372 lines
47 KiB
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372 lines
47 KiB
XML
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<mods:title id="97F247C2DCD4B630D42E202A195EB3DD">Historical and morphological review of the subgenus Indonthophagus Kabakov, 2006 of Onthophagus Latreille, 1802 (Coleoptera: Scarabaeidae: Scarabaeinae: Onthophagini)</mods:title>
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<mods:namePart id="7C5CCD342FE4EECC5D31C67CE1AE77D1">Ziani, Stefano</mods:namePart>
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<taxonomicName id="34967281D323473F93BFFC42FC67FC7E" ID-CoL="7XM9B" authority="d'Orbigny, 1902" authorityName="d'Orbigny" authorityYear="1902" box="[192,963,937,966]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="aeneopiceus" subGenus="Indonthophagus">
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(
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<emphasis id="C1E2D510D323473F9204FC42FDECFC7E" bold="true" box="[379,584,937,966]" italics="true" pageId="6" pageNumber="5">Indonthophagus</emphasis>
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<emphasis id="C1E2D510D323473F9126FC42FD54FC7D" bold="true" box="[601,752,937,965]" italics="true" pageId="6" pageNumber="5">aeneopiceus</emphasis>
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<bibRefCitation id="970774F3D323473F9188FC42FC67FC7E" author="d'Orbigny H." box="[759,963,937,966]" pageId="6" pageNumber="5" pagination="1 - 324" refId="ref27623" refString="d'Orbigny H. 1902. Memoire sur les Onthophagides d'Afrique. Annales de la Societe Entomologique de France 71 (1 - 2): 1 - 324." type="journal article" year="1902">d’Orbigny, 1902</bibRefCitation>
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(
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)
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<emphasis id="C1E2D510D323473F939FFC11FE46FBAA" box="[224,482,1018,1042]" italics="true" pageId="6" pageNumber="5">Onthophagus aeneopiceus</emphasis>
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<bibRefCitation id="970774F3D323473F9296FC11FD1DFBAA" author="d'Orbigny H." box="[489,697,1018,1042]" pageId="6" pageNumber="5" pagination="1 - 324" refId="ref27623" refString="d'Orbigny H. 1902. Memoire sur les Onthophagides d'Afrique. Annales de la Societe Entomologique de France 71 (1 - 2): 1 - 324." type="journal article" year="1902">d’Orbigny 1902: 156</bibRefCitation>
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<treatmentCitation id="72372F13D323473F91BBFC11FC31FBAA" author="d'Orbigny H." box="[708,917,1018,1042]" page="488" pageId="6" pageNumber="5" year="1905">
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<bibRefCitation id="970774F3D323473F91BBFC11FC31FBAA" author="d'Orbigny H." box="[708,917,1018,1042]" pageId="6" pageNumber="5" pagination="381 - 536" refId="ref27691" refString="d'Orbigny H. 1905. Onthophagides provenant du voyage de M. Alluaud dans l'Afrique orientale et descriptions d'autres especes africaines nouvelles. Annales de la Societe Entomologique de France 74 (3 - 4): 381 - 536." type="journal article" year="1905">d’Orbigny 1905: 488</bibRefCitation>
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;
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<treatmentCitation id="72372F13D323473F90E0FC11FBCBFBAA" author="d'Orbigny H." box="[927,1135,1018,1042]" page="376" pageId="6" pageNumber="5" year="1913">
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<bibRefCitation id="970774F3D323473F90E0FC11FBCBFBAA" author="d'Orbigny H." box="[927,1135,1018,1042]" pageId="6" pageNumber="5" pagination="1 - 608" refId="ref27773" refString="d'Orbigny H. 1913. Synopsis des Onthophagides d'Afrique. Annales de la Societe Entomologique de France 82: 1 - 608." type="journal article" year="1913">d’Orbigny 1913: 376</bibRefCitation>
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;
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<treatmentCitation id="72372F13D323473F9704FC11FAEFFBAA" author="d'Orbigny H." box="[1147,1355,1018,1042]" page="609" pageId="6" pageNumber="5" year="1922">
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<bibRefCitation id="970774F3D323473F9704FC11FAEFFBAA" author="d'Orbigny H." box="[1147,1355,1018,1042]" pageId="6" pageNumber="5" pagination="599 - 615" refId="ref27796" refString="d'Orbigny H. 1922. Coleopteres Onthophagines. p. 599 - 615. In: Rothschild MD (ed.). Voyage de M. le Baron Maurice de Rotschild en Ethiopie et en Afrique Orientale Anglaise (1904 - 1905). Resultats scientifiques. Animaux Articules. Imprimerie Nationale; Paris. 1041 p." type="book chapter" year="1922">d’Orbigny 1922: 609</bibRefCitation>
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<treatmentCitation id="72372F13D323473F9629FC11FDB5FB89" author="Boucomont A & Gillet JJE" page="151" pageId="6" pageNumber="5" year="1927">
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<bibRefCitation id="970774F3D323473F9629FC11FDB5FB89" author="Boucomont A & Gillet JJE" pageId="6" pageNumber="5" pagination="103 - 263" refId="ref26707" refString="Boucomont A, Gillet JJE. 1927. Fam. Scarabaeidae. Subfam. Coprinae II. p. 103 - 263. In: Gillet JJE, Boucomont A (eds.). Scarabaeidae: Coprinae, Termitotroginae. Partes 38 et 90, Vol. 19 (3). In: Schenkling S (ed.). Coleopterorum Catalogus. W. Junk; Berlin. 265 p." type="book chapter" year="1927">Boucomont and Gillet 1927: 151</bibRefCitation>
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<treatmentCitation id="72372F13D323473F9162FBF2FD4DFB89" author="Balthasar V." box="[541,745,1049,1073]" page="263" pageId="6" pageNumber="5" year="1963">
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<bibRefCitation id="970774F3D323473F9162FBF2FD4DFB89" author="Balthasar V." box="[541,745,1049,1073]" pageId="6" pageNumber="5" refId="ref25682" refString="Balthasar V. 1963. Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera Lamellicornia. Band 2. Coprinae (Onitini, Oniticellini, Onthophagini). Tschechoslowakischen Akademie der Wissenschaften; Prague. 627 p. + 16 pls." type="book" year="1963">Balthasar 1963: 263</bibRefCitation>
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;
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<treatmentCitation id="72372F13D323473F918AFBF2FC10FB89" author="Balthasar V." box="[757,948,1049,1073]" page="65" pageId="6" pageNumber="5" year="1969">
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<bibRefCitation id="970774F3D323473F918AFBF2FC10FB89" author="Balthasar V." box="[757,948,1049,1073]" pageId="6" pageNumber="5" pagination="59 - 73" refId="ref25804" refString="Balthasar V. 1969. Contributions a la connaissance de la faune entomologique de la Cote-d'Ivoire (J. Decelle, 1961 - 1964). XXXII. Coleoptera Scarabaeidae et Aphodiidae. Annales du Musee Royal de l'Afrique Centrale / Sciences Zoologiques 175: 59 - 73." type="journal article" year="1969">Balthasar 1969: 65</bibRefCitation>
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;
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<treatmentCitation id="72372F13D323473F90BFFBF2FBD9FB89" author="Ferreira MC" box="[960,1149,1049,1073]" page="557" pageId="6" pageNumber="5" year="1972">
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<bibRefCitation id="970774F3D323473F90BFFBF2FBD9FB89" author="Ferreira MC" box="[960,1149,1049,1073]" pageId="6" pageNumber="5" pagination="1968 - 1969" refId="ref28208" refString="Ferreira MC. 1972. Os Escarabideos de Africa (sul do Saara). I. Revista de Entomologia de Mocambique 11 [1968 - 1969]: 5 - 1088, 35 anns, 288 plates." type="book chapter" year="1972">Ferreira 1972: 557</bibRefCitation>
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;
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<treatmentCitation id="72372F13D323473F97F6FBF2FAECFB89" author="Balthasar V." box="[1161,1352,1049,1073]" page="22" pageId="6" pageNumber="5" year="1972">
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<bibRefCitation id="970774F3D323473F97F6FBF2FAECFB89" author="Balthasar V." box="[1161,1352,1049,1073]" pageId="6" pageNumber="5" pagination="18 - 32" refId="ref25853" refString="Balthasar V. 1972. Coleoptera aus Nordostafrika. Scarabaeoidea: Scarabaeidae und Aphodiidae. 135. Beitrag zur Kenntnis der Scarabaeoidea. Notulae Entomologicae 52: 18 - 32." type="journal article" year="1972">Balthasar 1972: 22</bibRefCitation>
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;
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<treatmentCitation id="72372F13D323473F962BFBF2FEC6FBE9" author="Zunino M." page="153" pageId="6" pageNumber="5" year="1975">
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<bibRefCitation id="970774F3D323473F962BFBF2FEC6FBE9" author="Zunino M." pageId="6" pageNumber="5" pagination="151 - 194" refId="ref35071" refString="Zunino M. 1975. Revisione delle specie paleartiche del sottogenere Onthophagus (sensu strictu) Latr. (Coleoptera, Scarabaeoidea). I tipi di H. d'Orbigny, A. Raffray e A. Boucomont nel Museum National d'Histoire Naturelle di Parigi. Bollettino del Museo di Zoologia dell'Universita di Torino 7: 151 - 194." type="journal article" year="1975">Zunino 1975: 153</bibRefCitation>
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<bibRefCitation id="970774F3D323473F9213FBD2FDFFFBE9" author="Barbero E & Borghesio L & Dellacasa G & Dellacasa M." box="[364,603,1081,1105]" pageId="6" pageNumber="5" pagination="233 - 254" refId="ref25960" refString="Barbero E, Borghesio L, Dellacasa G, Dellacasa M. 1998. Contribution to the knowledge of Scarabaeidae and Aphodiidae from southern Ethiopia (Coleoptera Scarabaeoidea). Bollettino della Societa Entomologica Italiana 130 (3): 233 - 254." type="journal article" year="1998">Barbero et al. 1998: 243</bibRefCitation>
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<bibRefCitation id="970774F3D323473F911AFBD2FC9EFBE9" author="Schoolmeesters P." box="[613,826,1081,1105]" pageId="6" pageNumber="5" refId="ref32957" refString="Schoolmeesters P. 2023. World Scarabaeidae Database. In: Banki O, Roskov Y, Doring M, Ower G, Hernandez Robles DR, Plata Corredor CA, Stjernegaard Jeppesen T, Orn A, Vandepitte L, Hobern D, Schalk P, DeWalt RE, Keping M, Miller J, Orrell T, Aalbu R, Abbott J, Adlard R, Adriaenssens EM, et al. (eds.). Catalogue of Life Checklist (Version 2023 - 05 - 30). https: // doi. org / 10.48580 / dfsr- 38 g" type="book" year="2023">Schoolmeesters 2023</bibRefCitation>
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<treatmentCitation id="72372F13D323473F903BFBD2FB95FBE9" author="Ziani S & Keith D." box="[836,1073,1081,1105]" httpUri="http://treatment.plazi.org/id/039487F9D415FFF3FF60257943F0B2BB" page="1" pageId="6" pageNumber="5" year="2023">
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<bibRefCitation id="970774F3D323473F903BFBD2FB95FBE9" author="Ziani S & Keith D." box="[836,1073,1081,1105]" pageId="6" pageNumber="5" pagination="1 - 4" refId="ref35001" refString="Ziani S, Keith D. 2023. 3 rd contribution to the knowledge of Arabian Onthophagini (Coleoptera: Scarabaeidae). Onthophagus aeneopiceus d'Orbigny, 1902, a new record for Yemen and for the Palaearctic ecozone. Insecta Mundi 0974: 1 - 4." type="book chapter" year="2023">Ziani and Keith 2023: 1</bibRefCitation>
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locality.
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“
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<collectingCountry id="8B814992D323473F9214FB89FE6AFBC4" box="[363,462,1122,1148]" name="Eritrea" pageId="6" pageNumber="5">Erythrée</collectingCountry>
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” [
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].
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<emphasis id="C1E2D510D323473F93BFFB67FECCFB1E" bold="true" box="[192,360,1164,1190]" pageId="6" pageNumber="5">Type material.</emphasis>
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||
, a minor
|
||
<specimenCount id="E590C28BD323473F913FFB65FDF1FB12" box="[576,597,1166,1194]" count="1" pageId="6" pageNumber="5" type="male">♂</specimenCount>
|
||
, fixed by monotypy, examined by photos, in d’Orbigny collection (
|
||
<collectionCode id="958791C7D323473F9649FB67FA30FB1E" box="[1334,1428,1164,1190]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="5">MNHN</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F3290902D323473F93BFFB5CFDF4FAAE" blockId="6.[192,1442,1207,1927]" pageId="6" pageNumber="5">
|
||
<emphasis id="C1E2D510D323473F93BFFB5CFE08FB69" bold="true" box="[192,428,1207,1233]" pageId="6" pageNumber="5">Diagnostic features.</emphasis>
|
||
Length 4.0 to
|
||
<quantity id="346EA4E7D323473F9128FB5CFD16FB6A" box="[599,690,1207,1234]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="6" pageNumber="5" unit="mm" value="6.5">6.5 mm</quantity>
|
||
. Colour dark brown, moderately shiny, with distinct isodiametric microreticulation, elytra sometimes with dark red irregular humeral spots, clypeal sides dark red. Antennae yellow. Dorsal pubescence pale-yellow.
|
||
</paragraph>
|
||
<paragraph id="F3290902D323473F927FFAF4FCA4F9B1" blockId="6.[192,1442,1207,1927]" pageId="6" pageNumber="5">
|
||
Head short, clearly wider than long, with clypeus broadly round, only very feebly sinuate anteriorly, sides not sinuate; clypeofrontal carina distinct and bent backward, slightly closer to the base of the horn than to the anterior margin of clypeus; occipital carina extended in a long horn arising between the eyes, where it is broad, broader than clypeofrontal carina, and flat, directed backward, tapering gradually to the extremity and following the curvature of pronotum, with moderately long yellow setae mainly at sides, in major males, reduced to a narrow straight lamina in minor males (
|
||
<typeStatus id="2C2DB7A0D323473F91D9FA27FCAAFA5E" box="[678,782,1484,1510]" pageId="6" pageNumber="5" type="holotype">holotype</typeStatus>
|
||
); clypeal surface with very transversely rugose, setigerous punctures, frontal surface doubly sparsely punctate.
|
||
</paragraph>
|
||
<paragraph id="F3290902D323473F927FF9F9FC54F961" blockId="6.[192,1442,1207,1927]" pageId="6" pageNumber="5">
|
||
Pronotum convex, declivous anteriorly, with a distinct anteromedian depressed area in major males (in minor male (
|
||
<typeStatus id="2C2DB7A0D323473F922DF9DFFE1CF9F6" box="[338,440,1588,1614]" pageId="6" pageNumber="5" type="holotype">holotype</typeStatus>
|
||
) without depression) and a small hardly distinguishable posterolateral dull area on either side near pronotal posterior angles; anterior angles distinctly produced, sides not sinuate behind them in dorsal view; dorsal surface setigerously punctate, punctures sub-regular in distribution, separated by 1-2 diameters on disc, smaller and more spaced on the anteromedian furrow; each puncture bears a small granule at its anterior margin, and a long pale-yellow seta, slightly longer that those on clypeus.
|
||
</paragraph>
|
||
<paragraph id="F3290902D323473F927FF90AFDFBF8DC" blockId="6.[192,1442,1207,1927]" pageId="6" pageNumber="5">Elytral striae shiny, distinctly impressed, with punctures slightly larger than strial width and barely crenulating interstrial sides; interstriae flat to barely convex, all rather regularly granulate; granules slightly smaller than strial punctures; posterior margin of each granule with a small, indistinct, setigerous puncture; setae pale-yellow, thin, as long as pronotal ones.</paragraph>
|
||
<paragraph id="F3290902D323473F927FF887FB7AF83E" blockId="6.[192,1442,1207,1927]" box="[256,1246,1900,1927]" pageId="6" pageNumber="5">Pygidium with rather regularly distributed, setigerous punctures; setae yellow, long, thin.</paragraph>
|
||
<caption id="A7E9598AD322473E93BFF8DAFBE1F83E" ID-DOI="http://doi.org/10.5281/zenodo.11450112" ID-Zenodo-Dep="11450112" httpUri="https://zenodo.org/record/11450112/files/figure.png" pageId="7" pageNumber="6" startId="7.[192,261,1841,1865]" targetBox="[219,1400,236,1780]" targetPageId="7" targetType="figure">
|
||
<paragraph id="F3290902D322473E93BFF8DAFBE1F83E" blockId="7.[192,1442,1841,1926]" pageId="7" pageNumber="6">
|
||
<emphasis id="C1E2D510D322473E93BFF8DAFEBBF8F1" bold="true" box="[192,287,1841,1865]" pageId="7" pageNumber="6">Figure 3.</emphasis>
|
||
<taxonomicName id="34967281D322473E9259F8DAFC2DF8F1" authority="d'Orbigny, 1902" authorityName="d'Orbigny" authorityYear="1902" box="[294,905,1841,1865]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="aeneopiceus" subGenus="Indonthophagus">
|
||
<emphasis id="C1E2D510D322473E9259F8DAFE0FF8F1" box="[294,427,1841,1865]" italics="true" pageId="7" pageNumber="6">Onthophagus</emphasis>
|
||
(
|
||
<emphasis id="C1E2D510D322473E92C5F8DAFDFEF8F1" box="[442,602,1841,1865]" italics="true" pageId="7" pageNumber="6">Indonthophagus</emphasis>
|
||
)
|
||
<emphasis id="C1E2D510D322473E9115F8D9FD44F8F1" box="[618,736,1842,1865]" italics="true" pageId="7" pageNumber="6">aeneopiceus</emphasis>
|
||
<bibRefCitation id="970774F3D322473E9198F8DAFC2DF8F1" author="d'Orbigny H." box="[743,905,1841,1865]" pageId="7" pageNumber="6" pagination="1 - 324" refId="ref27623" refString="d'Orbigny H. 1902. Memoire sur les Onthophagides d'Afrique. Annales de la Societe Entomologique de France 71 (1 - 2): 1 - 324." type="journal article" year="1902">d’Orbigny, 1902</bibRefCitation>
|
||
</taxonomicName>
|
||
[major ♂: Yemen, Ibb gov.].
|
||
<emphasis id="C1E2D510D322473E97CAF8DAFB72F8F1" bold="true" box="[1205,1238,1841,1865]" pageId="7" pageNumber="6">3a)</emphasis>
|
||
Dorsal habitus.
|
||
<emphasis id="C1E2D510D322473E9601F8DAFA06F8F1" bold="true" box="[1406,1442,1841,1865]" pageId="7" pageNumber="6">3b)</emphasis>
|
||
Parameres, lateral view.
|
||
<emphasis id="C1E2D510D322473E92C8F8A4FE73F8DF" bold="true" box="[439,471,1871,1895]" pageId="7" pageNumber="6">3c)</emphasis>
|
||
Parameres, dorsal view.
|
||
<emphasis id="C1E2D510D322473E91ACF8A4FD53F8DF" bold="true" box="[723,759,1871,1895]" pageId="7" pageNumber="6">3d)</emphasis>
|
||
Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Rs:
|
||
<emphasis id="C1E2D510D322473E939AF885FE9DF83E" box="[229,313,1902,1926]" italics="true" pageId="7" pageNumber="6">raspulae</emphasis>
|
||
. Photos by A. Degiovanni, edited by G. Fiumi [from
|
||
<bibRefCitation id="970774F3D322473E9030F885FB9FF83E" author="Ziani S & Keith D." box="[847,1083,1902,1926]" pageId="7" pageNumber="6" pagination="1 - 4" refId="ref35001" refString="Ziani S, Keith D. 2023. 3 rd contribution to the knowledge of Arabian Onthophagini (Coleoptera: Scarabaeidae). Onthophagus aeneopiceus d'Orbigny, 1902, a new record for Yemen and for the Palaearctic ecozone. Insecta Mundi 0974: 1 - 4." type="book chapter" year="2023">Ziani and Keith (2023)</bibRefCitation>
|
||
].
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F3290902D32D4731927FFF0DFD29FE9B" blockId="8.[192,1440,230,502]" pageId="8" pageNumber="7">Males with protibial spur clearly bent downward apically, and with a small denticle strongly curved downward on the inner angle of protibial apex.</paragraph>
|
||
<paragraph id="F3290902D32D4731927FFEC7FC83FE34" blockId="8.[192,1440,230,502]" pageId="8" pageNumber="7">
|
||
Female not examined. According to
|
||
<bibRefCitation id="970774F3D32D473191EEFEC7FCF6FEFE" author="d'Orbigny H." box="[657,850,300,326]" pageId="8" pageNumber="7" pagination="1 - 608" refId="ref27773" refString="d'Orbigny H. 1913. Synopsis des Onthophagides d'Afrique. Annales de la Societe Entomologique de France 82: 1 - 608." type="journal article" year="1913">d’Orbigny (1913)</bibRefCitation>
|
||
, females have the occipital carina extended in a backward bent lamina, placed between the eyes, in the middle of them, and a pronotal anteromedian transverse gibbosity, sometimes very slightly divided in two lobes.
|
||
</paragraph>
|
||
<paragraph id="F3290902D32D4731927FFE7EFB85FE4D" blockId="8.[192,1440,230,502]" pageId="8" pageNumber="7">
|
||
Male genital armature. Parameres short, apices bent ventrally, obviously diverging apically, without basolateral plate denticle, round at apex (
|
||
<figureCitation id="6BAD1587D32D47319120FE53FD68FE6A" box="[607,716,440,466]" captionStart="Figure 3" captionStartId="7.[192,261,1841,1865]" captionTargetBox="[219,1400,236,1780]" captionTargetId="figure-10@7.[194,1442,226,1792]" captionTargetPageId="7" captionText="Figure 3. Onthophagus (Indonthophagus) aeneopiceus d’Orbigny, 1902 [major ♂: Yemen, Ibb gov.]. 3a) Dorsal habitus. 3b) Parameres, lateral view. 3c) Parameres, dorsal view. 3d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Rs: raspulae. Photos by A. Degiovanni, edited by G. Fiumi [from Ziani and Keith (2023)]." figureDoi="http://doi.org/10.5281/zenodo.11450112" httpUri="https://zenodo.org/record/11450112/files/figure.png" pageId="8" pageNumber="7">Fig. 3b–c</figureCitation>
|
||
); endophallus with the presence of accessory endophallites, two evident raspulae and a bracket-shaped lamella copulatrix, very thin (
|
||
<figureCitation id="6BAD1587D32D473190BCFE30FBB0FE4D" box="[963,1044,475,501]" captionStart="Figure 3" captionStartId="7.[192,261,1841,1865]" captionTargetBox="[219,1400,236,1780]" captionTargetId="figure-10@7.[194,1442,226,1792]" captionTargetPageId="7" captionText="Figure 3. Onthophagus (Indonthophagus) aeneopiceus d’Orbigny, 1902 [major ♂: Yemen, Ibb gov.]. 3a) Dorsal habitus. 3b) Parameres, lateral view. 3c) Parameres, dorsal view. 3d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Rs: raspulae. Photos by A. Degiovanni, edited by G. Fiumi [from Ziani and Keith (2023)]." figureDoi="http://doi.org/10.5281/zenodo.11450112" httpUri="https://zenodo.org/record/11450112/files/figure.png" pageId="8" pageNumber="7">Fig. 3d</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BB8C5A89D32D473193BFFDEDFD44FDFC" pageId="8" pageNumber="7" type="distribution">
|
||
<paragraph id="F3290902D32D473193BFFDEDFD44FDFC" blockId="8.[192,1440,518,580]" pageId="8" pageNumber="7">
|
||
<emphasis id="C1E2D510D32D473193BFFDEDFEFFFD98" bold="true" box="[192,347,518,544]" pageId="8" pageNumber="7">Distribution.</emphasis>
|
||
<collectingCountry id="8B814992D32D4731921EFDEDFE0AFD98" box="[353,430,518,544]" name="Eritrea" pageId="8" pageNumber="7">Eritrea</collectingCountry>
|
||
(
|
||
<bibRefCitation id="970774F3D32D473192C2FDEDFDD5FD99" author="d'Orbigny H." box="[445,625,518,545]" pageId="8" pageNumber="7" pagination="1 - 324" refId="ref27623" refString="d'Orbigny H. 1902. Memoire sur les Onthophagides d'Afrique. Annales de la Societe Entomologique de France 71 (1 - 2): 1 - 324." type="journal article" year="1902">d’Orbigny 1902</bibRefCitation>
|
||
).
|
||
<collectingCountry id="8B814992D32D473191FBFDEDFD68FD98" box="[644,716,518,544]" name="Sudan" pageId="8" pageNumber="7">Sudan</collectingCountry>
|
||
,
|
||
<collectingCountry id="8B814992D32D473191A7FDEDFC9DFD98" box="[728,825,518,544]" name="Ethiopia" pageId="8" pageNumber="7">Ethiopia</collectingCountry>
|
||
,
|
||
<collectingCountry id="8B814992D32D4731903BFDECFC2EFD99" box="[836,906,519,545]" name="Kenya" pageId="8" pageNumber="7">Kenya</collectingCountry>
|
||
(
|
||
<bibRefCitation id="970774F3D32D473190E6FDEDFBE9FD99" author="d'Orbigny H." box="[921,1101,518,545]" pageId="8" pageNumber="7" pagination="1 - 608" refId="ref27773" refString="d'Orbigny H. 1913. Synopsis des Onthophagides d'Afrique. Annales de la Societe Entomologique de France 82: 1 - 608." type="journal article" year="1913">d’Orbigny 1913</bibRefCitation>
|
||
).
|
||
<collectingCountry id="8B814992D32D4731971FFDECFB47FD99" box="[1120,1251,519,545]" name="Ivory Coast" pageId="8" pageNumber="7">Ivory Coast</collectingCountry>
|
||
(
|
||
<bibRefCitation id="970774F3D32D4731978DFDEDFA38FD98" author="Balthasar V." box="[1266,1436,518,544]" pageId="8" pageNumber="7" pagination="59 - 73" refId="ref25804" refString="Balthasar V. 1969. Contributions a la connaissance de la faune entomologique de la Cote-d'Ivoire (J. Decelle, 1961 - 1964). XXXII. Coleoptera Scarabaeidae et Aphodiidae. Annales du Musee Royal de l'Afrique Centrale / Sciences Zoologiques 175: 59 - 73." type="journal article" year="1969">Balthasar 1969</bibRefCitation>
|
||
, doubtful record).
|
||
<collectingCountry id="8B814992D32D473192F8FDC1FE77FDFC" box="[391,467,554,580]" name="Yemen" pageId="8" pageNumber="7">Yemen</collectingCountry>
|
||
(
|
||
<bibRefCitation id="970774F3D32D4731929EFDC1FD70FDFC" author="Ziani S & Keith D." box="[481,724,554,580]" pageId="8" pageNumber="7" pagination="1 - 4" refId="ref35001" refString="Ziani S, Keith D. 2023. 3 rd contribution to the knowledge of Arabian Onthophagini (Coleoptera: Scarabaeidae). Onthophagus aeneopiceus d'Orbigny, 1902, a new record for Yemen and for the Palaearctic ecozone. Insecta Mundi 0974: 1 - 4." type="book chapter" year="2023">Ziani and Keith 2023</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BB8C5A89D32D473193BFFDBFFBFBFC41" pageId="8" pageNumber="7" type="materials_examined">
|
||
<paragraph id="F3290902D32D473193BFFDBFFB30FD0D" blockId="8.[192,1441,596,696]" pageId="8" pageNumber="7">
|
||
<emphasis id="C1E2D510D32D473193BFFDBFFE07FDD6" bold="true" box="[192,419,596,622]" pageId="8" pageNumber="7">Material examined.</emphasis>
|
||
<materialsCitation id="43FE035FD32D473192D5FDBFFA39FDD7" box="[426,1437,596,626]" country="Eritrea" location="Eritrea" pageId="8" pageNumber="7" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<emphasis id="C1E2D510D32D473192D5FDBFFE58FDD6" bold="true" box="[426,508,596,622]" pageId="8" pageNumber="7">
|
||
<collectingCountry id="8B814992D32D473192D5FDBFFE58FDD6" box="[426,508,596,622]" name="Eritrea" pageId="8" pageNumber="7">Eritrea</collectingCountry>
|
||
</emphasis>
|
||
: “Erythrée” [Eritrea], examined by photos (
|
||
<typeStatus id="2C2DB7A0D32D4731909CFDBEFBE3FDD7" box="[995,1095,597,623]" pageId="8" pageNumber="7" type="holotype">holotype</typeStatus>
|
||
<specimenCount id="E590C28BD32D47319730FDBDFBC0FDCA" box="[1103,1124,598,626]" count="1" pageId="8" pageNumber="7" type="male">♂</specimenCount>
|
||
of
|
||
<taxonomicName id="34967281D32D473197F7FDBEFA31FDD7" authority=", MNHN" authorityName="MNHN" box="[1160,1429,597,623]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="aeneopiceus">
|
||
<emphasis id="C1E2D510D32D473197F7FDBEFA8FFDD7" box="[1160,1323,597,623]" italics="true" pageId="8" pageNumber="7">O. aeneopiceus</emphasis>
|
||
, MNHN
|
||
</taxonomicName>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="43FE035FD32D473193BEFD93FBBEFD2A" box="[193,1050,632,661]" collectingDate="1976-03-30" collectionCode="DKCC" collectorName="R. Naviaux" country="Yemen" location="Jabal Hadish" pageId="8" pageNumber="7" specimenCount="1" specimenCount-male="1" stateProvince="Ibb" typeStatus="holotype">
|
||
<emphasis id="C1E2D510D32D473193BEFD93FEABFD2A" bold="true" box="[193,271,632,658]" pageId="8" pageNumber="7">
|
||
<collectingCountry id="8B814992D32D473193BEFD93FEABFD2A" box="[193,271,632,658]" name="Yemen" pageId="8" pageNumber="7">Yemen</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<collectingRegion id="3152C7E0D32D47319263FD93FEE6FD2A" box="[284,322,632,658]" country="Yemen" name="Ibb" pageId="8" pageNumber="7">Ibb</collectingRegion>
|
||
gov., W of
|
||
<location id="F6495FD9D32D473192C0FD93FDEBFD2A" LSID="urn:lsid:plazi:treatment:7B3FB814D323473193BFFC42FB06FB4F:F6495FD9D32D473192C0FD93FDEBFD2A" box="[447,591,632,658]" country="Yemen" name="Jabal Hadish" pageId="8" pageNumber="7" stateProvince="Ibb">Jabal Hadish</location>
|
||
,
|
||
<date id="87282FC2D32D47319125FD93FD6BFD2A" box="[602,719,632,658]" pageId="8" pageNumber="7" value="1976-03-30">
|
||
<collectingDate id="976CD62AD32D47319125FD93FD6BFD2A" box="[602,719,632,658]" pageId="8" pageNumber="7" value="1976-03-30">30.iii.1976</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="5E636CD4D32D473191A5FD93FCF1FD2A" box="[730,853,632,658]" pageId="8" pageNumber="7">R. Naviaux</collectorName>
|
||
leg.
|
||
<specimenCount id="E590C28BD32D473190F6FD93FC16FD2D" box="[905,946,632,661]" count="1" pageId="8" pageNumber="7" type="male">1 ♂</specimenCount>
|
||
(
|
||
<collectionCode id="958791C7D32D473190BFFD93FBB6FD2A" box="[960,1042,632,658]" pageId="8" pageNumber="7">DKCC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="43FE035FD32D4731975BFD93FB34FD0D" collectingDate="2005-10-26" collectingDateMax="2005-10-27" collectingDateMin="2005-10-26" collectionCode="NMPC" collectorName="D. Kral" country="Yemen" elevation="1600" latitude="13.874445" location="Jumai - Wadi Maytam" longLatPrecision="21" longitude="44.295834" pageId="8" pageNumber="7" specimenCount="1" specimenCount-male="1" stateProvince="Ibb" typeStatus="holotype">
|
||
<collectingRegion id="3152C7E0D32D4731975BFD93FBEEFD2A" box="[1060,1098,632,658]" country="Yemen" name="Ibb" pageId="8" pageNumber="7">Ibb</collectingRegion>
|
||
gov.,
|
||
<locationDeviation id="E29B1020D32D473197F6FD93FA8AFD2A" box="[1161,1326,632,659]" location="Najd" pageId="8" pageNumber="7">
|
||
<quantity id="346EA4E7D32D473197F6FD93FB66FD2A" box="[1161,1218,632,659]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="7" unit="km" value="5.0">5 km</quantity>
|
||
SW Najd
|
||
</locationDeviation>
|
||
al
|
||
<location id="F6495FD9D32D47319631FD93FEFAFD0E" LSID="urn:lsid:plazi:treatment:7B3FB814D323473193BFFC42FB06FB4F:F6495FD9D32D47319631FD93FEFAFD0E" country="Yemen" latitude="13.874445" longLatPrecision="21" longitude="44.295834" name="Jumai - Wadi Maytam" pageId="8" pageNumber="7" stateProvince="Ibb">Jumai - Wadi Maytam</location>
|
||
<quantity id="346EA4E7D32D4731921AFD70FE1EFD0E" box="[357,442,667,694]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="7" unit="m" value="1600.0">
|
||
<elevation id="78BBEE31D32D4731921AFD70FE1EFD0E" box="[357,442,667,694]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="7" unit="m" value="1600.0">1600 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="96A26FC5D32D473192BBFD70FDE1FD0E" box="[452,581,667,695]" degrees="13" direction="north" minutes="52" orientation="latitude" pageId="8" pageNumber="7" precision="15" seconds="28" value="13.874445">13°52’28″N</geoCoordinate>
|
||
<geoCoordinate id="96A26FC5D32D47319133FD70FD74FD0E" box="[588,720,667,694]" degrees="44" direction="east" minutes="17" orientation="longitude" pageId="8" pageNumber="7" precision="15" seconds="45" value="44.295834">44°17’45” E</geoCoordinate>
|
||
,
|
||
<date id="87282FC2D32D473191A5FD70FCC8FD0E" box="[730,876,667,694]" pageId="8" pageNumber="7" value="2005-10-26" valueMax="2005-10-27" valueMin="2005-10-26">
|
||
<collectingDate id="976CD62AD32D473191A5FD70FCC8FD0E" box="[730,876,667,694]" pageId="8" pageNumber="7" value="2005-10-26" valueMax="2005-10-27" valueMin="2005-10-26">26-27.x.2005</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="5E636CD4D32D47319008FD77FC6CFD0D" box="[887,968,667,694]" pageId="8" pageNumber="7">D. Král</collectorName>
|
||
leg.
|
||
<specimenCount id="E590C28BD32D47319084FD70FB87FD00" box="[1019,1059,667,696]" count="1" pageId="8" pageNumber="7" type="male">1 ♂</specimenCount>
|
||
(
|
||
<collectionCode id="958791C7D32D4731974DFD70FB2DFD0D" box="[1074,1161,667,693]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="8" pageNumber="7">NMPC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="F3290902D32D473193BFFD2DFBFBFC41" blockId="8.[190,1441,710,1017]" pageId="8" pageNumber="7">
|
||
<emphasis id="C1E2D510D32D473193BFFD2DFE2AFD58" bold="true" box="[192,398,710,736]" pageId="8" pageNumber="7">Historical review.</emphasis>
|
||
Despite
|
||
<taxonomicName id="34967281D32D4731928FFD2CFD37FD58" box="[496,659,711,737]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="aeneopiceus">
|
||
<emphasis id="C1E2D510D32D4731928FFD2CFD37FD58" box="[496,659,711,737]" italics="true" pageId="8" pageNumber="7">O. aeneopiceus</emphasis>
|
||
</taxonomicName>
|
||
being described more than a century ago, its nomenclatorial and systematic history is relatively poor. According to
|
||
<bibRefCitation id="970774F3D32D473191BCFD02FC27FCBB" author="d'Orbigny H." box="[707,899,745,771]" pageId="8" pageNumber="7" pagination="1 - 324" refId="ref27623" refString="d'Orbigny H. 1902. Memoire sur les Onthophagides d'Afrique. Annales de la Societe Entomologique de France 71 (1 - 2): 1 - 324." type="journal article" year="1902">d’Orbigny (1902)</bibRefCitation>
|
||
, the
|
||
<typeStatus id="2C2DB7A0D32D473190CAFD02FBBDFCBB" box="[949,1049,745,771]" pageId="8" pageNumber="7" type="holotype">holotype</typeStatus>
|
||
, a male, the sole available specimen for the description, was collected in
|
||
<collectingCountry id="8B814992D32D47319137FCE7FD3CFC9E" box="[584,664,780,806]" name="Eritrea" pageId="8" pageNumber="7">Eritrea</collectingCountry>
|
||
. Eleven years later,
|
||
<bibRefCitation id="970774F3D32D47319017FCE7FB8CFC9E" author="d'Orbigny H." box="[872,1064,780,806]" pageId="8" pageNumber="7" pagination="1 - 608" refId="ref27773" refString="d'Orbigny H. 1913. Synopsis des Onthophagides d'Afrique. Annales de la Societe Entomologique de France 82: 1 - 608." type="journal article" year="1913">d’Orbigny (1913)</bibRefCitation>
|
||
, evidently after examining further material, specified the presence of a long curved backward cephalic horn in major males, described the female and added other localities from three more countries.
|
||
<bibRefCitation id="970774F3D32D4731904BFCB9FC55FCD4" author="Balthasar V." box="[820,1009,850,876]" pageId="8" pageNumber="7" refId="ref25682" refString="Balthasar V. 1963. Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera Lamellicornia. Band 2. Coprinae (Onitini, Oniticellini, Onthophagini). Tschechoslowakischen Akademie der Wissenschaften; Prague. 627 p. + 16 pls." type="book" year="1963">Balthasar (1963)</bibRefCitation>
|
||
, whilst specifying that
|
||
<taxonomicName id="34967281D32D47319784FCB8FA04FCD4" box="[1275,1440,851,877]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="aeneopiceus">
|
||
<emphasis id="C1E2D510D32D47319784FCB8FA04FCD4" box="[1275,1440,851,877]" italics="true" pageId="8" pageNumber="7">O. aeneopiceus</emphasis>
|
||
</taxonomicName>
|
||
was an Afrotropical species, hypothesized its presence also in the Palaearctic Africa.
|
||
<bibRefCitation id="970774F3D32D47319701FC9DFA9CFC28" author="Balthasar V." box="[1150,1336,886,912]" pageId="8" pageNumber="7" pagination="59 - 73" refId="ref25804" refString="Balthasar V. 1969. Contributions a la connaissance de la faune entomologique de la Cote-d'Ivoire (J. Decelle, 1961 - 1964). XXXII. Coleoptera Scarabaeidae et Aphodiidae. Annales du Musee Royal de l'Afrique Centrale / Sciences Zoologiques 175: 59 - 73." type="journal article" year="1969">Balthasar (1969)</bibRefCitation>
|
||
reported the species for
|
||
<collectingCountry id="8B814992D32D47319218FC72FE4FFC0B" box="[359,491,921,947]" name="Ivory Coast" pageId="8" pageNumber="7">Ivory Coast</collectingCountry>
|
||
, but this record, completely off the known distribution area of the species, was rightly regarded dubiously by
|
||
<bibRefCitation id="970774F3D32D473192BCFC57FD61FC6E" author="Schoolmeesters P." box="[451,709,956,982]" pageId="8" pageNumber="7" refId="ref32957" refString="Schoolmeesters P. 2023. World Scarabaeidae Database. In: Banki O, Roskov Y, Doring M, Ower G, Hernandez Robles DR, Plata Corredor CA, Stjernegaard Jeppesen T, Orn A, Vandepitte L, Hobern D, Schalk P, DeWalt RE, Keping M, Miller J, Orrell T, Aalbu R, Abbott J, Adlard R, Adriaenssens EM, et al. (eds.). Catalogue of Life Checklist (Version 2023 - 05 - 30). https: // doi. org / 10.48580 / dfsr- 38 g" type="book" year="2023">Schoolmeesters (2023)</bibRefCitation>
|
||
. Lastly,
|
||
<bibRefCitation id="970774F3D32D4731905FFC57FB81FC6E" author="Ziani S & Keith D." box="[800,1061,956,982]" pageId="8" pageNumber="7" pagination="1 - 4" refId="ref35001" refString="Ziani S, Keith D. 2023. 3 rd contribution to the knowledge of Arabian Onthophagini (Coleoptera: Scarabaeidae). Onthophagus aeneopiceus d'Orbigny, 1902, a new record for Yemen and for the Palaearctic ecozone. Insecta Mundi 0974: 1 - 4." type="book chapter" year="2023">Ziani and Keith (2023)</bibRefCitation>
|
||
published the first record for the Palaearctic ecozone (
|
||
<emphasis id="C1E2D510D32D473192D5FC0BFE42FC41" box="[426,486,992,1017]" italics="true" pageId="8" pageNumber="7">sensu</emphasis>
|
||
<bibRefCitation id="970774F3D32D47319293FC34FD63FC41" author="Lobl I & Lobl D." box="[492,711,991,1017]" pageId="8" pageNumber="7" refId="ref31167" refString="Lobl I, Lobl D. 2016. Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition. Brill; Leiden, Boston. 983 p. https: // doi. org / 10.1163 / 9789004309142" type="book" year="2016">Löbl and Löbl 2016</bibRefCitation>
|
||
) after
|
||
<specimenCount id="E590C28BD32D47319072FC34FCDBFC41" box="[781,895,991,1017]" count="2" pageId="8" pageNumber="7" type="male">two males</specimenCount>
|
||
collected in
|
||
<collectingCountry id="8B814992D32D47319774FC34FBFFFC41" box="[1035,1115,991,1017]" name="Yemen" pageId="8" pageNumber="7">Yemen</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BB8C5A89D32D473193BFFBE1FB06FB4F" pageId="8" pageNumber="7" type="discussion">
|
||
<paragraph id="F3290902D32D473193BFFBE1FC80FBD3" blockId="8.[191,1440,1034,1271]" pageId="8" pageNumber="7">
|
||
<emphasis id="C1E2D510D32D473193BFFBE1FE8BFB9C" bold="true" box="[192,303,1034,1060]" pageId="8" pageNumber="7">Remarks.</emphasis>
|
||
<bibRefCitation id="970774F3D32D47319248FBE1FD9AFB9C" author="Ziani S & Keith D." box="[311,574,1034,1060]" pageId="8" pageNumber="7" pagination="1 - 4" refId="ref35001" refString="Ziani S, Keith D. 2023. 3 rd contribution to the knowledge of Arabian Onthophagini (Coleoptera: Scarabaeidae). Onthophagus aeneopiceus d'Orbigny, 1902, a new record for Yemen and for the Palaearctic ecozone. Insecta Mundi 0974: 1 - 4." type="book chapter" year="2023">Ziani and Keith (2023)</bibRefCitation>
|
||
, although preferring not to insert
|
||
<taxonomicName id="34967281D32D473190B6FBE1FBC9FB9C" box="[969,1133,1034,1060]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="aeneopiceus">
|
||
<emphasis id="C1E2D510D32D473190B6FBE1FBC9FB9C" box="[969,1133,1034,1060]" italics="true" pageId="8" pageNumber="7">O. aeneopiceus</emphasis>
|
||
</taxonomicName>
|
||
in any known subgenus of
|
||
<taxonomicName id="34967281D32D473193BDFBC6FEF7FBFF" authorityName="Latreille" authorityYear="1802" box="[194,339,1069,1095]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C1E2D510D32D473193BDFBC6FEF7FBFF" box="[194,339,1069,1095]" italics="true" pageId="8" pageNumber="7">Onthophagus</emphasis>
|
||
</taxonomicName>
|
||
, pointed out the similarity between the lamella copulatrix and the overall structures inside the endophallus of
|
||
<taxonomicName id="34967281D32D47319213FBBAFDAAFBD2" box="[364,526,1105,1131]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="aeneopiceus">
|
||
<emphasis id="C1E2D510D32D47319213FBBAFDAAFBD2" box="[364,526,1105,1131]" italics="true" pageId="8" pageNumber="7">O. aeneopiceus</emphasis>
|
||
</taxonomicName>
|
||
with those of
|
||
<taxonomicName id="34967281D32D473191D3FBBAFCBAFBD2" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[684,798,1105,1131]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="mopsus">
|
||
<emphasis id="C1E2D510D32D473191D3FBBAFCBAFBD2" box="[684,798,1105,1131]" italics="true" pageId="8" pageNumber="7">O. mopsus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="F3290902D32D4731927FFB9FFB06FB4F" blockId="8.[191,1440,1034,1271]" pageId="8" pageNumber="7">
|
||
It is noteworthy that
|
||
<taxonomicName id="34967281D32D47319299FB9FFD23FB35" box="[486,647,1140,1166]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="aeneopiceus">
|
||
<emphasis id="C1E2D510D32D47319299FB9FFD23FB35" box="[486,647,1140,1166]" italics="true" pageId="8" pageNumber="7">O. aeneopiceus</emphasis>
|
||
</taxonomicName>
|
||
is, so far, the sole species belonging to the group in which the male protibial spur is sinuate, clearly bent downward apically. This characteristic is common in those
|
||
<taxonomicName id="34967281D32D473197C7FB7DFA04FB08" box="[1208,1440,1174,1200]" pageId="8" pageNumber="7">
|
||
<emphasis id="C1E2D510D32D473197C7FB7DFAEFFB08" box="[1208,1355,1174,1200]" italics="true" pageId="8" pageNumber="7">Onthophagus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
somehow associated with burrows or nests of small mammals (
|
||
<bibRefCitation id="970774F3D32D47319006FB52FB2AFB6C" author="Ziani S & Gudenzi I." box="[889,1166,1209,1236]" pageId="8" pageNumber="7" pagination="29 - 44" refId="ref34954" refString="Ziani S, Gudenzi I. 2009. Studies on palearctic Onthophagus associated with burrows of small mammals. III. O. aerarius and closely related species (Coleoptera Scarabaeidae). Bollettino della Societa Entomologica Italiana 141 (1): 29 - 44." type="journal article" year="2009">Ziani and Gudenzi 2009</bibRefCitation>
|
||
). Only further investigation will be able to clarify whether the biology of this species is connected to pholeophily.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |