]" degrees="37" direction="north" minutes="7" orientation="latitude" pageId="10" pageNumber="11" precision="15" seconds="1" value="37.116943">37°7′1″ N</geoCoordinate>
, 
<geoCoordinate id="91D359F9A262FF8F9EF06D2AFDF9573C" box="[412,552,801,829]" degrees="8" direction="west" minutes="38" orientation="longitude" pageId="10" pageNumber="11" precision="15" seconds="37" value="-8.643611">8°38′37″ W</geoCoordinate>
, mounted in glycerol (slides 
<specimenCode id="A4419745A262FF8F9CEC6D29FB84573C" box="[896,1109,802,829]" pageId="10" pageNumber="11">C.IX-35–C.IX-66</specimenCode>
); 
<materialsCitation id="448F3563A262FF889B076D28FDEE5548" ID-GBIF-Occurrence="2610639307" collectionCode="C" lastPageId="13" lastPageNumber="14" latitude="41.59111" location="Afife Beach" longLatPrecision="19" longitude="-8.805" pageId="10" pageNumber="11" specimenCode="C.IX-3-C.IX-34" specimenCount="89" specimenCount-female="35" specimenCount-juvenile="39" specimenCount-male="14" typeStatus="paratype">
89 specimens (
<specimenCount id="E2E1F4B7A262FF8F9A706D29FAA1573C" box="[1308,1392,802,829]" count="35" pageId="10" pageNumber="11" type="female">35 ♀♀</specimenCount>
, 
<specimenCount id="E2E1F4B7A265FF889FD16F00FEDB5524" box="[189,266,265,293]" count="14" pageId="13" pageNumber="14" type="male">14 ♂♂</specimenCount>
, 
<specimenCount id="E2E1F4B7A265FF889E766F00FE795524" box="[282,424,267,293]" count="39" pageId="13" pageNumber="14" type="juvenile">39 juveniles</specimenCount>
and 
<specimenCount id="E2E1F4B7A265FF889EB36F00FD775527" box="[479,678,267,294]" count="1" pageId="13" pageNumber="14" type="larva">1 four-toed larva</specimenCount>
), collected at 
<location id="F13869E5A265FF889C2F6F00FC005524" LSID="urn:lsid:plazi:treatment:7C4E8E28A261FF959D196F98FC135549:F13869E5A265FF889C2F6F00FC005524" box="[835,977,267,293]" latitude="41.59111" longLatPrecision="19" longitude="-8.805" name="Afife Beach" pageId="13" pageNumber="14">Afife Beach</location>
, 
<geoCoordinate id="91D359F9A265FF889CB06F02FBA05527" box="[988,1137,265,294]" degrees="41" direction="north" minutes="35" orientation="latitude" pageId="13" pageNumber="14" precision="15" seconds="28" value="41.59111">41°35′28″ N</geoCoordinate>
, 
<geoCoordinate id="91D359F9A265FF889B106F02FAD45527" box="[1148,1285,265,294]" degrees="8" direction="west" minutes="48" orientation="longitude" pageId="13" pageNumber="14" precision="15" seconds="18" value="-8.805">8°48′18″ W</geoCoordinate>
, mounted in PVA (slides 
<specimenCode id="A4419745A265FF889E006F24FDE35548" box="[364,562,303,329]" pageId="13" pageNumber="14">C.IX-3–C.IX-34</specimenCode>
).
</materialsCitation>
</paragraph>
</materialsCitation>
</subSubSection>
<caption id="A0986FB6A262FF8F9FD1693DFDBA53DC" ID-DOI="http://doi.org/10.5281/zenodo.3806145" ID-Zenodo-Dep="3806145" httpUri="https://zenodo.org/record/3806145/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,232,1846,1872]" targetBox="[191,1397,933,1805]" targetPageId="10">
<paragraph id="F4583F3EA262FF8F9FD1693DFDBA53DC" blockId="10.[189,1399,1846,2013]" pageId="10" pageNumber="11">
<emphasis id="C693E32CA262FF8F9FD1693DFEDC5351" bold="true" box="[189,269,1846,1872]" pageId="10" pageNumber="11">Fig. 5.</emphasis>
<taxonomicName id="33E744BDA262FF8F9E74693DFE2D5351" authority="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura, 2018" authorityName="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura" authorityYear="2018" box="[280,508,1846,1872]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="10" pageNumber="11" phylum="Tardigrada" rank="species" species="lusitanus" status="sp. nov.">
<emphasis id="C693E32CA262FF8F9E74693DFE2D5351" box="[280,508,1846,1872]" italics="true" pageId="10" pageNumber="11">Batillipes lusitanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDA05E57A262FF8F9D6A693DFDB55351" box="[518,612,1846,1872]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="C693E32CA262FF8F9D02693DFD745351" bold="true" box="[622,677,1846,1872]" pageId="10" pageNumber="11">A–B</emphasis>
, 
<emphasis id="C693E32CA262FF8F9DDB693DFD1D5351" bold="true" box="[695,716,1846,1872]" pageId="10" pageNumber="11">D</emphasis>
. Holotype, ♀. 
<emphasis id="C693E32CA262FF8F9CE8693DFC6A5351" bold="true" box="[900,955,1846,1872]" pageId="10" pageNumber="11">A–B</emphasis>
. Detail of the head. 
<emphasis id="C693E32CA262FF8F9BD5693DFB1F5351" bold="true" box="[1209,1230,1846,1872]" pageId="10" pageNumber="11">A</emphasis>
. Dorsal view, showing internal cirrus (black arrowhead), lateral cirrus (asterisk), secondary clava (white arrowhead). 
<emphasis id="C693E32CA262FF8F9FD16976FF005396" bold="true" box="[189,209,1917,1943]" pageId="10" pageNumber="11">B</emphasis>
. Ventral view, showing primary clava (white arrowhead), external cirrus (black arrowhead). 
<emphasis id="C693E32CA262FF8F9A676977FAF15397" bold="true" box="[1291,1312,1916,1942]" pageId="10" pageNumber="11">C</emphasis>
. Detail of body lateral projections of a paratype (slide C.IX-46) (black arrowheads); 
<emphasis id="C693E32CA262FF8F9B3469ABFBBC53BB" bold="true" box="[1112,1133,1952,1978]" pageId="10" pageNumber="11">D</emphasis>
. Detail of leg sensory organ on leg IV. Scale bars = 10 µm.
</paragraph>
</caption>
<caption id="A0986FB6A263FF8E9FD06C3FFD905697" pageId="11" pageNumber="12" startId="11.[188,257,564,592]" targetBox="[199,1889,734,1719]" targetIsTable="true" targetPageId="11">
<paragraph id="F4583F3EA263FF8E9FD06C3FFD905697" blockId="11.[188,1942,564,662]" pageId="11" pageNumber="12">
<emphasis id="C693E32CA263FF8E9FD06C3FFECB5651" bold="true" box="[188,282,564,592]" pageId="11" pageNumber="12">Table 2</emphasis>
(continued on next page). Measurements (µm) of selected morphological structures of specimens of 
<taxonomicName id="33E744BDA263FF8E9AB66C3FF96F5651" authority="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura, 2018" authorityName="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura" authorityYear="2018" box="[1498,1726,564,592]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="12" phylum="Tardigrada" rank="species" species="lusitanus" status="sp. nov.">
<emphasis id="C693E32CA263FF8E9AB66C3FF96F5651" box="[1498,1726,564,592]" italics="true" pageId="11" pageNumber="12">Batillipes lusitanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDA05E57A263FF8E99A46C3FF8F45651" box="[1736,1829,564,592]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
(Holo = Holotype; Allo = Allotype; SD = Standard deviation; ‘Range’ refers to the smallest and largest measured specimen / structure; N = number of specimens / structures measured).
</paragraph>
</caption>
<paragraph id="F4583F3EA263FF8E9FAB6CD5F89E52B6" pageId="11" pageNumber="12">
<table id="86E7CD9EA263007A9FAB6CD5F8B052B6" box="[199,1889,734,1719]" colsContinueIn="12.[199,1889,699,1724]" gridcols="7" gridrows="19" pageId="11" pageNumber="12">
<tr id="4AD73D7CA263007A9FAB6CD5F8B056F9" box="[199,1889,734,760]" gridrow="0" pageId="11" pageNumber="12" rowspan-3="1">
<th id="09065400A263007A9FAB6CD5FE765758" box="[199,423,734,857]" gridcol="0" gridrow="0" pageId="11" pageNumber="12" rowspan="3" rowspanBelow="2">
<emphasis id="C693E32CA263FF8E9FAB6CD5FEEF56F9" bold="true" box="[199,318,734,760]" pageId="11" pageNumber="12">Structures</emphasis>
</th>
<th id="09065400A263007A9EA06CD5FDD85758" box="[460,521,734,857]" gridcol="1" gridrow="0" pageId="11" pageNumber="12" rowspan="3" rowspanBelow="2">
<emphasis id="C693E32CA263FF8E9EA36D34FDD75758" bold="true" box="[463,518,831,857]" pageId="11" pageNumber="12">Holo</emphasis>
</th>
<th id="09065400A263007A9D436CD5FC5156F9" box="[559,896,734,760]" gridcol="2" gridrow="0" pageId="11" pageNumber="12">
<emphasis id="C693E32CA263FF8E9D1D6CD5FD1F56F9" bold="true" box="[625,718,734,760]" pageId="11" pageNumber="12">Females</emphasis>
</th>
<th id="09065400A263007A9B666CD5FAB456F9" box="[1034,1381,734,760]" gridcol="4" gridrow="0" pageId="11" pageNumber="12">
<emphasis id="C693E32CA263FF8E9B306CD5FB7156F9" bold="true" box="[1116,1184,734,760]" pageId="11" pageNumber="12">Males</emphasis>
</th>
<th id="09065400A263007A9AE86CD5F90E56F9" box="[1412,1759,734,760]" gridcol="5" gridrow="0" pageId="11" pageNumber="12">
<emphasis id="C693E32CA263FF8E9A916CD5F9B756F9" bold="true" box="[1533,1638,734,760]" pageId="11" pageNumber="12">Juveniles</emphasis>
</th>
<th id="09065400A263007A98766CD5F8B056F9" box="[1818,1889,734,760]" gridcol="6" gridrow="0" pageId="11" pageNumber="12">
<emphasis id="C693E32CA263FF8E98776CD5F8B056F9" bold="true" box="[1819,1889,734,760]" pageId="11" pageNumber="12">Larva</emphasis>
</th>
</tr>
<tr id="4AD73D7CA263007A9FAB6D1CF8B05730" box="[199,1889,791,817]" gridrow="1" pageId="11" pageNumber="12" rowspan-3="1">
<td id="09065400A263007A9D436D1CFC515730" box="[559,896,791,817]" gridcol="2" gridrow="1" pageId="11" pageNumber="12">
<emphasis id="C693E32CA263FF8E9D166D1CFD175730" bold="true" box="[634,710,791,817]" pageId="11" pageNumber="12">Adults</emphasis>
</td>
<td id="09065400A263007A9B666D1CFAB45730" box="[1034,1381,791,817]" gridcol="4" gridrow="1" pageId="11" pageNumber="12">
<emphasis id="C693E32CA263FF8E9B356D1CFB745730" bold="true" box="[1113,1189,791,817]" pageId="11" pageNumber="12">Adults</emphasis>
</td>
<td id="09065400A263007A9AE86D1CF90E5730" box="[1412,1759,791,817]" gridcol="5" gridrow="1" pageId="11" pageNumber="12">
<emphasis id="C693E32CA263FF8E99626D1CF9855730" bold="true" box="[1550,1620,791,817]" pageId="11" pageNumber="12">6-toed</emphasis>
</td>
<td id="09065400A263007A98766D1CF8B05730" box="[1818,1889,791,817]" gridcol="6" gridrow="1" pageId="11" pageNumber="12">
<emphasis id="C693E32CA263FF8E98766D1CF8B05730" bold="true" box="[1818,1889,791,817]" pageId="11" pageNumber="12">4-toed</emphasis>
</td>
</tr>
<tr id="4AD73D7CA263007A9FAB6D34F8B05758" box="[199,1889,831,857]" gridrow="2" pageId="11" pageNumber="12" rowspan-6="1">
<td id="09065400A263007A9D436D34FC515758" box="[559,896,831,857]" gridcol="2" gridrow="2" pageId="11" pageNumber="12">
<emphasis id="C693E32CA263FF8E9D3A6D34FC885758" bold="true" box="[598,857,831,857]" pageId="11" pageNumber="12">Mean ± SD (Range); N</emphasis>
</td>
<td id="09065400A263007A9CCB6D34FC325758" box="[935,995,831,857]" gridcol="3" gridrow="2" pageId="11" pageNumber="12">
<emphasis id="C693E32CA263FF8E9CC26D34FC0C5758" bold="true" box="[942,989,831,857]" pageId="11" pageNumber="12">Allo</emphasis>
</td>
<td id="09065400A263007A9B666D34FAB45758" box="[1034,1381,831,857]" gridcol="4" gridrow="2" pageId="11" pageNumber="12">
<emphasis id="C693E32CA263FF8E9B5A6D34FAE95758" bold="true" box="[1078,1336,831,857]" pageId="11" pageNumber="12">Mean ± SD (Range); N</emphasis>
</td>
<td id="09065400A263007A9AE86D34F90E5758" box="[1412,1759,831,857]" gridcol="5" gridrow="2" pageId="11" pageNumber="12">
<emphasis id="C693E32CA263FF8E9ADC6D34F9635758" bold="true" box="[1456,1714,831,857]" pageId="11" pageNumber="12">Mean ± SD (Range); N</emphasis>
</td>
</tr>
<tr id="4AD73D7CA263007A9FAB6D6CF8B05780" box="[199,1889,871,897]" gridrow="3" pageId="11" pageNumber="12">
<th id="09065400A263007A9FAB6D6CFE765780" box="[199,423,871,897]" gridcol="0" gridrow="3" pageId="11" pageNumber="12">Body length</th>
<td id="09065400A263007A9EA06D6CFDD85780" box="[460,521,871,897]" gridcol="1" gridrow="3" pageId="11" pageNumber="12">248.4</td>
<td id="09065400A263007A9D436D6CFC515780" box="[559,896,871,897]" gridcol="2" gridrow="3" pageId="11" pageNumber="12">197.1 ± 35.62 (114.4–279.4); 55</td>
<td id="09065400A263007A9CCB6D6CFC325780" box="[935,995,871,897]" gridcol="3" gridrow="3" pageId="11" pageNumber="12">187.9</td>
<td id="09065400A263007A9B666D6CFAB45780" box="[1034,1381,871,897]" gridcol="4" gridrow="3" pageId="11" pageNumber="12">203.4 ± 38.58 (131.3–269.9); 36</td>
<td id="09065400A263007A9AE86D6CF90E5780" box="[1412,1759,871,897]" gridcol="5" gridrow="3" pageId="11" pageNumber="12">197.0 ± 35.86 (111.7–289.5); 76</td>
<td id="09065400A263007A98766D6CF8B05780" box="[1818,1889,871,897]" gridcol="6" gridrow="3" pageId="11" pageNumber="12">89.9</td>
</tr>
<tr id="4AD73D7CA263007A9FAB6D97F8B057D7" box="[199,1889,924,982]" gridrow="4" pageId="11" pageNumber="12">
<th id="09065400A263007A9FAB6D97FE7657D7" box="[199,423,924,982]" gridcol="0" gridrow="4" pageId="11" pageNumber="12">Body length without caudal apparatus</th>
<td id="09065400A263007A9EA06D97FDD857D7" box="[460,521,924,982]" gridcol="1" gridrow="4" pageId="11" pageNumber="12">236</td>
<td id="09065400A263007A9D436D97FC5157D7" box="[559,896,924,982]" gridcol="2" gridrow="4" pageId="11" pageNumber="12">185.4 ± 34.85 (105–266.5); 55</td>
<td id="09065400A263007A9CCB6D97FC3257D7" box="[935,995,924,982]" gridcol="3" gridrow="4" pageId="11" pageNumber="12">175.4</td>
<td id="09065400A263007A9B666D97FAB457D7" box="[1034,1381,924,982]" gridcol="4" gridrow="4" pageId="11" pageNumber="12">191.5 ± 37.25 (124.1–254.7); 36</td>
<td id="09065400A263007A9AE86D97F90E57D7" box="[1412,1759,924,982]" gridcol="5" gridrow="4" pageId="11" pageNumber="12">187.6 ± 35.45 (106.2–277.2); 76</td>
<td id="09065400A263007A98766D97F8B057D7" box="[1818,1889,924,982]" gridcol="6" gridrow="4" pageId="11" pageNumber="12">81.7</td>
</tr>
<tr id="4AD73D7CA263007A9FAB6DE6F8B05006" box="[199,1889,1005,1031]" gridrow="5" pageId="11" pageNumber="12">
<th id="09065400A263007A9FAB6DE6FE765006" box="[199,423,1005,1031]" gridcol="0" gridrow="5" pageId="11" pageNumber="12">Body width</th>
<td id="09065400A263007A9EA06DE6FDD85006" box="[460,521,1005,1031]" gridcol="1" gridrow="5" pageId="11" pageNumber="12">100.9</td>
<td id="09065400A263007A9D436DE6FC515006" box="[559,896,1005,1031]" gridcol="2" gridrow="5" pageId="11" pageNumber="12">75.4 ± 16.73 (40.5–112.6); 44</td>
<td id="09065400A263007A9CCB6DE6FC325006" box="[935,995,1005,1031]" gridcol="3" gridrow="5" pageId="11" pageNumber="12">67.6</td>
<td id="09065400A263007A9B666DE6FAB45006" box="[1034,1381,1005,1031]" gridcol="4" gridrow="5" pageId="11" pageNumber="12">78.9 ± 17.92 (44.3–109.1); 27</td>
<td id="09065400A263007A9AE86DE6F90E5006" box="[1412,1759,1005,1031]" gridcol="5" gridrow="5" pageId="11" pageNumber="12">77.9 ± 15.13 (51.5–111.2); 34</td>
<td id="09065400A263007A98766DE6F8B05006" box="[1818,1889,1005,1031]" gridcol="6" gridrow="5" pageId="11" pageNumber="12">28.9</td>
</tr>
<tr id="4AD73D7CA263007A9FAB6A29F8B0503D" box="[199,1889,1058,1084]" gridrow="6" pageId="11" pageNumber="12">
<th id="09065400A263007A9FAB6A29FE76503D" box="[199,423,1058,1084]" gridcol="0" gridrow="6" pageId="11" pageNumber="12">Median cirrus</th>
<td id="09065400A263007A9EA06A29FDD8503D" box="[460,521,1058,1084]" gridcol="1" gridrow="6" pageId="11" pageNumber="12">26.3</td>
<td id="09065400A263007A9D436A29FC51503D" box="[559,896,1058,1084]" gridcol="2" gridrow="6" pageId="11" pageNumber="12">26.0 ± 4.89 (15.2–35.2); 40</td>
<td id="09065400A263007A9CCB6A29FC32503D" box="[935,995,1058,1084]" gridcol="3" gridrow="6" pageId="11" pageNumber="12">23.2</td>
<td id="09065400A263007A9B666A29FAB4503D" box="[1034,1381,1058,1084]" gridcol="4" gridrow="6" pageId="11" pageNumber="12">25.1 ± 3.78 (17.7–32.1); 29</td>
<td id="09065400A263007A9AE86A29F90E503D" box="[1412,1759,1058,1084]" gridcol="5" gridrow="6" pageId="11" pageNumber="12">24.7 ± 4.29 (13.0–34.8); 56</td>
<td id="09065400A263007A98766A29F8B0503D" box="[1818,1889,1058,1084]" gridcol="6" gridrow="6" pageId="11" pageNumber="12">?</td>
</tr>
<tr id="4AD73D7CA263007A9FAB6A5CF8B05070" box="[199,1889,1111,1137]" gridrow="7" pageId="11" pageNumber="12">
<th id="09065400A263007A9FAB6A5CFE765070" box="[199,423,1111,1137]" gridcol="0" gridrow="7" pageId="11" pageNumber="12">Internal cirri</th>
<td id="09065400A263007A9EA06A5CFDD85070" box="[460,521,1111,1137]" gridcol="1" gridrow="7" pageId="11" pageNumber="12">29.5</td>
<td id="09065400A263007A9D436A5CFC515070" box="[559,896,1111,1137]" gridcol="2" gridrow="7" pageId="11" pageNumber="12">24.2 ± 4.10 (12.4–32.0); 52</td>
<td id="09065400A263007A9CCB6A5CFC325070" box="[935,995,1111,1137]" gridcol="3" gridrow="7" pageId="11" pageNumber="12">20.5</td>
<td id="09065400A263007A9B666A5CFAB45070" box="[1034,1381,1111,1137]" gridcol="4" gridrow="7" pageId="11" pageNumber="12">23.2 ± 3.98 (16.5–31.5); 35</td>
<td id="09065400A263007A9AE86A5CF90E5070" box="[1412,1759,1111,1137]" gridcol="5" gridrow="7" pageId="11" pageNumber="12">23.3 ± 3.72 (13.4–30.5); 70</td>
<td id="09065400A263007A98766A5CF8B05070" box="[1818,1889,1111,1137]" gridcol="6" gridrow="7" pageId="11" pageNumber="12">13.8</td>
</tr>
<tr id="4AD73D7CA263007A9FAB6A87F8B050A7" box="[199,1889,1164,1190]" gridrow="8" pageId="11" pageNumber="12">
<th id="09065400A263007A9FAB6A87FE7650A7" box="[199,423,1164,1190]" gridcol="0" gridrow="8" pageId="11" pageNumber="12">External cirri</th>
<td id="09065400A263007A9EA06A87FDD850A7" box="[460,521,1164,1190]" gridcol="1" gridrow="8" pageId="11" pageNumber="12">20.8</td>
<td id="09065400A263007A9D436A87FC5150A7" box="[559,896,1164,1190]" gridcol="2" gridrow="8" pageId="11" pageNumber="12">17.9 ± 2.72 (11.3–23.7); 52</td>
<td id="09065400A263007A9CCB6A87FC3250A7" box="[935,995,1164,1190]" gridcol="3" gridrow="8" pageId="11" pageNumber="12">15.3</td>
<td id="09065400A263007A9B666A87FAB450A7" box="[1034,1381,1164,1190]" gridcol="4" gridrow="8" pageId="11" pageNumber="12">16.7 ± 2.97 (9.0–23.6); 36</td>
<td id="09065400A263007A9AE86A87F90E50A7" box="[1412,1759,1164,1190]" gridcol="5" gridrow="8" pageId="11" pageNumber="12">17.2 ± 3.39 (9.0–25.2); 71</td>
<td id="09065400A263007A98766A87F8B050A7" box="[1818,1889,1164,1190]" gridcol="6" gridrow="8" pageId="11" pageNumber="12">9.4</td>
</tr>
<tr id="4AD73D7CA263007A9FAB6ACAF8B050DA" box="[199,1889,1217,1243]" gridrow="9" pageId="11" pageNumber="12">
<th id="09065400A263007A9FAB6ACAFE7650DA" box="[199,423,1217,1243]" gridcol="0" gridrow="9" pageId="11" pageNumber="12">
Lateral cirri 
<emphasis id="C693E32CA263FF8E9E206ACAFE8D50DA" box="[332,348,1217,1243]" italics="true" pageId="11" pageNumber="12">A</emphasis>
</th>
<td id="09065400A263007A9EA06ACAFDD850DA" box="[460,521,1217,1243]" gridcol="1" gridrow="9" pageId="11" pageNumber="12">40.2</td>
<td id="09065400A263007A9D436ACAFC5150DA" box="[559,896,1217,1243]" gridcol="2" gridrow="9" pageId="11" pageNumber="12">37.1 ± 4.94 (19.7–47.3); 50</td>
<td id="09065400A263007A9CCB6ACAFC3250DA" box="[935,995,1217,1243]" gridcol="3" gridrow="9" pageId="11" pageNumber="12">32.6</td>
<td id="09065400A263007A9B666ACAFAB450DA" box="[1034,1381,1217,1243]" gridcol="4" gridrow="9" pageId="11" pageNumber="12">36.2 ± 5.00 (27.2–47.8); 36</td>
<td id="09065400A263007A9AE86ACAF90E50DA" box="[1412,1759,1217,1243]" gridcol="5" gridrow="9" pageId="11" pageNumber="12">35.7 ± 5.10 (22.7–47.0); 71</td>
<td id="09065400A263007A98766ACAF8B050DA" box="[1818,1889,1217,1243]" gridcol="6" gridrow="9" pageId="11" pageNumber="12">26.0</td>
</tr>
<tr id="4AD73D7CA263007A9FAB6AFDF8B05111" box="[199,1889,1270,1296]" gridrow="10" pageId="11" pageNumber="12">
<th id="09065400A263007A9FAB6AFDFE765111" box="[199,423,1270,1296]" gridcol="0" gridrow="10" pageId="11" pageNumber="12">Primary clavae</th>
<td id="09065400A263007A9EA06AFDFDD85111" box="[460,521,1270,1296]" gridcol="1" gridrow="10" pageId="11" pageNumber="12">19.8</td>
<td id="09065400A263007A9D436AFDFC515111" box="[559,896,1270,1296]" gridcol="2" gridrow="10" pageId="11" pageNumber="12">17.2 ± 2.56 (11.3–21.5); 52</td>
<td id="09065400A263007A9CCB6AFDFC325111" box="[935,995,1270,1296]" gridcol="3" gridrow="10" pageId="11" pageNumber="12">13.8</td>
<td id="09065400A263007A9B666AFDFAB45111" box="[1034,1381,1270,1296]" gridcol="4" gridrow="10" pageId="11" pageNumber="12">16.8 ± 2.82 (12.2–22.2); 36</td>
<td id="09065400A263007A9AE86AFDF90E5111" box="[1412,1759,1270,1296]" gridcol="5" gridrow="10" pageId="11" pageNumber="12">16.8 ± 3.31 (5.5–22.6); 71</td>
<td id="09065400A263007A98766AFDF8B05111" box="[1818,1889,1270,1296]" gridcol="6" gridrow="10" pageId="11" pageNumber="12">8.8</td>
</tr>
<tr id="4AD73D7CA263007A9FAB6B21F8B05145" box="[199,1889,1322,1348]" gridrow="11" pageId="11" pageNumber="12">
<th id="09065400A263007A9FAB6B21FE765145" box="[199,423,1322,1348]" gridcol="0" gridrow="11" pageId="11" pageNumber="12">Secondary clavae</th>
<td id="09065400A263007A9EA06B21FDD85145" box="[460,521,1322,1348]" gridcol="1" gridrow="11" pageId="11" pageNumber="12">7.5</td>
<td id="09065400A263007A9D436B21FC515145" box="[559,896,1322,1348]" gridcol="2" gridrow="11" pageId="11" pageNumber="12">4.3 ± 0.93 (2.5–7.5); 52</td>
<td id="09065400A263007A9CCB6B21FC325145" box="[935,995,1322,1348]" gridcol="3" gridrow="11" pageId="11" pageNumber="12">4.0</td>
<td id="09065400A263007A9B666B21FAB45145" box="[1034,1381,1322,1348]" gridcol="4" gridrow="11" pageId="11" pageNumber="12">4.4 ± 0.95 (2.1–6.2); 34</td>
<td id="09065400A263007A9AE86B21F90E5145" box="[1412,1759,1322,1348]" gridcol="5" gridrow="11" pageId="11" pageNumber="12">4.3 ± 1.08 (2.0–7.1); 59</td>
<td id="09065400A263007A98766B21F8B05145" box="[1818,1889,1322,1348]" gridcol="6" gridrow="11" pageId="11" pageNumber="12">?</td>
</tr>
<tr id="4AD73D7CA263007A9FAB6B54F8B05178" box="[199,1889,1375,1401]" gridrow="12" pageId="11" pageNumber="12">
<th id="09065400A263007A9FAB6B54FE765178" box="[199,423,1375,1401]" gridcol="0" gridrow="12" pageId="11" pageNumber="12">
Cirri 
<emphasis id="C693E32CA263FF8E9E6C6B54FEC15178" box="[256,272,1375,1401]" italics="true" pageId="11" pageNumber="12">E</emphasis>
</th>
<td id="09065400A263007A9EA06B54FDD85178" box="[460,521,1375,1401]" gridcol="1" gridrow="12" pageId="11" pageNumber="12">26.2</td>
<td id="09065400A263007A9D436B54FC515178" box="[559,896,1375,1401]" gridcol="2" gridrow="12" pageId="11" pageNumber="12">19.5 ± 4.01 (9.1–26.2); 29</td>
<td id="09065400A263007A9CCB6B54FC325178" box="[935,995,1375,1401]" gridcol="3" gridrow="12" pageId="11" pageNumber="12">14.3</td>
<td id="09065400A263007A9B666B54FAB45178" box="[1034,1381,1375,1401]" gridcol="4" gridrow="12" pageId="11" pageNumber="12">20.3 ± 4.02 (10.3–27.1); 26</td>
<td id="09065400A263007A9AE86B54F90E5178" box="[1412,1759,1375,1401]" gridcol="5" gridrow="12" pageId="11" pageNumber="12">17.3 ± 3.39 (10.3–23.1); 28</td>
<td id="09065400A263007A98766B54F8B05178" box="[1818,1889,1375,1401]" gridcol="6" gridrow="12" pageId="11" pageNumber="12">?</td>
</tr>
<tr id="4AD73D7CA263007A9FAB6B9FF8B051AF" box="[199,1889,1428,1454]" gridrow="13" pageId="11" pageNumber="12">
<th id="09065400A263007A9FAB6B9FFE7651AF" box="[199,423,1428,1454]" gridcol="0" gridrow="13" pageId="11" pageNumber="12">Leg I spine</th>
<td id="09065400A263007A9EA06B9FFDD851AF" box="[460,521,1428,1454]" gridcol="1" gridrow="13" pageId="11" pageNumber="12">12.5</td>
<td id="09065400A263007A9D436B9FFC5151AF" box="[559,896,1428,1454]" gridcol="2" gridrow="13" pageId="11" pageNumber="12">10.5 ± 2.19 (6.1–15.0); 40</td>
<td id="09065400A263007A9CCB6B9FFC3251AF" box="[935,995,1428,1454]" gridcol="3" gridrow="13" pageId="11" pageNumber="12">11.0</td>
<td id="09065400A263007A9B666B9FFAB451AF" box="[1034,1381,1428,1454]" gridcol="4" gridrow="13" pageId="11" pageNumber="12">11.0 ± 2.87 (5.5–17.7); 29</td>
<td id="09065400A263007A9AE86B9FF90E51AF" box="[1412,1759,1428,1454]" gridcol="5" gridrow="13" pageId="11" pageNumber="12">10.9 ± 2.83 (4.9–17.7); 42</td>
<td id="09065400A263007A98766B9FF8B051AF" box="[1818,1889,1428,1454]" gridcol="6" gridrow="13" pageId="11" pageNumber="12">?</td>
</tr>
<tr id="4AD73D7CA263007A9FAB6BC2F8B051E2" box="[199,1889,1481,1507]" gridrow="14" pageId="11" pageNumber="12">
<th id="09065400A263007A9FAB6BC2FE7651E2" box="[199,423,1481,1507]" gridcol="0" gridrow="14" pageId="11" pageNumber="12">Leg II spine</th>
<td id="09065400A263007A9EA06BC2FDD851E2" box="[460,521,1481,1507]" gridcol="1" gridrow="14" pageId="11" pageNumber="12">16.1</td>
<td id="09065400A263007A9D436BC2FC5151E2" box="[559,896,1481,1507]" gridcol="2" gridrow="14" pageId="11" pageNumber="12">12.7 ± 3.82 (5.1–22.0); 30</td>
<td id="09065400A263007A9CCB6BC2FC3251E2" box="[935,995,1481,1507]" gridcol="3" gridrow="14" pageId="11" pageNumber="12">8.6</td>
<td id="09065400A263007A9B666BC2FAB451E2" box="[1034,1381,1481,1507]" gridcol="4" gridrow="14" pageId="11" pageNumber="12">13.0 ± 3.65 (7.1–22.3); 24</td>
<td id="09065400A263007A9AE86BC2F90E51E2" box="[1412,1759,1481,1507]" gridcol="5" gridrow="14" pageId="11" pageNumber="12">13.5 ± 3.50 (5.0–18.7); 22</td>
<td id="09065400A263007A98766BC2F8B051E2" box="[1818,1889,1481,1507]" gridcol="6" gridrow="14" pageId="11" pageNumber="12">?</td>
</tr>
<tr id="4AD73D7CA263007A9FAB6BF5F8B05219" box="[199,1889,1534,1560]" gridrow="15" pageId="11" pageNumber="12">
<th id="09065400A263007A9FAB6BF5FE765219" box="[199,423,1534,1560]" gridcol="0" gridrow="15" pageId="11" pageNumber="12">Leg III spine</th>
<td id="09065400A263007A9EA06BF5FDD85219" box="[460,521,1534,1560]" gridcol="1" gridrow="15" pageId="11" pageNumber="12">18.0</td>
<td id="09065400A263007A9D436BF5FC515219" box="[559,896,1534,1560]" gridcol="2" gridrow="15" pageId="11" pageNumber="12">12.8 ± 3.61 (4.6–19.9); 32</td>
<td id="09065400A263007A9CCB6BF5FC325219" box="[935,995,1534,1560]" gridcol="3" gridrow="15" pageId="11" pageNumber="12">8.0</td>
<td id="09065400A263007A9B666BF5FAB45219" box="[1034,1381,1534,1560]" gridcol="4" gridrow="15" pageId="11" pageNumber="12">12.8 ± 3.19 (8.0–19.1); 25</td>
<td id="09065400A263007A9AE86BF5F90E5219" box="[1412,1759,1534,1560]" gridcol="5" gridrow="15" pageId="11" pageNumber="12">13.0 ± 3.69 (7.6–21.4); 25</td>
<td id="09065400A263007A98766BF5F8B05219" box="[1818,1889,1534,1560]" gridcol="6" gridrow="15" pageId="11" pageNumber="12">?</td>
</tr>
<tr id="4AD73D7CA263007A9FAB6838F8B0524E" box="[199,1889,1587,1615]" gridrow="16" pageId="11" pageNumber="12">
<th id="09065400A263007A9FAB6838FE76524E" box="[199,423,1587,1615]" gridcol="0" gridrow="16" pageId="11" pageNumber="12">Leg IV sensory organ</th>
<td id="09065400A263007A9EA06838FDD8524E" box="[460,521,1587,1615]" gridcol="1" gridrow="16" pageId="11" pageNumber="12">21.6</td>
<td id="09065400A263007A9D436838FC51524E" box="[559,896,1587,1615]" gridcol="2" gridrow="16" pageId="11" pageNumber="12">18.8 ± 3.70 (9.1–27.9); 43</td>
<td id="09065400A263007A9CCB6838FC32524E" box="[935,995,1587,1615]" gridcol="3" gridrow="16" pageId="11" pageNumber="12">18.6</td>
<td id="09065400A263007A9B666838FAB4524E" box="[1034,1381,1587,1615]" gridcol="4" gridrow="16" pageId="11" pageNumber="12">19.3 ± 4.09 (8.6–25.4); 33</td>
<td id="09065400A263007A9AE86838F90E524E" box="[1412,1759,1587,1615]" gridcol="5" gridrow="16" pageId="11" pageNumber="12">19.1 ± 3.06 (10.4–26.5); 62</td>
<td id="09065400A263007A98766838F8B0524E" box="[1818,1889,1587,1615]" gridcol="6" gridrow="16" pageId="11" pageNumber="12">6.6</td>
</tr>
<tr id="4AD73D7CA263007A9FAB6863F8B05283" box="[199,1889,1640,1666]" gridrow="17" pageId="11" pageNumber="12">
<th id="09065400A263007A9FAB6863FE765283" box="[199,423,1640,1666]" gridcol="0" gridrow="17" pageId="11" pageNumber="12">Leg IV base</th>
<td id="09065400A263007A9EA06863FDD85283" box="[460,521,1640,1666]" gridcol="1" gridrow="17" pageId="11" pageNumber="12">7.1</td>
<td id="09065400A263007A9D436863FC515283" box="[559,896,1640,1666]" gridcol="2" gridrow="17" pageId="11" pageNumber="12">8.1 ± 2.22 (3.2–16.8); 42</td>
<td id="09065400A263007A9CCB6863FC325283" box="[935,995,1640,1666]" gridcol="3" gridrow="17" pageId="11" pageNumber="12">9.4</td>
<td id="09065400A263007A9B666863FAB45283" box="[1034,1381,1640,1666]" gridcol="4" gridrow="17" pageId="11" pageNumber="12">8.8 ± 2.43 (2.6–13.2); 32</td>
<td id="09065400A263007A9AE86863F90E5283" box="[1412,1759,1640,1666]" gridcol="5" gridrow="17" pageId="11" pageNumber="12">8.1 ± 1.99 (3.5–12.0); 61</td>
<td id="09065400A263007A98766863F8B05283" box="[1818,1889,1640,1666]" gridcol="6" gridrow="17" pageId="11" pageNumber="12">3.6</td>
</tr>
<tr id="4AD73D7CA263007A9FAB6896F8B052B6" box="[199,1889,1693,1719]" gridrow="18" pageId="11" pageNumber="12">
<th id="09065400A263007A9FAB6896FE7652B6" box="[199,423,1693,1719]" gridcol="0" gridrow="18" pageId="11" pageNumber="12">Leg IV spine</th>
<td id="09065400A263007A9EA06896FDD852B6" box="[460,521,1693,1719]" gridcol="1" gridrow="18" pageId="11" pageNumber="12">14.5</td>
<td id="09065400A263007A9D436896FC5152B6" box="[559,896,1693,1719]" gridcol="2" gridrow="18" pageId="11" pageNumber="12">10.9 ± 2.12 (5.5–15.3); 42</td>
<td id="09065400A263007A9CCB6896FC3252B6" box="[935,995,1693,1719]" gridcol="3" gridrow="18" pageId="11" pageNumber="12">9.2</td>
<td id="09065400A263007A9B666896FAB452B6" box="[1034,1381,1693,1719]" gridcol="4" gridrow="18" pageId="11" pageNumber="12">10.7 ± 1.81 (6.0–14.6); 32</td>
<td id="09065400A263007A9AE86896F90E52B6" box="[1412,1759,1693,1719]" gridcol="5" gridrow="18" pageId="11" pageNumber="12">11.0 ± 1.74 (6.4–15.4); 61</td>
<td id="09065400A263007A98766896F8B052B6" box="[1818,1889,1693,1719]" gridcol="6" gridrow="18" pageId="11" pageNumber="12">3.0</td>
</tr>
</table>
</paragraph>
<caption id="A0986FB6A264FF899FD06C6FFE7C5681" box="[188,429,612,640]" pageId="12" pageNumber="13">
<paragraph id="F4583F3EA264FF899FD06C6FFE7C5681" blockId="12.[188,429,612,640]" box="[188,429,612,640]" pageId="12" pageNumber="13">
<emphasis id="C693E32CA264FF899FD06C6FFEC65681" bold="true" box="[188,279,612,640]" pageId="12" pageNumber="13">Table 2</emphasis>
(continued).
</paragraph>
</caption>
<paragraph id="F4583F3EA264FF899FAB6CB0F895529C" pageId="12" pageNumber="13">
<table id="86E7CD9EA264007A9FAB6CB0F8B052BD" box="[199,1889,699,1724]" colsContinueFrom="11.[199,1889,734,1719]" gridcols="7" gridrows="18" pageId="12" pageNumber="13">
<tr id="4AD73D7CA264007A9FAB6CB0F8B056D4" box="[199,1889,699,725]" gridrow="0" pageId="12" pageNumber="13" rowspan-1="1" rowspan-3="1">
<th id="09065400A264007A9FAB6CB0FE5656D4" box="[199,391,699,725]" gridcol="0" gridrow="0" pageId="12" pageNumber="13">
<emphasis id="C693E32CA264FF899FAB6CB0FEEF56D4" bold="true" box="[199,318,699,725]" pageId="12" pageNumber="13">Structures</emphasis>
</th>
<th id="09065400A264007A9D2A6CB0FCBA56D4" box="[582,875,699,725]" gridcol="2" gridrow="0" pageId="12" pageNumber="13">
<emphasis id="C693E32CA264FF899D1D6CB0FD1F56D4" bold="true" box="[625,718,699,725]" pageId="12" pageNumber="13">Females</emphasis>
</th>
<th id="09065400A264007A9B496CB0FA9A56D4" box="[1061,1355,699,725]" gridcol="4" gridrow="0" pageId="12" pageNumber="13">
<emphasis id="C693E32CA264FF899B306CB0FB7156D4" bold="true" box="[1116,1184,699,725]" pageId="12" pageNumber="13">Males</emphasis>
</th>
<th id="09065400A264007A9AF36CB0F91456D4" box="[1439,1733,699,725]" gridcol="5" gridrow="0" pageId="12" pageNumber="13">
<emphasis id="C693E32CA264FF899A916CB0F9B756D4" bold="true" box="[1533,1638,699,725]" pageId="12" pageNumber="13">Juveniles</emphasis>
</th>
<th id="09065400A264007A98766CB0F8B056D4" box="[1818,1889,699,725]" gridcol="6" gridrow="0" pageId="12" pageNumber="13">
<emphasis id="C693E32CA264FF8998776CB0F8B056D4" bold="true" box="[1819,1889,699,725]" pageId="12" pageNumber="13">Larva</emphasis>
</th>
</tr>
<tr id="4AD73D7CA264007A9FAB6CFFF8B0570F" box="[199,1889,756,782]" gridrow="1" pageId="12" pageNumber="13" rowspan-0="1" rowspan-1="1" rowspan-3="1">
<td id="09065400A264007A9D2A6CFFFCBA570F" box="[582,875,756,782]" gridcol="2" gridrow="1" pageId="12" pageNumber="13">
<emphasis id="C693E32CA264FF899D166CFFFD17570F" bold="true" box="[634,710,756,782]" pageId="12" pageNumber="13">Adults</emphasis>
</td>
<td id="09065400A264007A9B496CFFFA9A570F" box="[1061,1355,756,782]" gridcol="4" gridrow="1" pageId="12" pageNumber="13">
<emphasis id="C693E32CA264FF899B356CFFFB74570F" bold="true" box="[1113,1189,756,782]" pageId="12" pageNumber="13">Adults</emphasis>
</td>
<td id="09065400A264007A9AF36CFFF914570F" box="[1439,1733,756,782]" gridcol="5" gridrow="1" pageId="12" pageNumber="13">
<emphasis id="C693E32CA264FF8999626CFFF985570F" bold="true" box="[1550,1620,756,782]" pageId="12" pageNumber="13">6-toed</emphasis>
</td>
<td id="09065400A264007A98766CFFF8B0570F" box="[1818,1889,756,782]" gridcol="6" gridrow="1" pageId="12" pageNumber="13">
<emphasis id="C693E32CA264FF8998766CFFF8B0570F" bold="true" box="[1818,1889,756,782]" pageId="12" pageNumber="13">4-toed</emphasis>
</td>
</tr>
<tr id="4AD73D7CA264007A9FAB6D17F8B05737" box="[199,1889,796,822]" gridrow="2" pageId="12" pageNumber="13" rowspan-0="1" rowspan-6="1">
<td id="09065400A264007A9EA36D17FDD75737" box="[463,518,796,822]" gridcol="1" gridrow="2" pageId="12" pageNumber="13">
<emphasis id="C693E32CA264FF899EA36D17FDD75737" bold="true" box="[463,518,796,822]" pageId="12" pageNumber="13">Holo</emphasis>
</td>
<td id="09065400A264007A9D2A6D17FCBA5737" box="[582,875,796,822]" gridcol="2" gridrow="2" pageId="12" pageNumber="13">
<emphasis id="C693E32CA264FF899D3A6D17FC885737" bold="true" box="[598,857,796,822]" pageId="12" pageNumber="13">Mean ± SD (Range); N</emphasis>
</td>
<td id="09065400A264007A9CC26D17FC0C5737" box="[942,989,796,822]" gridcol="3" gridrow="2" pageId="12" pageNumber="13">
<emphasis id="C693E32CA264FF899CC26D17FC0C5737" bold="true" box="[942,989,796,822]" pageId="12" pageNumber="13">Allo</emphasis>
</td>
<td id="09065400A264007A9B496D17FA9A5737" box="[1061,1355,796,822]" gridcol="4" gridrow="2" pageId="12" pageNumber="13">
<emphasis id="C693E32CA264FF899B5A6D17FAE95737" bold="true" box="[1078,1336,796,822]" pageId="12" pageNumber="13">Mean ± SD (Range); N</emphasis>
</td>
<td id="09065400A264007A9AF36D17F9145737" box="[1439,1733,796,822]" gridcol="5" gridrow="2" pageId="12" pageNumber="13">
<emphasis id="C693E32CA264FF899ADC6D17F9635737" bold="true" box="[1456,1714,796,822]" pageId="12" pageNumber="13">Mean ± SD (Range); N</emphasis>
</td>
</tr>
<tr id="4AD73D7CA264007A9FAB6D4FF8B0575F" box="[199,1889,836,862]" gridrow="3" pageId="12" pageNumber="13">
<th id="09065400A264007A9FAB6D4FFE56575F" box="[199,391,836,862]" gridcol="0" gridrow="3" pageId="12" pageNumber="13">Body projection 1</th>
<td id="09065400A264007A9EA36D4FFDD7575F" box="[463,518,836,862]" gridcol="1" gridrow="3" pageId="12" pageNumber="13">5.5</td>
<td id="09065400A264007A9D2A6D4FFCBA575F" box="[582,875,836,862]" gridcol="2" gridrow="3" pageId="12" pageNumber="13">4.6 ± 1.43 (1.9–7.2); 46</td>
<td id="09065400A264007A9CC26D4FFC0C575F" box="[942,989,836,862]" gridcol="3" gridrow="3" pageId="12" pageNumber="13">6.2</td>
<td id="09065400A264007A9B496D4FFA9A575F" box="[1061,1355,836,862]" gridcol="4" gridrow="3" pageId="12" pageNumber="13">4.7 ± 1.51 (2.0–7.8); 32</td>
<td id="09065400A264007A9AF36D4FF914575F" box="[1439,1733,836,862]" gridcol="5" gridrow="3" pageId="12" pageNumber="13">4.4 ± 1.19 (2.0–6.5); 45</td>
<td id="09065400A264007A98766D4FF8B0575F" box="[1818,1889,836,862]" gridcol="6" gridrow="3" pageId="12" pageNumber="13">1.4</td>
</tr>
<tr id="4AD73D7CA264007A9FAB6D72F8B05792" box="[199,1889,889,915]" gridrow="4" pageId="12" pageNumber="13">
<th id="09065400A264007A9FAB6D72FE565792" box="[199,391,889,915]" gridcol="0" gridrow="4" pageId="12" pageNumber="13">Body projection 2</th>
<td id="09065400A264007A9EA36D72FDD75792" box="[463,518,889,915]" gridcol="1" gridrow="4" pageId="12" pageNumber="13">?</td>
<td id="09065400A264007A9D2A6D72FCBA5792" box="[582,875,889,915]" gridcol="2" gridrow="4" pageId="12" pageNumber="13">4.6 ± 1.06 (3.5–6.5); 5</td>
<td id="09065400A264007A9CC26D72FC0C5792" box="[942,989,889,915]" gridcol="3" gridrow="4" pageId="12" pageNumber="13">?</td>
<td id="09065400A264007A9B496D72FA9A5792" box="[1061,1355,889,915]" gridcol="4" gridrow="4" pageId="12" pageNumber="13">5.8 ± 1.95 (3.0–9.3); 7</td>
<td id="09065400A264007A9AF36D72F9145792" box="[1439,1733,889,915]" gridcol="5" gridrow="4" pageId="12" pageNumber="13">5.1 ± 1.44 (2.9–7.1); 7</td>
<td id="09065400A264007A98766D72F8B05792" box="[1818,1889,889,915]" gridcol="6" gridrow="4" pageId="12" pageNumber="13">?</td>
</tr>
<tr id="4AD73D7CA264007A9FAB6DA6F8B057C6" box="[199,1889,941,967]" gridrow="5" pageId="12" pageNumber="13">
<th id="09065400A264007A9FAB6DA6FE5657C6" box="[199,391,941,967]" gridcol="0" gridrow="5" pageId="12" pageNumber="13">Body projection 3</th>
<td id="09065400A264007A9EA36DA6FDD757C6" box="[463,518,941,967]" gridcol="1" gridrow="5" pageId="12" pageNumber="13">?</td>
<td id="09065400A264007A9D2A6DA6FCBA57C6" box="[582,875,941,967]" gridcol="2" gridrow="5" pageId="12" pageNumber="13">5.7 ± 1.33 (4.3–8.6); 8</td>
<td id="09065400A264007A9CC26DA6FC0C57C6" box="[942,989,941,967]" gridcol="3" gridrow="5" pageId="12" pageNumber="13">?</td>
<td id="09065400A264007A9B496DA6FA9A57C6" box="[1061,1355,941,967]" gridcol="4" gridrow="5" pageId="12" pageNumber="13">6.4 ± 1.57 (3.5–8.5); 10</td>
<td id="09065400A264007A9AF36DA6F91457C6" box="[1439,1733,941,967]" gridcol="5" gridrow="5" pageId="12" pageNumber="13">6.4 ± 3.09 (2.8–11.8); 9</td>
<td id="09065400A264007A98766DA6F8B057C6" box="[1818,1889,941,967]" gridcol="6" gridrow="5" pageId="12" pageNumber="13">3.2</td>
</tr>
<tr id="4AD73D7CA264007A9FAB6DE9F8B057FD" box="[199,1889,994,1020]" gridrow="6" pageId="12" pageNumber="13">
<th id="09065400A264007A9FAB6DE9FE5657FD" box="[199,391,994,1020]" gridcol="0" gridrow="6" pageId="12" pageNumber="13">Body projection 4</th>
<td id="09065400A264007A9EA36DE9FDD757FD" box="[463,518,994,1020]" gridcol="1" gridrow="6" pageId="12" pageNumber="13">6.8</td>
<td id="09065400A264007A9D2A6DE9FCBA57FD" box="[582,875,994,1020]" gridcol="2" gridrow="6" pageId="12" pageNumber="13">8.6 ±2.05 (5.3–14.6); 46</td>
<td id="09065400A264007A9CC26DE9FC0C57FD" box="[942,989,994,1020]" gridcol="3" gridrow="6" pageId="12" pageNumber="13">10.6</td>
<td id="09065400A264007A9B496DE9FA9A57FD" box="[1061,1355,994,1020]" gridcol="4" gridrow="6" pageId="12" pageNumber="13">8.7 ±2.60 (4.4–17.2); 30</td>
<td id="09065400A264007A9AF36DE9F91457FD" box="[1439,1733,994,1020]" gridcol="5" gridrow="6" pageId="12" pageNumber="13">7.5 ±1.95 (4.3–13.2); 48</td>
<td id="09065400A264007A98766DE9F8B057FD" box="[1818,1889,994,1020]" gridcol="6" gridrow="6" pageId="12" pageNumber="13">4.9</td>
</tr>
<tr id="4AD73D7CA264007A9FAB6A1CF8B05030" box="[199,1889,1047,1073]" gridrow="7" pageId="12" pageNumber="13">
<th id="09065400A264007A9FAB6A1CFE565030" box="[199,391,1047,1073]" gridcol="0" gridrow="7" pageId="12" pageNumber="13">Caudal apparatus</th>
<td id="09065400A264007A9EA36A1CFDD75030" box="[463,518,1047,1073]" gridcol="1" gridrow="7" pageId="12" pageNumber="13">12.4</td>
<td id="09065400A264007A9D2A6A1CFCBA5030" box="[582,875,1047,1073]" gridcol="2" gridrow="7" pageId="12" pageNumber="13">11.9 ± 2.89 (4.8–18.6); 54</td>
<td id="09065400A264007A9CC26A1CFC0C5030" box="[942,989,1047,1073]" gridcol="3" gridrow="7" pageId="12" pageNumber="13">12.5</td>
<td id="09065400A264007A9B496A1CFA9A5030" box="[1061,1355,1047,1073]" gridcol="4" gridrow="7" pageId="12" pageNumber="13">12.6 ± 2.90 (6.0–19.7); 34</td>
<td id="09065400A264007A9AF36A1CF9145030" box="[1439,1733,1047,1073]" gridcol="5" gridrow="7" pageId="12" pageNumber="13">10.3 ± 3.44 (4.5–19.0); 69</td>
<td id="09065400A264007A98766A1CF8B05030" box="[1818,1889,1047,1073]" gridcol="6" gridrow="7" pageId="12" pageNumber="13">8.2</td>
</tr>
<tr id="4AD73D7CA264007A9FAB6A47F8B05067" box="[199,1889,1100,1126]" gridrow="8" pageId="12" pageNumber="13">
<th id="09065400A264007A9FAB6A47FE565067" box="[199,391,1100,1126]" gridcol="0" gridrow="8" pageId="12" pageNumber="13">Toe 2</th>
<td id="09065400A264007A9EA36A47FDD75067" box="[463,518,1100,1126]" gridcol="1" gridrow="8" pageId="12" pageNumber="13">7.4</td>
<td id="09065400A264007A9D2A6A47FCBA5067" box="[582,875,1100,1126]" gridcol="2" gridrow="8" pageId="12" pageNumber="13">5.5 ± 1.23 (2.5–8.4); 39</td>
<td id="09065400A264007A9CC26A47FC0C5067" box="[942,989,1100,1126]" gridcol="3" gridrow="8" pageId="12" pageNumber="13">4.5</td>
<td id="09065400A264007A9B496A47FA9A5067" box="[1061,1355,1100,1126]" gridcol="4" gridrow="8" pageId="12" pageNumber="13">5.6 ± 1.15 (3.7–8.4); 29</td>
<td id="09065400A264007A9AF36A47F9145067" box="[1439,1733,1100,1126]" gridcol="5" gridrow="8" pageId="12" pageNumber="13">5.4 ± 1.27 (3.0–7.8); 42</td>
<td id="09065400A264007A98766A47F8B05067" box="[1818,1889,1100,1126]" gridcol="6" gridrow="8" pageId="12" pageNumber="13">–</td>
</tr>
<tr id="4AD73D7CA264007A9FAB6A8AF8B0509A" box="[199,1889,1153,1179]" gridrow="9" pageId="12" pageNumber="13">
<th id="09065400A264007A9FAB6A8AFE56509A" box="[199,391,1153,1179]" gridcol="0" gridrow="9" pageId="12" pageNumber="13">Toe 3</th>
<td id="09065400A264007A9EA36A8AFDD7509A" box="[463,518,1153,1179]" gridcol="1" gridrow="9" pageId="12" pageNumber="13">24.1</td>
<td id="09065400A264007A9D2A6A8AFCBA509A" box="[582,875,1153,1179]" gridcol="2" gridrow="9" pageId="12" pageNumber="13">18.0 ± 3.51 (9.7–24.5); 40</td>
<td id="09065400A264007A9CC26A8AFC0C509A" box="[942,989,1153,1179]" gridcol="3" gridrow="9" pageId="12" pageNumber="13">13.7</td>
<td id="09065400A264007A9B496A8AFA9A509A" box="[1061,1355,1153,1179]" gridcol="4" gridrow="9" pageId="12" pageNumber="13">17.7 ± 3.44 (11.5–24.9); 29</td>
<td id="09065400A264007A9AF36A8AF914509A" box="[1439,1733,1153,1179]" gridcol="5" gridrow="9" pageId="12" pageNumber="13">17.4 ± 2.81 (9.3–22.0); 42</td>
<td id="09065400A264007A98766A8AF8B0509A" box="[1818,1889,1153,1179]" gridcol="6" gridrow="9" pageId="12" pageNumber="13">12</td>
</tr>
<tr id="4AD73D7CA264007A9FAB6ABDF8B050D1" box="[199,1889,1206,1232]" gridrow="10" pageId="12" pageNumber="13">
<th id="09065400A264007A9FAB6ABDFE5650D1" box="[199,391,1206,1232]" gridcol="0" gridrow="10" pageId="12" pageNumber="13">Toe 4</th>
<td id="09065400A264007A9EA36ABDFDD750D1" box="[463,518,1206,1232]" gridcol="1" gridrow="10" pageId="12" pageNumber="13">15.0</td>
<td id="09065400A264007A9D2A6ABDFCBA50D1" box="[582,875,1206,1232]" gridcol="2" gridrow="10" pageId="12" pageNumber="13">11.4 ± 2.26 (6.9–15.7); 40</td>
<td id="09065400A264007A9CC26ABDFC0C50D1" box="[942,989,1206,1232]" gridcol="3" gridrow="10" pageId="12" pageNumber="13">7.9</td>
<td id="09065400A264007A9B496ABDFA9A50D1" box="[1061,1355,1206,1232]" gridcol="4" gridrow="10" pageId="12" pageNumber="13">11.6 ± 2.18 (7.9–15.6); 29</td>
<td id="09065400A264007A9AF36ABDF91450D1" box="[1439,1733,1206,1232]" gridcol="5" gridrow="10" pageId="12" pageNumber="13">11.2 ± 2.03 (5.6–14.6);42</td>
<td id="09065400A264007A98766ABDF8B050D1" box="[1818,1889,1206,1232]" gridcol="6" gridrow="10" pageId="12" pageNumber="13">7.3</td>
</tr>
<tr id="4AD73D7CA264007A9FAB6AE0F8B05104" box="[199,1889,1259,1285]" gridrow="11" pageId="12" pageNumber="13">
<th id="09065400A264007A9FAB6AE0FE565104" box="[199,391,1259,1285]" gridcol="0" gridrow="11" pageId="12" pageNumber="13">Toe 5</th>
<td id="09065400A264007A9EA36AE0FDD75104" box="[463,518,1259,1285]" gridcol="1" gridrow="11" pageId="12" pageNumber="13">28.4</td>
<td id="09065400A264007A9D2A6AE0FCBA5104" box="[582,875,1259,1285]" gridcol="2" gridrow="11" pageId="12" pageNumber="13">20.9 ± 3.70 (13.8–28.4); 40</td>
<td id="09065400A264007A9CC26AE0FC0C5104" box="[942,989,1259,1285]" gridcol="3" gridrow="11" pageId="12" pageNumber="13">17.7</td>
<td id="09065400A264007A9B496AE0FA9A5104" box="[1061,1355,1259,1285]" gridcol="4" gridrow="11" pageId="12" pageNumber="13">20.8 ± 3.77 (14.4–28.2); 29</td>
<td id="09065400A264007A9AF36AE0F9145104" box="[1439,1733,1259,1285]" gridcol="5" gridrow="11" pageId="12" pageNumber="13">20.3 ± 3.42 (10.6–26.8);42</td>
<td id="09065400A264007A98766AE0F8B05104" box="[1818,1889,1259,1285]" gridcol="6" gridrow="11" pageId="12" pageNumber="13">–</td>
</tr>
<tr id="4AD73D7CA264007A9FAB6B2BF8B0513B" box="[199,1889,1312,1338]" gridrow="12" pageId="12" pageNumber="13">
<th id="09065400A264007A9FAB6B2BFE56513B" box="[199,391,1312,1338]" gridcol="0" gridrow="12" pageId="12" pageNumber="13">Toe 6</th>
<td id="09065400A264007A9EA36B2BFDD7513B" box="[463,518,1312,1338]" gridcol="1" gridrow="12" pageId="12" pageNumber="13">15.3</td>
<td id="09065400A264007A9D2A6B2BFCBA513B" box="[582,875,1312,1338]" gridcol="2" gridrow="12" pageId="12" pageNumber="13">13.2 ± 2.26 (7.8–16.4); 40</td>
<td id="09065400A264007A9CC26B2BFC0C513B" box="[942,989,1312,1338]" gridcol="3" gridrow="12" pageId="12" pageNumber="13">10.6</td>
<td id="09065400A264007A9B496B2BFA9A513B" box="[1061,1355,1312,1338]" gridcol="4" gridrow="12" pageId="12" pageNumber="13">13.2 ± 2.58 (8.8–18.3); 29</td>
<td id="09065400A264007A9AF36B2BF914513B" box="[1439,1733,1312,1338]" gridcol="5" gridrow="12" pageId="12" pageNumber="13">13.2 ± 2.41 (7.9–18.3); 42</td>
<td id="09065400A264007A98766B2BF8B0513B" box="[1818,1889,1312,1338]" gridcol="6" gridrow="12" pageId="12" pageNumber="13">11.3</td>
</tr>
<tr id="4AD73D7CA264007A9FAB6B5EF8B0516E" box="[199,1889,1365,1391]" gridrow="13" pageId="12" pageNumber="13">
<th id="09065400A264007A9FAB6B5EFE56516E" box="[199,391,1365,1391]" gridcol="0" gridrow="13" pageId="12" pageNumber="13">Leg IV: Toe 1</th>
<td id="09065400A264007A9EA36B5EFDD7516E" box="[463,518,1365,1391]" gridcol="1" gridrow="13" pageId="12" pageNumber="13">17.9</td>
<td id="09065400A264007A9D2A6B5EFCBA516E" box="[582,875,1365,1391]" gridcol="2" gridrow="13" pageId="12" pageNumber="13">16.6 ± 2.71 (11.0–21.6); 45</td>
<td id="09065400A264007A9CC26B5EFC0C516E" box="[942,989,1365,1391]" gridcol="3" gridrow="13" pageId="12" pageNumber="13">14.9</td>
<td id="09065400A264007A9B496B5EFA9A516E" box="[1061,1355,1365,1391]" gridcol="4" gridrow="13" pageId="12" pageNumber="13">16.4 ± 3.04 (9.2–22.2); 33</td>
<td id="09065400A264007A9AF36B5EF914516E" box="[1439,1733,1365,1391]" gridcol="5" gridrow="13" pageId="12" pageNumber="13">16.4 ± 2.83 (10.0–20.1); 51</td>
<td id="09065400A264007A98766B5EF8B0516E" box="[1818,1889,1365,1391]" gridcol="6" gridrow="13" pageId="12" pageNumber="13">10.2</td>
</tr>
<tr id="4AD73D7CA264007A9FAB6B81F8B051A5" box="[199,1889,1418,1444]" gridrow="14" pageId="12" pageNumber="13">
<th id="09065400A264007A9FAB6B81FE5651A5" box="[199,391,1418,1444]" gridcol="0" gridrow="14" pageId="12" pageNumber="13">Toe 2</th>
<td id="09065400A264007A9EA36B81FDD751A5" box="[463,518,1418,1444]" gridcol="1" gridrow="14" pageId="12" pageNumber="13">27.1</td>
<td id="09065400A264007A9D2A6B81FCBA51A5" box="[582,875,1418,1444]" gridcol="2" gridrow="14" pageId="12" pageNumber="13">23.5 ± 3.84 (15.5–30.2); 44</td>
<td id="09065400A264007A9CC26B81FC0C51A5" box="[942,989,1418,1444]" gridcol="3" gridrow="14" pageId="12" pageNumber="13">20.8</td>
<td id="09065400A264007A9B496B81FA9A51A5" box="[1061,1355,1418,1444]" gridcol="4" gridrow="14" pageId="12" pageNumber="13">23.0 ± 4.11 (15.7–29.8); 33</td>
<td id="09065400A264007A9AF36B81F91451A5" box="[1439,1733,1418,1444]" gridcol="5" gridrow="14" pageId="12" pageNumber="13">23.4 ± 3.57 (14.5–30.3); 54</td>
<td id="09065400A264007A98766B81F8B051A5" box="[1818,1889,1418,1444]" gridcol="6" gridrow="14" pageId="12" pageNumber="13">17</td>
</tr>
<tr id="4AD73D7CA264007A9FAB6BB4F8B051D8" box="[199,1889,1471,1497]" gridrow="15" pageId="12" pageNumber="13">
<th id="09065400A264007A9FAB6BB4FE5651D8" box="[199,391,1471,1497]" gridcol="0" gridrow="15" pageId="12" pageNumber="13">Toe 3</th>
<td id="09065400A264007A9EA36BB4FDD751D8" box="[463,518,1471,1497]" gridcol="1" gridrow="15" pageId="12" pageNumber="13">13.2</td>
<td id="09065400A264007A9D2A6BB4FCBA51D8" box="[582,875,1471,1497]" gridcol="2" gridrow="15" pageId="12" pageNumber="13">13.2 ± 2.41 (7.6–18.0); 45</td>
<td id="09065400A264007A9CC26BB4FC0C51D8" box="[942,989,1471,1497]" gridcol="3" gridrow="15" pageId="12" pageNumber="13">11.5</td>
<td id="09065400A264007A9B496BB4FA9A51D8" box="[1061,1355,1471,1497]" gridcol="4" gridrow="15" pageId="12" pageNumber="13">12.8 ± 2.36 (7.8–17.6); 33</td>
<td id="09065400A264007A9AF36BB4F91451D8" box="[1439,1733,1471,1497]" gridcol="5" gridrow="15" pageId="12" pageNumber="13">13.2 ± 2.31 (7.0–16.9); 48</td>
<td id="09065400A264007A98766BB4F8B051D8" box="[1818,1889,1471,1497]" gridcol="6" gridrow="15" pageId="12" pageNumber="13">–</td>
</tr>
<tr id="4AD73D7CA264007A9FAB6BF8F8B0520C" box="[199,1889,1523,1549]" gridrow="16" pageId="12" pageNumber="13">
<th id="09065400A264007A9FAB6BF8FE56520C" box="[199,391,1523,1549]" gridcol="0" gridrow="16" pageId="12" pageNumber="13">Toe 4</th>
<td id="09065400A264007A9EA36BF8FDD7520C" box="[463,518,1523,1549]" gridcol="1" gridrow="16" pageId="12" pageNumber="13">13.7</td>
<td id="09065400A264007A9D2A6BF8FCBA520C" box="[582,875,1523,1549]" gridcol="2" gridrow="16" pageId="12" pageNumber="13">13.1 ± 2.35 (7.8–18.2); 46</td>
<td id="09065400A264007A9CC26BF8FC0C520C" box="[942,989,1523,1549]" gridcol="3" gridrow="16" pageId="12" pageNumber="13">11.3</td>
<td id="09065400A264007A9B496BF8FA9A520C" box="[1061,1355,1523,1549]" gridcol="4" gridrow="16" pageId="12" pageNumber="13">12.8 ± 2.44 (7.7–17.4); 33</td>
<td id="09065400A264007A9AF36BF8F914520C" box="[1439,1733,1523,1549]" gridcol="5" gridrow="16" pageId="12" pageNumber="13">12.9 ± 2.13 (7.4–16.0); 48</td>
<td id="09065400A264007A98766BF8F8B0520C" box="[1818,1889,1523,1549]" gridcol="6" gridrow="16" pageId="12" pageNumber="13">–</td>
</tr>
<tr id="4AD73D7CA264007A9FAB6823F8B052BD" box="[199,1889,1576,1724]" gridrow="17" pageId="12" pageNumber="13">
<th id="09065400A264007A9FAB6823FE5652BD" box="[199,391,1576,1724]" gridcol="0" gridrow="17" pageId="12" pageNumber="13">Toe 5 Toe 6 Gonopore-Anus distance</th>
<td id="09065400A264007A9EA36823FDD752BD" box="[463,518,1576,1724]" gridcol="1" gridrow="17" pageId="12" pageNumber="13">30.7 19.5 16.1</td>
<td id="09065400A264007A9D2A6823FCBA52BD" box="[582,875,1576,1724]" gridcol="2" gridrow="17" pageId="12" pageNumber="13">25.6 ± 4.04 (16.4–32.0); 46 17.0 ± 2.91 (9.1–21.5); 45 10.8 ± 3.00 (6.1–17.1); 55</td>
<td id="09065400A264007A9CC26823FC0C52BD" box="[942,989,1576,1724]" gridcol="3" gridrow="17" pageId="12" pageNumber="13">22.1 12.8 4.2</td>
<td id="09065400A264007A9B496823FA9A52BD" box="[1061,1355,1576,1724]" gridcol="4" gridrow="17" pageId="12" pageNumber="13">24.7 ± 4.21 (15.6–31.7); 33 16.5 ± 3.14 (10.6–22.4); 33 5.0 ± 1.09 (3.0–7.8); 36</td>
<td id="09065400A264007A9AF36823F91452BD" box="[1439,1733,1576,1724]" gridcol="5" gridrow="17" pageId="12" pageNumber="13">25.1 ± 4.33 (14.0–32.5); 50 16.3 ± 3.14 (9.4–21.3); 52 –</td>
<td id="09065400A264007A98766823F8B052BD" box="[1818,1889,1576,1724]" gridcol="6" gridrow="17" pageId="12" pageNumber="13">14.3 8.6 –</td>
</tr>
</table>
</paragraph>
<subSubSection id="BCFD6CB5A265FF8A9FD16F7BFB7A558A" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="description">
<paragraph id="F4583F3EA265FF889FD16F7BFE9F558B" blockId="13.[189,334,368,394]" box="[189,334,368,394]" pageId="13" pageNumber="14">
<heading id="AF108852A265FF889FD16F7BFE9F558B" bold="true" box="[189,334,368,394]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="C693E32CA265FF889FD16F7BFE9F558B" bold="true" box="[189,334,368,394]" pageId="13" pageNumber="14">Description</emphasis>
</heading>
</paragraph>
<paragraph id="F4583F3EA265FF8B9FD16F95FA865548" blockId="13.[189,1399,414,829]" lastBlockId="14.[189,1399,267,329]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<typeStatus id="2B5C819CA265FF889FD16F95FEFC55B9" box="[189,301,414,440]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
, female of robust body 
<quantity id="331F92DBA265FF889D2D6F95FD4D55B8" box="[577,668,414,441]" metricMagnitude="-1" metricUnit="m" metricValue="2.36" pageId="13" pageNumber="14" unit="mm" value="236.0">236 µm</quantity>
long (
<quantity id="331F92DBA265FF889D896F94FC8955B8" box="[741,856,415,441]" metricMagnitude="-1" metricUnit="m" metricValue="2.484" pageId="13" pageNumber="14" unit="mm" value="248.4">248.4 µm</quantity>
including the caudal apparatus) and 
<quantity id="331F92DBA265FF889A6A6F94FAA655B8" box="[1286,1399,415,441]" metricMagnitude="-1" metricUnit="m" metricValue="1.009" pageId="13" pageNumber="14" unit="mm" value="100.9">100.9 µm</quantity>
wide between the third and fourth pair of legs (
<figureCitation id="6CDC23BBA265FF889C626FC9FC8F55DD" box="[782,862,450,476]" captionStart="Fig" captionStartId="9.[189,232,1960,1986]" captionTargetBox="[193,1397,1034,1917]" captionTargetId="figure@9.[187,1397,1034,1917]" captionTargetPageId="9" captionText="Fig. 4. Batillipes lusitanus sp. nov. A. Schematic drawing, dorsal view. B. Habitus of the holotype, ♀, ventral view. Scale bars = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3806143" httpUri="https://zenodo.org/record/3806143/files/figure.png" pageId="13" pageNumber="14">Figs 4</figureCitation>
A–B, 
<tableCitation id="B9650A85A265FF889CC46FC9FBD255DD" box="[936,1027,450,476]" captionStart="Table 2" captionStartId="11.[188,257,564,592]" captionTargetPageId="11" captionText="Table 2 (continued on next page). Measurements (µm) of selected morphological structures of specimens of Batillipes lusitanus sp. nov. (Holo = Holotype; Allo = Allotype; SD = Standard deviation; ‘Range’ refers to the smallest and largest measured specimen / structure; N = number of specimens/structures measured)." pageId="13" pageNumber="14">Table 2</tableCitation>
for morphometrics). Eyes not observed in mounted specimens. Trapezoid head (
<figureCitation id="6CDC23BBA265FF889C4A6FEEFCA55601" box="[806,884,485,512]" captionStart="Fig" captionStartId="10.[189,232,1846,1872]" captionTargetBox="[191,1397,933,1805]" captionTargetId="figure@10.[191,1397,933,1805]" captionTargetPageId="10" captionText="Fig. 5. Batillipes lusitanus sp. nov. A–B, D. Holotype, ♀. A–B. Detail of the head. A. Dorsal view, showing internal cirrus (black arrowhead), lateral cirrus (asterisk), secondary clava (white arrowhead). B. Ventral view, showing primary clava (white arrowhead), external cirrus (black arrowhead). C. Detail of body lateral projections of a paratype (slide C.IX-46) (black arrowheads); D. Detail of leg sensory organ on leg IV. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3806145" httpUri="https://zenodo.org/record/3806145/files/figure.png" pageId="13" pageNumber="14">Figs 5</figureCitation>
A–B) separated from the body by a neck constriction followed by well-developed lateral processes (body projection 1). Head with eleven cephalic appendages: Internal cephalic cirri with cirrophores (ca 
<quantity id="331F92DBA265FF889C296C20FC465647" box="[837,919,555,582]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="13" pageNumber="14" unit="mm" value="3.4">3.4 µm</quantity>
long), are inserted dorsally on the frontal edge of the head (
<figureCitation id="6CDC23BBA265FF889EFC6C45FE3E5668" box="[400,495,590,617]" captionStart="Fig" captionStartId="10.[189,232,1846,1872]" captionTargetBox="[191,1397,933,1805]" captionTargetId="figure@10.[191,1397,933,1805]" captionTargetPageId="10" captionText="Fig. 5. Batillipes lusitanus sp. nov. A–B, D. Holotype, ♀. A–B. Detail of the head. A. Dorsal view, showing internal cirrus (black arrowhead), lateral cirrus (asterisk), secondary clava (white arrowhead). B. Ventral view, showing primary clava (white arrowhead), external cirrus (black arrowhead). C. Detail of body lateral projections of a paratype (slide C.IX-46) (black arrowheads); D. Detail of leg sensory organ on leg IV. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3806145" httpUri="https://zenodo.org/record/3806145/files/figure.png" pageId="13" pageNumber="14">Fig. 5A</figureCitation>
). External cephalic cirri, with indistinct cirrophores, inserted more ventrally, near the lateral cirri 
<emphasis id="C693E32CA265FF889EC96C79FE67568D" box="[421,438,626,652]" italics="true" pageId="13" pageNumber="14">A</emphasis>
and primary clavae (
<figureCitation id="6CDC23BBA265FF889DC56C79FCD5568D" box="[681,772,625,652]" captionStart="Fig" captionStartId="10.[189,232,1846,1872]" captionTargetBox="[191,1397,933,1805]" captionTargetId="figure@10.[191,1397,933,1805]" captionTargetPageId="10" captionText="Fig. 5. Batillipes lusitanus sp. nov. A–B, D. Holotype, ♀. A–B. Detail of the head. A. Dorsal view, showing internal cirrus (black arrowhead), lateral cirrus (asterisk), secondary clava (white arrowhead). B. Ventral view, showing primary clava (white arrowhead), external cirrus (black arrowhead). C. Detail of body lateral projections of a paratype (slide C.IX-46) (black arrowheads); D. Detail of leg sensory organ on leg IV. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3806145" httpUri="https://zenodo.org/record/3806145/files/figure.png" pageId="13" pageNumber="14">Fig. 5B</figureCitation>
). The median cirrus with cirrophore. The lateral cirrus 
<emphasis id="C693E32CA265FF889FD16C9EFF1E56AE" box="[189,207,661,687]" italics="true" pageId="13" pageNumber="14">A</emphasis>
located dorsally in relation to the unconstricted tubular primary clava. These two appendages share a common pedestal. A van der Land’s organ present at the base of primary clavae, which have a wrinkled surface and exhibit a terminal hole (
<figureCitation id="6CDC23BBA265FF889D0A6CD0FD6156F7" box="[614,688,731,758]" captionStart="Fig" captionStartId="10.[189,232,1846,1872]" captionTargetBox="[191,1397,933,1805]" captionTargetId="figure@10.[191,1397,933,1805]" captionTargetPageId="10" captionText="Fig. 5. Batillipes lusitanus sp. nov. A–B, D. Holotype, ♀. A–B. Detail of the head. A. Dorsal view, showing internal cirrus (black arrowhead), lateral cirrus (asterisk), secondary clava (white arrowhead). B. Ventral view, showing primary clava (white arrowhead), external cirrus (black arrowhead). C. Detail of body lateral projections of a paratype (slide C.IX-46) (black arrowheads); D. Detail of leg sensory organ on leg IV. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3806145" httpUri="https://zenodo.org/record/3806145/files/figure.png" pageId="13" pageNumber="14">Figs 5</figureCitation>
A–B). An indentation present in the frontal edge of the head between the external cephalic cirrus and the pedestal with the primary clava and lateral cirrus. Papillar secondary clavae (
<figureCitation id="6CDC23BBA265FF889EF36D29FE2F573D" box="[415,510,801,828]" captionStart="Fig" captionStartId="10.[189,232,1846,1872]" captionTargetBox="[191,1397,933,1805]" captionTargetId="figure@10.[191,1397,933,1805]" captionTargetPageId="10" captionText="Fig. 5. Batillipes lusitanus sp. nov. A–B, D. Holotype, ♀. A–B. Detail of the head. A. Dorsal view, showing internal cirrus (black arrowhead), lateral cirrus (asterisk), secondary clava (white arrowhead). B. Ventral view, showing primary clava (white arrowhead), external cirrus (black arrowhead). C. Detail of body lateral projections of a paratype (slide C.IX-46) (black arrowheads); D. Detail of leg sensory organ on leg IV. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3806145" httpUri="https://zenodo.org/record/3806145/files/figure.png" pageId="13" pageNumber="14">Fig. 5A</figureCitation>
) well visible (major diameter ca 
<quantity id="331F92DBA265FF889CF06D2AFC25573D" box="[924,1012,801,828]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="13" pageNumber="14" unit="mm" value="7.5">7.5 µm</quantity>
long). Cephalic cirri, including the external cirri and the lateral cirri with swollen tips. Ventral mouth opening in a protruded circular structure. Ovoid pharyngeal bulb, 
<quantity id="331F92DBA266FF8B9D236F24FD635548" box="[591,690,303,329]" metricMagnitude="-2" metricUnit="m" metricValue="2.81" pageId="14" pageNumber="15" unit="mm" value="28.1">28.1 µm</quantity>
long and 
<quantity id="331F92DBA266FF8B9C4A6F25FC585548" box="[806,905,302,329]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" pageId="14" pageNumber="15" unit="mm" value="25.5">25.5 µm</quantity>
wide, with placoids (ca 
<quantity id="331F92DBA266FF8B9BCB6F24FADB5548" box="[1191,1290,303,329]" metricMagnitude="-2" metricUnit="m" metricValue="1.72" pageId="14" pageNumber="15" unit="mm" value="17.2">17.2 µm</quantity>
long).
</paragraph>
<caption id="A0986FB6A265FF889FD1693CFD1B53DE" ID-DOI="http://doi.org/10.5281/zenodo.3806149" ID-Zenodo-Dep="3806149" httpUri="https://zenodo.org/record/3806149/files/figure.png" pageId="13" pageNumber="14" startId="13.[189,232,1847,1873]" targetBox="[190,1398,902,1805]" targetPageId="13">
<paragraph id="F4583F3EA265FF889FD1693CFD1B53DE" blockId="13.[189,1398,1847,2015]" pageId="13" pageNumber="14">
<emphasis id="C693E32CA265FF889FD1693CFED85350" bold="true" box="[189,265,1847,1873]" pageId="13" pageNumber="14">Fig. 6.</emphasis>
<taxonomicName id="33E744BDA265FF889E7C693CFE215350" authority="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura, 2018" authorityName="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura" authorityYear="2018" box="[272,496,1847,1873]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="lusitanus" status="sp. nov.">
<emphasis id="C693E32CA265FF889E7C693CFE215350" box="[272,496,1847,1873]" italics="true" pageId="13" pageNumber="14">Batillipes lusitanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDA05E57A265FF889E9B6933FD815353" box="[503,592,1848,1874]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="C693E32CA265FF889D3A693CFDBA5350" bold="true" box="[598,619,1847,1873]" pageId="13" pageNumber="14">A</emphasis>
. Fourth leg of a paratype (slide C.IX-46) with details of toes (1–6), note the shape of the ventro-lateral projection between legs III–IV (black arrowhead). 
<emphasis id="C693E32CA265FF889BD56950FB1C5374" bold="true" box="[1209,1229,1883,1909]" pageId="13" pageNumber="14">B</emphasis>
. Fourth leg of a paratype (slide C.IX-1), showing the rigid process on the distal extreme (black arrowhead). 
<emphasis id="C693E32CA265FF889B916976FAC35396" bold="true" box="[1277,1298,1917,1943]" pageId="13" pageNumber="14">C</emphasis>
. Female gonopore (white arrowhead) and anus (black arrowhead). 
<emphasis id="C693E32CA265FF889C1269AAFC4253BA" bold="true" box="[894,915,1953,1979]" pageId="13" pageNumber="14">D</emphasis>
. Male gonopore (white arrowhead) and anus (black arrowhead). Scale bars = 10 µm.
</paragraph>
</caption>
<paragraph id="F4583F3EA266FF8B9FD16F7BFA9855AC" blockId="14.[189,1398,368,430]" pageId="14" pageNumber="15">
Well-developed blunt ventro-lateral processes between legs III and IV (body projection 4) (
<figureCitation id="6CDC23BBA266FF8B9A636F7BFAA3558A" box="[1295,1394,368,395]" captionStart="Fig" captionStartId="10.[189,232,1846,1872]" captionTargetBox="[191,1397,933,1805]" captionTargetId="figure@10.[191,1397,933,1805]" captionTargetPageId="10" captionText="Fig. 5. Batillipes lusitanus sp. nov. A–B, D. Holotype, ♀. A–B. Detail of the head. A. Dorsal view, showing internal cirrus (black arrowhead), lateral cirrus (asterisk), secondary clava (white arrowhead). B. Ventral view, showing primary clava (white arrowhead), external cirrus (black arrowhead). C. Detail of body lateral projections of a paratype (slide C.IX-46) (black arrowheads); D. Detail of leg sensory organ on leg IV. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3806145" httpUri="https://zenodo.org/record/3806145/files/figure.png" pageId="14" pageNumber="15">Figs 5C</figureCitation>
, 
<figureCitation id="6CDC23BBA266FF8B9FD16F98FF3455AF" box="[189,229,403,430]" captionStart="Fig" captionStartId="13.[189,232,1847,1873]" captionTargetBox="[190,1398,902,1805]" captionTargetId="figure@13.[189,1399,902,1805]" captionTargetPageId="13" captionText="Fig. 6. Batillipes lusitanus sp. nov. A. Fourth leg of a paratype (slide C.IX-46) with details of toes (1–6), note the shape of the ventro-lateral projection between legs III–IV (black arrowhead). B. Fourth leg of a paratype (slide C.IX-1), showing the rigid process on the distal extreme (black arrowhead). C. Female gonopore (white arrowhead) and anus (black arrowhead). D. Male gonopore (white arrowhead) and anus (black arrowhead). Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3806149" httpUri="https://zenodo.org/record/3806149/files/figure.png" pageId="14" pageNumber="15">6A</figureCitation>
). The caudal apparatus consists of a prominent semicircular projection (
<figureCitation id="6CDC23BBA266FF8B9B5C6F98FBAA55AC" box="[1072,1147,403,429]" captionStart="Fig" captionStartId="9.[189,232,1960,1986]" captionTargetBox="[193,1397,1034,1917]" captionTargetId="figure@9.[187,1397,1034,1917]" captionTargetPageId="9" captionText="Fig. 4. Batillipes lusitanus sp. nov. A. Schematic drawing, dorsal view. B. Habitus of the holotype, ♀, ventral view. Scale bars = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3806143" httpUri="https://zenodo.org/record/3806143/files/figure.png" pageId="14" pageNumber="15">Figs 4</figureCitation>
A–B, 6C, 7A–B).
</paragraph>
<paragraph id="F4583F3EA266FF8B9FD16FDEFECB567D" blockId="14.[189,1399,469,636]" pageId="14" pageNumber="15">
Sensory spines present on all legs, increasing in length from leg I to IV. First and fourth leg sensory organs with swollen tips. The sensory organ on leg IV divided into a cirrophore (not measurable in the 
<typeStatus id="2B5C819CA266FF8B9FD16C10FEF55634" box="[189,292,539,565]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
but observed in 
<typeStatus id="2B5C819CA266FF8B9E816C17FD8E5637" box="[493,607,540,566]" pageId="14" pageNumber="15" type="paratype">paratypes</typeStatus>
) and a basal portion (
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long) separated from a distal portion (
<quantity id="331F92DBA266FF8B9FAA6C35FEFD5658" box="[198,300,574,601]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="14" pageNumber="15" unit="mm" value="14.5">14.5 µm</quantity>
long) by a van der Land’s organ (
<figureCitation id="6CDC23BBA266FF8B9DBD6C35FCE35658" box="[721,818,574,601]" captionStart="Fig" captionStartId="10.[189,232,1846,1872]" captionTargetBox="[191,1397,933,1805]" captionTargetId="figure@10.[191,1397,933,1805]" captionTargetPageId="10" captionText="Fig. 5. Batillipes lusitanus sp. nov. A–B, D. Holotype, ♀. A–B. Detail of the head. A. Dorsal view, showing internal cirrus (black arrowhead), lateral cirrus (asterisk), secondary clava (white arrowhead). B. Ventral view, showing primary clava (white arrowhead), external cirrus (black arrowhead). C. Detail of body lateral projections of a paratype (slide C.IX-46) (black arrowheads); D. Detail of leg sensory organ on leg IV. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3806145" httpUri="https://zenodo.org/record/3806145/files/figure.png" pageId="14" pageNumber="15">Fig. 5D</figureCitation>
). Sharply pointed cirri 
<emphasis id="C693E32CA266FF8B9B216C34FB8E5659" box="[1101,1119,575,600]" italics="true" pageId="14" pageNumber="15">E</emphasis>
with small cirrophores present.
</paragraph>
<paragraph id="F4583F3EA266FF8B9FD16CA8FE5D576F" blockId="14.[189,1399,675,878]" pageId="14" pageNumber="15">
Telescopic legs with toes with the distal stalk enlarged distally (ca 
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wide), ovoid suction discs (5.2 × 
<quantity id="331F92DBA266FF8B9E636CCDFEB656E0" box="[271,359,710,737]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="14" pageNumber="15" unit="mm" value="5.1">5.1 µm</quantity>
) and conspicuous braces (
<figureCitation id="6CDC23BBA266FF8B9DC96CCDFCD756E0" box="[677,774,710,737]" captionStart="Fig" captionStartId="13.[189,232,1847,1873]" captionTargetBox="[190,1398,902,1805]" captionTargetId="figure@13.[189,1399,902,1805]" captionTargetPageId="13" captionText="Fig. 6. Batillipes lusitanus sp. nov. A. Fourth leg of a paratype (slide C.IX-46) with details of toes (1–6), note the shape of the ventro-lateral projection between legs III–IV (black arrowhead). B. Fourth leg of a paratype (slide C.IX-1), showing the rigid process on the distal extreme (black arrowhead). C. Female gonopore (white arrowhead) and anus (black arrowhead). D. Male gonopore (white arrowhead) and anus (black arrowhead). Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3806149" httpUri="https://zenodo.org/record/3806149/files/figure.png" pageId="14" pageNumber="15">Fig. 6A</figureCitation>
). On feet of the first three pairs of legs, toe 2 is the shortest, toes 3 and 5 are the longest and toes 1, 4 and 6 are medium sized. On feet of the fourth pair of legs, (
<figureCitation id="6CDC23BBA266FF8B9E686D07FE985726" box="[260,329,780,807]" captionStart="Fig" captionStartId="13.[189,232,1847,1873]" captionTargetBox="[190,1398,902,1805]" captionTargetId="figure@13.[189,1399,902,1805]" captionTargetPageId="13" captionText="Fig. 6. Batillipes lusitanus sp. nov. A. Fourth leg of a paratype (slide C.IX-46) with details of toes (1–6), note the shape of the ventro-lateral projection between legs III–IV (black arrowhead). B. Fourth leg of a paratype (slide C.IX-1), showing the rigid process on the distal extreme (black arrowhead). C. Female gonopore (white arrowhead) and anus (black arrowhead). D. Male gonopore (white arrowhead) and anus (black arrowhead). Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3806149" httpUri="https://zenodo.org/record/3806149/files/figure.png" pageId="14" pageNumber="15">Fig. 6</figureCitation>
A–B) the medial toes 3 and 4 are equal in length. Toes 1 and 6 are medium sized; toe 2 and especially toe 5 are the longest. A small rigid process (ca 
<quantity id="331F92DBA266FF8B9C0F6D3BFC70574B" box="[867,929,816,842]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="15" unit="mm" value="3.0">3 µm</quantity>
long) is present in the distal extreme of legs IV (
<figureCitation id="6CDC23BBA266FF8B9E4E6D58FEAF576F" box="[290,382,851,878]" captionStart="Fig" captionStartId="13.[189,232,1847,1873]" captionTargetBox="[190,1398,902,1805]" captionTargetId="figure@13.[189,1399,902,1805]" captionTargetPageId="13" captionText="Fig. 6. Batillipes lusitanus sp. nov. A. Fourth leg of a paratype (slide C.IX-46) with details of toes (1–6), note the shape of the ventro-lateral projection between legs III–IV (black arrowhead). B. Fourth leg of a paratype (slide C.IX-1), showing the rigid process on the distal extreme (black arrowhead). C. Female gonopore (white arrowhead) and anus (black arrowhead). D. Male gonopore (white arrowhead) and anus (black arrowhead). Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3806149" httpUri="https://zenodo.org/record/3806149/files/figure.png" pageId="14" pageNumber="15">Fig. 6B</figureCitation>
).
</paragraph>
<paragraph id="F4583F3EA266FF8B9FD16D9FFAA75019" blockId="14.[189,1399,916,1048]" pageId="14" pageNumber="15">
The punctation of the dorsal cuticle is very peculiar. It is constituted by large pillars (about 5 pillars / 
<quantity id="331F92DBA266FF8B9E7A6DB3FEB857D3" box="[278,361,952,978]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="15" unit="mm" value="10.0">10 µm</quantity>
, each pillar with ca 
<quantity id="331F92DBA266FF8B9D056DBCFD1057D3" box="[617,705,951,978]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="14" pageNumber="15" unit="mm" value="3.6">3.6 µm</quantity>
high) that, when observed under PCM and DIC, appear as large tubercles, especially if observed laterally (
<figureCitation id="6CDC23BBA266FF8B9C766DD0FCB157F4" box="[794,864,987,1013]" captionStart="Fig" captionStartId="14.[189,232,1886,1912]" captionTargetBox="[193,1395,1122,1838]" captionTargetId="figure@14.[192,1395,1122,1838]" captionTargetPageId="14" captionText="Fig. 7. Batillipes lusitanus sp. nov. A. Posterior portion of the body, showing the dorsal sculpture. B. Caudal apparatus, showing details of cuticular pillars. C. SEM photo showing the dorsal (white arrowhead) and ventral (black arrowhead) aspect of the cuticle. Pillars are also visible (black arrow). Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3806153" httpUri="https://zenodo.org/record/3806153/files/figure.png" pageId="14" pageNumber="15">Fig. 7</figureCitation>
A–B). When observed under SEM, tubercles showed to be depressions that correspond to the wide top of pillars, surrounded by a epicuticular fold
</paragraph>
<caption id="A0986FB6A266FF8B9FD16955FE7953E2" ID-DOI="http://doi.org/10.5281/zenodo.3806153" ID-Zenodo-Dep="3806153" httpUri="https://zenodo.org/record/3806153/files/figure.png" pageId="14" pageNumber="15" startId="14.[189,232,1886,1912]" targetBox="[193,1395,1122,1838]" targetPageId="14">
<paragraph id="F4583F3EA266FF8B9FD16955FE7953E2" blockId="14.[189,1399,1886,2019]" pageId="14" pageNumber="15">
<emphasis id="C693E32CA266FF8B9FD16955FEDE5378" bold="true" box="[189,271,1886,1913]" pageId="14" pageNumber="15">Fig. 7.</emphasis>
<taxonomicName id="33E744BDA266FF8B9E766955FDD15379" authority="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura, 2018" authorityName="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura" authorityYear="2018" box="[282,512,1886,1912]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="14" pageNumber="15" phylum="Tardigrada" rank="species" species="lusitanus" status="sp. nov.">
<emphasis id="C693E32CA266FF8B9E766955FDD15379" box="[282,512,1886,1912]" italics="true" pageId="14" pageNumber="15">Batillipes lusitanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDA05E57A266FF8B9D606954FDBB5378" box="[524,618,1887,1913]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="C693E32CA266FF8B9D196955FD5B5379" bold="true" box="[629,650,1886,1912]" pageId="14" pageNumber="15">A</emphasis>
. Posterior portion of the body, showing the dorsal sculpture. 
<emphasis id="C693E32CA266FF8B9FD16989FF00539D" bold="true" box="[189,209,1922,1948]" pageId="14" pageNumber="15">B</emphasis>
. Caudal apparatus, showing details of cuticular pillars. 
<emphasis id="C693E32CA266FF8B9C11698AFC43539A" bold="true" box="[893,914,1921,1947]" pageId="14" pageNumber="15">C</emphasis>
. SEM photo showing the dorsal (white arrowhead) and ventral (black arrowhead) aspect of the cuticle. Pillars are also visible (black arrow). Scale bars = 10 µm.
</paragraph>
</caption>
<paragraph id="F4583F3EA267FF8A9FD16F00FC3D5548" blockId="15.[189,1399,267,329]" pageId="15" pageNumber="16">
(
<figureCitation id="6CDC23BBA267FF8A9FAA6F00FEF05524" box="[198,289,267,293]" captionStart="Fig" captionStartId="14.[189,232,1886,1912]" captionTargetBox="[193,1395,1122,1838]" captionTargetId="figure@14.[192,1395,1122,1838]" captionTargetPageId="14" captionText="Fig. 7. Batillipes lusitanus sp. nov. A. Posterior portion of the body, showing the dorsal sculpture. B. Caudal apparatus, showing details of cuticular pillars. C. SEM photo showing the dorsal (white arrowhead) and ventral (black arrowhead) aspect of the cuticle. Pillars are also visible (black arrow). Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3806153" httpUri="https://zenodo.org/record/3806153/files/figure.png" pageId="15" pageNumber="16">Fig. 7C</figureCitation>
white arrowhead). Faint transverse cuticular folds are visible. Ventrally, the cuticle is finely punctuated (15–16 pillars / 
<quantity id="331F92DBA267FF8A9E946F24FD975548" box="[504,582,303,329]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="16" unit="mm" value="10.0">10 µm</quantity>
) (
<figureCitation id="6CDC23BBA267FF8A9D326F25FD765549" box="[606,679,302,328]" captionStart="Fig" captionStartId="13.[189,232,1847,1873]" captionTargetBox="[190,1398,902,1805]" captionTargetId="figure@13.[189,1399,902,1805]" captionTargetPageId="13" captionText="Fig. 6. Batillipes lusitanus sp. nov. A. Fourth leg of a paratype (slide C.IX-46) with details of toes (1–6), note the shape of the ventro-lateral projection between legs III–IV (black arrowhead). B. Fourth leg of a paratype (slide C.IX-1), showing the rigid process on the distal extreme (black arrowhead). C. Female gonopore (white arrowhead) and anus (black arrowhead). D. Male gonopore (white arrowhead) and anus (black arrowhead). Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3806149" httpUri="https://zenodo.org/record/3806149/files/figure.png" pageId="15" pageNumber="16">Figs 6</figureCitation>
C–D, 7C black arrowhead).
</paragraph>
<paragraph id="F4583F3EA267FF8A9FD16F7BFB7A558A" blockId="15.[189,1195,368,395]" box="[189,1195,368,395]" pageId="15" pageNumber="16">
Rosette-shaped gonopore separated from the anus by a well-defined groove (
<figureCitation id="6CDC23BBA267FF8A9B2D6F7BFB4F558A" box="[1089,1182,368,395]" captionStart="Fig" captionStartId="13.[189,232,1847,1873]" captionTargetBox="[190,1398,902,1805]" captionTargetId="figure@13.[189,1399,902,1805]" captionTargetPageId="13" captionText="Fig. 6. Batillipes lusitanus sp. nov. A. Fourth leg of a paratype (slide C.IX-46) with details of toes (1–6), note the shape of the ventro-lateral projection between legs III–IV (black arrowhead). B. Fourth leg of a paratype (slide C.IX-1), showing the rigid process on the distal extreme (black arrowhead). C. Female gonopore (white arrowhead) and anus (black arrowhead). D. Male gonopore (white arrowhead) and anus (black arrowhead). Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3806149" httpUri="https://zenodo.org/record/3806149/files/figure.png" pageId="15" pageNumber="16">Fig. 6C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="BCFD6CB5A267FF8A9FD16FBAFAA750B7" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="F4583F3EA267FF8A9FD16FBAFE1855CA" blockId="15.[189,457,433,459]" box="[189,457,433,459]" pageId="15" pageNumber="16">
<heading id="AF108852A267FF8A9FD16FBAFE1855CA" bold="true" box="[189,457,433,459]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="C693E32CA267FF8A9FD16FBAFE1855CA" bold="true" box="[189,457,433,459]" pageId="15" pageNumber="16">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="F4583F3EA267FF8A9FD16FEBFC52575A" blockId="15.[189,1399,480,859]" pageId="15" pageNumber="16">
Excluding the new species, there are 26 species of 
<taxonomicName id="33E744BDA267FF8A9C7A6FEBFC5755FB" authorityName="Richters" authorityYear="1909" box="[790,902,480,506]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="15" pageNumber="16" phylum="Tardigrada" rank="genus">
<emphasis id="C693E32CA267FF8A9C7A6FEBFC5755FB" box="[790,902,480,506]" italics="true" pageId="15" pageNumber="16">Batillipes</emphasis>
</taxonomicName>
with the same toe arrangement pattern on feet of fourth legs as 
<taxonomicName id="33E744BDA267FF8A9EDD6C0FFDEA561C" authority="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura, 2018" authorityName="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura" authorityYear="2018" box="[433,571,515,541]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="15" pageNumber="16" phylum="Tardigrada" rank="species" species="lusitanus" status="sp. nov.">
<emphasis id="C693E32CA267FF8A9EDD6C0FFDEA561C" box="[433,571,515,541]" italics="true" pageId="15" pageNumber="16">B. lusitanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDA05E57A267FF8A9D2D6C0FFD4F561F" box="[577,670,516,542]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
, which has the middle toes (3 and 4) on the fourth feet equal in length. However, only two of those species, 
<taxonomicName id="33E744BDA267FF8A9DAC6C2CFBE95641" authority="Kristensen, 1978" authorityName="Kristensen" authorityYear="1978" box="[704,1080,550,577]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="15" pageNumber="16" phylum="Tardigrada" rank="species" species="roscoffensis">
<emphasis id="C693E32CA267FF8A9DAC6C2CFCBA5640" box="[704,875,551,577]" italics="true" pageId="15" pageNumber="16">B. roscoffensis</emphasis>
<bibRefCitation id="907642CFA267FF8A9C1E6C2DFBE95641" author="Kristensen R. M." box="[882,1080,550,576]" pageId="15" pageNumber="16" pagination="1 - 17" refId="ref17216" refString="Kristensen R. M. 1978. Notes on marine heterotardigrades. 1. Description of two Batillipes species, using the electron microscope. Zoologischer Anzeiger 200: 1 - 17." type="journal article" year="1978">Kristensen, 1978</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="33E744BDA267FF8A9B1C6C2CFAD65641" authorityName="Grimaldi de Zio, Morone De Lucia, D'Addabbo Gallo &amp; Grimaldi" authorityYear="1979" box="[1136,1287,550,576]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="15" pageNumber="16" phylum="Tardigrada" rank="species" species="adriaticus">
<emphasis id="C693E32CA267FF8A9B1C6C2CFAD65641" box="[1136,1287,550,576]" italics="true" pageId="15" pageNumber="16">B. adriaticus</emphasis>
</taxonomicName>
, have the dorsal cuticular ornamentation, constituted by large pillars (ca 3.0 µm high). 
<taxonomicName id="33E744BDA267FF8A9B4E6C41FAF35665" authorityName="Kristensen" authorityYear="1978" box="[1058,1314,586,612]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="15" pageNumber="16" phylum="Tardigrada" rank="species" species="roscoffensis">
<emphasis id="C693E32CA267FF8A9B4E6C41FAF35665" box="[1058,1314,586,612]" italics="true" pageId="15" pageNumber="16">Batillipes roscoffensis</emphasis>
</taxonomicName>
clearly differs from the new species in having scattered and randomly distributed dots (“small pox” according the terminology of 
<bibRefCitation id="907642CFA267FF8A9EC66C9BFDA356AB" author="Kristensen R. M." box="[426,626,656,682]" pageId="15" pageNumber="16" pagination="1 - 17" refId="ref17216" refString="Kristensen R. M. 1978. Notes on marine heterotardigrades. 1. Description of two Batillipes species, using the electron microscope. Zoologischer Anzeiger 200: 1 - 17." type="journal article" year="1978">Kristensen 1978</bibRefCitation>
). Those dots lack in all the other known 
<taxonomicName id="33E744BDA267FF8A9B1D6C9BFB3056AB" authorityName="Richters" authorityYear="1909" box="[1137,1249,656,682]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="15" pageNumber="16" phylum="Tardigrada" rank="genus">
<emphasis id="C693E32CA267FF8A9B1D6C9BFB3056AB" box="[1137,1249,656,682]" italics="true" pageId="15" pageNumber="16">Batillipes</emphasis>
</taxonomicName>
species and they were described by 
<bibRefCitation id="907642CFA267FF8A9EBC6CB8FD7256CC" author="Kristensen R. M." box="[464,675,691,717]" pageId="15" pageNumber="16" pagination="1 - 17" refId="ref17216" refString="Kristensen R. M. 1978. Notes on marine heterotardigrades. 1. Description of two Batillipes species, using the electron microscope. Zoologischer Anzeiger 200: 1 - 17." type="journal article" year="1978">Kristensen (1978)</bibRefCitation>
under TEM (transmission electron microscopy), as elevations (of spongy electron dense mass, see 
<bibRefCitation id="907642CFA267FF8A9D126CDDFC9456F1" author="Kristensen R. M." box="[638,837,726,752]" pageId="15" pageNumber="16" pagination="1 - 17" refId="ref17216" refString="Kristensen R. M. 1978. Notes on marine heterotardigrades. 1. Description of two Batillipes species, using the electron microscope. Zoologischer Anzeiger 200: 1 - 17." type="journal article" year="1978">Kristensen 1978</bibRefCitation>
: figs 13, 17). Moreover, in 
<taxonomicName id="33E744BDA267FF8A9BF56CDCFA9956F0" authorityName="Kristensen" authorityYear="1978" box="[1177,1352,727,753]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="15" pageNumber="16" phylum="Tardigrada" rank="species" species="roscoffensis">
<emphasis id="C693E32CA267FF8A9BF56CDCFA9956F0" box="[1177,1352,727,753]" italics="true" pageId="15" pageNumber="16">B. roscoffensis</emphasis>
</taxonomicName>
the caudal apparatus totally lacks and the lateral cirri 
<emphasis id="C693E32CA267FF8A9C656CF1FCCA5715" box="[777,795,762,788]" italics="true" pageId="15" pageNumber="16">A</emphasis>
is much longer than in 
<taxonomicName id="33E744BDA267FF8A9B586CF1FB6E5715" authority="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura, 2018" authorityName="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura" authorityYear="2018" box="[1076,1215,762,788]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="15" pageNumber="16" phylum="Tardigrada" rank="species" species="lusitanus" status="sp. nov.">
<emphasis id="C693E32CA267FF8A9B586CF1FB6E5715" box="[1076,1215,762,788]" italics="true" pageId="15" pageNumber="16">B. lusitanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDA05E57A267FF8A9BAB6CF1FAF35715" box="[1223,1314,762,788]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
(cirrus 
<emphasis id="C693E32CA267FF8A9FD16D16FF1E5736" box="[189,207,797,823]" italics="true" pageId="15" pageNumber="16">A</emphasis>
ca 
<quantity id="331F92DBA267FF8A9F966D16FE875736" box="[250,342,797,823]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="16" unit="mm" value="100.0">100 µm</quantity>
long, corresponding to 40% of body length in 
<taxonomicName id="33E744BDA267FF8A9CEA6D15FBE25736" authorityName="Kristensen" authorityYear="1978" box="[902,1075,797,823]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="15" pageNumber="16" phylum="Tardigrada" rank="species" species="roscoffensis">
<emphasis id="C693E32CA267FF8A9CEA6D15FBE25736" box="[902,1075,797,823]" italics="true" pageId="15" pageNumber="16">B. roscoffensis</emphasis>
</taxonomicName>
, and less than 
<quantity id="331F92DBA267FF8A9B896D16FAE25736" box="[1253,1331,797,824]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="16" unit="mm" value="50.0">50 µm</quantity>
long, corresponding to ca 20% of body length in the new species).
</paragraph>
<paragraph id="F4583F3EA267FF8A9FD16D8AFAA750B7" blockId="15.[189,1399,897,1206]" pageId="15" pageNumber="16">
<taxonomicName id="33E744BDA267FF8A9FD16D8AFE72579A" authority="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura, 2018" authorityName="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura" authorityYear="2018" box="[189,419,897,923]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="15" pageNumber="16" phylum="Tardigrada" rank="species" species="lusitanus" status="sp. nov.">
<emphasis id="C693E32CA267FF8A9FD16D8AFE72579A" box="[189,419,897,923]" italics="true" pageId="15" pageNumber="16">Batillipes lusitanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDA05E57A267FF8A9EC36D89FDDF579D" box="[431,526,898,924]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
shares with 
<taxonomicName id="33E744BDA267FF8A9DC16D89FC98579A" authorityName="Grimaldi de Zio, Morone De Lucia, D'Addabbo Gallo &amp; Grimaldi" authorityYear="1979" box="[685,841,897,923]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="15" pageNumber="16" phylum="Tardigrada" rank="species" species="adriaticus">
<emphasis id="C693E32CA267FF8A9DC16D89FC98579A" box="[685,841,897,923]" italics="true" pageId="15" pageNumber="16">B. adriaticus</emphasis>
</taxonomicName>
, the most similar species, the peculiar dorsal cuticle punctation constituted by large pillars, appearing as large tubercles. These two species can be clearly distinguished by the shape of the caudal apparatus that consists in a roundish cuticular expansion in 
<taxonomicName id="33E744BDA267FF8A9FB36DE7FEBF5004" authority="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura, 2018" authorityName="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura" authorityYear="2018" box="[223,366,1003,1029]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="15" pageNumber="16" phylum="Tardigrada" rank="species" species="lusitanus" status="sp. nov.">
<emphasis id="C693E32CA267FF8A9FB36DE7FEBF5004" box="[223,366,1003,1029]" italics="true" pageId="15" pageNumber="16">B. lusitanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDA05E57A267FF8A9E156DE7FE095007" box="[377,472,1004,1030]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
and in a sharp and long spine in 
<taxonomicName id="33E744BDA267FF8A9C126DE7FBCB5004" authorityName="Grimaldi de Zio, Morone De Lucia, D'Addabbo Gallo &amp; Grimaldi" authorityYear="1979" box="[894,1050,1003,1029]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="15" pageNumber="16" phylum="Tardigrada" rank="species" species="adriaticus">
<emphasis id="C693E32CA267FF8A9C126DE7FBCB5004" box="[894,1050,1003,1029]" italics="true" pageId="15" pageNumber="16">B. adriaticus</emphasis>
</taxonomicName>
. The lateral body projection between legs III–IV is also blunt or slightly pointed in the new species while it is sharply pointed in 
<taxonomicName id="33E744BDA267FF8A9FD16A39FE87504A" authorityName="Grimaldi de Zio, Morone De Lucia, D'Addabbo Gallo &amp; Grimaldi" authorityYear="1979" box="[189,342,1073,1099]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="15" pageNumber="16" phylum="Tardigrada" rank="species" species="adriaticus">
<emphasis id="C693E32CA267FF8A9FD16A39FE87504A" box="[189,342,1073,1099]" italics="true" pageId="15" pageNumber="16">B. adriaticus</emphasis>
</taxonomicName>
. Moreover, contrary to 
<taxonomicName id="33E744BDA267FF8A9D1F6A39FCDD504A" authorityName="Grimaldi de Zio, Morone De Lucia, D'Addabbo Gallo &amp; Grimaldi" authorityYear="1979" box="[627,780,1073,1099]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="15" pageNumber="16" phylum="Tardigrada" rank="species" species="adriaticus">
<emphasis id="C693E32CA267FF8A9D1F6A39FCDD504A" box="[627,780,1073,1099]" italics="true" pageId="15" pageNumber="16">B. adriaticus</emphasis>
</taxonomicName>
, in the new species, a small rigid process is present in the distal extreme of legs IV and the surface of the primary clava is wrinkled. In addition, although with overlapping values, in adults of 
<taxonomicName id="33E744BDA267FF8A9D076A72FD245093" authority="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura, 2018" authorityName="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura" authorityYear="2018" box="[619,757,1144,1170]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="15" pageNumber="16" phylum="Tardigrada" rank="species" species="lusitanus" status="sp. nov.">
<emphasis id="C693E32CA267FF8A9D076A72FD245093" box="[619,757,1144,1170]" italics="true" pageId="15" pageNumber="16">B. lusitanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDA05E57A267FF8A9D976A73FC845093" box="[763,853,1144,1170]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
the sensory organ on legs IV is slightly shorter than in 
<taxonomicName id="33E744BDA267FF8A9E786A97FE7A50B4" authorityName="Grimaldi de Zio, Morone De Lucia, D'Addabbo Gallo &amp; Grimaldi" authorityYear="1979" box="[276,427,1179,1205]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="15" pageNumber="16" phylum="Tardigrada" rank="species" species="adriaticus">
<emphasis id="C693E32CA267FF8A9E786A97FE7A50B4" box="[276,427,1179,1205]" italics="true" pageId="15" pageNumber="16">B. adriaticus</emphasis>
</taxonomicName>
(range: 
<quantity id="331F92DBA267FF8A9D606A90FDA450B7" box="[524,629,1179,1206]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.7" metricValueMin="0.9" pageId="15" pageNumber="16" unit="mm" value="18.0" valueMax="27.0" valueMin="9.0">9–27 µm</quantity>
long in 
<taxonomicName id="33E744BDA267FF8A9DB86A97FC8E50B4" authority="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura, 2018" authorityName="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura" authorityYear="2018" box="[724,863,1179,1205]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="15" pageNumber="16" phylum="Tardigrada" rank="species" species="lusitanus" status="sp. nov.">
<emphasis id="C693E32CA267FF8A9DB86A97FC8E50B4" box="[724,863,1179,1205]" italics="true" pageId="15" pageNumber="16">B. lusitanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDA05E57A267FF8A9C096A97FC6E50B7" box="[869,959,1180,1206]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
and 
<quantity id="331F92DBA267FF8A9C9B6A90FBBE50B7" box="[1015,1135,1179,1206]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="15" pageNumber="16" unit="mm" value="30.0" valueMax="40.0" valueMin="20.0">20–40 µm</quantity>
long in 
<taxonomicName id="33E744BDA267FF8A9BA26A97FAB450B4" authorityName="Grimaldi de Zio, Morone De Lucia, D'Addabbo Gallo &amp; Grimaldi" authorityYear="1979" box="[1230,1381,1179,1205]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="15" pageNumber="16" phylum="Tardigrada" rank="species" species="adriaticus">
<emphasis id="C693E32CA267FF8A9BA26A97FAB450B4" box="[1230,1381,1179,1205]" italics="true" pageId="15" pageNumber="16">B. adriaticus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="BCFD6CB5A267FF8A9FD16AD7FBCE5127" pageId="15" pageNumber="16" type="description">
<paragraph id="F4583F3EA267FF8A9FD16AD7FE7150F7" blockId="15.[189,416,1244,1270]" box="[189,416,1244,1270]" pageId="15" pageNumber="16">
<heading id="AF108852A267FF8A9FD16AD7FE7150F7" bold="true" box="[189,416,1244,1270]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="C693E32CA267FF8A9FD16AD7FE7150F7" bold="true" box="[189,416,1244,1270]" pageId="15" pageNumber="16">Associated species</emphasis>
</heading>
</paragraph>
<paragraph id="F4583F3EA267FF8A9FD16B00FBCE5127" blockId="15.[189,1055,1291,1318]" box="[189,1055,1291,1318]" pageId="15" pageNumber="16">
<taxonomicName id="33E744BDA267FF8A9FD16B00FE1C5124" authority="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura, 2018" authorityName="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura" authorityYear="2018" box="[189,461,1291,1317]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="15" pageNumber="16" phylum="Tardigrada" rank="species" species="algharbensis" status="sp. nov.">
<emphasis id="C693E32CA267FF8A9FD16B00FE1C5124" box="[189,461,1291,1317]" italics="true" pageId="15" pageNumber="16">Batillipes algharbensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDA05E57A267FF8A9EB86B07FDE05127" box="[468,561,1292,1318]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="33E744BDA267FF8A9D506B07FD6B5124" authorityName="Marcus" authorityYear="1946" box="[572,698,1291,1317]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="15" pageNumber="16" phylum="Tardigrada" rank="species" species="pennaki">
<emphasis id="C693E32CA267FF8A9D506B07FD6B5124" box="[572,698,1291,1317]" italics="true" pageId="15" pageNumber="16">B. pennaki</emphasis>
</taxonomicName>
, 
<taxonomicName id="33E744BDA267FF8A9DA56B07FCB55124" authorityName="Renaud-Debyser" authorityYear="1959" box="[713,868,1291,1317]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="15" pageNumber="16" phylum="Tardigrada" rank="species" species="phreaticus">
<emphasis id="C693E32CA267FF8A9DA56B07FCB55124" box="[713,868,1291,1317]" italics="true" pageId="15" pageNumber="16">B. phreaticus</emphasis>
</taxonomicName>
and 
<taxonomicName id="33E744BDA267FF8A9CF16B07FBC95124" authorityName="Renaud-Mornant &amp; Deroux" authorityYear="1976" box="[925,1048,1292,1317]" class="Heterotardigrada" family="Halechiniscidae" genus="Halechiniscus" kingdom="Animalia" order="Arthrotardigrada" pageId="15" pageNumber="16" phylum="Tardigrada" rank="species" species="greveni">
<emphasis id="C693E32CA267FF8A9CF16B07FBC95124" box="[925,1048,1292,1317]" italics="true" pageId="15" pageNumber="16">H. greveni</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="BCFD6CB5A267FF959FD16B46FC135549" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="discussion">
<paragraph id="F4583F3EA267FF8A9FD16B46FEFE5166" blockId="15.[189,303,1357,1383]" box="[189,303,1357,1383]" pageId="15" pageNumber="16">
<heading id="AF108852A267FF8A9FD16B46FEFE5166" bold="true" box="[189,303,1357,1383]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="C693E32CA267FF8A9FD16B46FEFE5166" bold="true" box="[189,303,1357,1383]" pageId="15" pageNumber="16">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="F4583F3EA267FF8A9FD16B77FCF152F4" blockId="15.[189,1399,1403,1782]" pageId="15" pageNumber="16">
Measurements of structures obtained from specimens of the new species are provided in 
<tableCitation id="B9650A85A267FF8A9BAF6B70FACD5194" box="[1219,1308,1403,1429]" captionStart="Table 2" captionStartId="11.[188,257,564,592]" captionTargetPageId="11" captionText="Table 2 (continued on next page). Measurements (µm) of selected morphological structures of specimens of Batillipes lusitanus sp. nov. (Holo = Holotype; Allo = Allotype; SD = Standard deviation; ‘Range’ refers to the smallest and largest measured specimen / structure; N = number of specimens/structures measured)." pageId="15" pageNumber="16">Table 2</tableCitation>
. Sexual dimorphism is not evident. Males are similar to females in both qualitative and quantitative characters, except for their circular gonopore with a cuticular crescent shaped fold and located nearer the anus (
<figureCitation id="6CDC23BBA267FF8A9FAA6BEFFEF551FE" box="[198,292,1508,1535]" captionStart="Fig" captionStartId="13.[189,232,1847,1873]" captionTargetBox="[190,1398,902,1805]" captionTargetId="figure@13.[189,1399,902,1805]" captionTargetPageId="13" captionText="Fig. 6. Batillipes lusitanus sp. nov. A. Fourth leg of a paratype (slide C.IX-46) with details of toes (1–6), note the shape of the ventro-lateral projection between legs III–IV (black arrowhead). B. Fourth leg of a paratype (slide C.IX-1), showing the rigid process on the distal extreme (black arrowhead). C. Female gonopore (white arrowhead) and anus (black arrowhead). D. Male gonopore (white arrowhead) and anus (black arrowhead). Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3806149" httpUri="https://zenodo.org/record/3806149/files/figure.png" pageId="15" pageNumber="16">Fig. 6D</figureCitation>
). Juveniles, with six toes on each leg but without a visible gonopore, are also similar to adults (
<tableCitation id="B9650A85A267FF8A9FA96803FECE5223" box="[197,287,1544,1570]" captionStart="Table 2" captionStartId="11.[188,257,564,592]" captionTargetPageId="11" captionText="Table 2 (continued on next page). Measurements (µm) of selected morphological structures of specimens of Batillipes lusitanus sp. nov. (Holo = Holotype; Allo = Allotype; SD = Standard deviation; ‘Range’ refers to the smallest and largest measured specimen / structure; N = number of specimens/structures measured)." pageId="15" pageNumber="16">Table 2</tableCitation>
for morphometrics). In the 
<typeStatus id="2B5C819CA267FF8A9D056803FD025223" box="[617,723,1544,1570]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
, ventro-lateral body projections between legs I–III (body projections 2–3, respectively) are indistinct, but in some 
<typeStatus id="2B5C819CA267FF8A9C196827FC395247" box="[885,1000,1580,1606]" pageId="15" pageNumber="16" type="paratype">paratypes</typeStatus>
, although small, they are visible between all leg pairs (
<figureCitation id="6CDC23BBA267FF8A9ED66845FDC75268" box="[442,534,1614,1641]" captionStart="Fig" captionStartId="10.[189,232,1846,1872]" captionTargetBox="[191,1397,933,1805]" captionTargetId="figure@10.[191,1397,933,1805]" captionTargetPageId="10" captionText="Fig. 5. Batillipes lusitanus sp. nov. A–B, D. Holotype, ♀. A–B. Detail of the head. A. Dorsal view, showing internal cirrus (black arrowhead), lateral cirrus (asterisk), secondary clava (white arrowhead). B. Ventral view, showing primary clava (white arrowhead), external cirrus (black arrowhead). C. Detail of body lateral projections of a paratype (slide C.IX-46) (black arrowheads); D. Detail of leg sensory organ on leg IV. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3806145" httpUri="https://zenodo.org/record/3806145/files/figure.png" pageId="15" pageNumber="16">Fig. 5C</figureCitation>
). This variability could be attributed to the effect of microslide preparation highlighted by the ventral position of lateral projections. Among adults, the size and morphology of the caudal apparatus and lateral body projections, especially those between legs III and IV, show some variation, being perfectly semicircular, as in the 
<typeStatus id="2B5C819CA267FF8A9D8068B3FC8752D3" box="[748,854,1720,1746]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
, or slightly pointed. In four-toed larvae, caudal apparatus and lateral projections are clearly conical.
</paragraph>
<paragraph id="F4583F3EA267FF8A9FD16917FAA653C2" blockId="15.[189,1399,1820,1988]" pageId="15" pageNumber="16">
The variability in shape of lateral projections and caudal apparatus was responsible for the misidentification of three specimens collected in 
<quantity id="331F92DBA267FF8A9C75694BFCA4535B" box="[793,885,1856,1882]" metricMagnitude="1" metricUnit="m" metricValue="5.10794" pageId="15" pageNumber="16" unit="in" value="2011.0">2011 in</quantity>
<collectingCountry id="8CF07FAEA267FF8A9CED694BFC35535B" box="[897,996,1856,1882]" name="Portugal" pageId="15" pageNumber="16">Portugal</collectingCountry>
at Memória Beach, Matosinhos, (
<geoCoordinate id="91D359F9A267FF8A9FAA696AFE8D537C" box="[198,348,1889,1917]" degrees="41" direction="north" minutes="13" orientation="latitude" pageId="15" pageNumber="16" precision="15" seconds="49" value="41.230278">41°13′49″ N</geoCoordinate>
, 
<geoCoordinate id="91D359F9A267FF8A9E04696AFE22537C" box="[360,499,1889,1917]" degrees="8" direction="west" minutes="43" orientation="longitude" pageId="15" pageNumber="16" precision="15" seconds="21" value="-8.7225">8°43′21″ W</geoCoordinate>
) (
<bibRefCitation id="907642CFA267FF8A9D676968FD38537F" author="Rubal M. &amp; Veiga P. &amp; Fontoura P. &amp; Sousa-Pinto I." box="[523,745,1891,1918]" pageId="15" pageNumber="16" pagination="8 - 14" refId="ref17974" refString="Rubal M., Veiga P., Fontoura P. &amp; Sousa-Pinto I. 2013 b. A new intertidal arthrotardigrade, Prostygarctus aculeatus gen. nov., sp. nov. (Tardigrada: Heterotardigrada) from the North of Portugal (Atlantic Ocean). Journal of Limnology 72: 8 - 14. https: // doi. org / 10.4081 / jlimnol. 2013. s 1. e 2" type="journal article" year="2013">
Rubal 
<emphasis id="C693E32CA267FF8A9D36696FFD5C537C" box="[602,653,1891,1917]" italics="true" pageId="15" pageNumber="16">et al</emphasis>
. 2013b
</bibRefCitation>
). These specimens were wrongly attributed to 
<taxonomicName id="33E744BDA267FF8A9A65696FFAA6537C" box="[1289,1399,1891,1917]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="15" pageNumber="16" phylum="Tardigrada" rank="species" species="similis">
<emphasis id="C693E32CA267FF8A9A65696FFAA6537C" box="[1289,1399,1891,1917]" italics="true" pageId="15" pageNumber="16">B. similis</emphasis>
</taxonomicName>
on account of the conical-shaped cuticular projections, evidenced by microslide preparation. A deeper examination showed that the middle toes on legs IV are equal in length and that the cuticle exhibits the
</paragraph>
<paragraph id="F4583F3EA278FF959FD16F07FC135549" blockId="16.[189,1399,267,329]" pageId="16" pageNumber="17">
same peculiar pattern of 
<taxonomicName id="33E744BDA278FF959E8D6F07FDBD5524" authority="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura, 2018" authorityName="Santos &amp; Rubal &amp; Veiga &amp; da Rocha &amp; Fontoura" authorityYear="2018" box="[481,620,267,293]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="16" pageNumber="17" phylum="Tardigrada" rank="species" species="lusitanus" status="sp. nov.">
<emphasis id="C693E32CA278FF959E8D6F07FDBD5524" box="[481,620,267,293]" italics="true" pageId="16" pageNumber="17">B. lusitanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDA05E57A278FF959D186F07FD1E5527" box="[628,719,268,294]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
Therefore, as suggested by 
<bibRefCitation id="907642CFA278FF959B766F00FB225524" author="Rubal M. &amp; Veiga P. &amp; Fontoura P. &amp; Santos E. &amp; Sousa-Pinto I." box="[1050,1267,267,294]" pageId="16" pageNumber="17" pagination="747 - 754" refId="ref18050" refString="Rubal M., Veiga P., Fontoura P., Santos E. &amp; Sousa-Pinto I. 2016. Biodiversity of marine tardigrades from the northern coast of Portugal (Iberian Peninsula). Zoological Journal of the Linnean Society 178: 747 - 754. https: // doi. org / 10.1111 / zoj. 12462" type="journal article" year="2016">
Rubal 
<emphasis id="C693E32CA278FF959B046F07FB4D5524" box="[1128,1180,267,293]" italics="true" pageId="16" pageNumber="17">et al</emphasis>
. (2016
</bibRefCitation>
, 
<bibRefCitation id="907642CFA278FF959B936F00FAEF5524" author="Rubal M. &amp; Veiga P. &amp; Fontoura P. &amp; Sousa-Pinto I." box="[1279,1342,267,293]" pageId="16" pageNumber="17" pagination="921 - 928" refId="ref18110" refString="Rubal M., Veiga P., Fontoura P. &amp; Sousa-Pinto I. 2017. A new Batillipes (Tardigrada, Heterotardigrada, Batillipedidae) from North Portugal (Atlantic Ocean). Marine Biodiversity 47 (3): 921 - 928. https: // doi. org / 10.1007 / s 12526 - 016 - 0526 - x" type="journal article" year="2017">2017</bibRefCitation>
), the presence of 
<taxonomicName id="33E744BDA278FF959E276F24FE695548" box="[331,440,303,329]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="16" pageNumber="17" phylum="Tardigrada" rank="species" species="similis">
<emphasis id="C693E32CA278FF959E276F24FE695548" box="[331,440,303,329]" italics="true" pageId="16" pageNumber="17">B. similis</emphasis>
</taxonomicName>
on the Portuguese coast cannot be assumed.
</paragraph>
</subSubSection>
</treatment>
</document>