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<document id="580CD3D8D08C15B1152F4F04A9BD02D6" ID="10.11646/zootaxa.4268.4.3SLASH11070" ID-CLB-Dataset="33024" ID-DOI="10.11646/zootaxa.4268.4.3" ID-GBIF-Dataset="dc91aa5e-ab23-4599-8df9-56ebda03c098" ID-ISSN="1175-5326" ID-Zenodo-Dep="580926" ID-ZooBank="71929131-1CB7-4FF0-8A80-E062F1C00960" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1495095107143" checkinUser="plazi" docAuthor="Lee, Seung-Gyu &amp; Ahn, Kee-Jeong" docDate="2017" docId="7B3A87E8FFD7C96DFF2BF9F0FEE05EB2" docLanguage="en" docName="zootaxa.4268.4.3SLASH11070.pdf" docOrigin="Zootaxa 4268 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Atheta (Datomicra) pyongangsani Pasnik 2001" docType="treatment" docVersion="7" lastPageNumber="517" masterDocId="8703FF90FFDFC964FFBCFFB5FFCB5C36" masterDocTitle="A taxonomic review of the Korean species of the subgenus Datomicra Mulsant &amp; Rey of the genus Atheta Thomson, with description of a new species (Coleoptera, Staphylinidae, Aleocharinae)" masterLastPageNumber="522" masterPageNumber="508" pageNumber="516" updateTime="1698439703284" updateUser="ExternalLinkService">
<mods:mods id="A4F587E70AB2FA47E3D6B08E3E2E6CB2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="81415AC36BB2394E98F100AD1155A53A">A taxonomic review of the Korean species of the subgenus Datomicra Mulsant &amp; Rey of the genus Atheta Thomson, with description of a new species (Coleoptera, Staphylinidae, Aleocharinae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="873A987386DFAB0A3675748606F4ADAB">Lee, Seung-Gyu</mods:namePart>
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<mods:namePart id="FE7EC154159C85003E5F4AC62A7FE4F4">Ahn, Kee-Jeong</mods:namePart>
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<mods:typeOfResource id="87CA6A90190305883DA3C180F138C8F5">text</mods:typeOfResource>
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<mods:title id="6872AA6B18FC8C159EFC0EC0B88EE8D7">Zootaxa</mods:title>
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<mods:part id="D812DB2281DF330FA6B09DB33C4506AB">
<mods:date id="7B432108343254B1C5D957E5D9294C3E">2017</mods:date>
<mods:detail id="61CA4E38A62F88BC03C5E4672E948030" type="volume">
<mods:number id="D0E88DCCDF12184279AFDD644D829FC8">4268</mods:number>
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<mods:classification id="FAF4DB79D6EE1F0C21950D83ECBC87C2">journal article</mods:classification>
<mods:identifier id="368C76F34D3AF5BDB468627F3D2A3F8B" type="CLB-Dataset">33024</mods:identifier>
<mods:identifier id="37467446114AD6BF17C46D6ABDF4616B" type="DOI">10.11646/zootaxa.4268.4.3</mods:identifier>
<mods:identifier id="BB4F51CED7D36C185B2339689633983B" type="GBIF-Dataset">dc91aa5e-ab23-4599-8df9-56ebda03c098</mods:identifier>
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<mods:identifier id="736A5C2D085C49BAEDBB8E2BD41154AC" type="Zenodo-Dep">580926</mods:identifier>
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<treatment id="7B3A87E8FFD7C96DFF2BF9F0FEE05EB2" ID-DOI="http://doi.org/10.5281/zenodo.6038295" ID-GBIF-Taxon="130810281" ID-Zenodo-Dep="6038295" LSID="urn:lsid:plazi:treatment:7B3A87E8FFD7C96DFF2BF9F0FEE05EB2" httpUri="http://treatment.plazi.org/id/7B3A87E8FFD7C96DFF2BF9F0FEE05EB2" lastPageId="9" lastPageNumber="517" pageId="8" pageNumber="516">
<subSubSection id="BB896575FFD7C96CFF2BF9F0FF315AB6" pageId="8" pageNumber="516" type="nomenclature">
<paragraph id="F32C36FEFFD7C96CFF2BF9F0FD135A69" blockId="8.[151,728,1605,1664]" box="[151,728,1605,1631]" pageId="8" pageNumber="516">
<heading id="A8648192FFD7C96CFF2BF9F0FD135A69" bold="true" box="[151,728,1605,1631]" fontSize="11" level="1" pageId="8" pageNumber="516" reason="1">
<taxonomicName id="34934D7DFFD7C96CFF2BF9F0FD135A69" ID-CoL="JC8C" authority="Pasnik, 2001" authorityName="Pasnik" authorityYear="2001" box="[151,728,1605,1631]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="516" phylum="Arthropoda" rank="species" species="pyongangsani" subGenus="Datomicra">
<emphasis id="C1E7EAECFFD7C96CFF2BF9F0FD135A69" bold="true" box="[151,728,1605,1631]" pageId="8" pageNumber="516">
<emphasis id="C1E7EAECFFD7C96CFF2BF9F0FF2D5A69" bold="true" box="[151,230,1605,1631]" italics="true" pageId="8" pageNumber="516">Atheta</emphasis>
(
<emphasis id="C1E7EAECFFD7C96CFF4BF9F0FEB35A69" bold="true" box="[247,376,1605,1631]" italics="true" pageId="8" pageNumber="516">Datomicra</emphasis>
)
<emphasis id="C1E7EAECFFD7C96CFE35F9F0FDF95A69" bold="true" box="[393,562,1605,1631]" italics="true" pageId="8" pageNumber="516">pyongangsani</emphasis>
<bibRefCitation id="97024B0FFFD7C96CFD86F9F0FD135A69" author="Pasnik" box="[570,728,1605,1631]" pageId="8" pageNumber="516" refString="Pasnik, G. (2001) The North Korean Aleocharinae (Coleoptera, Staphylinidae): diversity and biogeography. Acta Zoologica Cracoviensia, 44, 185 - 234." type="journal article" year="2001">Paśnik, 2001</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="F32C36FEFFD7C96CFF2BF9D2FF315AB6" blockId="8.[151,728,1605,1664]" box="[151,250,1639,1664]" pageId="8" pageNumber="516">
(
<figureCitation id="6BA82A7BFFD7C96CFF23F9D2FF155AB6" box="[159,222,1639,1664]" captionStart="FIGURE 1" captionStartId="2.[151,250,1706,1728]" captionTargetBox="[387,1202,741,1685]" captionTargetId="figure@2.[384,1203,737,1685]" captionTargetPageId="2" captionText="FIGURE 1. Habitus. A, Atheta (Datomicra) celata, 1.8 mm; B, A. (D.) dadopora, 1.7 mm; C, A. (D.) dentiventris, 2.3 mm; D, A. (D.) lewisiana, 1.8 mm; E, Atheta (D.) nigra, 1.7 mm; F, A. (D.) pyongangsani, 2.0 mm; G, A. (D.) semidentiventris sp. nov., 2.1 mm; H, A. (D.) sordidula, 1.6 mm." httpUri="https://zenodo.org/record/580927/files/figure.png" pageId="8" pageNumber="516">Fig. 1</figureCitation>
F)
</paragraph>
</subSubSection>
<subSubSection id="BB896575FFD7C96CFF2BF905FB1D5AF0" box="[151,1238,1712,1734]" pageId="8" pageNumber="516" type="reference_group">
<paragraph id="F32C36FEFFD7C96CFF2BF905FB1D5AF0" blockId="8.[151,1238,1712,1734]" box="[151,1238,1712,1734]" pageId="8" pageNumber="516">
<treatmentCitationGroup id="D38311D0FFD7C96CFF2BF905FB1D5AF0" box="[151,1238,1712,1734]" pageId="8" pageNumber="516">
<taxonomicName id="34934D7DFFD7C96CFF2BF905FD795AF0" ID-CoL="JC8C" authority="Pasnik, 2001: 215" authorityName="Pasnik" authorityPageNumber="215" authorityYear="2001" box="[151,690,1712,1734]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="516" phylum="Arthropoda" rank="species" species="pyongangsani" subGenus="Datomicra">
<emphasis id="C1E7EAECFFD7C96CFF2BF905FF115AF0" box="[151,218,1712,1734]" italics="true" pageId="8" pageNumber="516">Atheta</emphasis>
(
<emphasis id="C1E7EAECFFD7C96CFF54F904FE9E5AF0" box="[232,341,1713,1734]" italics="true" pageId="8" pageNumber="516">Datomicra</emphasis>
)
<emphasis id="C1E7EAECFFD7C96CFED8F904FE395AF0" box="[356,498,1713,1734]" italics="true" pageId="8" pageNumber="516">pyongangsani</emphasis>
<treatmentCitation id="723210EFFFD7C96CFE44F905FD795AF0" author="Pasnik" box="[504,690,1712,1734]" page="215" pageId="8" pageNumber="516" year="2001">
<bibRefCitation id="97024B0FFFD7C96CFE44F905FDB45AF0" author="Pasnik" box="[504,639,1712,1734]" pageId="8" pageNumber="516" refString="Pasnik, G. (2001) The North Korean Aleocharinae (Coleoptera, Staphylinidae): diversity and biogeography. Acta Zoologica Cracoviensia, 44, 185 - 234." type="journal article" year="2001">Paśnik, 2001</bibRefCitation>
: 215
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="723210EFFFD7C96CFD00F905FC425AF0" author="Smetana" box="[700,905,1712,1734]" page="379" pageId="8" pageNumber="516" year="2004">
<bibRefCitation id="97024B0FFFD7C96CFD00F905FC9C5AF0" author="Smetana" box="[700,855,1712,1734]" pageId="8" pageNumber="516" refString="Smetana, A. (2004) Family Staphylinidae: subfamily Aleocharinae Fleming, 1821. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Histeroidea - Staphylinoidea. Apollo Books, Stenstrup, pp. 353 - 494." type="book chapter" year="2004">Smetana, 2004</bibRefCitation>
: 379
</treatmentCitation>
;
<treatmentCitation id="723210EFFFD7C96CFC2FF905FB195AF0" author="Schulke" box="[915,1234,1712,1734]" page="528" pageId="8" pageNumber="516" year="2015">
<bibRefCitation id="97024B0FFFD7C96CFC2FF905FB555AF0" author="Schulke" box="[915,1182,1712,1734]" pageId="8" pageNumber="516" refString="Schulke, M. &amp; Smetana, A. (2015) Family Staphylinidae: subfamily Aleocharinae Fleming, 1821. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2 / 1. Revised and updated edition. Hydrophiloidea - Staphylinoidea. Brill, Leiden, pp. 490 - 729." type="journal article" year="2015">Schülke &amp; Smetana, 2015</bibRefCitation>
: 528
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BB896575FFD7C96DFF2BF947FEC55D9B" lastPageId="9" lastPageNumber="517" pageId="8" pageNumber="516" type="diagnosis">
<paragraph id="F32C36FEFFD7C96DFF2BF947FEC55D9B" blockId="8.[151,1437,1778,2019]" lastBlockId="9.[151,1437,151,645]" lastPageId="9" lastPageNumber="517" pageId="8" pageNumber="516">
<emphasis id="C1E7EAECFFD7C96CFF2BF947FEDA5B3D" bold="true" box="[151,273,1778,1803]" pageId="8" pageNumber="516">Diagnosis.</emphasis>
Length 2.0–
<quantity id="346B9B1BFFD7C96CFE18F946FE345B3C" box="[420,511,1779,1803]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="516" unit="mm" value="2.2">2.2 mm</quantity>
. Body (
<figureCitation id="6BA82A7BFFD7C96CFDE6F947FD515B3D" box="[602,666,1778,1803]" captionStart="FIGURE 1" captionStartId="2.[151,250,1706,1728]" captionTargetBox="[387,1202,741,1685]" captionTargetId="figure@2.[384,1203,737,1685]" captionTargetPageId="2" captionText="FIGURE 1. Habitus. A, Atheta (Datomicra) celata, 1.8 mm; B, A. (D.) dadopora, 1.7 mm; C, A. (D.) dentiventris, 2.3 mm; D, A. (D.) lewisiana, 1.8 mm; E, Atheta (D.) nigra, 1.7 mm; F, A. (D.) pyongangsani, 2.0 mm; G, A. (D.) semidentiventris sp. nov., 2.1 mm; H, A. (D.) sordidula, 1.6 mm." httpUri="https://zenodo.org/record/580927/files/figure.png" pageId="8" pageNumber="516">Fig. 1</figureCitation>
F) dark brown; elytra paler than other parts; legs yellowish brown.
<emphasis id="C1E7EAECFFD7C96CFF2BF8A3FF1F5B19" box="[151,212,1814,1839]" italics="true" pageId="8" pageNumber="516">Head</emphasis>
. Slightly transverse, approximately 1.1–1.2 times as wide as long, widest behind eyes, slightly narrower than pronotum; eyes large and prominent, about 1.3–1.4 times as long as temples; infraorbital carina complete. Antennae slightly longer than head and pronotum combined; antennomeres 1–3 elongate, 1 longest, 4–10 subquadrate to slightly transverse, 11 longer than wide, about as long as preceding two combined.
<emphasis id="C1E7EAECFFD7C96CFAF8F837FA5E5BAD" box="[1348,1429,1922,1947]" italics="true" pageId="8" pageNumber="516">Thorax</emphasis>
. Pronotum approximately 1.3–1.4 times as wide as long, widest in apical third; pubescence directed anteriorly in midline; hypomera partially visible in lateral aspect. Elytra slightly transverse, slightly wider than pronotum; elytron approximately 1.6–1.7 times as long as wide, postero-lateral margin almost straight.
<emphasis id="C1E7EAECFFD6C96DFB7FFF22FAE65C86" box="[1219,1325,151,176]" italics="true" pageId="9" pageNumber="517">Abdomen</emphasis>
. Parallelsided, widest at middle; surface glossy and densely pubescent, with coarse punctures and transverse microsculpture; male tergite VIII with four macrosetae on each side of midline, posterior margin broadly rounded, slightly emarginate in median region; male sternite VIII with eight macrosetae on each side of midline, posterior margin convex, convergent apically; posterior margin of female tergite VIII truncate in median region; posterior margin of female sternite VIII broadly rounded, slightly emarginate in median region.
<emphasis id="C1E7EAECFFD6C96DFBE1FEF9FB015D53" box="[1117,1226,332,357]" italics="true" pageId="9" pageNumber="517">Aedeagus</emphasis>
. Apical process of median lobe narrowly elongate, slightly pointed apically in ventral aspect.
<emphasis id="C1E7EAECFFD6C96DFC63FEDAFBBA5DBE" box="[991,1137,367,392]" italics="true" pageId="9" pageNumber="517">Spermatheca</emphasis>
.
<collectingRegion id="3157F81CFFD6C96DFB3CFEC5FB0A5DB1" box="[1152,1217,368,391]" country="Turkey" name="Bursa" pageId="9" pageNumber="517">Bursa</collectingRegion>
without umbilicus; duct short.
</paragraph>
</subSubSection>
<subSubSection id="BB896575FFD6C96DFF7BFE02FC395E2E" pageId="9" pageNumber="517" type="materials_examined">
<paragraph id="F32C36FEFFD6C96DFF7BFE02FC395E2E" blockId="9.[151,1437,151,645]" pageId="9" pageNumber="517">
<materialsCitation id="43FB3CA3FFD6C96DFF7BFE02FA535DE6" ID-GBIF-Occurrence="1503373047" box="[199,1432,439,464]" collectingDate="1974-06-21" country="South Korea" location="Korea" pageId="9" pageNumber="517" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="C1E7EAECFFD6C96DFF7BFE02FE4A5DE6" bold="true" box="[199,385,439,464]" pageId="9" pageNumber="517">
<typeStatus id="2C28885CFFD6C96DFF7BFE02FEC95DE6" box="[199,258,439,464]" pageId="9" pageNumber="517">Type</typeStatus>
material.
</emphasis>
<typeStatus id="2C28885CFFD6C96DFE28FE0DFE345DE6" box="[404,511,440,464]" pageId="9" pageNumber="517" type="holotype">Holotype</typeStatus>
,
<specimenCount id="E595FD77FFD6C96DFDABFE0DFDE55DF9" box="[535,558,440,463]" pageId="9" pageNumber="517" type="male">♂</specimenCount>
, labeled as follows: ‘
<collectingCountry id="8B84766EFFD6C96DFCF6FE0DFC585DE6" box="[842,915,440,464]" name="South Korea" pageId="9" pageNumber="517">Korea</collectingCountry>
<collectingDate id="9769E9D6FFD6C96DFC1BFE0DFBDC5DE6" box="[935,1047,440,464]" pageId="9" pageNumber="517" value="1974-06-21">
<date id="872D103EFFD6C96DFC1BFE0DFBDC5DE6" box="[935,1047,440,464]" pageId="9" pageNumber="517" value="1974-06-21">21.6.1974</date>
</collectingDate>
Pnjongjang Exp. Inst.Zool.Cr.
</materialsCitation>
;
<materialsCitation id="43FB3CA3FFD6C96DFF2BFE69FDD75E2E" ID-GBIF-Occurrence="1503373033" collectingDate="1974-06-21" collectionCode="ISEA" country="North Korea" determinerName="G. Pasnik" location="Pyeongannam Prov." municipality="Pyeongyang-si" pageId="9" pageNumber="517" specimenCount="1" typeStatus="holotype">
<typeStatus id="2C28885CFFD6C96DFF2BFE69FEE05DC2" box="[151,299,476,500]" pageId="9" pageNumber="517" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="34934D7DFFD6C96DFE89FE69FD715DC3" box="[309,698,476,501]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="517" phylum="Arthropoda" rank="species" species="pyongangsani" status="sp. nov." subGenus="Datomicra">Atheta (Datomicra) pyongangsani</taxonomicName>
<taxonomicNameLabel id="DAD45797FFD6C96DFD78FE68FCCF5DC2" box="[708,772,477,500]" pageId="9" pageNumber="517" rank="species">sp. n.</taxonomicNameLabel>
det.
<determinerName id="06F0DA3AFFD6C96DFCFEFE69FC7B5DC3" box="[834,944,476,501]" pageId="9" pageNumber="517">G. Paśnik</determinerName>
2000’ [
<collectingCountry id="8B84766EFFD6C96DFBB0FE69FB685DC2" box="[1036,1187,476,501]" name="North Korea" pageId="9" pageNumber="517">North Korea</collectingCountry>
,
<location id="F64C6025FFD6C96DFB0EFE68FA535DC2" LSID="urn:lsid:plazi:treatment:7B3A87E8FFD7C96DFF2BF9F0FEE05EB2:F64C6025FFD6C96DFB0EFE68FA535DC2" box="[1202,1432,477,501]" country="North Korea" municipality="Pyeongyang-si" name="Pyeongannam Prov." pageId="9" pageNumber="517">Pyeongannam Prov.</location>
,
<collectingMunicipality id="1348AC84FFD6C96DFF2BFE4AFE8B5E2E" box="[151,320,511,536]" pageId="9" pageNumber="517">Pyeongyang-si</collectingMunicipality>
,
<collectingDate id="9769E9D6FFD6C96DFEF7FDB5FE0D5E2E" box="[331,454,511,536]" pageId="9" pageNumber="517" value="1974-06-21">
<date id="872D103EFFD6C96DFEF7FDB5FE0D5E2E" box="[331,454,511,536]" pageId="9" pageNumber="517" value="1974-06-21">21 vi 1974</date>
</collectingDate>
,
<collectionCode id="9582AE3BFFD6C96DFE6DFDB5FDDE5E2E" box="[465,533,512,536]" pageId="9" pageNumber="517">ISEA</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="43FB3CA3FFD6C96DFD9AFDB5FC395E2E" ID-GBIF-Occurrence="1503373051" box="[550,1010,511,536]" collectingDate="1974-06-21" collectionCode="ISEA" country="North Korea" determinerName="G. Pasnik" location="Pyeongannam Prov." municipality="Pyeongyang-si" pageId="9" pageNumber="517" specimenCount="1" typeStatus="paratype">
<typeStatus id="2C28885CFFD6C96DFD9AFDB5FD5D5E2E" box="[550,662,512,536]" pageId="9" pageNumber="517" type="paratype">Paratypes</typeStatus>
, 2 exx., same data as holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="BB896575FFD6C96DFF7BFD91FDC55E0B" box="[199,526,548,573]" pageId="9" pageNumber="517" type="distribution">
<paragraph id="F32C36FEFFD6C96DFF7BFD91FDC55E0B" blockId="9.[151,1437,151,645]" box="[199,526,548,573]" pageId="9" pageNumber="517">
<emphasis id="C1E7EAECFFD6C96DFF7BFD91FEAB5E0B" bold="true" box="[199,352,548,573]" pageId="9" pageNumber="517">Distribution.</emphasis>
<collectingCountry id="8B84766EFFD6C96DFEDBFD90FE605E0A" box="[359,427,549,572]" name="South Korea" pageId="9" pageNumber="517">Korea</collectingCountry>
(North).
</paragraph>
</subSubSection>
<subSubSection id="BB896575FFD6C96DFF7BFDF2FEE05EB2" pageId="9" pageNumber="517" type="discussion">
<paragraph id="F32C36FEFFD6C96DFF7BFDF2FEE05EB2" blockId="9.[151,1437,151,645]" pageId="9" pageNumber="517">
<emphasis id="C1E7EAECFFD6C96DFF7BFDF2FEF05E56" bold="true" box="[199,315,583,608]" pageId="9" pageNumber="517">Remarks.</emphasis>
This species was described by
<bibRefCitation id="97024B0FFFD6C96DFD26FDF2FCFC5E56" author="Pasnik" box="[666,823,583,608]" pageId="9" pageNumber="517" refString="Pasnik, G. (2001) The North Korean Aleocharinae (Coleoptera, Staphylinidae): diversity and biogeography. Acta Zoologica Cracoviensia, 44, 185 - 234." type="journal article" year="2001">Paśnik (2001)</bibRefCitation>
in North Korea and no specimen has been collected in South Korea.
</paragraph>
</subSubSection>
</treatment>
</document>
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