238 lines
29 KiB
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238 lines
29 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/vz.74.e110674" ID-Pensoft-Pub="2625-8498-74-23" ID-Pensoft-UUID="41B30B26C7D754A6847973C6FED0710C" ID-ZooBank="335FBFDDE54340CA80140E16A7977586" ModsDocID="2625-8498-74-23" checkinTime="1705718409033" checkinUser="pensoft" docAuthor="Agarwal, Ishan, Thackeray, Tejas & Khandekar, Akshay" docDate="2024" docId="7BFE4CBE2FC65EE1A45AA5AFADBFED26" docLanguage="en" docName="VertZool 74: 23-83" docOrigin="Vertebrate Zoology 74" docPubDate="2024-01-19" docSource="http://dx.doi.org/10.3897/vz.74.e110674" docTitle="Dravidoseps Agarwal & Thackeray & Khandekar 2024, gen. nov." docType="treatment" docUuid="D8825033-C908-4C22-B246-6DE3AD538D30" docUuidSource="ZooBank" docVersion="1" id="41B30B26C7D754A6847973C6FED0710C" lastPageNumber="23" masterDocId="41B30B26C7D754A6847973C6FED0710C" masterDocTitle="A non-adaptive radiation of viviparous skinks from the seasonal tropics of India: Systematics of Subdoluseps (Squamata: Scincidae), with description of a new genus and five cryptic new species" masterLastPageNumber="83" masterPageNumber="23" pageNumber="23" updateTime="1705718409033" updateUser="pensoft">
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<mods:titleInfo id="806F0945376E87FC53E1DCC4F3468519">
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<mods:title id="88A72D03D19D9FDD704EDB9B6FD589C7">A non-adaptive radiation of viviparous skinks from the seasonal tropics of India: Systematics of Subdoluseps (Squamata: Scincidae), with description of a new genus and five cryptic new species</mods:title>
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<mods:name id="44E589052DB97E114BA8AB485F49ECB8" type="personal">
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<mods:namePart id="AA5EE9E6019ABD16D37406F39CD78FE3">Agarwal, Ishan</mods:namePart>
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<mods:nameIdentifier id="33996E943F849777FC31AC36D24325E0" type="ORCID">https://orcid.org/0000-0001-9734-5379</mods:nameIdentifier>
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<mods:affiliation id="FB2E209CD7F23216A342DC2F9C941910">Thackeray Wildlife Foundation, Mumbai, 400051, India</mods:affiliation>
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<mods:nameIdentifier id="C77A0A3737AFA5ED8ADD44E0F0F37348" type="email">ishan.agarwal@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="BF94EC61E6EE623904F1E9126D7655DA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="10BB1C77ADF08A33AC05677F27BED975">Thackeray, Tejas</mods:namePart>
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<mods:nameIdentifier id="B927861710E9B4D8506DFB6E0A15D0DB" type="ORCID">https://orcid.org/0000-0002-9981-8763</mods:nameIdentifier>
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<mods:affiliation id="9CA072E71C9B46547158F4593F5B6870">Thackeray Wildlife Foundation, Mumbai, 400051, India</mods:affiliation>
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<mods:namePart id="F664C15143657807E630747F745FA248">Khandekar, Akshay</mods:namePart>
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<mods:nameIdentifier id="9590B33C5BE12FAC703B1A26451BD958" type="ORCID">https://orcid.org/0000-0002-7956-089X</mods:nameIdentifier>
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<mods:affiliation id="099EB9A86B37E29C1CB236F91057C0DF">Thackeray Wildlife Foundation, Mumbai, 400051, India & Department of Zoology, Shivaji University, Kolhapur, 416004, India</mods:affiliation>
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<mods:title id="670720E190B1915DFE235C62108C4480">Vertebrate Zoology</mods:title>
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<mods:part id="25B8AFFF1D8EFC565FA0BA3F7472B929">
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<mods:date id="AAB8E3AEF776E9FB9C449668F246B6AF">2024</mods:date>
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<mods:number id="AEBD7E1F28D265D77C88B97007553746">2024-01-19</mods:number>
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<mods:detail id="29EF5CAD456139CE4E626D7549005101" type="volume">
