188 lines
22 KiB
XML
188 lines
22 KiB
XML
<document ID-CLB-Dataset="21253" ID-DOI="http://dx.doi.org/10.3897/zookeys.961.54846" ID-GBIF-Dataset="a5c61646-70dc-471b-9bd8-0b407e1cedc4" ID-PMC="PMC7449986" ID-Pensoft-Pub="1313-2970-961-1" ID-Pensoft-UUID="634A9CDA65075FEE8D7C196F5E37F6FC" ID-PubMed="32904058" ID-ZooBank="C5E8BED7F08549A999C60C3AA8492D09" ModsDocID="1313-2970-961-1" checkinTime="1597940209253" checkinUser="pensoft" docAuthor="Breedy, Odalisca & Guzman, Hector M." docDate="2020" docId="7BA59E16FC785DA9882937EE56C2FFD8" docLanguage="en" docName="ZooKeys 961: 1-30" docOrigin="ZooKeys 961" docSource="http://dx.doi.org/10.3897/zookeys.961.54846" docTitle="Psammogorgia gracilis Verrill 1868" docType="treatment" docVersion="6" id="634A9CDA65075FEE8D7C196F5E37F6FC" lastPageNumber="1" masterDocId="634A9CDA65075FEE8D7C196F5E37F6FC" masterDocTitle="A revision of the genus Psammogorgia Verrill, 1868 (Cnidaria, Anthozoa, Octocorallia) in the tropical eastern Pacific Ocean" masterLastPageNumber="30" masterPageNumber="1" pageNumber="1" updateTime="1732748309243" updateUser="ExternalLinkService">
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<mods:title id="FF785757F8EAD442D9ABBC0C7769A2DD">A revision of the genus Psammogorgia Verrill, 1868 (Cnidaria, Anthozoa, Octocorallia) in the tropical eastern Pacific Ocean</mods:title>
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<mods:namePart id="62FED5D08FE6DA50BDB53E735FBE50BC">Breedy, Odalisca</mods:namePart>
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<mods:affiliation id="81D74922D6940D593104D60CB6351543">Centro de Investigacion en Estructuras Microscopicas, Centro de Investigacion en Ciencias del Mar y Limnologia, Escuela de Biologia, Universidad de Costa Rica. P. O. Box 11501 - 2060, Universidad de Costa Rica, San Jose, Costa Rica & Smithsonian Tropical Research Institute, P. O. Box 0843 - 03092, Panama, Republic of Panama</mods:affiliation>
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<mods:nameIdentifier id="ABFC46DAA51FB33991005D8D93396AEA" type="email">odaliscab@gmail.com</mods:nameIdentifier>
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<mods:namePart id="C85250B6EAA920D47E75E3DB32AEE969">Guzman, Hector M.</mods:namePart>
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<mods:affiliation id="F3718B5FF4C20CD50EE2CD02D7EC8EFF">Smithsonian Tropical Research Institute, P. O. Box 0843 - 03092, Panama, Republic of Panama</mods:affiliation>
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<mods:date id="CDCE70B7B85BEAB121CD048CD789F3BE">2020</mods:date>
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<mods:url id="AC68B2B2D37243E834B6DFE1B899A894">http://dx.doi.org/10.3897/zookeys.961.54846</mods:url>
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<mods:identifier id="D1910F5898ABB51FD21A106D05079D45" type="DOI">http://dx.doi.org/10.3897/zookeys.961.54846</mods:identifier>
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<treatment id="7BA59E16FC785DA9882937EE56C2FFD8" ID-GBIF-Taxon="167284436" LSID="urn:lsid:plazi:treatment:7BA59E16FC785DA9882937EE56C2FFD8" httpUri="http://treatment.plazi.org/id/7BA59E16FC785DA9882937EE56C2FFD8" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Anthozoa" scope_family="Plexauridae" scope_order="Alcyonacea" scope_phylum="Cnidaria" scope_subClass="Octocorallia">
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<subSubSection id="50BC086BC1655EA21A8F7C143331D66D" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="3D2852496282981C73F02CBF5D9CD643" pageId="0" pageNumber="1">
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<taxonomicName id="01F2EEF9A789A01007E7A77341171BDC" ID-CoL="78BSM" LSID="7BA59E16-FC78-5DA9-8829-37EE56C2FFD8" authority="Verrill, 1868" authorityName="Verrill" authorityYear="1868" class="Anthozoa" family="Plexauridae" genus="Psammogorgia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammogorgia gracilis" order="Alcyonacea" pageId="0" pageNumber="1" phylum="Cnidaria" rank="species" species="gracilis">Psammogorgia gracilis Verrill, 1868</taxonomicName>
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<figureCitation id="0AE38D5BC1CEF62D0F5FF79C7ABE6DA3" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Psammogorgia gracilis Verrill, 1868. YPM 813 a A colony B detail of branches showing a tumor C coenenchymal and anthocodial sclerites." figureDoi="10.3897/zookeys.961.54846.figure12" httpUri="https://binary.pensoft.net/fig/442685" pageId="0" pageNumber="1">Figures 12</figureCitation>
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<figureCitation id="6CA61445194343B250C3418FB70516CD" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Psammogorgia gracilis Verrill, 1868. YPM 813 a A spindles B wart clubs C radiates D anthocodial spindles." figureDoi="10.3897/zookeys.961.54846.figure13" httpUri="https://binary.pensoft.net/fig/442686" pageId="0" pageNumber="1">, 13</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="480151E571D96AAABDB90E8FEC3472CB" pageId="0" pageNumber="1" type="reference_group">
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<paragraph id="BC40BAF891E0E07803DD932A5F3AD5EA" pageId="0" pageNumber="1">
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<taxonomicName id="A2F7F32773ACEBA1644B270C9153A1B6" authorityName="Verrill" authorityYear="1868" class="Anthozoa" family="Plexauridae" genus="Psammogorgia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammogorgia gracilis" order="Alcyonacea" pageId="0" pageNumber="1" phylum="Cnidaria" rank="species" species="gracilis">Psammogorgia gracilis</taxonomicName>
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Verrill, 1868b: 417-418;
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<bibRefCitation id="276604CF0F4C39BEC9FE5A59F605CDD9" author="Kuekenthal, W" journalOrPublisher="Proceedings of the Zoological Society, London" pageId="0" pageNumber="1" refId="B22" refString="Kuekenthal, W, 1919. Gorgonaria. Wissenschaftliche Ergebnisse der deutsche Tiefsee-Expeditionen "Valdivia" 1898-99, 13(2): 1-946." title="Gorgonaria. Wissenschaftliche Ergebnisse der deutsche Tiefsee-Expeditionen " Valdivia " 1898 - 99, 13 (2): 1 - 946." year="1919">
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<normalizedToken id="9D7F8CA5B46017DDE7F82F233F1C0560" originalValue="Kükenthal">Kuekenthal</normalizedToken>
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1919
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</bibRefCitation>
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: 238;
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<bibRefCitation id="B030178D3A4644D9E92D5A303DBA9D03" author="Kuekenthal, W" journalOrPublisher="Walter de Gruyter and Company, Berlin, und Leipzig" pageId="0" pageNumber="1" refId="B23" refString="Kuekenthal, W, 1924. Gorgonaria. Das Tierreich, Vol. 47. Walter de Gruyter and Company, Berlin, und Leipzig" title="Gorgonaria. Das Tierreich, Vol. 47." year="1924">
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<normalizedToken id="6089B3C1BC0E066AAD57082C67AA2496" originalValue="Kükenthal">Kuekenthal</normalizedToken>
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1924
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</bibRefCitation>
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: 108; Breedy & Guzman 2011: 29.
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</paragraph>
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<paragraph id="7C7C576E095DE38143906DAD702F70E9" pageId="0" pageNumber="1">
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<taxonomicName id="7E21C626FCA7936F9697A947980EFDC8" baseAuthorityName="Verrill" baseAuthorityYear="1868" class="Anthozoa" family="Plexauridae" genus="Heterogorgia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heterogorgia gracilis" order="Alcyonacea" pageId="0" pageNumber="1" phylum="Cnidaria" rank="species" species="gracilis">Heterogorgia gracilis</taxonomicName>
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Harden, 1979: 112-113.
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</paragraph>
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</subSubSection>
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<subSubSection id="3763762E92A7C56BD1DC08DB59D408CD" pageId="0" pageNumber="1" type="type material">
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<paragraph id="771253547C2272ADF1FE584A65846A31" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="A8393755E944ADD95C97A296401D2C7D" pageId="0" pageNumber="1">
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<emphasis id="03D08F8EAE09C2E8D940BC547E57D7D0" bold="true" italics="true" pageId="0" pageNumber="1">Lectotype</emphasis>
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(designated herein): YPM 813a, dry, Pearl Islands,
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<normalizedToken id="881FC7D59F42B6BCDC0C8E1B6FE866FB" originalValue="Panamá">Panama</normalizedToken>
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, F.H. Bradley, 1866-1867, no additional data.
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</paragraph>
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<paragraph id="9C4B409C305CD72C2B3759C9C881C1A9" pageId="0" pageNumber="1">
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<emphasis id="6F01F33B6339001A1846340DF93BC310" bold="true" italics="true" pageId="0" pageNumber="1">Paralectotype</emphasis>
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: YPM 813b, dry small fragment, same data as the lectotype.
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</paragraph>
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</subSubSection>
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<subSubSection id="7FDECE0F7475DA0C1689A8DA3637FBDD" pageId="0" pageNumber="1" type="type locality">
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<paragraph id="DAA5BED8CBEC5ED24071EAAA15698657" pageId="0" pageNumber="1">Type locality.</paragraph>
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<paragraph id="9EC80AF4C76C4F1D50563C2B10EE94D6" pageId="0" pageNumber="1">
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Pearl Islands,
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<normalizedToken id="7DFE45B6C595EAB18D8A8B399FED3610" originalValue="Panamá">Panama</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="B855ED168B2AF6AF26601BA030343FC1" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="FAC512A449EF2F3AF4A524591E20A0A3" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="448B40D69D2BD23E02286A975B3F9DC8" pageId="0" pageNumber="1">Colonies red, tall, flabelliform, branches subparallel and elongated. Stem a few centimetres long. Calyces close together and all around the branches. Calyces slightly raised and swollen with concentration of wart-clubs up to 0.25 mm long, and acute spindles around the calyx rim. Coenenchymal sclerites: irregular spindles with acute or bifurcated ends up to 0.20 mm long; warty and irregular radiates and some crosses. Anthocodial spindles up to 0.24 mm long in collaret and points arrangements. Sclerites red and orange or of lighter hues.</paragraph>
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</subSubSection>
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<subSubSection id="89A0FCC2784779B93E0E5D90ED868DF9" pageId="0" pageNumber="1" type="description">
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<paragraph id="52090C28A0FBCA15BE31492C02A7EB66" pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph id="17A6C6203EC3A2F9C65E8618E61C63FA" pageId="0" pageNumber="1">
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The lectotype colony is red and 10 cm long and ~ 6 cm wide. Branching is irregularly dichotomous. The holdfast is absent. The branches emerge at angles of 45-90°, ascend parallel and slightly curve. The main branch of the colony is 2.5 mm in diameter subdividing into long, slender, ascending branchlets, 1.5-1.6 mm in diameter. Branches are round, slender, some extending up to 6.3 cm, undivided or bifurcating at the ends. Branches subdivide up to nine times while terminal branchlets are up to 60 mm long with rounded slightly tapered tips. (Fig.
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<figureCitation id="54BC33F783BFDFEE8002C437E05C54B7" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Psammogorgia gracilis Verrill, 1868. YPM 813 a A colony B detail of branches showing a tumor C coenenchymal and anthocodial sclerites." figureDoi="10.3897/zookeys.961.54846.figure12" httpUri="https://binary.pensoft.net/fig/442685" pageId="0" pageNumber="1">12A</figureCitation>
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). The calyces are densely arranged around the branches, slightly raised, swollen and are around 0.3 mm tall and 1 mm in diameter (Fig.
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<figureCitation id="0309F3E6A993C4F91C05A0B310FA24D0" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Psammogorgia gracilis Verrill, 1868. YPM 813 a A colony B detail of branches showing a tumor C coenenchymal and anthocodial sclerites." figureDoi="10.3897/zookeys.961.54846.figure12" httpUri="https://binary.pensoft.net/fig/442685" pageId="0" pageNumber="1">12B</figureCitation>
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). The calyces have wart-clubs and some irregular spindles around the calyx rim and the base of the anthocodiae. Coenenchymal sclerites are red, orange or of lighter hues (Fig.
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<figureCitation id="C159A8707D16CE9BB308E60F198C0495" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Psammogorgia gracilis Verrill, 1868. YPM 813 a A colony B detail of branches showing a tumor C coenenchymal and anthocodial sclerites." figureDoi="10.3897/zookeys.961.54846.figure12" httpUri="https://binary.pensoft.net/fig/442685" pageId="0" pageNumber="1">12A-C</figureCitation>
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); including slim spindles with acute ends that may bifurcate (Figs
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<figureCitation id="D3E880847BE65FE2DA6E6A117BA24B7E" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Psammogorgia gracilis Verrill, 1868. YPM 813 a A colony B detail of branches showing a tumor C coenenchymal and anthocodial sclerites." figureDoi="10.3897/zookeys.961.54846.figure12" httpUri="https://binary.pensoft.net/fig/442685" pageId="0" pageNumber="1">12C</figureCitation>
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,
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<figureCitation id="BA0D0DC31732B4688EBA7A69E0210FD9" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Psammogorgia gracilis Verrill, 1868. YPM 813 a A spindles B wart clubs C radiates D anthocodial spindles." figureDoi="10.3897/zookeys.961.54846.figure13" httpUri="https://binary.pensoft.net/fig/442686" pageId="0" pageNumber="1">13A</figureCitation>
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). Spindles are 0.12-0.24 mm long and 0.04-0.07 mm wide. Wart-clubs are 0.11-0.25 mm long and 0.04-0.08 mm wide at the expanded head (Fig.
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<figureCitation id="21F42F7B07478F83C63E2E561ADE99E2" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Psammogorgia gracilis Verrill, 1868. YPM 813 a A spindles B wart clubs C radiates D anthocodial spindles." figureDoi="10.3897/zookeys.961.54846.figure13" httpUri="https://binary.pensoft.net/fig/442686" pageId="0" pageNumber="1">13B</figureCitation>
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). Warty radiates and crosses are 0.08-0.10 mm long and 0.055-0.07 mm wide (Fig.
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<figureCitation id="995F5236BBBD5CF0B92CC0C7A139035C" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Psammogorgia gracilis Verrill, 1868. YPM 813 a A spindles B wart clubs C radiates D anthocodial spindles." figureDoi="10.3897/zookeys.961.54846.figure13" httpUri="https://binary.pensoft.net/fig/442686" pageId="0" pageNumber="1">13C</figureCitation>
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); and some warty crosses up to 0.10 mm by 0.09 mm. Anthocodial armature is well developed with orange and pale yellow spiny spindles arranged in collaret and points (Fig.
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<figureCitation id="78C2A4FADC46DAC3AE704BAE571DABE8" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Psammogorgia gracilis Verrill, 1868. YPM 813 a A colony B detail of branches showing a tumor C coenenchymal and anthocodial sclerites." figureDoi="10.3897/zookeys.961.54846.figure12" httpUri="https://binary.pensoft.net/fig/442685" pageId="0" pageNumber="1">12A-C</figureCitation>
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), measuring 0.11-0.20 mm long and 0.03-0.04 mm wide (Fig.
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<figureCitation id="2C2B2E897E82EF6BE6FBDA7C09A829A6" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Psammogorgia gracilis Verrill, 1868. YPM 813 a A spindles B wart clubs C radiates D anthocodial spindles." figureDoi="10.3897/zookeys.961.54846.figure13" httpUri="https://binary.pensoft.net/fig/442686" pageId="0" pageNumber="1">13D</figureCitation>
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) and flat spindles with small tubercles and scattered warts.
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</paragraph>
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<caption id="ABCBCDB51ED1066A99A501436977DD51" doi="10.3897/zookeys.961.54846.figure12" httpUri="https://binary.pensoft.net/fig/442685" pageId="0" pageNumber="1" start="Figure 12" startId="F12">
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<paragraph id="2BBDDE4E2D5D02061355CF04AB5304F7" pageId="0" pageNumber="1">
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<emphasis id="4ABCE829FD1C7186D482D7BCFF608C8F" bold="true" pageId="0" pageNumber="1">Figure 12.</emphasis>
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<taxonomicName id="2838EFB8709E3CA06696AF8CA6FAAF1D" authorityName="Verrill" authorityYear="1868" class="Anthozoa" family="Plexauridae" genus="Psammogorgia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammogorgia gracilis" order="Alcyonacea" pageId="0" pageNumber="1" phylum="Cnidaria" rank="species" species="gracilis">
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<emphasis id="C95B18F578B4760F5D42D36C3F727D24" italics="true" pageId="0" pageNumber="1">Psammogorgia gracilis</emphasis>
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</taxonomicName>
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Verrill, 1868. YPM 813a
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<emphasis id="7D407EAA1666A1F8B2C7F21818221DB1" bold="true" pageId="0" pageNumber="1">A</emphasis>
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colony
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<emphasis id="9EA85978787B814AFB25EA755BE339B5" bold="true" pageId="0" pageNumber="1">B</emphasis>
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detail of branches showing a tumor
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<emphasis id="36225012ED38CC6C08D511C48E27C184" bold="true" pageId="0" pageNumber="1">C</emphasis>
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coenenchymal and anthocodial sclerites.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="6CD04DEB99615C58F6B3A8301534196A" pageId="0" pageNumber="1" type="remarks">
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<paragraph id="F09BA1DEA45F4D7A7F550282A025BBCC" pageId="0" pageNumber="1">Remarks and comparison.</paragraph>
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<paragraph id="FBCB529129DBDB43681F567E0784127C" pageId="0" pageNumber="1">
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This species differs from the others by having long, slender and ascending branchlets, which are thinner than in the other species of the genus (Table
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<tableCitation id="5BC0AF03A700EDE9E5E05156D2242731" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparative features of Psammogorgia colonies from the tropical eastern Pacific, according to analyses of type material from museums (YPM, MCZ, MNHN), and taxonomic descriptions by Verrill (1868 b, 1870) and Bayer (1958). Diameter of the branches includes calyces. Measurements in millimetres." httpUri="http://table.plazi.org/id/B8D52D66E392F1F5E140A65F2133FE82" pageId="0" pageNumber="1" tableUuid="B8D52D66E392F1F5E140A65F2133FE82">1</tableCitation>
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).
|
||
<bibRefCitation id="182310769B93E5FE2E5464F1456B875A" author="Verrill, AE" journalOrPublisher="Transactions of the Connecticut Academy of Arts and Sciences, (Second Edition)" pageId="0" pageNumber="1" pagination="377 - 558" publicationUrl="https://biodiversitylibrary.org/page/13465394" refId="B41" refString="Verrill, AE, 1868b. Notes on Radiata in the Museum of Yale College, with descriptions of new genera and species. No. 6. Review of the corals and polyps of the West Coast of America. Transactions of the Connecticut Academy of Arts and Sciences, (Second Edition) 1 (2): 377 - 558, https://biodiversitylibrary.org/page/13465394" title="Notes on Radiata in the Museum of Yale College, with descriptions of new genera and species. No. 6. Review of the corals and polyps of the West Coast of America." url="https://biodiversitylibrary.org/page/13465394" volume="1" year="1868 b">Verrill (1868b)</bibRefCitation>
|
||
pointed out the abundance of wart-clubs in this species when compared to
|
||
<taxonomicName id="20D5031B62F0C376DDBC1A163DA6B494" lsidName="P. arbuscula" pageId="0" pageNumber="1" rank="species" species="arbuscula">
|
||
<emphasis id="820C8BF447243CC7B435D601DC6D36DE" italics="true" pageId="0" pageNumber="1">P. arbuscula</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="9B752928080441EA9B6F83D4B05B3691" lsidName="P. teres" pageId="0" pageNumber="1" rank="species" species="teres">
|
||
<emphasis id="C7D49912C7886415380BAE828E1BDC07" italics="true" pageId="0" pageNumber="1">P. teres</emphasis>
|
||
</taxonomicName>
|
||
. According to Verrill, the specimen he described was slender, flabelliform with subparallel and elongated branchlets, measuring12.7 cm long and 10.2 cm wide. The material left (YPM 813a) matches
|
||
<normalizedToken id="8AFF2FF0336585B7A1CE50DD0779E5F6" originalValue="Verrill´s">Verrill's</normalizedToken>
|
||
description and illustration. Therefore, we designate this specimen, YPM 813a, as the lectotype of
|
||
<taxonomicName id="06D53295E14F0C261F0CA7F6D8EDA57E" lsidName="P. gracilis" pageId="0" pageNumber="1" rank="species" species="gracilis">
|
||
<emphasis id="1A9BD8C2103E8ACC1BB7CEBFB738D1E7" italics="true" pageId="0" pageNumber="1">P. gracilis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="15A04BEAC9C9209F8CE542D501D2ACED" doi="10.3897/zookeys.961.54846.figure13" httpUri="https://binary.pensoft.net/fig/442686" pageId="0" pageNumber="1" start="Figure 13" startId="F13">
|
||
<paragraph id="A92BC386DC6CCBAEE83DAF7960623165" pageId="0" pageNumber="1">
|
||
<emphasis id="E8A12DDA851FF48C4419106A3C1AC4BA" bold="true" pageId="0" pageNumber="1">Figure 13.</emphasis>
|
||
<taxonomicName id="9538C574173D720185A059B31F7F4B7F" authorityName="Verrill" authorityYear="1868" class="Anthozoa" family="Plexauridae" genus="Psammogorgia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammogorgia gracilis" order="Alcyonacea" pageId="0" pageNumber="1" phylum="Cnidaria" rank="species" species="gracilis">
|
||
<emphasis id="55F5FED3B6A19CE9C556857C5FBF3A13" italics="true" pageId="0" pageNumber="1">Psammogorgia gracilis</emphasis>
|
||
</taxonomicName>
|
||
Verrill, 1868. YPM 813a
|
||
<emphasis id="17C1FB36AB511E1600D0FD62ACBA698D" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
spindles
|
||
<emphasis id="99201DD06C71B3B93AC965EADF729C0E" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
wart clubs
|
||
<emphasis id="38A125CAE1CF9E36F8E948A376F15760" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
radiates
|
||
<emphasis id="A4446F6FB777875DC38F2BFFE13DE184" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
anthocodial spindles.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="428BD6A6F247E20644FFE2C713578D02" pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph id="B6AD93A789F2C8A14666205168C8E993" pageId="0" pageNumber="1">Distribution.</paragraph>
|
||
<paragraph id="BFB3E4593867BD984771312515708873" pageId="0" pageNumber="1">
|
||
Tropical eastern Pacific: only recorded at the type locality Pearl Islands,
|
||
<normalizedToken id="C645E3DAB4DC45416AB900B765B26C50" originalValue="Panamá">Panama</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |