treatments-xml/data/7B/8D/6F/7B8D6F5AA0A85C0095BDA6EB9A0B4075.xml

<document ID-CLB-Dataset="20595" ID-DOI="http://dx.doi.org/10.3897/phytokeys.190.72740" ID-GBIF-Dataset="05a5cd96-b4e8-4790-aad7-ce23ecd5f576" ID-PMC="PMC8873181" ID-Pensoft-Pub="1314-2003-190-1" ID-Pensoft-UUID="EE4B5CE540D758DD83B5A745F4792F5D" ID-PubMed="35437379" ModsDocID="1314-2003-190-1" checkinTime="1645139658121" checkinUser="pensoft" docAuthor="Clark, John L. &amp; Clavijo, Laura" docDate="2022" docId="7B8D6F5AA0A85C0095BDA6EB9A0B4075" docLanguage="en" docName="PhytoKeys 190: 1-14" docOrigin="PhytoKeys 190" docPubDate="2022-02-17" docSource="http://dx.doi.org/10.3897/phytokeys.190.72740" docTitle="Drymonia longiflora J. L. Clark &amp; Clavijo 2022, sp. nov." docType="treatment" docVersion="7" id="EE4B5CE540D758DD83B5A745F4792F5D" lastPageNumber="1" masterDocId="EE4B5CE540D758DD83B5A745F4792F5D" masterDocTitle="Two new species of Drymonia (Gesneriaceae) from Northwestern South America, including the discovery of the longest flower known in the genus" masterLastPageNumber="14" masterPageNumber="1" pageNumber="1" updateTime="1720559167030" updateUser="ExternalLinkService">
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<mods:title id="B64A82A4B6021B9ADAEB1EE04E0C7B11">Two new species of Drymonia (Gesneriaceae) from Northwestern South America, including the discovery of the longest flower known in the genus</mods:title>
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<mods:namePart id="AB2EAC1C38421B31DF70325020CCC87D">Clark, John L.</mods:namePart>
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<mods:affiliation id="CC1544CDA62B7CE34BCF304973C5FCEC">Science Department, The Lawrenceville School, Lawrenceville, NJ 08648, US A</mods:affiliation>
<mods:nameIdentifier id="480B290C6C7D19A56E7DAFAE5ED17A68" type="email">phinaea@gmail.com</mods:nameIdentifier>
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<mods:namePart id="F027B70C1DCF10AC101928B486562B82">Clavijo, Laura</mods:namePart>
<mods:nameIdentifier id="FAA6961199E4800498FEA424EA05F534" type="ORCID">https://orcid.org/0000-0002-3009-9158</mods:nameIdentifier>
<mods:affiliation id="61FF2CDC10C1FF3439D61F9A9D9D3470">Universidad Nacional de Colombia - Sede Bogota, Facultad de Ciencias, Instituto de Ciencias Naturales, Bogota, D. C., 111321, Colombia</mods:affiliation>
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<mods:date id="239C6CA82D05DE7755025913F7F1E971">2022</mods:date>
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<mods:identifier id="106090BF527F8E7C300998766CEA70E4" type="DOI">http://dx.doi.org/10.3897/phytokeys.190.72740</mods:identifier>
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<subSection id="78B92A36EDC4898916ABD50CEAFF4397" pageId="0" pageNumber="1" type="multiple">
<treatment id="7B8D6F5AA0A85C0095BDA6EB9A0B4075" ID-GBIF-Taxon="193397204" LSID="urn:lsid:plazi:treatment:7B8D6F5AA0A85C0095BDA6EB9A0B4075" httpUri="http://treatment.plazi.org/id/7B8D6F5AA0A85C0095BDA6EB9A0B4075" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection id="20B18122D2BE8912869773E447F88EB3" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="24B3D116BE7A03286ADA1648B0149543" pageId="0" pageNumber="1">
<taxonomicName id="7AF28BCE5807AA1A247FB4DF30F586AD" ID-CoL="98XWD" LSID="7B8D6F5A-A0A8-5C00-95BD-A6EB9A0B4075" authority="J. L. Clark &amp; Clavijo" authorityName="J. L. Clark &amp; Clavijo" authorityYear="2022" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Drymonia longiflora" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="longiflora" status="sp. nov.">Drymonia longiflora J.L.Clark &amp; Clavijo</taxonomicName>
<taxonomicNameLabel id="EAC2F7976BFAC3689E042D53B1A3C2B8" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="0C56F101C8ED70AD2CFD9B926297FAC8" pageId="0" pageNumber="1" type="description">
<paragraph id="A9C01E31311E67C39D20492EC0237806" pageId="0" pageNumber="1">
<figureCitation id="3A4672A1DDE4FF2A35336DA552F2BA5D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Drymonia longiflora J. L. Clark &amp; Clavijo A erect subwoody habit B mature calyx lobes C front view of corolla D lateral view of flower (A from J. L. Clark et al. 7196 B from J. L. Clark et al. 13417 C, D from J. L. Clark et al. 10343). Photos by J. L. Clark." figureDoi="10.3897/phytokeys.190.72740.figure4" httpUri="https://binary.pensoft.net/fig/647436" pageId="0" pageNumber="1">Fig. 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="7EAAA312F91FB43A9DF42E0743756850" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="4BC360050B2C5EC1FAEA5EAC6E2A9A77" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="80AED3F4040CC9EB5641BA21B91CBE0A" pageId="0" pageNumber="1">
Differs from all 
<taxonomicName id="63A51FA2BD3E45B01928E00C3F695E15" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="Drymonia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="9FFC23BCE2C8D820A570F5DD0619415B" italics="true" pageId="0" pageNumber="1">Drymonia</emphasis>
</taxonomicName>
by an elongate corolla that exceeds 6.5 cm in length, the longest corolla known in the genus. The subshrub habit with elongate shoots and corolla shape are similar to 
<taxonomicName id="E88FC4CFD6032D90A737EE3678B0C599" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="Drymonia macrophylla" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="macrophylla">
<emphasis id="9C2E598CC576E0144612D4F6F46F1D4B" italics="true" pageId="0" pageNumber="1">Drymonia macrophylla</emphasis>
</taxonomicName>
and 
<emphasis id="F741BF3285C9AB223FFA9C158B53E927" italics="true" pageId="0" pageNumber="1">
<taxonomicName id="4465B1939CC36816113586BA4155B608" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="D. peponifera" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="peponifera">D. peponifera</taxonomicName>
. 
<taxonomicName id="859249B7139C97CB32567435E27ABE9E" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="Drymonia macrophylla" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="macrophylla">Drymonia macrophylla</taxonomicName>
</emphasis>
has a corolla that rarely exceeds 3.5 cm in length and 
<taxonomicName id="7C393497F20990D98DAFD0B3D1270C08" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="D. peponifera" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="peponifera">
<emphasis id="B09D641278556B7AEBA8225F60AA39B8" italics="true" pageId="0" pageNumber="1">D. peponifera</emphasis>
</taxonomicName>
has a corolla that rarely exceeds 4.5 cm in length.
</paragraph>
</subSubSection>
<subSubSection id="52E8497677F5CBEC86BDC2F67B4642F9" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="38F40E48202ABD872D04CA751ED31B12" pageId="0" pageNumber="1">
<typeStatus id="4AF6D4B18A9D23BEEA7888D8539B8987">Type</typeStatus>
.
</paragraph>
<paragraph id="F6855B7DB105A1874CBA7CAF194B6AB9" pageId="0" pageNumber="1">
<materialsCitation id="DF3B3845A5512C38D724592CF645575E" accessionNumber="III 10442" collectingDate="2008-05-29" collectorName="J. L. Clark, B. Bisvicuth, S. Ginzbarg, J. Melton" country="Ecuador" county="San Lorenzo" elevation="695" latitude="0.95527774" location="San Lorenzo-Ibarra" longLatPrecision="21" longitude="-78.558334" municipality="Alto Tambo" specimenCount="1" stateProvince="Esmeraldas" typeStatus="holotype">
<collectingCountry id="3043DD7D0EB52B4A77B49BA5AC63D9B5" name="Ecuador">Ecuador</collectingCountry>
. 
<collectingRegion id="F9A99EF955B3537283634E8E3A186F78" country="Ecuador" name="Esmeraldas">Esmeraldas</collectingRegion>
: 
<normalizedToken id="1D1AC556213C08513F0CFEC7DF910954" originalValue="cantón">canton</normalizedToken>
<collectingCounty id="1E5D0567ECE3FD10D0A2BDF42565103B">San Lorenzo</collectingCounty>
, parroquia 
<collectingMunicipality id="B53A50F43B4DFC1FE2C3C01867C6D13D">Alto Tambo</collectingMunicipality>
, remnant patch of primary forest on north side of road 
<locationDeviation id="D2A0EAD87D4D4EDA68A43C25D5010CCD" firstLocation="Durango" secondLocation="Alto Tambo">between Durango and Alto Tambo</locationDeviation>
on highway 
<location id="01F3419B00C2D222D5B3C34EE4E721D8" LSID="urn:lsid:plazi:treatment:7B8D6F5AA0A85C0095BDA6EB9A0B4075:01F3419B00C2D222D5B3C34EE4E721D8" country="Ecuador" county="San Lorenzo" latitude="0.95527774" longLatPrecision="21" longitude="-78.558334" municipality="Alto Tambo" name="San Lorenzo-Ibarra" stateProvince="Esmeraldas">San Lorenzo-Ibarra</location>
, 
<geoCoordinate id="0842B61648371DA11CA5E17384BE5084" degrees="0" direction="north" minutes="57" orientation="latitude" precision="15" seconds="19" value="0.95527774">0°57'19&quot;N</geoCoordinate>
, 
<geoCoordinate id="9EB27FB44CCB2F5401B4F597D8F69600" degrees="78" direction="west" minutes="33" orientation="longitude" precision="15" seconds="30" value="-78.558334">78°33'30&quot;W</geoCoordinate>
, 
<quantity id="30079EA11F0E695A969963054B6C329E" metricMagnitude="2" metricUnit="m" metricValue="6.95" unit="m" value="695.0">
<elevation id="D8A45E665FB950FBD68C8E0CFBD5117D" metricMagnitude="2" metricUnit="m" metricValue="6.95" unit="m" value="695.0">695 m</elevation>
</quantity>
, 
<collectingDate id="BC2774662505D0EF45E9A18E6F6E16B5" value="2008-05-29">29 May 2008</collectingDate>
, 
<emphasis id="B53540F74A6DFD8E1B7C2E562170FCA4" italics="true" pageId="0" pageNumber="1">
<collectorName id="5EF72AADC2F94D3761DA2128A74E64EE">J.L. Clark</collectorName>
</emphasis>
, 
<emphasis id="AA500182C99C74D9CF2373D779C99FB9" italics="true" pageId="0" pageNumber="1">
<collectorName id="9F58B07D6AA26A1A1F37A568984B5B24">B. Bisvicuth</collectorName>
</emphasis>
, 
<emphasis id="BFF22451A66E69AA23543327308CFBF4" italics="true" pageId="0" pageNumber="1">
<collectorName id="040DCAD78541E90F7B93D65153C43A5C">S. Ginzbarg</collectorName>
&amp; 
<collectorName id="7D9249AAD376DE1A70C431FEE4917F30">J. Melton</collectorName>
<accessionNumber id="A157CCF4DF0D8A139976105821A024DB" httpUri="http://www.ncbi.nlm.nih.gov/protein/III10442">III 10442</accessionNumber>
</emphasis>
(
<typeStatus id="599B78CEE1D486CE607D2C95138A1630">holotype</typeStatus>
:
</materialsCitation>
<collectingCountry id="933BB29364A746DE256FCFC33F3E2D0F" name="United States of America">US</collectingCountry>
; isotypes: COL, E, G, MO, NY, QCNE, SEL).
</paragraph>
</subSubSection>
<subSubSection id="662F079655F1DFAB7289147109E4CB39" pageId="0" pageNumber="1" type="description">
<paragraph id="B85D0B1B7BEC643FD712F7A62F6A84B5" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="B8E38B8A6003E5F8A0F7699766CF4C43" pageId="0" pageNumber="1">
Terrestrial, hemiepiphytic or epiphytic herb or subshrub, with scandent to horizontal shoots to 1 m long. Stems subquadrangular in cross-section, 0.3-0.6 cm in diameter, strigose to glabrate, internodes 1.0-8 cm long. 
<emphasis id="D5F48D3430FF518A0502136184D7FDFB" bold="true" pageId="0" pageNumber="1">Leaves</emphasis>
opposite, decussate, usually evenly spaced and becoming clustered near apex, subequal to unequal in a pair; petioles 0.7-3.0 cm long, basal enations present, strigillose to strigose, terete in cross-section; blade elliptic to ovate, 8.0-28 
<normalizedToken id="F394071A40E214F2D59F7F90CD7E3C01" originalValue="×">x</normalizedToken>
3.5-13 cm, membranous to subcoriaceous, the base rounded to acute, sometimes oblique, the apex acute to long-acuminate, the margin serrulate, the upper surface glossy-green, glabrescent or strigose, the lower surface light green with red venation, glabrescent or strigillose, lateral secondary veins 5-7 pairs, prominent abaxially, strigose abaxially. 
<emphasis id="C99A5CD61B0246DD8C5B09666B6EF816" bold="true" pageId="0" pageNumber="1">Inflorescence</emphasis>
reduced to a single flower in the upper or lower leaf axils; peduncle absent; bracts small, lanceolate, 3.0-7.0 
<normalizedToken id="7C76B648DEC5AE21BB979C2E13A7D780" originalValue="×">x</normalizedToken>
1.1-1.3 mm, green, lanceolate, strigose; pedicels short, 3-7 mm long, green, strigose. 
<emphasis id="03CBF4324BB232106CB09A271C0D92B6" bold="true" pageId="0" pageNumber="1">Flowers</emphasis>
with calyx green, dark red with green margins, or uniformly dark red, with enations at base; lobes 5, nearly free, 4 equal, the upper lobe slightly smaller and more narrow, membranous, lanceolate with an elongate acute apex, the margins deeply serrate to slightly laciniate, lobes covered with sparsely pilose trichomes, 1.5-2.7 
<normalizedToken id="63F497A93766FDA4524674CDA54697E5" originalValue="×">x</normalizedToken>
0.3-0.5 cm. Corolla zygomorphic, tubular, elongate, 6.5-8.0 cm long, with slight spur (nectar chamber) at base, 0.5-0.7 cm long; the corolla tube perpendicular relative to calyx, ampliate toward the limb, not contracted above, 5.0-6.5 cm long, 0.5-1.1 cm wide at the mid portion, uniformly pale yellow and sericeous outside, usually darker yellow inside; throat to 1.4 cm in diameter, the inside yellow, sometimes with brown spotting on lower portion, sparsely pilose or glandular; limb with 5 spreading lobes, subequal, globose, rounded at apex, margins erose, glabrous abaxially and adaxially, lobes 5-10 
<normalizedToken id="6FD123CA35D5E807AB29443518D5CA6E" originalValue="×">x</normalizedToken>
4-11 mm, the lower lobe slightly larger. Androecium of 4 didynamous stamens, included; filaments 5-6 cm long, adnate to the base of the corolla tube for 2-3 mm, glabrous, coiled after anthesis; the anthers at first coherent, after anthesis separating, dehiscent by basal pores, 3-5 
<normalizedToken id="3988D7704787353A0F16C08AB8441EF0" originalValue="×">x</normalizedToken>
2-1.8 mm. Gynoecium with a single dorsal nectary gland, thick, ovate, 2-3 mm long, glabrous; the ovary superior, sericeous, green; style included, 5-6 cm long, white, puberulent; stigma stomatomorphic. 
<emphasis id="31791F7C9D8AF58958DB20B9A9C2EA38" bold="true" pageId="0" pageNumber="1">Fruit</emphasis>
and seeds not observed.
</paragraph>
</subSubSection>
<subSubSection id="1FCB37F0C9E41B6A8A6E69B72087BE14" pageId="0" pageNumber="1" type="phenology">
<paragraph id="2EF8C713AF78360EBCA36CE4C9C7AB87" pageId="0" pageNumber="1">Phenology.</paragraph>
<paragraph id="C4718D62714CB05A919D78D99031F879" pageId="0" pageNumber="1">This species has been found with flowers in two periods: February to May and August to October.</paragraph>
</subSubSection>
<subSubSection id="B0493692F063D00862EC7177A73D785C" pageId="0" pageNumber="1" type="etymology">
<paragraph id="46167C8B68A0153FCEC06CF90505D209" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="FDBA99C3C7AE74166B4FCEE5AF5392BE" pageId="0" pageNumber="1">
The specific epithet is in reference to the elongate corolla tube, the longest of any known species of 
<taxonomicName id="0D254D73CF4EB1249E5ADF293F36CDAD" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="Drymonia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="CCB8B2B3AADF1DD7EA48FB28570707CD" italics="true" pageId="0" pageNumber="1">Drymonia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C7606B4EAA2816903068FD580485BE7F" pageId="0" pageNumber="1" type="distribution">
<paragraph id="95715E14945FA1C1E858469627825D3E" pageId="0" pageNumber="1">Distribution and preliminary conservation assessment.</paragraph>
<paragraph id="C8AD4E79771760182BBD1D2DA0A64155" pageId="0" pageNumber="1">
<taxonomicName id="6536A5D5519AC8ACDAE7978710023D9D" authorityName="J. L. Clark &amp; Clavijo" authorityYear="2022" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="Drymonia longiflora" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="longiflora">
<emphasis id="B5097AD266BF73D3BF9C69D667920D99" italics="true" pageId="0" pageNumber="1">Drymonia longiflora</emphasis>
</taxonomicName>
is locally abundant in forests along the western slopes of the northern Andes in Ecuador (Provinces Carchi and Esmeraldas) and Colombia (Departments 
<normalizedToken id="19FCCC3254B75228989DE29031472862" originalValue="Chocó">Choco</normalizedToken>
, Valle del Cauca, and 
<normalizedToken id="D40A3ED754FEF4E5942447AFD7AB95BA" originalValue="Nariño">Narino</normalizedToken>
). One disjunct population is documented from southern Ecuador (Province Morona-Santiago). 
<taxonomicName id="D6FC44FBB7E9871048879E832FBF7C72" authorityName="J. L. Clark &amp; Clavijo" authorityYear="2022" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="Drymonia longiflora" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="longiflora">
<emphasis id="3EC6B58B1290B577A612AADAECA10E60" italics="true" pageId="0" pageNumber="1">Drymonia longiflora</emphasis>
</taxonomicName>
grows in mature shaded forests, from 150 to 900 m in elevation. We provisionally assess this new species as Vulnerable (VU), according to the IUCN Red List criteria (
<bibRefCitation id="CD0A064C4F93BB125F68AEF7DF3FDED1" author="IUCN" journalOrPublisher="Rheedea" pageId="0" pageNumber="1" refId="B4" refString="IUCN, 2022. Guidelines for Using the IUCN Red List Categories and Criteria. Version 15. Prepared by the Standards and Petitions Committee. [accessed 20.01.2022]" title="Guidelines for Using the IUCN Red List Categories and Criteria. Version 15. Prepared by the Standards and Petitions Committee. [accessed 20.01.2022]" year="2022">IUCN 2022</bibRefCitation>
) for limited geographic range (EOO &lt;20,000 km2 and AOO &lt;2,000 km2) and associated subcriteria, including occurrence at fewer than ten locations (B2a) and continuing decline of Andean forests (B2b).
</paragraph>
<caption id="9A85C4882C0118BACD4FE0CF09822DD9" doi="10.3897/phytokeys.190.72740.figure3" httpUri="https://binary.pensoft.net/fig/647435" pageId="0" pageNumber="1" start="Figure 3" startId="F3">
<paragraph id="62A240B2CA2C50AC8E93828FCB28E95C" pageId="0" pageNumber="1">
<emphasis id="2EB8079A261C34D6A33D1E868CA6907D" bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
<taxonomicName id="74F0250285AE4C98F3BD2D87FE3E5E33" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="Drymonia macrophylla" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="macrophylla">
<emphasis id="E9F704594A41A22FEACE1CC0894F1EAD" italics="true" pageId="0" pageNumber="1">Drymonia macrophylla</emphasis>
</taxonomicName>
(Oerst.) H.E.Moore 
<emphasis id="D2BD6B6984EEA2ABAA5D1B40700C1291" bold="true" pageId="0" pageNumber="1">A</emphasis>
erect subwoody habit 
<emphasis id="C33F56D6BDF0E4EB3AEE8A4D7DBBCEC4" bold="true" pageId="0" pageNumber="1">B</emphasis>
mature fleshy bivalved capsule 
<emphasis id="D65DF4B942535576361FCE9F642A9BA4" bold="true" pageId="0" pageNumber="1">C</emphasis>
front view of corolla 
<emphasis id="4D2ECFED62AFEDE590E9A561C72D0A4D" bold="true" pageId="0" pageNumber="1">D</emphasis>
lateral view of flower (
<emphasis id="BE058638309225505EAA9B1B36EB4A38" bold="true" pageId="0" pageNumber="1">A</emphasis>
from 
<emphasis id="2E2985C40BAD5698BF1909313434BC4E" italics="true" pageId="0" pageNumber="1">P. Pedraza et al. 850</emphasis>
<emphasis id="65207B516BBCC5BFD4C466AABC69F978" bold="true" pageId="0" pageNumber="1">B</emphasis>
from 
<emphasis id="757F2326B162AE7F4D1974FC494FFE6C" italics="true" pageId="0" pageNumber="1">J.L. Clark et al. 12119</emphasis>
<emphasis id="436AA6090E832F6E535A7CBA9B7DEBAD" bold="true" pageId="0" pageNumber="1">C, D</emphasis>
from 
<emphasis id="E53ACAD764D93646E00AA15B183BEDF8" italics="true" pageId="0" pageNumber="1">J.L. Clark et al. 10044</emphasis>
). Photo 
<emphasis id="16FE0C6C4FC6CB3736974E75FDFB709F" bold="true" pageId="0" pageNumber="1">A</emphasis>
by P. Pedraza and photos 
<emphasis id="9DC613026CAB3A540B6EE42334C7B07B" bold="true" pageId="0" pageNumber="1">B-D</emphasis>
by J.L. Clark.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="63E327375B3DA3743CB3137A0A522388" pageId="0" pageNumber="1" type="comments">
<paragraph id="4BA3F459DDA46A2B403BCF38BB80EE4F" pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph id="321C56B1F656D8AA699450124793CD9E" pageId="0" pageNumber="1">
<taxonomicName id="B33D82BB4222D68C462B85CF4CD93A2C" authorityName="J. L. Clark &amp; Clavijo" authorityYear="2022" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="Drymonia longiflora" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="longiflora">
<emphasis id="601801E8E00809A2F3396F59AB517055" italics="true" pageId="0" pageNumber="1">Drymonia longiflora</emphasis>
</taxonomicName>
is distinguished by the longest known corolla recorded for 
<taxonomicName id="2286ECB55A3AFB33D1FF8A73D043CDF8" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="Drymonia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="831C258CBB9AD05E2B10A867D35B56D1" italics="true" pageId="0" pageNumber="1">Drymonia</emphasis>
</taxonomicName>
(&gt; 6.5 cm). The unbranched subwoody epiphytic or terrestrial subshrub habit and the foliage are similar to 
<taxonomicName id="F139B33C6BA18C8CD1259D3DE5A79FBA" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="Drymonia macrophylla" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="macrophylla">
<emphasis id="B32E81BAB0A856C29F5AFB076AFA054E" italics="true" pageId="0" pageNumber="1">Drymonia macrophylla</emphasis>
</taxonomicName>
, 
<taxonomicName id="30CCF9265900C9988494FBE3B3EC88C9" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="D. fimbriata" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="fimbriata">
<emphasis id="DDCCBC4937009164ABAB9A7722ADAE48" italics="true" pageId="0" pageNumber="1">D. fimbriata</emphasis>
</taxonomicName>
, and 
<taxonomicName id="DD54E4A892C198B6FFBA7361287835A2" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="D. peponifera" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="peponifera">
<emphasis id="33DD74AFA2913132FF5ECE42AFF28220" italics="true" pageId="0" pageNumber="1">D. peponifera</emphasis>
</taxonomicName>
. All four species are vegetatively similar, but readily differentiated by corolla tube length and calyx lobes (Fig. 
<figureCitation id="DE93894B1696E8FBBF6B6D6907477BF6" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Drymonia peponifera J. L. Clark &amp; Clavijo A habit B upper view of mature fruit C lateral view of immature calyx lobes D lateral view of axillary inflorescence with a single mature flower (A-D from J. L. Clark &amp; L. Jost 6957). Photos by J. L. Clark." figureDoi="10.3897/phytokeys.190.72740.figure5" httpUri="https://binary.pensoft.net/fig/647437" pageId="0" pageNumber="1">5C, D</figureCitation>
). The calyx lobes of 
<taxonomicName id="D18AC0B669BB67703E5D391107AE0350" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="Drymonia peponifera" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="peponifera">
<emphasis id="A1AC75926094BE324381D1A73ED418C8" italics="true" pageId="0" pageNumber="1">Drymonia peponifera</emphasis>
</taxonomicName>
appear crispate from the reflexed and undulating lobes (Fig. 
<figureCitation id="33359E793E5493D4D3B65E7CF56561A0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Drymonia peponifera J. L. Clark &amp; Clavijo A habit B upper view of mature fruit C lateral view of immature calyx lobes D lateral view of axillary inflorescence with a single mature flower (A-D from J. L. Clark &amp; L. Jost 6957). Photos by J. L. Clark." figureDoi="10.3897/phytokeys.190.72740.figure5" httpUri="https://binary.pensoft.net/fig/647437" pageId="0" pageNumber="1">5C, D</figureCitation>
). In contrast, the calyx lobes of 
<taxonomicName id="8498A4325BC9AF78BB72E30B357BC762" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="D. macrophylla" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="macrophylla">
<emphasis id="59F0D755945F77E0144706E948CA34B3" italics="true" pageId="0" pageNumber="1">D. macrophylla</emphasis>
</taxonomicName>
(Fig. 
<figureCitation id="D62DA3C207A4F8683E03AC3098519A58" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Drymonia macrophylla (Oerst.) H. E. Moore A erect subwoody habit B mature fleshy bivalved capsule C front view of corolla D lateral view of flower (A from P. Pedraza et al. 850 B from J. L. Clark et al. 12119 C, D from J. L. Clark et al. 10044). Photo A by P. Pedraza and photos B-D by J. L. Clark." figureDoi="10.3897/phytokeys.190.72740.figure3" httpUri="https://binary.pensoft.net/fig/647435" pageId="0" pageNumber="1">3D</figureCitation>
) and 
<taxonomicName id="44CC8B49B6EC03A7C302B9D64362D480" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="D. longiflora" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="longiflora">
<emphasis id="7355D3D2794E2A7D78926181DFA886CC" italics="true" pageId="0" pageNumber="1">D. longiflora</emphasis>
</taxonomicName>
(Fig. 
<figureCitation id="B7D360DBEA112C2F5B00EF7D249196F9" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Drymonia longiflora J. L. Clark &amp; Clavijo A erect subwoody habit B mature calyx lobes C front view of corolla D lateral view of flower (A from J. L. Clark et al. 7196 B from J. L. Clark et al. 13417 C, D from J. L. Clark et al. 10343). Photos by J. L. Clark." figureDoi="10.3897/phytokeys.190.72740.figure4" httpUri="https://binary.pensoft.net/fig/647436" pageId="0" pageNumber="1">4D</figureCitation>
) are flat. The margins of the calyx lobes in 
<taxonomicName id="2204AE2A39BBD6BAEEE82E9603621A93" authorityName="J. L. Clark &amp; Clavijo" authorityYear="2022" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="Drymonia longiflora" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="longiflora">
<emphasis id="300E664A41D31B428076481B2DB1CF6C" italics="true" pageId="0" pageNumber="1">Drymonia longiflora</emphasis>
</taxonomicName>
are serrate to slightly laciniate, in contrast to the strongly laciniate with unbranched filiform teeth of 
<taxonomicName id="60C929E59BB4265407B98DDAE4FBB673" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="D. fimbriata" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="fimbriata">
<emphasis id="F97CCB12DC24DB9D2FAF80C478BD05F9" italics="true" pageId="0" pageNumber="1">D. fimbriata</emphasis>
</taxonomicName>
. The corolla in 
<taxonomicName id="3396ABDEA00E1943906FC1EAA8C21D0D" authorityName="J. L. Clark &amp; Clavijo" authorityYear="2022" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="Drymonia longiflora" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="longiflora">
<emphasis id="9956175A1D9DB485E0F59B12B3665093" italics="true" pageId="0" pageNumber="1">Drymonia longiflora</emphasis>
</taxonomicName>
exceeds 6.5 cm in length (Fig. 
<figureCitation id="53296D8E2EFD364C7D34D7A215F082E8" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Drymonia longiflora J. L. Clark &amp; Clavijo A erect subwoody habit B mature calyx lobes C front view of corolla D lateral view of flower (A from J. L. Clark et al. 7196 B from J. L. Clark et al. 13417 C, D from J. L. Clark et al. 10343). Photos by J. L. Clark." figureDoi="10.3897/phytokeys.190.72740.figure4" httpUri="https://binary.pensoft.net/fig/647436" pageId="0" pageNumber="1">4D</figureCitation>
). In contrast, the corolla rarely exceeds 3.5 cm in 
<taxonomicName id="353FAAE5C211103E52B5D72B0530176B" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="D. macrophylla" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="macrophylla">
<emphasis id="6223153F9546B37CC8DEBAB3A88A0873" italics="true" pageId="0" pageNumber="1">D. macrophylla</emphasis>
</taxonomicName>
(Fig. 
<figureCitation id="B90E47F45A8F8AE2F5F70A5B18707D68" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Drymonia macrophylla (Oerst.) H. E. Moore A erect subwoody habit B mature fleshy bivalved capsule C front view of corolla D lateral view of flower (A from P. Pedraza et al. 850 B from J. L. Clark et al. 12119 C, D from J. L. Clark et al. 10044). Photo A by P. Pedraza and photos B-D by J. L. Clark." figureDoi="10.3897/phytokeys.190.72740.figure3" httpUri="https://binary.pensoft.net/fig/647435" pageId="0" pageNumber="1">3D</figureCitation>
) and 
<taxonomicName id="0CE4151DCC33FE89879EC27281ECC462" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="D. fimbriata" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="fimbriata">
<emphasis id="B578E8CE068754A64DFDF5E3337BDC24" italics="true" pageId="0" pageNumber="1">D. fimbriata</emphasis>
</taxonomicName>
. The corolla is less than 4.5 cm in 
<taxonomicName id="71BDAB3A3E828B59FF67C85F9C35C5B6" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="D. peponifera" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="peponifera">
<emphasis id="274F95BA25248B9A190598361BDF448E" italics="true" pageId="0" pageNumber="1">D. peponifera</emphasis>
</taxonomicName>
(Fig. 
<figureCitation id="CE5198A173B3AB5BD7A95ED5E2DF74E8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Drymonia peponifera J. L. Clark &amp; Clavijo A habit B upper view of mature fruit C lateral view of immature calyx lobes D lateral view of axillary inflorescence with a single mature flower (A-D from J. L. Clark &amp; L. Jost 6957). Photos by J. L. Clark." figureDoi="10.3897/phytokeys.190.72740.figure5" httpUri="https://binary.pensoft.net/fig/647437" pageId="0" pageNumber="1">5D</figureCitation>
). 
<taxonomicName id="3A770C82FF3F8078F42B75AE5ECF99D1" authorityName="J. L. Clark &amp; Clavijo" authorityYear="2022" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="Drymonia longiflora" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="longiflora">
<emphasis id="43E73C081DF3FE5ABB597C073967B459" italics="true" pageId="0" pageNumber="1">Drymonia longiflora</emphasis>
</taxonomicName>
and 
<taxonomicName id="2FD92FAF57CDDCBEB9F910F994AD90E3" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="D. intermedia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="intermedia">
<emphasis id="0BE5BFC08BE9588556EC659D848B3F62" italics="true" pageId="0" pageNumber="1">D. intermedia</emphasis>
</taxonomicName>
share similar shapes for corolla tubes and calyx lobes, but differ in size, with 
<taxonomicName id="552A3E17BDDE526E376CDA0A64EFF0E1" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="D. intermedia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="intermedia">
<emphasis id="85654712BD2BA4B2A40A750BE73142E0" italics="true" pageId="0" pageNumber="1">D. intermedia</emphasis>
</taxonomicName>
much smaller. A summary of diagnostic characters is provided in Table 
<tableCitation id="D6ED9DEBC5E1A0796BE0C75C538AE452" captionStart="Table 1" captionStartId="T1" captionText="Table 1. General geographic distribution (names in parentheses indicate provinces for Ecuador or departments for Colombia) and comparison of morphological characters to differentiate Drymonia fimbriata, D. intermedia, D. laciniosa, D. longiflora, D. macrophylla, and D. peponifera." httpUri="http://table.plazi.org/id/CE026658719071926293EA1436B853EB" pageId="0" pageNumber="1" tableUuid="CE026658719071926293EA1436B853EB">1</tableCitation>
.
</paragraph>
<caption id="36645CBCAF4DE338E454C1DCBE900EF6" doi="10.3897/phytokeys.190.72740.figure4" httpUri="https://binary.pensoft.net/fig/647436" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
<paragraph id="1BAC8419804FC053CFD095A49464ACA1" pageId="0" pageNumber="1">
<emphasis id="6B8DF77CC5D9096844E801BC439A676F" bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
<taxonomicName id="8B7F9F1B1D0DD29233A3133500525C0D" authorityName="J. L. Clark &amp; Clavijo" authorityYear="2022" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="Drymonia longiflora" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="longiflora">
<emphasis id="8A911B3D8E97DA0800B06926478F7424" italics="true" pageId="0" pageNumber="1">Drymonia longiflora</emphasis>
</taxonomicName>
J.L.Clark &amp; Clavijo 
<emphasis id="C7A61DCE7241C2FB20A595F30922942A" bold="true" pageId="0" pageNumber="1">A</emphasis>
erect subwoody habit 
<emphasis id="41970989DD53D6BD0BA82F837CD9F7D8" bold="true" pageId="0" pageNumber="1">B</emphasis>
mature calyx lobes 
<emphasis id="D9B025DCCA3891A56DE28F8A35F90D2C" bold="true" pageId="0" pageNumber="1">C</emphasis>
front view of corolla 
<emphasis id="C92C4F23D65F610CBA4387173D3C9CBD" bold="true" pageId="0" pageNumber="1">D</emphasis>
lateral view of flower (
<emphasis id="4D6FB5FF053A383CA99FF4AC9B2F2314" bold="true" pageId="0" pageNumber="1">A</emphasis>
from 
<emphasis id="4EB073EBBAEDA4EE94F574558C51FA3D" italics="true" pageId="0" pageNumber="1">J.L. Clark et al. 7196</emphasis>
<emphasis id="02F3A261C2CC72BCD3F94786B9DDCA50" bold="true" pageId="0" pageNumber="1">B</emphasis>
from 
<emphasis id="6D6D55C24A04A8DE9153609225ED27EE" italics="true" pageId="0" pageNumber="1">J.L. Clark et al. 13417</emphasis>
<emphasis id="AED2FC958547FED79E0D5102EBCF6CEF" bold="true" pageId="0" pageNumber="1">C, D</emphasis>
from 
<emphasis id="77396AE40E791DE5AE09D51F6FB364B8" italics="true" pageId="0" pageNumber="1">J.L. Clark et al. 10343</emphasis>
). Photos by J.L. Clark.
</paragraph>
</caption>
<paragraph id="8E8E48C3E59726440F95DA631B135AF0" pageId="0" pageNumber="1">
Our preliminary DNA sequence data strongly support a close relationship in a clade that includes 
<taxonomicName id="4F0B9687729BB5D9FAB851DBBB6A209A" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="D. intermedia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="intermedia">
<emphasis id="54BDB2496663F8B7606D29C14D747891" italics="true" pageId="0" pageNumber="1">D. intermedia</emphasis>
</taxonomicName>
, 
<taxonomicName id="9F5CE5C3D1D6BD25DD9AACB8BCD88A41" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="D. laciniosa" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="laciniosa">
<emphasis id="9037224ECBC5FFAB3ADB0CC5DF1B19B1" italics="true" pageId="0" pageNumber="1">D. laciniosa</emphasis>
</taxonomicName>
, 
<taxonomicName id="1DBF66FC970A91527DAD561F6E8492DC" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="D. longiflora" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="longiflora">
<emphasis id="34E93783C447E492CC5E7BF59B1EBDC6" italics="true" pageId="0" pageNumber="1">D. longiflora</emphasis>
</taxonomicName>
, 
<taxonomicName id="949CBC71B9B81C8373BE2F0A2C8D049F" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="D. macrophylla" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="macrophylla">
<emphasis id="5F463B3B21A2FDA1E1BE84048E8F95F6" italics="true" pageId="0" pageNumber="1">D. macrophylla</emphasis>
</taxonomicName>
, and 
<taxonomicName id="1B73E5BA71DA7C43D43FAF4CFBE9947F" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="D. peponifera" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="peponifera">
<emphasis id="D79B9EF723EB5D06A2597B316436CE4B" italics="true" pageId="0" pageNumber="1">D. peponifera</emphasis>
</taxonomicName>
. This clade is defined by fleshy capsular fruits with tardily dehiscent endocarps. A photographic guide and description of fruit types is summarized in 
<bibRefCitation id="D78079EEE953B281FAB98D69CE71AEEF" DOI="https://doi.org/10.1007/s12228-021-09688-3" author="Clark, JL" journalOrPublisher="Evolutionary Ecology" pageId="0" pageNumber="1" refId="B2" refString="Clark, JL, Clavijo, L, 2022. Drymonia peponifera, a new species of Gesneriaceae from Ecuador with an overview of Drymonia fruit traits. Brittonia. https://doi.org/10.1007/s12228-021-09688-3" title="Drymonia peponifera, a new species of Gesneriaceae from Ecuador with an overview of Drymonia fruit traits. Brittonia." url="https://doi.org/10.1007/s12228-021-09688-3" year="2022">Clark and Clavijo (2022)</bibRefCitation>
. Most 
<taxonomicName id="17DA40D50618674BF840B5BBEC1A3B12" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="Drymonia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="589724901DDDBC5C9A1B0CFD8E8730A4" italics="true" pageId="0" pageNumber="1">Drymonia</emphasis>
</taxonomicName>
capsules have endocarps that dehisce when mature. In contrast, the clade that includes 
<taxonomicName id="83D90E9651A806B3E28360E3007113E6" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="D. intermedia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="intermedia">
<emphasis id="70479399E6EE01FFB825A5778A1AE265" italics="true" pageId="0" pageNumber="1">D. intermedia</emphasis>
</taxonomicName>
and 
<taxonomicName id="AA3E4C288E9BC82179C447DECDED1E48" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="D. longiflora" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="longiflora">
<emphasis id="12727B3BEA6FAF31D532332D7A6760C3" italics="true" pageId="0" pageNumber="1">D. longiflora</emphasis>
</taxonomicName>
is defined by endocarps that remain attached and surround the placentae and mass of funiculi and seeds (Figs 
<figureCitation id="B6222C6C24ED5E2977544E8D5DCC7FED" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Drymonia intermedia Clavijo &amp; J. L. Clark A holotype specimen B upper view of mature fruit C front view of corolla D lateral view of flower (A holotype (SEL) of J. L. Clark et al. 10442 B from J. L. Clark et al. 7148 C, D from J. L. Clark et al. 10344). Photos by J. L. Clark." figureDoi="10.3897/phytokeys.190.72740.figure1" httpUri="https://binary.pensoft.net/fig/647433" pageId="0" pageNumber="1">1B</figureCitation>
, 
<figureCitation id="B2B0841D24934A4F39198150829EF270" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Drymonia peponifera J. L. Clark &amp; Clavijo A habit B upper view of mature fruit C lateral view of immature calyx lobes D lateral view of axillary inflorescence with a single mature flower (A-D from J. L. Clark &amp; L. Jost 6957). Photos by J. L. Clark." figureDoi="10.3897/phytokeys.190.72740.figure5" httpUri="https://binary.pensoft.net/fig/647437" pageId="0" pageNumber="1">5B</figureCitation>
). The endocarp eventually becomes dehiscent at a later stage when it detaches from the reflexed outer layers of the fruit wall (Fig. 
<figureCitation id="A7C293B692CE3CD215218B0CFF3679B3" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Drymonia peponifera J. L. Clark &amp; Clavijo A habit B upper view of mature fruit C lateral view of immature calyx lobes D lateral view of axillary inflorescence with a single mature flower (A-D from J. L. Clark &amp; L. Jost 6957). Photos by J. L. Clark." figureDoi="10.3897/phytokeys.190.72740.figure5" httpUri="https://binary.pensoft.net/fig/647437" pageId="0" pageNumber="1">5B</figureCitation>
). Although we have not observed mature fruits of 
<taxonomicName id="248BC0A4BEA828F41D75B7F19BDDA8CF" authorityName="J. L. Clark &amp; Clavijo" authorityYear="2022" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="Drymonia longiflora" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="longiflora">
<emphasis id="B1AC83358CCC3C25B143405356E83B3C" italics="true" pageId="0" pageNumber="1">Drymonia longiflora</emphasis>
</taxonomicName>
, we predict that the fruit type of this species shares the same characters with closely related congeners.
</paragraph>
<caption id="E266ADC373C4AA317F80353AAC2F3194" doi="10.3897/phytokeys.190.72740.figure5" httpUri="https://binary.pensoft.net/fig/647437" pageId="0" pageNumber="1" start="Figure 5" startId="F5">
<paragraph id="842E0442A9D51CC991529967580A937E" pageId="0" pageNumber="1">
<emphasis id="AA0238D10885D5B1BF76372A46DC63B8" bold="true" pageId="0" pageNumber="1">Figure 5.</emphasis>
<taxonomicName id="6722ED2C6EFF133FD82C8C681AE0BC40" class="Magnoliopsida" family="Gesneriaceae" genus="Drymonia" kingdom="Plantae" lsidName="Drymonia peponifera" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="peponifera">
<emphasis id="D9F17F467DFF546F9CD1A1E4F596E497" italics="true" pageId="0" pageNumber="1">Drymonia peponifera</emphasis>
</taxonomicName>
J.L.Clark &amp; Clavijo 
<emphasis id="B42A0862E30241197E0884FF8546CBD7" bold="true" pageId="0" pageNumber="1">A</emphasis>
habit 
<emphasis id="A67EB10F45BC21C5DB8EBCB59DF8F4B7" bold="true" pageId="0" pageNumber="1">B</emphasis>
upper view of mature fruit 
<emphasis id="EE918A7FEFCBD007888A58890A487E1D" bold="true" pageId="0" pageNumber="1">C</emphasis>
lateral view of immature calyx lobes 
<emphasis id="D867586C1B6AD7999E1DF67463C28CD9" bold="true" pageId="0" pageNumber="1">D</emphasis>
lateral view of axillary inflorescence with a single mature flower (
<emphasis id="C8D50999A64BC060323FEEE2D17A7BE3" bold="true" pageId="0" pageNumber="1">A-D</emphasis>
from 
<emphasis id="D99BFB76FDE12BA066B5BBEC7CB3313F" italics="true" pageId="0" pageNumber="1">J.L. Clark &amp; L. Jost 6957</emphasis>
). Photos by J.L. Clark.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="FBE91CC4135C322E469A025F9338428B" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="E3B9A5B70C04463C2A1591D2D6657B5C" pageId="0" pageNumber="1">Specimens examined.</paragraph>
<paragraph id="1AACA3687912D496F8036C17EFF3A503" pageId="0" pageNumber="1">
<emphasis id="FD6AC3A3802CC27105ACC7D307F0245A" bold="true" pageId="0" pageNumber="1">
<collectingCountry id="E82760F0CCD9D93D8EC19B75320CD390" name="Colombia">Colombia</collectingCountry>
. 
<normalizedToken id="07C5103B6E3DC63F5897FE1705BB59A2" originalValue="Chocó">
<collectingRegion id="C931CD96756A3141E61700FB1DC41319" country="Colombia" name="Choco">Choco</collectingRegion>
</normalizedToken>
</emphasis>
: 
<quantity id="FC8FCB0A4D1425B72F085877B0C38CAD" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
south of Las Animas on road to Istmina, 
<quantity id="8AC55F60A85588DA3551AAEC91B9C120" metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="m" value="150.0">150 m</quantity>
, 
<date id="0BB07E448BFD6B698DD689F61483E85A" value="1976-08-13">13 Aug 1976</date>
, 
<emphasis id="12A6A553841BAE1B84DC0D0B61801FA3" italics="true" pageId="0" pageNumber="1">A. Gentry &amp; M. Fallen 17620</emphasis>
(COL); 
<emphasis id="1F4471218B778A3368E1FDDB164F63AA" bold="true" pageId="0" pageNumber="1">
<collectingRegion id="1CA18BEA03AED859CB7E45AA41EA71EB" country="Colombia" name="Valle del Cauca">Valle del Cauca</collectingRegion>
</emphasis>
: municipio Buenaventura, carretera vieja a Benaventura, 
<quantity id="B4FA4831D26F261847B138074FFE5DE9" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.5" metricValueMin="0.9" unit="m" value="1200.0" valueMax="1500.0" valueMin="900.0">900-1500 m</quantity>
<date id="7EC584779075C7E640F19EB015A3CD4C" value="1998-04">Apr 1998</date>
, 
<emphasis id="974CA504D942C4396B5198C862CE2CA8" italics="true" pageId="0" pageNumber="1">M. Amaya-M. &amp; J.F. Smith 594</emphasis>
(COL, US); municipio Buenaventura, Los Tubos, Pericos, vertiente occidental de cordillera occidental, km 43 Cali-Buenaventura, 
<geoCoordinate id="5D8399CFC83C46DE819AF5B1D9FB6278" degrees="3" direction="north" minutes="51" orientation="latitude" precision="15" seconds="00" value="3.85">3°51'00&quot;N</geoCoordinate>
, 
<geoCoordinate id="032A4FA7316D155E83E9241432FC5141" degrees="76" direction="west" minutes="47" orientation="longitude" precision="15" seconds="19" value="-76.78861">76°47'19&quot;W</geoCoordinate>
, 
<quantity id="B322DAB71E58BA5A474C2508605BB7CE" metricMagnitude="2" metricUnit="m" metricValue="5.21" unit="m" value="521.0">521 m</quantity>
, 
<date id="49D7A788F0620294AF9663B88306D02F" value="2011-08-30">30 Aug 2011</date>
, 
<emphasis id="20C3592E4C161F5664E086782EFEE63F" italics="true" pageId="0" pageNumber="1">J. Home 228</emphasis>
(CUVC); Bajo Calima, 
<normalizedToken id="3C3E5CBD8A0CF4D613B94DB6C9398CAD" originalValue="concesión">concesion</normalizedToken>
Pulpapel/Buenaventura, carretera a San Isidro, 
<geoCoordinate id="D2EAC8D0ADDFCF57295CA1C4D59F36BB" degrees="3" direction="north" minutes="55" orientation="latitude" precision="925" value="3.9166667">3°55'N</geoCoordinate>
, 
<geoCoordinate id="EF00018154BBFB3FF56A3A0B66D79D15" degrees="77" direction="west" orientation="longitude" precision="55555" value="-77.0">77°W</geoCoordinate>
, 
<quantity id="9FCE099316974F3CF3BDB79F35A1D0FC" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</quantity>
, 
<date id="B70DE3F2D3A5198220C4016674F3694A" value="1987-09-30">30 Sep 1987</date>
, 
<emphasis id="672B02624FB8E806A2B1AD6AB8E48323" italics="true" pageId="0" pageNumber="1">M. Monsalve B. 1865</emphasis>
(CUVC, MO); road Cali-Buenaventura, further down the road towards Buenaventura, about 
<quantity id="E48AAD2B382CAA25EC3B0396D65A15D7" metricMagnitude="3" metricUnit="m" metricValue="4.0" unit="km" value="4.0">4 km</quantity>
, 
<date id="DDF91BFD1DE48B8BE5531F685EAA4F2C" value="1972-05-01">1 May 1972</date>
, 
<emphasis id="2B08139C90B788672E35F822D58D87CF" italics="true" pageId="0" pageNumber="1">H. Wiehler</emphasis>
, 
<emphasis id="836CD1FE72CA9A9D03A85210BBE07AD4" italics="true" pageId="0" pageNumber="1">R.L. Dressler</emphasis>
, 
<emphasis id="1907DCD8E766DE52892613F8D62FCDFE" italics="true" pageId="0" pageNumber="1">N.H. Williams &amp; N.F. Williams 72106</emphasis>
(SEL); municipio Dagua, Vereda Yatacue, sitio La Riqueza, camino desde el tunel a las torres, 
<geoCoordinate id="8A5F2023BB0C3170B8BCA8DAD1212CB3" degrees="3" direction="north" minutes="35" orientation="latitude" precision="1" seconds="40.6" value="3.594611">3°35'40.6&quot;N</geoCoordinate>
, 
<geoCoordinate id="68B99DF66C2901392C6424508D4757AC" degrees="76" direction="west" minutes="53" orientation="longitude" precision="1" seconds="31.8" value="-76.892166">76°53'31.8&quot;W</geoCoordinate>
. 
<quantity id="BDA520C9DF415B09B46901904827C0F4" metricMagnitude="2" metricUnit="m" metricValue="7.05" metricValueMax="7.8" metricValueMin="6.3" unit="m" value="705.0" valueMax="780.0" valueMin="630.0">630-780 m</quantity>
, 
<date id="1A5D8A1C668C55A7A0E4E922EF956583" value="2019-01-20">20 Jan 2019</date>
, 
<emphasis id="2473A814718960CD27D0F823C0B9AAA1" italics="true" pageId="0" pageNumber="1">L. Clavijo</emphasis>
, 
<emphasis id="586F2799D8ED4B72303910EA6483B2B2" italics="true" pageId="0" pageNumber="1">M. Perret</emphasis>
, 
<emphasis id="F695E53770819A7DD543D0085179A6CC" italics="true" pageId="0" pageNumber="1">K. Arango</emphasis>
, 
<emphasis id="4F5C372FCE32720DE1D694D0BB255998" italics="true" pageId="0" pageNumber="1">A. Zuluaga &amp; W. Villar 2231</emphasis>
(COL, CUVC, G). 
<emphasis id="A92B753E85A7265EA2F262DD2A439DEA" bold="true" pageId="0" pageNumber="1">
<normalizedToken id="5A10FD94D4303F1B9569FBE3B56BD317" originalValue="Nariño">
<collectingRegion id="D403EE094FC54AD78148654A9010E6B5" country="Colombia" name="Narino">Narino</collectingRegion>
</normalizedToken>
</emphasis>
: municipio Barbacoas, corregimiento El Diviso, vereda El Verde, western slopes of the Cordillera Occidental, remnant forest along highway between Altaquer and El Diviso, 
<geoCoordinate id="87C0E7C78F88105695D5D517750C5515" degrees="1" direction="north" minutes="21" orientation="latitude" precision="15" seconds="46" value="1.3627778">1°21'46&quot;N</geoCoordinate>
, 
<geoCoordinate id="78D1F30F238A52F4FBFFF4823D65961D" degrees="78" direction="west" minutes="10" orientation="longitude" precision="15" seconds="32" value="-78.17555">78°10'32&quot;W</geoCoordinate>
, 
<quantity id="ABC4089861D18C21CEF9EB9ECD16F6A7" metricMagnitude="2" metricUnit="m" metricValue="7.95" unit="m" value="795.0">795 m</quantity>
, 
<date id="0DF04B19EB0061997EC97BC7E6CCDD04" value="2013-05-13">13 May 2013</date>
, 
<emphasis id="9F6C6A70AEB866B4E820848FA9F600E8" italics="true" pageId="0" pageNumber="1">J.L. Clark</emphasis>
, 
<emphasis id="FC4F0A9F4D0149FFAEF06DF04627FA1F" italics="true" pageId="0" pageNumber="1">L. Clavijo</emphasis>
, 
<emphasis id="DE08754182F019D32C86CA9EEAE5807C" italics="true" pageId="0" pageNumber="1">
O. 
<normalizedToken id="C528267DA2E792BD7D749AEC65357446" originalValue="Marín">Marin</normalizedToken>
&amp; M. Flores 13417
</emphasis>
(COL, CUVC, SEL, US); municipio de Ricaurte, Resguardo 
<normalizedToken id="E06D599FDFF2D9744FB6B1ECF25B1F78" originalValue="Indígena">Indigena</normalizedToken>
Nulpe Medio, camino a la quebrada La Conga, 
<geoCoordinate id="E682C2FFE2FEEC106E187A9808206431" degrees="1" direction="north" minutes="6" orientation="latitude" precision="925" value="1.1">1°6'N</geoCoordinate>
, 
<geoCoordinate id="FDB9003A2151DEDA9CE3A16BD7174499" degrees="78" direction="west" minutes="13" orientation="longitude" precision="925" value="-78.21667">78°13'W</geoCoordinate>
, 
<quantity id="83442668DE3275583B4EC82CE17D6692" metricMagnitude="2" metricUnit="m" metricValue="7.5" unit="m" value="750.0">750 m</quantity>
, 
<date id="0F572FDFB04F7BAD549BA0BFEB0F76BB" value="1996-01-08">8 Jan 1996</date>
, 
<emphasis id="AB3092E921C4E9B4574A69E3691F0B76" italics="true" pageId="0" pageNumber="1">
M.S. 
<normalizedToken id="AA499AA927C7EC2FDAAFEC9F69498E73" originalValue="González">Gonzalez</normalizedToken>
&amp; B.R. 
<normalizedToken id="4D39216A7A3BEF120CD30E0E0B08D411" originalValue="Ramírez-P">Ramirez-P</normalizedToken>
. 1606
</emphasis>
(PSO); municipio de Ricaurte, Resguardo Indigena Nulpe Medio 
<normalizedToken id="F3DDCB81869889D0CD25144701462A00" originalValue="Andalucía">Andalucia</normalizedToken>
, quebrada La Babosa, 
<geoCoordinate id="F64F955AA7BD3B618F9306730D5A1356" degrees="1" direction="north" minutes="5" orientation="latitude" precision="925" value="1.0833334">1°5'N</geoCoordinate>
, 
<geoCoordinate id="C35CB97EE3F6D9C38D87DDD094E78436" degrees="78" direction="west" minutes="14" orientation="longitude" precision="925" value="-78.23333">78°14'W</geoCoordinate>
, 
<quantity id="EB430175EF4DAACA3872E124465F40EF" metricMagnitude="2" metricUnit="m" metricValue="7.975" metricValueMax="8.15" metricValueMin="7.8" unit="m" value="797.5" valueMax="815.0" valueMin="780.0">780-815 m</quantity>
, 
<date id="0A42E1C20D3DD42AB37E973EF5971E89" value="1996-01-12">12 Jan 1996</date>
, 
<emphasis id="7FA3D2B5843A5362BB2F625911A14E2B" italics="true" pageId="0" pageNumber="1">
M.S. 
<normalizedToken id="EC3C33D952FB8944ABF7D68D02807E19" originalValue="González">Gonzalez</normalizedToken>
&amp; B.R. 
<normalizedToken id="2A20A6113D51785C48DE6A9D46CFD516" originalValue="Ramírez-P">Ramirez-P</normalizedToken>
. 1678
</emphasis>
(PSO). 
<emphasis id="437C2CA80C1321F8E2E3A03ECEEDD15F" bold="true" pageId="0" pageNumber="1">
<collectingCountry id="2AAA0DC4F6C07B2E580DE76CAC62FF6F" name="Ecuador">Ecuador</collectingCountry>
. 
<collectingRegion id="F8D5A57C680D199ECCF65730C0BC59C2" country="Ecuador" name="Carchi">Carchi</collectingRegion>
</emphasis>
: border area between 
<collectingRegion id="29DCEFA6D5420311285D25B6CF36B19B" country="Ecuador" name="Carchi">Carchi</collectingRegion>
and 
<collectingRegion id="68754BEC704D1BDBE24153B01330C670" country="Ecuador" name="Esmeraldas">Esmeraldas</collectingRegion>
, about 
<quantity id="00D44B2B6F67064D3E9530E83201B529" metricMagnitude="4" metricUnit="m" metricValue="3.0" unit="km" value="30.0">30 km</quantity>
past Lita on road Lita-Alto Tambo, 
<quantity id="53AD50BFA2834B818734C9F52F019346" metricMagnitude="2" metricUnit="m" metricValue="4.5" unit="m" value="450.0">450 m</quantity>
, 
<date id="73CCC1838019B6BF4C625001A94B92D5" value="1991-06-28">28 Jun 1991</date>
, 
<emphasis id="91BD470221F3833FD9135DD0BB651FF5" italics="true" pageId="0" pageNumber="1">H. van der Werff</emphasis>
, 
<emphasis id="E13BDF59293274E7CB022615AD7CB1ED" italics="true" pageId="0" pageNumber="1">B. Gray &amp; G. Tipas 12113</emphasis>
(MO, QCNE, SEL, US). 
<emphasis id="838C76F31ACC0313F1A4E6944B5E8BC3" bold="true" pageId="0" pageNumber="1">
<collectingRegion id="DB917242E9714E042D141D822119D6B9" country="Ecuador" name="Esmeraldas">Esmeraldas</collectingRegion>
</emphasis>
: 
<normalizedToken id="84D21B031A69A7B41B26C90517D9D59D" originalValue="cantón">canton</normalizedToken>
San Lorenzo, parroquia Alta Tambo, 
<normalizedToken id="A0930B0B85ECEC7EE2647BA738E173D4" originalValue="Awá">Awa</normalizedToken>
Indigenous Territory, 
<normalizedToken id="509329C44C28188664E79CD55E0CC303" originalValue="Río">Rio</normalizedToken>
<normalizedToken id="624C0E553506CE7D4A3310BCEC4167F6" originalValue="Bogotá">Bogota</normalizedToken>
community (future biological research station), 
<quantity id="FB8195653A877E93F3E426C4B8CD25E9" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
S of Lita-San Lorenzo road, near Quebrada Pambilar, 
<geoCoordinate id="07B4F168870B08866E877E9DB1DEDE49" degrees="0" direction="north" minutes="58" orientation="latitude" precision="15" seconds="57" value="0.98249996">0°58'57&quot;N</geoCoordinate>
, 
<geoCoordinate id="C72F56FA098ACBFD63458ABC06455353" degrees="78" direction="west" minutes="35" orientation="longitude" precision="15" seconds="60" value="-78.600006">78°35'60&quot;W</geoCoordinate>
, 
<quantity id="C57A1167C80BD7BCB0E605148C961783" metricMagnitude="2" metricUnit="m" metricValue="4.75" metricValueMax="6.0" metricValueMin="3.5" unit="m" value="475.0" valueMax="600.0" valueMin="350.0">350-600 m</quantity>
, 
<date id="6AA0E8A6E22F539F83D6314DC245D49D" value="2003-02-12">12 Feb 2003</date>
, 
<emphasis id="FCE3E5DA562956FABF66DB0B045060BD" italics="true" pageId="0" pageNumber="1">J.L. Clark</emphasis>
, 
<emphasis id="AA675532450634E7CC64442A646B16A0" italics="true" pageId="0" pageNumber="1">G. Zapata &amp; G. Toasa 7129</emphasis>
(MO, QCNE, SEL, UNA); 
<normalizedToken id="2F8062885D9BDED1CFB8ADF6453D041B" originalValue="cantón">canton</normalizedToken>
San Lorenzo, parroquia Alta Tambo, border region of 
<normalizedToken id="08407248BC21CABECE76FD79F5D13697" originalValue="Awá">Awa</normalizedToken>
Indigenous Territory, entrance to the 
<normalizedToken id="82B6DC3E605764680848554D36577DC9" originalValue="Río">Rio</normalizedToken>
<normalizedToken id="029DB3A38088C417FBFD54AA4FC858FD" originalValue="Bogotá">Bogota</normalizedToken>
community (future biological research station), near Quebrada Pambilar, 
<geoCoordinate id="F1CAE62568266DBEF03A17CFCBC949CB" degrees="0" direction="north" minutes="58" orientation="latitude" precision="15" seconds="57" value="0.98249996">0°58'57&quot;N</geoCoordinate>
, 
<geoCoordinate id="D1B7A580D25E249C45318A39D3214D89" degrees="78" direction="west" minutes="35" orientation="longitude" precision="15" seconds="50" value="-78.59722">78°35'50&quot;W</geoCoordinate>
, 
<quantity id="AC96AA377039CA583CB51F91C27E1997" metricMagnitude="2" metricUnit="m" metricValue="4.75" metricValueMax="6.0" metricValueMin="3.5" unit="m" value="475.0" valueMax="600.0" valueMin="350.0">350-600 m</quantity>
, 
<date id="D68B5BDFC0C88F13523454A4A3CA8870" value="2003-02-13">13 Feb 2003</date>
, 
<emphasis id="1C68C2B43811DD3B65DF7D1BED189C41" italics="true" pageId="0" pageNumber="1">J.L. Clark 7197</emphasis>
(F, MO, QCNE, SEL, UNA, US); parroquia Alto Tambo, trail from the community Durango to 
<normalizedToken id="56A29A8C69BDF3FA8F79336FA21C873E" originalValue="Río">Rio</normalizedToken>
Tululbi via trail north of San Lorenzo-Ibarra highway, forest managed by Fundacion Sirua, 
<geoCoordinate id="99E44020F2E88EAB5410465D6AE1203A" degrees="1" direction="north" minutes="2" orientation="latitude" precision="15" seconds="50" value="1.0472221">1°2'50&quot;N</geoCoordinate>
, 
<geoCoordinate id="CE9E0D78EFE0902C09B3F77FE9F89032" degrees="78" direction="west" minutes="36" orientation="longitude" precision="15" seconds="54" value="-78.615">78°36'54&quot;W</geoCoordinate>
, 
<quantity id="214B4871047085147B2C14957CA3C5F2" metricMagnitude="2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" unit="m" value="175.0" valueMax="200.0" valueMin="150.0">150-200 m</quantity>
, 
<date id="5F6DCC9CE20DCD98954932E0ED0B45F8" value="2008-05-28">28 May 2008</date>
, 
<emphasis id="1EFBBCC0B763702DD243DF69ED6187ED" italics="true" pageId="0" pageNumber="1">J.L. Clark</emphasis>
, 
<emphasis id="73804388E543713A41ED76DEFDD24194" italics="true" pageId="0" pageNumber="1">
B. Bisvicuth &amp; J. Melton 
<accessionNumber id="90733E9F6F635137FF6B325E90C86E1C" httpUri="http://www.ncbi.nlm.nih.gov/protein/III10343">III 10343</accessionNumber>
</emphasis>
(MO, NY, QCNE, SEL, US); 
<normalizedToken id="1E1A3B37F7DCC4F9B786EA243ACA2585" originalValue="cantón">canton</normalizedToken>
San Lorenzo, remnant patch of forest along the Ibarra-San Lorenzo highway, between Durango and Alto Tambo, 
<geoCoordinate id="064593483F08EB8AD3E8EEF93C2C5140" degrees="1" direction="north" minutes="0" orientation="latitude" precision="15" seconds="33" value="1.0091667">1°0'33&quot;N</geoCoordinate>
, 
<geoCoordinate id="8C4944E7F08A8FAA2FE3099B79381088" degrees="78" direction="west" minutes="35" orientation="longitude" precision="15" seconds="58" value="-78.59945">78°35'58&quot;W</geoCoordinate>
, 
<quantity id="4A6CC0E0683E513605809F52AC20E70C" metricMagnitude="2" metricUnit="m" metricValue="5.16" unit="m" value="516.0">516 m</quantity>
, 
<date id="5FC73EFEC2399BFFC8BBDE88E761111D" value="2009-06-03">3 Jun 2009</date>
, 
<emphasis id="B301A9CAE23A030C232CA42A23817685" italics="true" pageId="0" pageNumber="1">J.L. Clark J.L. Clark &amp; The 2009 Gesneriad Research Expedition Participants 11113</emphasis>
(MO, NY, QCNE, SEL, US); 
<normalizedToken id="C5BB0E2F50B53885E51DF2D62491BBCA" originalValue="cantón">canton</normalizedToken>
San Lorenzo, remnant patch of forest along highway Ibarra - San Lorenzo, between Durango and Alto Tambo, 
<geoCoordinate id="C4D7A27621092B3B54BA9C78416F619C" degrees="1" direction="north" minutes="0" orientation="latitude" precision="15" seconds="33" value="1.0091667">1°0'33&quot;N</geoCoordinate>
, 
<geoCoordinate id="CBF2E1B8DA050CDA2DB1257F550CF051" degrees="78" direction="west" minutes="35" orientation="longitude" precision="15" seconds="58" value="-78.59945">78°35'58&quot;W</geoCoordinate>
, 
<quantity id="0B2DE141F2AE479A71F7E7FF3AFC59B4" metricMagnitude="2" metricUnit="m" metricValue="5.16" unit="m" value="516.0">516 m</quantity>
, 
<emphasis id="203AEED5E797300C405131FC2891C20A" italics="true" pageId="0" pageNumber="1">J.L. Clark &amp; The 2009 Gesneriad Research Expedition Participants 11127</emphasis>
(QCNE, SEL, US); 
<normalizedToken id="4E598A3F3B40CC6C6839469CDBEAFD99" originalValue="cantón">canton</normalizedToken>
Eloy Alfaro, Reserva 
<normalizedToken id="BCC2B291928EBF1C2884468D9DCA203F" originalValue="Ecológica">Ecologica</normalizedToken>
Cotachi-Cayapas, parroquia Luis Vargas Torres, 
<normalizedToken id="06457673022B13E10C787269988A2DBC" originalValue="Río">Rio</normalizedToken>
Santiago, 
<geoCoordinate id="AEE4F9AEC5C7B91DC080DF7F9D8697FB" degrees="0" direction="south" minutes="49" orientation="latitude" precision="925" value="-0.81666666">0°49'S</geoCoordinate>
, 
<geoCoordinate id="CB41412EC267E8ECCBC7E497AA919E26" degrees="78" direction="west" minutes="45" orientation="longitude" precision="925" value="-78.75">78°45'W</geoCoordinate>
, 
<quantity id="A163480B6A410120972C576F89FD43B1" metricMagnitude="2" metricUnit="m" metricValue="2.5" unit="m" value="250.0">250 m</quantity>
, 
<date id="8B4288C123EEE405B2CD3FA292DEDCFA" value="1993-12-08">8 Dec 1993</date>
, 
<emphasis id="B880B00374DFBDB80DC87EA52787D4A8" italics="true" pageId="0" pageNumber="1">M. Tirado</emphasis>
, 
<emphasis id="7691E28816ED54B8ADD29F8D34BA95AB" italics="true" pageId="0" pageNumber="1">P. Asimbaya</emphasis>
, 
<emphasis id="FA03FE671E15D96DAB43B48D68A93DD9" italics="true" pageId="0" pageNumber="1">M.I. Corosa &amp; V. Arroyo 776</emphasis>
(MO, QCNE, US); 
<normalizedToken id="897F7D967000B80227BF2A506126A8EA" originalValue="cantón">canton</normalizedToken>
San Lorenzo, San Francisco, recinto Durango, sector Colinado, terrenos propiedad Sr. Demetrio Paez, 
<quantity id="3DB51FD0FECA8F70D4DB09102C9B34C6" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="km" value="1.0">1 km</quantity>
al este de la carretera Lita-San Lorenzo, 
<geoCoordinate id="52DAD00A8DA4D3BBE25B3D2876AF53E3" degrees="1" direction="north" minutes="2" orientation="latitude" precision="925" value="1.0333333">1°2'N</geoCoordinate>
, 
<geoCoordinate id="B073798E7994C279883314C0216BCE48" degrees="78" direction="west" minutes="36" orientation="longitude" precision="925" value="-78.6">78°36'W</geoCoordinate>
, 
<quantity id="4C28E98496FC2557E4D2764244E4A039" metricMagnitude="2" metricUnit="m" metricValue="2.56" unit="m" value="256.0">256 m</quantity>
, 
<date id="DDFFE4EBA4AC1759F3443F5B10510614" value="1999-10-17">17 Oct 1999</date>
, 
<emphasis id="220987D33C243FF063E1597234F5914E" italics="true" pageId="0" pageNumber="1">J.C. Valenzuela &amp; E. Freire 474</emphasis>
(QCNE, MO, US). 
<emphasis id="24012BD180A2A19F14A65E5A8222430B" bold="true" pageId="0" pageNumber="1">
<collectingRegion id="377749CD7307C040BEA8F5077062DB00" country="Ecuador" name="Morona-Santiago">Morona-Santiago</collectingRegion>
</emphasis>
: 
<normalizedToken id="A772CACD35E0AAD551E73C3D71272CB1" originalValue="Región">Region</normalizedToken>
de la Cordillera del 
<normalizedToken id="DD0A1696935B741B16B5A920DBE46D1D" originalValue="Cóndor">Condor</normalizedToken>
y Cordillera de 
<normalizedToken id="D18BE1FD8E86854D09BCE0D4DBFD26AF" originalValue="Cutucú">Cutucu</normalizedToken>
, Cordillera de Shaime, al norte del 
<normalizedToken id="2A940993D0C95BA966F0AF485421154F" originalValue="Río">Rio</normalizedToken>
Santiago, Centro Shuar Jempekat, 
<geoCoordinate id="8D0B865F5867E95ADE01519E2FAC3BB9" degrees="2" direction="south" minutes="57" orientation="latitude" precision="925" value="-2.95">2°57'S</geoCoordinate>
, 
<geoCoordinate id="81FF4B0293CD6FCF38DAF6C0CCB89911" degrees="77" direction="west" minutes="50" orientation="longitude" precision="925" value="-77.833336">77°50'W</geoCoordinate>
, 
<quantity id="1702F01B560877F22115B3383DA0B2F9" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</quantity>
, 
<date id="8D7DC73EA4A15C198A8E75382A282697" value="2003-10-15">15 Oct 2003</date>
, 
<emphasis id="AE01B7EDC9C9B2B970F431FC0259BE13" italics="true" pageId="0" pageNumber="1">G. Toasa 9343</emphasis>
(MO, QCNE).
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>