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<mods:number id="B7D7F4A7A482051FDDA0A94847FC80CE">74</mods:number>
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<mods:start id="C99683C838B0C37B4B29F2B522D40DEC">23</mods:start>
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<mods:end id="F3152B4E8ED619EDE6058FF44C59F44A">83</mods:end>
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<mods:url id="33B35948A5B5BC3A7B25B3D95C2C272B">http://dx.doi.org/10.3897/vz.74.e110674</mods:url>
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<mods:identifier id="D5C1112496D5736314E5CFF29922CEDC" type="DOI">http://dx.doi.org/10.3897/vz.74.e110674</mods:identifier>
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<treatment id="7BFE4CBE2FC65EE1A45AA5AFADBFED26" LSID="urn:lsid:zoobank.org:act:D8825033-C908-4C22-B246-6DE3AD538D30" httpUri="http://treatment.plazi.org/id/7BFE4CBE2FC65EE1A45AA5AFADBFED26" lastPageNumber="23" pageId="0" pageNumber="23">
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<subSubSection id="CB52FF03BE08FE1B14E7B15BDCFD46E5" pageId="0" pageNumber="23" type="nomenclature">
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<paragraph id="D42DEECE1DF88DF378C18F852274BD51" pageId="0" pageNumber="23">
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<taxonomicName id="B408ED6076B997B2F56445193EA28AE6" LSID="https://zoobank.org/D8825033-C908-4C22-B246-6DE3AD538D30" authority="Agarwal & Thackeray & Khandekar, 2024" authorityName="Agarwal & Thackeray & Khandekar" authorityYear="2024" family="Scincidae" genus="Dravidoseps" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Dravidoseps" order="Squamata" pageId="0" pageNumber="23" rank="genus" status="gen. nov.">Dravidoseps</taxonomicName>
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<taxonomicNameLabel id="C97D3F3398C2D91A16528B38DB891283" pageId="0" pageNumber="23">gen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="E9B846D64871014F4F099A8C5D73EB54" pageId="0" pageNumber="23" type="type species">
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<paragraph id="29C81A8B501E4819A65A72F0630D17C2" pageId="0" pageNumber="23">Type species.</paragraph>
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<paragraph id="02ECBC7AB0228112B13B761DCBB71EB7" pageId="0" pageNumber="23">
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<taxonomicName id="382D53C53794C99B2C5FDB4138097510" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygosoma pruthi" order="Squamata" pageId="0" pageNumber="23" phylum="Chordata" rank="species" species="pruthi">
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<emphasis id="32B9D6A67A5AEBEF4B14196D691613F0" italics="true" pageId="0" pageNumber="23">Lygosoma pruthi</emphasis>
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</taxonomicName>
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Sharma, 1977
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</paragraph>
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</subSubSection>
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<subSubSection id="61C31F3E38C11CBBCDD4B22C78C9675E" pageId="0" pageNumber="23" type="chresonymy">
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<paragraph id="AB4C33DAB9E5010078F43D4F3D2F60D8" pageId="0" pageNumber="23">Chresonymy.</paragraph>
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<paragraph id="8693E28EB47E69F18F605229D01ED929" pageId="0" pageNumber="23">
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<taxonomicName id="B941D78CAAF432AB588FFE1B3FEEF12B" class="Reptilia" family="Scincidae" genus="Riopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Riopa" order="Squamata" pageId="0" pageNumber="23" phylum="Chordata" rank="genus">
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<emphasis id="F6D90E6F320A9BAD0E34F27B5BAF1F44" italics="true" pageId="0" pageNumber="23">Riopa</emphasis>
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</taxonomicName>
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-
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<bibRefCitation id="33FF4194023ECC686D0375C3CFE6A942" author="Sharma, RC" journalOrPublisher="Records of the Zoological Survey of India" pageId="0" pageNumber="23" pagination="149 - 167" refId="B94" refString="Sharma, RC, 1976. Records of the reptiles of Goa. Records of the Zoological Survey of India 71: 149 - 167" title="Records of the reptiles of Goa." volume="71" year="1976">Sharma (1976</bibRefCitation>
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, 1978)
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</paragraph>
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<paragraph id="C06C2894AFA91288B8CB8E81A4CC68B6" pageId="0" pageNumber="23">
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<taxonomicName id="48BA45A0D6BC65A3CD35DFAE4F82AF49" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygosoma" order="Squamata" pageId="0" pageNumber="23" phylum="Chordata" rank="genus">
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<emphasis id="300926F4C41988A412FDE40AB028512C" italics="true" pageId="0" pageNumber="23">Lygosoma</emphasis>
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</taxonomicName>
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- Das (1996)
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</paragraph>
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<paragraph id="EA78B6483EC33A40A50CACC9D9485EE7" pageId="0" pageNumber="23">
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<taxonomicName id="8547D00B45CD1DDCD4C382CB5B6A5AA4" authorityName="Freitas, Datta-Roy, Karanth, L.Grismer & Siler" authorityYear="2019" class="Reptilia" family="Scincidae" genus="Subdoluseps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Subdoluseps" order="Squamata" pageId="0" pageNumber="23" phylum="Chordata" rank="genus">
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<emphasis id="2665B6C39F885CEB39F0D9B2F2A242F6" italics="true" pageId="0" pageNumber="23">Subdoluseps</emphasis>
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</taxonomicName>
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-
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<bibRefCitation id="79682ED2C06D2F4FB62F7E65851AACF1" DOI="https://doi.org/10.1093/zoolinnean/zlz001" author="Freitas, ES" journalOrPublisher="Zoological Journal of the Linnaean Society" pageId="0" pageNumber="23" pagination="1067 - 1096" refId="B38" refString="Freitas, ES, Datta-Roy, A, Karanth, P, Grismer, LL, Siler, CD, 2019. Multilocus phylogeny and a new classification for African, Asian and Indian supple and writhing skinks (Scincidae: Lygosominae). Zoological Journal of the Linnaean Society 186: 1067 - 1096, DOI: https://doi.org/10.1093/zoolinnean/zlz001" title="Multilocus phylogeny and a new classification for African, Asian and Indian supple and writhing skinks (Scincidae: Lygosominae)." url="https://doi.org/10.1093/zoolinnean/zlz001" volume="186" year="2019">Freitas et al. (2019)</bibRefCitation>
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,
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<bibRefCitation id="044D6046FDE13DD31FB6196A6447EB27" DOI="https://doi.org/10.11646/zootaxa.4950.2.7" author="Ganesh, SR" journalOrPublisher="Zootaxa" pageId="0" pageNumber="23" pagination="361 - 376" refId="B44" refString="Ganesh, SR, Srikanthan, AN, Ghosh, A, Adhikari, OD, Kumar, SV, Datta-Roy, A, 2021. A new species of Asian gracile skink (Scincidae: Lygosominae: Subdoluseps) from the rain-shadow belts of Nilgiri hills, Western Ghats, India. Zootaxa 4950: 361 - 376, DOI: https://doi.org/10.11646/zootaxa.4950.2.7" title="A new species of Asian gracile skink (Scincidae: Lygosominae: Subdoluseps) from the rain-shadow belts of Nilgiri hills, Western Ghats, India." url="https://doi.org/10.11646/zootaxa.4950.2.7" volume="4950" year="2021">Ganesh et al. (2021)</bibRefCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="EAFFD8C1FACBCDF5A7956DCC0F3E4584" pageId="0" pageNumber="23" type="diagnosis">
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<paragraph id="7D09D83AAE5FE978B6FD78FD3C31851D" pageId="0" pageNumber="23">Diagnosis.</paragraph>
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<paragraph id="90F12ED567FD93BF6C9C3898FEC23D74" pageId="0" pageNumber="23">
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Medium-sized skinks (adult SVL <58 mm; n = 89), original tail equal to or slightly longer than body. Dorsal scales on body and tail smooth, cycloid, imbricate; ventrals similar except marginally larger on pectoral and precloacal region; scales on lateral tail base smooth or tricarinate; 62-70 scales in paravertebral rows; 26-32 scales around mid-body; 61-73 ventral scales (rarely 76, n = 1/89); 8-12 enlarged precloacal scales (rarely 13, n = 1/89); and 18-23 scales round the tail. Supranasals in contact with each other behind rostral (rarely not in contact, n = 1/89); single frontonasal; prefrontals relatively small, widely separated on midline; frontal elongate, bell-shaped; four supraoculars; three supraoculars in contact with frontoparietal (rarely two, n = 4/89); frontoparietal divided; interparietal diamond-shaped, eyespot in posterior projection; parietals large, in medial contact posterior to interparietal; 2-4 nuchals, either in contact behind parietals or separated medially by 1-3 paravertebral scales. Nasal divided; two loreals; a single pre-supraocular; two preoculars (rarely three, n = 4/89); and a single sub-preocular (rarely absent, n = 5/89); 6-8 supraciliaries (rarely nine, n = 1/89); lower eyelid with enlarged, transparent central window; a single post-supraocular and postocular; and three or four sub-postoculars (rarely five, n = 3/89); a single primary, two secondary (rarely three, n = 1/89), and three tertiary (rarely four, n = 1/89) temporals. Six or seven supralabials and infralabials; fourth or fifth supralabial elongate, below eye; one or two post-supralabials; 1-3 ear lobules; three enlarged pairs of chin shields. Pentadactyl; limbs well-developed; subdigital lamellae unpaired, smooth to weakly keeled; 4-7 lamellae under digit I of manus and pes, 9-12 lamellae under digit IV of manus and 12-17 lamellae under digit IV of pes (rarely 18, n = 1/89). Viviparous, litter size 2-4. Dorsum light coconut to dark chocolate brown; thick dark band from rostrum to tail speckled with light spots; supralabials with a white streak; males with yellow on lower parts of forebody and flanks, sometimes extending onto belly; venter white with some darker markings (Fig.
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<figureCitation id="20B1EFEBB31F1361057D3CE592EB91B4" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Dravidoseps gen. nov. species in life: A D. pruthi comb. nov. (neotype, NRC-AA- 1291); B D. goaensis comb. nov. (neotype, BNHS 2567); C D. nilgiriensis comb. nov. (ZSI-R- 28604); D D. gingeeensis sp. nov. (holotype, NRC-AA- 8273); E D. jawadhuensis sp. nov. (holotype, NRC-AA- 8274); F D. kalakadensis sp. nov. (holotype, NRC-AA- 8275); G D. srivilliputhurensis sp. nov. (holotype, NRC-AA- 8279); and H D. tamilnaduensis sp. nov. (holotype, NRC-AA- 8286). Photos by Akshay Khandekar (B-D, F-H) and Ishan Agarwal (A, E)." figureDoi="10.3897/vz.74.e110674.figure9" httpUri="https://binary.pensoft.net/fig/970068" pageId="0" pageNumber="23">9</figureCitation>
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).
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</paragraph>
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<caption id="9A05167540E0650CA8943D534F2A1B01" doi="10.3897/vz.74.e110674.figure9" httpUri="https://binary.pensoft.net/fig/970068" pageId="0" pageNumber="23" start="Figure 9" startId="F9">
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<paragraph id="406E875918A63F3DF2263AAF5E6C9683" pageId="0" pageNumber="23">
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<emphasis id="A6188F2F51D57299F60A7F54593AAAA6" bold="true" pageId="0" pageNumber="23">Figure 9.</emphasis>
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<taxonomicName id="DDD1CEEEDD286C8F53B3093ADE3AC0B5" authorityName="Agarwal & Thackeray & Khandekar" authorityYear="2024" family="Scincidae" genus="Dravidoseps" kingdom="Animalia" lsidName="Dravidoseps" order="Squamata" pageId="0" pageNumber="23" rank="genus">
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<emphasis id="71B87D5385A745FC4DB7BCE9BB86492E" italics="true" pageId="0" pageNumber="23">Dravidoseps</emphasis>
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</taxonomicName>
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<emphasis id="DA31F0D0EB39204BF6A494B336ED7B5B" bold="true" pageId="0" pageNumber="23">gen. nov.</emphasis>
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species in life:
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<emphasis id="8600D874FE5C82118774D60C495E0311" bold="true" pageId="0" pageNumber="23">A</emphasis>
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<taxonomicName id="64FE2A32C2DF1CC9F75FA0A4E389DAB9" lsidName="D. pruthi" pageId="0" pageNumber="23" rank="species" species="pruthi">
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<emphasis id="CC95A47AA1445146B3DD80F739301730" italics="true" pageId="0" pageNumber="23">D. pruthi</emphasis>
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</taxonomicName>
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<emphasis id="CFDC6A306A6A7521114A58334391ED97" bold="true" pageId="0" pageNumber="23">comb. nov.</emphasis>
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(neotype, NRC-AA-1291);
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<emphasis id="C681203A25C690A0E2194589C6D2197A" bold="true" pageId="0" pageNumber="23">B</emphasis>
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<taxonomicName id="03B9F2E149BBFD921E3FB8632BAC1341" lsidName="D. goaensis" pageId="0" pageNumber="23" rank="species" species="goaensis">
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<emphasis id="C3299E1A9C06D66E040FCA67EB710AEE" italics="true" pageId="0" pageNumber="23">D. goaensis</emphasis>
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</taxonomicName>
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<emphasis id="895664AA36BFB078D9A4F7018CB8D1E2" bold="true" pageId="0" pageNumber="23">comb. nov.</emphasis>
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(neotype, BNHS 2567);
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<emphasis id="4617E79679406103BD145E53C50C53A8" bold="true" pageId="0" pageNumber="23">C</emphasis>
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<taxonomicName id="9C293199E8DF14BDA2BF8AF703E2CFB2" lsidName="D. nilgiriensis" pageId="0" pageNumber="23" rank="species" species="nilgiriensis">
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<emphasis id="5300682CDEE7C9BB586F14E369E409B4" italics="true" pageId="0" pageNumber="23">D. nilgiriensis</emphasis>
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</taxonomicName>
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<emphasis id="04323323FD6B092F9441B57184F5D44B" bold="true" pageId="0" pageNumber="23">comb. nov.</emphasis>
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(ZSI-R-28604);
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<emphasis id="AA0C18F067707A9DB7A9A7122DBC427D" bold="true" pageId="0" pageNumber="23">D</emphasis>
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<taxonomicName id="1D876F4216C982072F408CD1BF00265F" lsidName="D. gingeeensis" pageId="0" pageNumber="23" rank="species" species="gingeeensis">
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<emphasis id="C865712ECC933ECA8B1F5B9F0D1ABA99" italics="true" pageId="0" pageNumber="23">D. gingeeensis</emphasis>
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</taxonomicName>
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<emphasis id="4AAED37500694140D20F97B5FFC18934" bold="true" pageId="0" pageNumber="23">sp. nov.</emphasis>
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(holotype, NRC-AA-8273);
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<emphasis id="5D525C46DF7FD7C4C886098F31AEC90A" bold="true" pageId="0" pageNumber="23">E</emphasis>
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<taxonomicName id="7D261D2679D0857B45B84B7F567B049D" lsidName="D. jawadhuensis" pageId="0" pageNumber="23" rank="species" species="jawadhuensis">
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<emphasis id="DA68F4750431B5B0CD5EA2214E4EE03C" italics="true" pageId="0" pageNumber="23">D. jawadhuensis</emphasis>
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</taxonomicName>
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<emphasis id="85AA505A5B2E1683D6FA3F358B0E8153" bold="true" pageId="0" pageNumber="23">sp. nov.</emphasis>
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(holotype, NRC-AA-8274);
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<emphasis id="12F85F52523731F2E636FC22FF2494E0" bold="true" pageId="0" pageNumber="23">F</emphasis>
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<taxonomicName id="7C09FED82E54F9C5B21798982A911AED" lsidName="D. kalakadensis" pageId="0" pageNumber="23" rank="species" species="kalakadensis">
|
||
<emphasis id="F680725A05AF025FB68E8F966D403125" italics="true" pageId="0" pageNumber="23">D. kalakadensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C4AC3B0B170B744192D226D46EBCD7CA" bold="true" pageId="0" pageNumber="23">sp. nov.</emphasis>
|
||
(holotype, NRC-AA-8275);
|
||
<emphasis id="3B1DF7EB5340D54CC65E749B7697BEF4" bold="true" pageId="0" pageNumber="23">G</emphasis>
|
||
<taxonomicName id="080F6354FE0DD2CA4C6BC31F21D33EBF" lsidName="D. srivilliputhurensis" pageId="0" pageNumber="23" rank="species" species="srivilliputhurensis">
|
||
<emphasis id="CB429512D6A5D1D6FE8E0E0EFCFA5736" italics="true" pageId="0" pageNumber="23">D. srivilliputhurensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D6E6FB9A622DBC9F3773ED2ED4F9014F" bold="true" pageId="0" pageNumber="23">sp. nov.</emphasis>
|
||
(holotype, NRC-AA-8279); and
|
||
<emphasis id="2572BF9A8D9132771EBB339DC0D53C17" bold="true" pageId="0" pageNumber="23">H</emphasis>
|
||
<taxonomicName id="B3958C46FB7797A635CD2E5ACB0FA8C3" lsidName="D. tamilnaduensis" pageId="0" pageNumber="23" rank="species" species="tamilnaduensis">
|
||
<emphasis id="2D090144BAE3E426AE425CBA9A7A9D31" italics="true" pageId="0" pageNumber="23">D. tamilnaduensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D03EF6F67C3EC0E2B7319BCBDE7EE36A" bold="true" pageId="0" pageNumber="23">sp. nov.</emphasis>
|
||
(holotype, NRC-AA-8286). Photos by Akshay Khandekar (B-D, F-H) and Ishan Agarwal (A, E).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="CC33075FFDEFA5CA12E1376AE98A0401" pageId="0" pageNumber="23">
|
||
<taxonomicName id="FEA0E29534720EEEFB0BD51D64A4B6D1" authorityName="Agarwal & Thackeray & Khandekar" authorityYear="2024" family="Scincidae" genus="Dravidoseps" kingdom="Animalia" lsidName="Dravidoseps" order="Squamata" pageId="0" pageNumber="23" rank="genus">
|
||
<emphasis id="AFB3D14D53A25252D1794419503D114F" italics="true" pageId="0" pageNumber="23">Dravidoseps</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="196D99A0E8B89D75FA971A1684D8DD0A" bold="true" pageId="0" pageNumber="23">gen. nov.</emphasis>
|
||
differs from
|
||
<taxonomicName id="82AD42B1A6A52ACBB9C66528E12815F2" authorityName="Freitas, Datta-Roy, Karanth, L.Grismer & Siler" authorityYear="2019" class="Reptilia" family="Scincidae" genus="Subdoluseps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Subdoluseps" order="Squamata" pageId="0" pageNumber="23" phylum="Chordata" rank="genus">
|
||
<emphasis id="155DC9AD2BB23F7EB6871A3EA5B37AC7" italics="true" pageId="0" pageNumber="23">Subdoluseps</emphasis>
|
||
</taxonomicName>
|
||
by the presence of a transparent central window in the lower eyelid (versus no transparent central window in the lower eyelid), by the presence of an open secondary palate (versus a closed secondary palate) and by being viviparous (versus oviparous) (
|
||
<bibRefCitation id="6C92D7166EF59451481E8886EC391869" DOI="https://doi.org/10.1093/zoolinnean/zlz001" author="Freitas, ES" journalOrPublisher="Zoological Journal of the Linnaean Society" pageId="0" pageNumber="23" pagination="1067 - 1096" refId="B38" refString="Freitas, ES, Datta-Roy, A, Karanth, P, Grismer, LL, Siler, CD, 2019. Multilocus phylogeny and a new classification for African, Asian and Indian supple and writhing skinks (Scincidae: Lygosominae). Zoological Journal of the Linnaean Society 186: 1067 - 1096, DOI: https://doi.org/10.1093/zoolinnean/zlz001" title="Multilocus phylogeny and a new classification for African, Asian and Indian supple and writhing skinks (Scincidae: Lygosominae)." url="https://doi.org/10.1093/zoolinnean/zlz001" volume="186" year="2019">Freitas et al. 2019</bibRefCitation>
|
||
; Zimin et al. 2021).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="16081B4BC4469194343941D3391116D7" pageId="0" pageNumber="23" type="content">
|
||
<paragraph id="1A2D94051756DDC1BA90C5A06DCC962F" pageId="0" pageNumber="23">Content.</paragraph>
|
||
<paragraph id="09BD165CCEB2FEFDBC0B3C4173B6979F" pageId="0" pageNumber="23">
|
||
<taxonomicName id="C9F8196D5C3850DBFBF78C1CB96E0971" authorityName="Sharma" authorityYear="1976" baseAuthorityName="Sharma" baseAuthorityYear="1976" family="Scincidae" genus="Dravidoseps" kingdom="Animalia" lsidName="Dravidoseps goaensis" order="Squamata" pageId="0" pageNumber="23" rank="species" species="goaensis">
|
||
<emphasis id="024548A52F53F306E608B7958932C1E6" italics="true" pageId="0" pageNumber="23">Dravidoseps goaensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0BC32923941E08E7E9AB938F6EB5F0D4" bold="true" pageId="0" pageNumber="23">comb. nov.</emphasis>
|
||
,
|
||
<taxonomicName id="91020A5C23E317359BFE8235ADF3DD5B" authorityName="Sharma" authorityYear="1977" baseAuthorityName="Sharma" baseAuthorityYear="1977" family="Scincidae" genus="Dravidoseps" kingdom="Animalia" lsidName="Dravidoseps pruthi" order="Squamata" pageId="0" pageNumber="23" rank="species" species="pruthi">
|
||
<emphasis id="BABFA6D45AAADCBAA9AB19C3AA49772B" italics="true" pageId="0" pageNumber="23">Dravidoseps pruthi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7BC5F120F05C479C226731CE87F875C8" bold="true" pageId="0" pageNumber="23">comb. nov</emphasis>
|
||
.,
|
||
<taxonomicName id="22C8DEF4DD836673CA239FE3259F413B" family="Scincidae" genus="Dravidoseps" kingdom="Animalia" lsidName="Dravidoseps nilgiriensis" order="Squamata" pageId="0" pageNumber="23" rank="species" species="nilgiriensis">
|
||
<emphasis id="65590CE6EC39746996EE5EE42CF3B225" italics="true" pageId="0" pageNumber="23">Dravidoseps nilgiriensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="8DA829ECB0255173C34AAA9C956FE055" bold="true" pageId="0" pageNumber="23">comb. nov.</emphasis>
|
||
, and five species described below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2C7A06419E6791AEB9BA559859EE7410" pageId="0" pageNumber="23" type="phylogenetic definition">
|
||
<paragraph id="5EC8ECCC6FB37BA028AC8E9558CCDA86" pageId="0" pageNumber="23">Phylogenetic definition.</paragraph>
|
||
<paragraph id="DF5AC0355378D5A7FDBE89E91913B0D1" pageId="0" pageNumber="23">
|
||
This genus comprises species that share a more recent common ancestor with
|
||
<taxonomicName id="F0B52F31C305934D2D648D8A180DC7A9" authorityName="Sharma" authorityYear="1977" baseAuthorityName="Sharma" baseAuthorityYear="1977" family="Scincidae" genus="Dravidoseps" kingdom="Animalia" lsidName="Dravidoseps pruthi" order="Squamata" pageId="0" pageNumber="23" rank="species" species="pruthi">
|
||
<emphasis id="651CBA6BA1983181CBA46412E4847A12" italics="true" pageId="0" pageNumber="23">Dravidoseps pruthi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B4846829BCD1F5C6E5ECD88E59810D36" bold="true" pageId="0" pageNumber="23">comb. nov.</emphasis>
|
||
than with
|
||
<taxonomicName id="11D40B7A5EB2940209D044FD637BCB02" baseAuthorityName="Gunther" baseAuthorityYear="1864" class="Reptilia" family="Scincidae" genus="Subdoluseps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Subdoluseps bowringii" order="Squamata" pageId="0" pageNumber="23" phylum="Chordata" rank="species" species="bowringii">
|
||
<emphasis id="7535FCEEE49583D0D160312FA9BC6ED7" italics="true" pageId="0" pageNumber="23">Subdoluseps bowringii</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32F8072663A173BE64D8D9E65EE60E1" pageId="0" pageNumber="23" type="etymology">
|
||
<paragraph id="C80D049A658A2553A0A4103611C33292" pageId="0" pageNumber="23">Etymology.</paragraph>
|
||
<paragraph id="7D7FDA46DCD32E6E8AC7E2F9C9EAE5EF" pageId="0" pageNumber="23">
|
||
A combination of the Sanskrit
|
||
<normalizedToken id="7DE657662427FCD28711986B3E84A7FC" originalValue="‘Dravida’">'Dravida'</normalizedToken>
|
||
, referring to the original inhabitants of southern India and Sri Lanka, and the Ancient Greek
|
||
<normalizedToken id="E3A4182C14588B76E5F5B118F205EC86" originalValue="‘seps’">'seps'</normalizedToken>
|
||
, for a snake-like creature that has been previously used in skink generic names (e.g.
|
||
<bibRefCitation id="CC1AEEE7778435DC371FB6F65A95F56E" DOI="https://doi.org/10.1016/j.ympev.2016.12.008" author="Erens, J" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="23" pagination="466 - 472" refId="B35" refString="Erens, J, Miralles, A, Glaw, F, Chatrou, LW, Vences, M, 2017. Extended molecular phylogenetics and revised systematics of Malagasy scincine lizards. Molecular Phylogenetics and Evolution 107: 466 - 472, DOI: https://doi.org/10.1016/j.ympev.2016.12.008" title="Extended molecular phylogenetics and revised systematics of Malagasy scincine lizards." url="https://doi.org/10.1016/j.ympev.2016.12.008" volume="107" year="2017">Erens et al. 2017</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="F15B4762A3D83880A94B360DBAC80194" DOI="https://doi.org/10.1093/zoolinnean/zlz001" author="Freitas, ES" journalOrPublisher="Zoological Journal of the Linnaean Society" pageId="0" pageNumber="23" pagination="1067 - 1096" refId="B38" refString="Freitas, ES, Datta-Roy, A, Karanth, P, Grismer, LL, Siler, CD, 2019. Multilocus phylogeny and a new classification for African, Asian and Indian supple and writhing skinks (Scincidae: Lygosominae). Zoological Journal of the Linnaean Society 186: 1067 - 1096, DOI: https://doi.org/10.1093/zoolinnean/zlz001" title="Multilocus phylogeny and a new classification for African, Asian and Indian supple and writhing skinks (Scincidae: Lygosominae)." url="https://doi.org/10.1093/zoolinnean/zlz001" volume="186" year="2019">Freitas et al. 2019</bibRefCitation>
|
||
). The gender of the genus is masculine and the suggested common name is Indian leaf-litter skinks.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F720C56676001158DE023579E41B5805" pageId="0" pageNumber="23" type="distribution">
|
||
<paragraph id="C9B311B73A3BC88540012CCE43180492" pageId="0" pageNumber="23">Distribution and natural history.</paragraph>
|
||
<paragraph id="57E03DB9E227D7AD6BA80F5E1F8A7BDD" pageId="0" pageNumber="23">
|
||
The three known species and five unnamed lineages (described below) are distributed in peninsular India, including the northern Western Ghats, central and southern Western Ghats, the edge of the Mysore Plateau and isolated massifs in Tamil Nadu (Fig.
|
||
<figureCitation id="56EDDFE1F1E4A1E5751BAB4A1887964B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Elevation map showing sampling localities from peninsular India. Stars indicate type localities (neotype locality shown for Dravidoseps goaensis comb. nov., original type locality is close to the southernmost point), light blue D. jawadhuensis sp. nov., dark blue D. srivilliputhurensis sp. nov., light green D. kalakadensis sp. nov., dark green D. pruthi comb. nov., red D. gingeeensis sp. nov., yellow D. nilgiriensis comb. nov., pink D. tamilnaduensis sp. nov., and grey, D. sp. Kalrayan (colour scheme representing species is the same as Figs 3 - 6), black squares unsampled previously known localities. The approximate extent of the Central Western Ghats (CWG), Northern Western Ghats (NWG), and Southern Western Ghats (SWG) are shown; major hill ranges and features are marked by bold white text, within the Western Ghats: AG, Agasthyamalai; AN, Anaimalai; DV, Devarmalai; N, Nilgiris; PG, Palghat Gap; PL, Palani; and outside the Western Ghats: BR, Biligirangan; J, Jawadhu; K, Kollimalai; KR, Kalrayan; mm, Male Mahadeshwara; MP, Mysore Plateau; PC, Pachaimalai; PM, Palamalai; S, Sitteri; SR, Sirumalai; T, Tirupati; V, Velikonda; Y, Yercaud; YL, Yelagiri. Inset, global distribution of Dravidoseps gen. nov. (brown) and Subdoluseps sensu stricto (green). Dravidoseps goaensis = Riopa goaensis (see main text)." figureDoi="10.3897/vz.74.e110674.figure1" httpUri="https://binary.pensoft.net/fig/970060" pageId="0" pageNumber="23">1</figureCitation>
|
||
). Members of the genus are diurnal (sometimes seen moving in the litter soon after dark) and are terrestrial and partially fossorial.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |