treatments-xml/data/7B/2F/87/7B2F87B2F621FA52D18171CFFCE2FD4C.xml

<document id="0F2B74703022B363721E99B4A86CAB99" ID="10.11646/zootaxa.4254.3.4SLASH10848" ID-CLB-Dataset="33208" ID-DOI="10.11646/zootaxa.4254.3.4" ID-GBIF-Dataset="a9d4b1f4-fa63-42bb-881e-53514379d22c" ID-ISSN="1175-5326" ID-Zenodo-Dep="546840" ID-ZooBank="CA94CE44-D17E-4501-801B-734A427663AF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1492502379480" checkinUser="plazi" docAuthor="Joya, Daniel Chirivi" docDate="2017" docId="7B2F87B2F621FA52D18171CFFCE2FD4C" docLanguage="en" docName="zootaxa.4254.3.4SLASH10848.pdf" docOrigin="Zootaxa 4254 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phrynus calypso Joya, 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="365" masterDocId="8716FFCAF623FA5AD1167158FFABFFC8" masterDocTitle="A new species of Phrynus Lamarck, 1801 (Arachnida: Amblypygi), from Trinidad and Tobago, and Venezuela, with a redescription of Phrynus pulchripes (Pocock, 1894)" masterLastPageNumber="374" masterPageNumber="357" pageNumber="359" updateTime="1698438918520" updateUser="ExternalLinkService">
<mods:mods id="592030EC7C21F4F946C295B9EBDEF69A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="8C8BEBB9C63C051B38D64A239BD3042E">A new species of Phrynus Lamarck, 1801 (Arachnida: Amblypygi), from Trinidad and Tobago, and Venezuela, with a redescription of Phrynus pulchripes (Pocock, 1894)</mods:title>
</mods:titleInfo>
<mods:name id="DA50FE320A3C7E4D163738BE78498990" type="personal">
<mods:role id="065C41A6BBD9FF55D7946C35766CE8D2">
<mods:roleTerm id="EA249C13DD0006094A1312BA3BD0ACE9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C6132AB62A83B28E7077E7559BA31613">Joya, Daniel Chirivi</mods:namePart>
</mods:name>
<mods:typeOfResource id="33C2644B3897FC6BA3167C74F4AF4CAE">text</mods:typeOfResource>
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<mods:titleInfo id="6529A3BC859505740AD353DC39742DE9">
<mods:title id="8A89BB86F026A956044F820384523ACA">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B18F1AF2971094C79951549E32EDBBFE">
<mods:date id="8E3A1D0F4BF29ECDEDF3204E9F525D3C">2017</mods:date>
<mods:detail id="5C10864168470606B120DFFF17E15449" type="volume">
<mods:number id="CD566AD3F1F6D915AD898E0C7BA6E0BB">4254</mods:number>
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<mods:detail id="8D64AEA1C6E1CF0A92B7A49E3BA3F4CD" type="issue">
<mods:number id="402EC11AD2056D341199135329CE2029">3</mods:number>
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<mods:start id="E311F3CF162AAD14576B3D9FC7D41CD0">357</mods:start>
<mods:end id="3ABDA361BD24CCD0C99D8D57BF5FDFA0">374</mods:end>
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<mods:classification id="E2ECDCEE96EAF52559EFBD5ABF7FCDA9">journal article</mods:classification>
<mods:identifier id="95C0B6158DA9A1C05D535AF4B4891CA2" type="CLB-Dataset">33208</mods:identifier>
<mods:identifier id="6C6C75371FA4715C8768FCE332E80731" type="DOI">10.11646/zootaxa.4254.3.4</mods:identifier>
<mods:identifier id="FE18C114BA537805B133F417C447AA33" type="GBIF-Dataset">a9d4b1f4-fa63-42bb-881e-53514379d22c</mods:identifier>
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<treatment id="7B2F87B2F621FA52D18171CFFCE2FD4C" ID-DOI="http://doi.org/10.5281/zenodo.6004321" ID-GBIF-Taxon="128535142" ID-Zenodo-Dep="6004321" LSID="urn:lsid:plazi:treatment:7B2F87B2F621FA52D18171CFFCE2FD4C" httpUri="http://treatment.plazi.org/id/7B2F87B2F621FA52D18171CFFCE2FD4C" lastPageId="8" lastPageNumber="365" pageId="2" pageNumber="359">
<subSubSection id="BB9C652FF621FA58D18171CFFD7DFF1B" pageId="2" pageNumber="359" type="nomenclature">
<paragraph id="F33936A4F621FA58D18171CFFE68FF7A" blockId="2.[151,726,151,211]" box="[151,451,151,178]" pageId="2" pageNumber="359">
<heading id="A87181C8F621FA58D18171CFFE68FF7A" bold="true" box="[151,451,151,178]" fontSize="11" level="1" pageId="2" pageNumber="359" reason="1">
<emphasis id="C1F2EAB6F621FA58D18171CFFE68FF7A" bold="true" box="[151,451,151,178]" pageId="2" pageNumber="359">
<taxonomicName id="34864D27F621FA58D18171CFFEF4FF79" ID-CoL="6VDPW" box="[151,351,151,177]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="359" phylum="Arthropoda" rank="species" species="calypso" status="sp. nov.">
<emphasis id="C1F2EAB6F621FA58D18171CFFEF4FF79" bold="true" box="[151,351,151,177]" italics="true" pageId="2" pageNumber="359">Phrynus calypso</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAC157CDF621FA58D07371C0FE68FF7A" box="[357,451,152,178]" pageId="2" pageNumber="359" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="F33936A4F621FA58D18171E2FD7DFF1B" blockId="2.[151,726,151,211]" box="[151,726,186,211]" pageId="2" pageNumber="359">
<emphasis id="C1F2EAB6F621FA58D18171E2FF5BFF1B" bold="true" box="[151,240,186,211]" pageId="2" pageNumber="359">Figures</emphasis>
: 1A; 2E–F; 3–4; 7C, D, F; 8 A, C.; Table 1
</paragraph>
</subSubSection>
<subSubSection id="BB9C652FF621FA58D181705AFCA7FC23" pageId="2" pageNumber="359" type="materials_examined">
<paragraph id="F33936A4F621FA58D181705AFE2CFEAB" blockId="2.[151,1437,258,1111]" pageId="2" pageNumber="359">
<emphasis id="C1F2EAB6F621FA58D181705AFD84FED3" bold="true" box="[151,559,258,283]" pageId="2" pageNumber="359">
<typeStatus id="2C3D8806F621FA58D181705AFF79FED3" box="[151,210,258,283]" pageId="2" pageNumber="359">Type</typeStatus>
material: 
<typeStatus id="2C3D8806F621FA58D059705AFE17FED3" box="[335,444,258,283]" pageId="2" pageNumber="359" type="holotype">Holotype</typeStatus>
(female):
</emphasis>
<collectingCountry id="8B917634F621FA58D320705AFC8DFED3" box="[566,806,258,283]" name="Trinidad and Tobago" pageId="2" pageNumber="359">
Trinidad and 
<collectingRegion id="3142F846F621FA58D3D8705AFC8DFED3" box="[718,806,258,283]" country="Trinidad and Tobago" name="Tobago" pageId="2" pageNumber="359">Tobago</collectingRegion>
</collectingCountry>
: 
<collectingCountry id="8B917634F621FA58D227705AFC3EFED3" box="[817,917,258,283]" name="Trinidad and Tobago" pageId="2" pageNumber="359">Trinidad</collectingCountry>
: Laja Trace (10.741194, -61.271472): tropical rain forest of low mountain, 180–550 masl, Victor Townsend, Paul Reslar, Dan Proud, 
<date id="87381064F621FA58D5B5707FFAEBFEF7" box="[1187,1344,294,319]" pageId="2" pageNumber="359" value="2006-07-22">22 July 2006</date>
, female (CASENT 9060553).
</paragraph>
<paragraph id="F33936A4F621FA58D1D17036FCA7FC23" blockId="2.[151,1437,258,1111]" pageId="2" pageNumber="359">
<materialsCitation id="43EE3CF9F621FA58D1D17036FD79FE07" ID-GBIF-Occurrence="1456819543" collectingDate="2006-07-22" collectorName="Trinidad &amp; Laja Trace &amp; Victor Townsend &amp; Paul Reslar &amp; Dan Proud" country="Trinidad and Tobago" location="Trinidad and Tobago" pageId="2" pageNumber="359" specimenCount="43" specimenCount-female="17" specimenCount-juvenile="8" specimenCount-male="18" typeStatus="paratype">
<emphasis id="C1F2EAB6F621FA58D1D17036FD51FE4F" bold="true" box="[199,762,366,391]" pageId="2" pageNumber="359">
<typeStatus id="2C3D8806F621FA58D1D17036FE94FE4F" box="[199,319,366,391]" pageId="2" pageNumber="359" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="E580FD2DF621FA58D0447036FE7DFE4F" box="[338,470,366,391]" pageId="2" pageNumber="359" type="female">16 females</specimenCount>
, 
<specimenCount id="E580FD2DF621FA58D0F07036FDFFFE4F" box="[486,596,366,391]" pageId="2" pageNumber="359" type="male">17 males</specimenCount>
, 
<specimenCount id="E580FD2DF621FA58D3727036FD46FE4F" box="[612,749,366,391]" pageId="2" pageNumber="359" type="juvenile">8 juveniles</specimenCount>
):
</emphasis>
<collectingCountry id="8B917634F621FA58D2137036FC52FE4F" box="[773,1017,366,391]" name="Trinidad and Tobago" pageId="2" pageNumber="359">Trinidad and Tobago</collectingCountry>
, 
<collectorName id="5E735372F621FA58D51F7036FBC7FE4F" box="[1033,1132,366,391]" pageId="2" pageNumber="359">Trinidad</collectorName>
, 
<collectorName id="5E735372F621FA58D56C7036FB5DFE4E" box="[1146,1270,366,391]" pageId="2" pageNumber="359">Laja Trace</collectorName>
(10.741194, - 61.271472): tropical rain forest of low mountain, 180–550 masl, 
<collectorName id="5E735372F621FA58D27A70CAFB85FE63" box="[876,1070,402,427]" pageId="2" pageNumber="359">Victor Townsend</collectorName>
, 
<collectorName id="5E735372F621FA58D52F70CAFB11FE63" box="[1081,1210,402,427]" pageId="2" pageNumber="359">Paul Reslar</collectorName>
, 
<collectorName id="5E735372F621FA58D5D370CBFAEBFE63" box="[1221,1344,402,427]" pageId="2" pageNumber="359">Dan Proud</collectorName>
, 
<collectingDate id="977CE98CF621FA58D45D70CBFF79FE07" pageId="2" pageNumber="359" value="2006-07-22">
<date id="87381064F621FA58D45D70CBFF79FE07" pageId="2" pageNumber="359" value="2006-07-22">22 July 2006</date>
</collectingDate>
, 
<specimenCount id="E580FD2DF621FA58D1F770EFFEECFE07" box="[225,327,438,463]" pageId="2" pageNumber="359" type="female">1 female</specimenCount>
and 
<specimenCount id="E580FD2DF621FA58D09070EFFE7EFE07" box="[390,469,438,463]" pageId="2" pageNumber="359" type="male">1 male</specimenCount>
(CASENT 9060553)
</materialsCitation>
. 
<materialsCitation id="43EE3CF9F621FA58D3F670EEFDD6FDDF" ID-GBIF-Occurrence="1456819545" collectingDate="2005-07-12" country="Trinidad and Tobago" location="Trinidad and Tobago" pageId="2" pageNumber="359" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<collectingCountry id="8B917634F621FA58D3F670EEFC7EFE07" box="[736,981,438,463]" name="Trinidad and Tobago" pageId="2" pageNumber="359">Trinidad and Tobago</collectingCountry>
, Trinidad, Petite Tacarib (10.741194, - 61.225750): Crapo-cocorite sessional forest-jungle, 0–10 masl, Victor Townsend, Dan Proud, Rebeca Hunter, 
<collectingDate id="977CE98CF621FA58D4967083FEA1FDDF" pageId="2" pageNumber="359" value="2005-07-12">
<date id="87381064F621FA58D4967083FEA1FDDF" pageId="2" pageNumber="359" value="2005-07-12">12 July 2005</date>
</collectingDate>
, 
<specimenCount id="E580FD2DF621FA58D00170A7FE2EFDDF" box="[279,389,510,535]" pageId="2" pageNumber="359" type="female">2 females</specimenCount>
(CASENT 9060537)
</materialsCitation>
. 
<materialsCitation id="43EE3CF9F621FA58D39F70A6FEF3FD97" ID-GBIF-Occurrence="1456819541" collectingDate="2014-10-01" collectionCode="MZUSP" collectorName="Trinidad &amp; Fernando Marques" country="Trinidad and Tobago" location="Maracas Bay" municipality="North Road" pageId="2" pageNumber="359" specimenCode="MZUSP 70892" specimenCount="3" specimenCount-juvenile="1" specimenCount-male="2" stateProvince="Port of Spain" typeStatus="paratype">
<collectingCountry id="8B917634F621FA58D39F70A6FCD2FDDF" box="[649,889,510,535]" name="Trinidad and Tobago" pageId="2" pageNumber="359">Trinidad and Tobago</collectingCountry>
, 
<collectorName id="5E735372F621FA58D29170A6FC42FDDF" box="[903,1001,510,535]" pageId="2" pageNumber="359">Trinidad</collectorName>
, 
<collectingMunicipality id="135DACDEF621FA58D2E170A6FBD4FDDF" box="[1015,1151,510,535]" pageId="2" pageNumber="359">North Road</collectingMunicipality>
, 
<quantity id="347E9B41F621FA58D59A70A6FB15FDDF" box="[1164,1214,510,535]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="359" unit="km" value="5.0">5km</quantity>
from 
<location id="F659607FF621FA58D41170A7FA37FDDF" LSID="urn:lsid:plazi:treatment:7B2F87B2F621FA52D18171CFFCE2FD4C:F659607FF621FA58D41170A7FA37FDDF" box="[1287,1436,511,535]" country="Trinidad and Tobago" municipality="North Road" name="Maracas Bay" pageId="2" pageNumber="359" stateProvince="Port of Spain">Maracas Bay</location>
<collectingRegion id="3142F846F621FA58D181737BFE80FDF3" box="[151,299,546,571]" country="Trinidad and Tobago" name="Port of Spain" pageId="2" pageNumber="359">Port of Spain</collectingRegion>
(10.750593, -61.478858): 
<collectorName id="5E735372F621FA58D341737AFC86FDF2" box="[599,813,546,571]" pageId="2" pageNumber="359">Fernando Marques</collectorName>
, manual collection, 
<collectingDate id="977CE98CF621FA58D51B737BFB15FDF3" box="[1037,1214,546,571]" pageId="2" pageNumber="359" value="2014-10-01">
<date id="87381064F621FA58D51B737BFB15FDF3" box="[1037,1214,546,571]" pageId="2" pageNumber="359" value="2014-10-01">1 October 2014</date>
</collectingDate>
, 
<specimenCount id="E580FD2DF621FA58D5DF737BFA90FDF3" box="[1225,1339,546,571]" pageId="2" pageNumber="359" type="juvenile">1 juvenile</specimenCount>
, 
<specimenCount id="E580FD2DF621FA58D450737BFA37FDF3" box="[1350,1436,546,571]" pageId="2" pageNumber="359" type="male">2 males</specimenCount>
(
<specimenCode id="A3209EDFF621FA58D188731FFEFBFD97" box="[158,336,583,607]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="359">
<collectionCode id="9597AE61F621FA58D188731FFF56FD97" box="[158,253,583,607]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="359">MZUSP</collectionCode>
70892
</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="43EE3CF9F621FA58D075731EFC19FD4B" ID-GBIF-Occurrence="1456819521" collectingDate="2002-12-08" collectionCode="MNRJ" collectorName="Cave over Cueva &amp; Guacharo &amp; Abel Perez &amp; Alessandro Giupponi" country="Venezuela" location="Monagas" pageId="2" pageNumber="359" specimenCode="MNRJ 09310" specimenCount="1" stateProvince="Monagas" typeStatus="paratype">
<collectingCountry id="8B917634F621FA58D075731EFE72FD97" box="[355,473,582,607]" name="Venezuela" pageId="2" pageNumber="359">Venezuela</collectingCountry>
, Estado de 
<collectingRegion id="3142F846F621FA58D34C731FFD68FD97" box="[602,707,583,607]" country="Venezuela" name="Monagas" pageId="2" pageNumber="359">Monagas</collectingRegion>
, 
<collectorName id="5E735372F621FA58D3D9731FFC3BFD97" box="[719,912,583,607]" pageId="2" pageNumber="359">Cave over Cueva</collectorName>
del 
<collectorName id="5E735372F621FA58D2D4731EFB84FD97" box="[962,1071,582,607]" pageId="2" pageNumber="359">Guacharo</collectorName>
(10.200000, -63.639722): 
<collectorName id="5E735372F621FA58D470731EFF73FD4B" pageId="2" pageNumber="359">Abel Pérez</collectorName>
, 
<collectorName id="5E735372F621FA58D1F57332FE7FFD4B" box="[227,468,618,643]" pageId="2" pageNumber="359">Alessandro Giupponi</collectorName>
, 
<collectingDate id="977CE98CF621FA58D0C87333FD0CFD4B" box="[478,679,618,643]" pageId="2" pageNumber="359" value="2002-12-08">
<date id="87381064F621FA58D0C87333FD0CFD4B" box="[478,679,618,643]" pageId="2" pageNumber="359" value="2002-12-08">8 December 2002</date>
</collectingDate>
, female (
<specimenCode id="A3209EDFF621FA58D21A7333FC01FD4B" box="[780,938,619,643]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="2" pageNumber="359">
<collectionCode id="9597AE61F621FA58D21A7333FCF2FD4B" box="[780,857,619,643]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="2" pageNumber="359">MNRJ</collectionCode>
09310
</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="43EE3CF9F621FA58D2D47332FE7AFD03" ID-GBIF-Occurrence="1456819526" collectingDate="2002-12-08" collectionCode="MNRJ" collectorName="Cave over Cueva &amp; Guacharo &amp; Abel Perez &amp; Alessandro Giupponi" country="Venezuela" location="Monagas" pageId="2" pageNumber="359" specimenCode="MNRJ 09315" specimenCount="11" specimenCount-female="5" specimenCount-male="6" stateProvince="Monagas" typeStatus="paratype">
<collectingCountry id="8B917634F621FA58D2D47332FB93FD4B" box="[962,1080,618,643]" name="Venezuela" pageId="2" pageNumber="359">Venezuela</collectingCountry>
, Estado de 
<collectingRegion id="3142F846F621FA58D5A07333FAB4FD4B" box="[1206,1311,619,643]" country="Venezuela" name="Monagas" pageId="2" pageNumber="359">Monagas</collectingRegion>
, 
<collectorName id="5E735372F621FA58D43C7333FF74FD6F" pageId="2" pageNumber="359">Cave over Cueva</collectorName>
del 
<collectorName id="5E735372F621FA58D00273D6FE2AFD6F" box="[276,385,654,679]" pageId="2" pageNumber="359">Guacharo</collectorName>
(10.200000, -63.639722): 
<collectorName id="5E735372F621FA58D3D673D6FCEAFD6F" box="[704,833,654,679]" pageId="2" pageNumber="359">Abel Pérez</collectorName>
, 
<collectorName id="5E735372F621FA58D25973D6FBE8FD6F" box="[847,1091,654,679]" pageId="2" pageNumber="359">Alessandro Giupponi</collectorName>
, 
<collectingDate id="977CE98CF621FA58D54773D7FA8BFD6F" box="[1105,1312,654,679]" pageId="2" pageNumber="359" value="2002-12-08">
<date id="87381064F621FA58D54773D7FA8BFD6F" box="[1105,1312,654,679]" pageId="2" pageNumber="359" value="2002-12-08">8 December 2002</date>
</collectingDate>
, 
<specimenCount id="E580FD2DF621FA58D43B73D6FA37FD6F" box="[1325,1436,654,679]" pageId="2" pageNumber="359" type="female">5 females</specimenCount>
and 
<specimenCount id="E580FD2DF621FA58D1D073EAFEB7FD03" box="[198,284,690,715]" pageId="2" pageNumber="359" type="male">6 males</specimenCount>
(
<specimenCode id="A3209EDFF621FA58D03D73EBFE62FD03" box="[299,457,690,715]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="2" pageNumber="359">
<collectionCode id="9597AE61F621FA58D03D73EBFED3FD03" box="[299,376,691,715]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="2" pageNumber="359">MNRJ</collectionCode>
09315
</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="43EE3CF9F621FA58D0F573EAFB3FFD27" ID-GBIF-Occurrence="1456819523" collectingDate="2002-12-08" collectionCode="MNRJ" collectorName="Cueva &amp; Guacharo &amp; Abel Perez &amp; Alessandro Giupponi" country="Venezuela" location="Monagas" pageId="2" pageNumber="359" specimenCode="MNRJ 09312" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Monagas" typeStatus="paratype">
<collectingCountry id="8B917634F621FA58D0F573EAFDF2FD03" box="[483,601,690,715]" name="Venezuela" pageId="2" pageNumber="359">Venezuela</collectingCountry>
, Estado de 
<collectingRegion id="3142F846F621FA58D3CE73EBFCEAFD03" box="[728,833,691,715]" country="Venezuela" name="Monagas" pageId="2" pageNumber="359">Monagas</collectingRegion>
, 
<collectorName id="5E735372F621FA58D25B73EBFC3EFD03" box="[845,917,691,715]" pageId="2" pageNumber="359">Cueva</collectorName>
del 
<collectorName id="5E735372F621FA58D2D273EAFB9FFD03" box="[964,1076,690,715]" pageId="2" pageNumber="359">Guacharo</collectorName>
, (10.200000, -63.639722): 
<collectorName id="5E735372F621FA58D47073EAFF73FD27" pageId="2" pageNumber="359">Abel Pérez</collectorName>
, 
<collectorName id="5E735372F621FA58D1FE738EFE74FD27" box="[232,479,726,751]" pageId="2" pageNumber="359">Alessandro Giupponi</collectorName>
, 
<collectingDate id="977CE98CF621FA58D0F9738FFD68FD27" box="[495,707,726,751]" pageId="2" pageNumber="359" value="2002-12-08">
<date id="87381064F621FA58D0F9738FFD68FD27" box="[495,707,726,751]" pageId="2" pageNumber="359" value="2002-12-08">8 December 2002</date>
</collectingDate>
, 
<specimenCount id="E580FD2DF621FA58D3C5738FFC92FD27" box="[723,825,726,751]" pageId="2" pageNumber="359" type="female">1 female</specimenCount>
and 
<specimenCount id="E580FD2DF621FA58D26C738FFC7EFD27" box="[890,981,726,751]" pageId="2" pageNumber="359" type="male">3 males</specimenCount>
(
<specimenCode id="A3209EDFF621FA58D2FF738FFB27FD27" box="[1001,1164,727,751]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="2" pageNumber="359">
<collectionCode id="9597AE61F621FA58D2FF738FFB9EFD27" box="[1001,1077,727,751]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="2" pageNumber="359">MNRJ</collectionCode>
09312
</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="43EE3CF9F621FA58D5B5738EFDECFCFF" ID-GBIF-Occurrence="1456819534" collectingDate="2002-12-08" collectionCode="MNRJ" collectorName="Quebrada Cambural &amp; Abel Perez &amp; Alessandro Giupponi" country="Venezuela" location="Monagas" pageId="2" pageNumber="359" specimenCode="MNRJ 09317" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Monagas" typeStatus="paratype">
<collectingCountry id="8B917634F621FA58D5B5738EFAB2FD27" box="[1187,1305,726,751]" name="Venezuela" pageId="2" pageNumber="359">Venezuela</collectingCountry>
, Estado de 
<collectingRegion id="3142F846F621FA58D18173A3FEAAFCDB" box="[151,257,763,787]" country="Venezuela" name="Monagas" pageId="2" pageNumber="359">Monagas</collectingRegion>
, 
<collectorName id="5E735372F621FA58D01B73A2FE5AFCDB" box="[269,497,762,787]" pageId="2" pageNumber="359">Quebrada Cambural</collectorName>
(10.042500, -63.558861): 
<collectorName id="5E735372F621FA58D23373A2FC0DFCDB" box="[805,934,762,787]" pageId="2" pageNumber="359">Abel Pérez</collectorName>
, 
<collectorName id="5E735372F621FA58D2A573A2FB0CFCDB" box="[947,1191,762,787]" pageId="2" pageNumber="359">Alessandro Giupponi</collectorName>
, 
<collectingDate id="977CE98CF621FA58D5A573A3FA2AFCDB" box="[1203,1409,762,787]" pageId="2" pageNumber="359" value="2002-12-08">
<date id="87381064F621FA58D5A573A3FA2AFCDB" box="[1203,1409,762,787]" pageId="2" pageNumber="359" value="2002-12-08">8 December 2002</date>
</collectingDate>
, 
<specimenCount id="E580FD2DF621FA58D49873A3FF45FCFF" pageId="2" pageNumber="359" type="female">2 females</specimenCount>
and 
<specimenCount id="E580FD2DF621FA58D03B7247FE23FCFF" box="[301,392,798,823]" pageId="2" pageNumber="359" type="male">4 males</specimenCount>
(
<specimenCode id="A3209EDFF621FA58D08A7247FD94FCFF" box="[412,575,799,823]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="2" pageNumber="359">
<collectionCode id="9597AE61F621FA58D08A7247FE42FCFF" box="[412,489,799,823]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="2" pageNumber="359">MNRJ</collectionCode>
09317
</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="43EE3CF9F621FA58D3407246FDA4FCB7" ID-GBIF-Occurrence="1456819539" collectingDate="2002-12-03" collectionCode="MNRJ" collectorName="Way &amp; Cueva &amp; Guacharo &amp; Caripe &amp; Abel Perez &amp; Alessandro Giupponi" country="Venezuela" location="Monagas" pageId="2" pageNumber="359" specimenCode="MNRJ 09316" specimenCount="7" specimenCount-female="1" specimenCount-juvenile="5" specimenCount-male="1" stateProvince="Monagas" typeStatus="paratype">
<collectingCountry id="8B917634F621FA58D3407246FD67FCFF" box="[598,716,798,823]" name="Venezuela" pageId="2" pageNumber="359">Venezuela</collectingCountry>
, Estado de 
<collectingRegion id="3142F846F621FA58D24F7247FC68FCFF" box="[857,963,799,823]" country="Venezuela" name="Monagas" pageId="2" pageNumber="359">Monagas</collectingRegion>
, 
<collectorName id="5E735372F621FA58D2C57247FBADFCFF" box="[979,1030,799,823]" pageId="2" pageNumber="359">Way</collectorName>
between 
<collectorName id="5E735372F621FA58D56C7247FB69FCFF" box="[1146,1218,799,823]" pageId="2" pageNumber="359">Cueva</collectorName>
del 
<collectorName id="5E735372F621FA58D5ED7246FAC3FCFF" box="[1275,1384,798,823]" pageId="2" pageNumber="359">Guacharo</collectorName>
and 
<collectorName id="5E735372F621FA58D181721AFF49FC93" box="[151,226,834,859]" pageId="2" pageNumber="359">Caripe</collectorName>
(10.169900, -63.505800): 1000 masl, 
<collectorName id="5E735372F621FA58D3B5721AFC8EFC93" box="[675,805,834,859]" pageId="2" pageNumber="359">Abel Pérez</collectorName>
, 
<collectorName id="5E735372F621FA58D222721AFB81FC93" box="[820,1066,834,859]" pageId="2" pageNumber="359">Alessandro Giupponi</collectorName>
, 
<collectingDate id="977CE98CF621FA58D52E721BFAA2FC93" box="[1080,1289,834,859]" pageId="2" pageNumber="359" value="2002-12-03">
<date id="87381064F621FA58D52E721BFAA2FC93" box="[1080,1289,834,859]" pageId="2" pageNumber="359" value="2002-12-03">3 December 2002</date>
</collectingDate>
, 
<specimenCount id="E580FD2DF621FA58D40E721BFAD4FC93" box="[1304,1407,834,859]" pageId="2" pageNumber="359" type="female">1 female</specimenCount>
, 
<specimenCount id="E580FD2DF621FA58D498721BFF7BFCB7" pageId="2" pageNumber="359" type="male">1 male</specimenCount>
, 
<specimenCount id="E580FD2DF621FA58D1C9723EFEF9FCB7" box="[223,338,870,895]" pageId="2" pageNumber="359" type="juvenile">5 juvenile</specimenCount>
(
<specimenCode id="A3209EDFF621FA58D072723FFDACFCB7" box="[356,519,870,895]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="2" pageNumber="359">
<collectionCode id="9597AE61F621FA58D072723FFE1AFCB7" box="[356,433,871,895]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="2" pageNumber="359">MNRJ</collectionCode>
09316
</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="43EE3CF9F621FA58D30B723EFA8CFCB7" ID-GBIF-Occurrence="1456819537" box="[541,1319,870,895]" country="Venezuela" location="Sucre" pageId="2" pageNumber="359" specimenCount="1" stateProvince="Sucre" typeStatus="paratype">
<collectingCountry id="8B917634F621FA58D30B723EFD38FCB7" box="[541,659,870,895]" name="Venezuela" pageId="2" pageNumber="359">Venezuela</collectingCountry>
, Estado de 
<collectingRegion id="3142F846F621FA58D208723FFCCAFCB7" box="[798,865,871,895]" country="Venezuela" name="Sucre" pageId="2" pageNumber="359">Sucre</collectingRegion>
, Santa Maria (10.631944, -63.077777)
</materialsCitation>
: 
<materialsCitation id="43EE3CF9F621FA58D423723FFAB8FC6B" ID-GBIF-Occurrence="1456819532" collectingDate="2002-12-06" collectionCode="MNRJ" collectorName="Quebrada Macha &amp; Abel Perez &amp; Alessandro Giupponi" country="Venezuela" location="La Vega" pageId="2" pageNumber="359" specimenCode="MNRJ09313" specimenCount="3" specimenCount-female="3" stateProvince="La Vega" typeStatus="paratype">
<collectingRegion id="3142F846F621FA58D423723FFA33FCB7" box="[1333,1432,871,895]" country="Dominican Republic" name="La Vega" pageId="2" pageNumber="359">La Vega</collectingRegion>
, 
<collectorName id="5E735372F621FA58D18172D2FEF4FC6B" box="[151,351,906,931]" pageId="2" pageNumber="359">Quebrada Macha</collectorName>
, 
<collectorName id="5E735372F621FA58D07972D2FE58FC6B" box="[367,499,906,931]" pageId="2" pageNumber="359">Abel Pérez</collectorName>
, 
<collectorName id="5E735372F621FA58D31572D2FD51FC6B" box="[515,762,906,931]" pageId="2" pageNumber="359">Alessandro Giupponi</collectorName>
, 
<collectingDate id="977CE98CF621FA58D21C72D2FC75FC6B" box="[778,990,906,931]" pageId="2" pageNumber="359" value="2002-12-06">
<date id="87381064F621FA58D21C72D2FC75FC6B" box="[778,990,906,931]" pageId="2" pageNumber="359" value="2002-12-06">6 December 2002</date>
</collectingDate>
, 
<specimenCount id="E580FD2DF621FA58D2F872D3FBCBFC6B" box="[1006,1120,906,931]" pageId="2" pageNumber="359" type="female">3 females</specimenCount>
(
<specimenCode id="A3209EDFF621FA58D56572D3FAA0FC6B" box="[1139,1291,907,931]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="2" pageNumber="359">
<collectionCode id="9597AE61F621FA58D56572D3FB69FC6B" box="[1139,1218,907,931]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="2" pageNumber="359">MNRJ</collectionCode>
09313
</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="43EE3CF9F621FA58D43472D2FC83FC0F" ID-GBIF-Occurrence="1456819524" country="Venezuela" location="Sucre" pageId="2" pageNumber="359" specimenCount="1" stateProvince="Sucre" typeStatus="paratype">
<collectingCountry id="8B917634F621FA58D43472D2FA33FC6B" box="[1314,1432,906,931]" name="Venezuela" pageId="2" pageNumber="359">Venezuela</collectingCountry>
, Estado de 
<collectingRegion id="3142F846F621FA58D00172F7FEF1FC0F" box="[279,346,943,967]" country="Venezuela" name="Sucre" pageId="2" pageNumber="359">Sucre</collectingRegion>
, Santa Maria (10.631944, -63.077777)
</materialsCitation>
: 
<materialsCitation id="43EE3CF9F621FA58D22E72F7FCA3FC23" ID-GBIF-Occurrence="1456819536" collectingDate="2002-12-06" collectionCode="MNRJ" collectorName="Quebrada Macha &amp; Abel Perez &amp; Alessandro Giupponi" country="Venezuela" location="La Vega" pageId="2" pageNumber="359" specimenCode="MNRJ 09314" specimenCount="2" specimenCount-juvenile="2" stateProvince="La Vega" typeStatus="paratype">
<collectingRegion id="3142F846F621FA58D22E72F7FC35FC0F" box="[824,926,943,967]" country="Dominican Republic" name="La Vega" pageId="2" pageNumber="359">La Vega</collectingRegion>
, 
<collectorName id="5E735372F621FA58D2B972F6FBDCFC0F" box="[943,1143,942,967]" pageId="2" pageNumber="359">Quebrada Macha</collectorName>
, 
<collectorName id="5E735372F621FA58D59F72F6FAA6FC0F" box="[1161,1293,942,967]" pageId="2" pageNumber="359">Abel Pérez</collectorName>
, 
<collectorName id="5E735372F621FA58D40872F6FEAFFC23" pageId="2" pageNumber="359">Alessandro Giupponi</collectorName>
, 
<collectingDate id="977CE98CF621FA58D018728AFE72FC23" box="[270,473,978,1003]" pageId="2" pageNumber="359" value="2002-12-06">
<date id="87381064F621FA58D018728AFE72FC23" box="[270,473,978,1003]" pageId="2" pageNumber="359" value="2002-12-06">6 December 2002</date>
</collectingDate>
, 
<specimenCount id="E580FD2DF621FA58D0F2728BFDF8FC23" box="[484,595,978,1003]" pageId="2" pageNumber="359" type="juvenile">2 juvenile</specimenCount>
(
<specimenCode id="A3209EDFF621FA58D374728BFCABFC23" box="[610,768,979,1003]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="2" pageNumber="359">
<collectionCode id="9597AE61F621FA58D374728BFD05FC23" box="[610,686,979,1003]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="2" pageNumber="359">MNRJ</collectionCode>
09314
</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BB9C652FF621FA58D1D172AEFB78FBC7" box="[199,1235,1014,1039]" pageId="2" pageNumber="359" type="distribution">
<paragraph id="F33936A4F621FA58D1D172AEFB78FBC7" blockId="2.[151,1437,258,1111]" box="[199,1235,1014,1039]" pageId="2" pageNumber="359">
<emphasis id="C1F2EAB6F621FA58D1D172AEFECBFBC7" bold="true" box="[199,352,1014,1039]" pageId="2" pageNumber="359">Distribution.</emphasis>
<collectingCountry id="8B917634F621FA58D07172AEFE6DFBC7" box="[359,454,1014,1039]" name="Trinidad and Tobago" pageId="2" pageNumber="359">Trinidad</collectingCountry>
<collectingCountry id="8B917634F621FA58D0DB72AEFDBFFBC7" box="[461,532,1014,1039]" name="Iceland" pageId="2" pageNumber="359">Island</collectingCountry>
, in 
<collectingCountry id="8B917634F621FA58D32B72AEFC82FBC7" box="[573,809,1014,1039]" name="Trinidad and Tobago" pageId="2" pageNumber="359">
Trinidad and 
<collectingRegion id="3142F846F621FA58D3C572AEFC82FBC7" box="[723,809,1014,1039]" country="Trinidad and Tobago" name="Tobago" pageId="2" pageNumber="359">Tobago</collectingRegion>
</collectingCountry>
, and Coast of 
<collectingCountry id="8B917634F621FA58D2DF72AEFB97FBC7" box="[969,1084,1014,1039]" name="Venezuela" pageId="2" pageNumber="359">Venezuela</collectingCountry>
(
<figureCitation id="6BBD2A21F621FA58D55D72AEFB0FFBC7" box="[1099,1188,1014,1039]" captionStart="FIGURE 8" captionStartId="14.[151,250,1299,1321]" captionTargetBox="[158,1424,203,1271]" captionTargetId="figure@14.[151,1436,193,1277]" captionTargetPageId="14" captionText="FIGURE 8. A. Distribution of Phrynus calypso sp. nov and Phrynus pulchripes. B. Distribution of P. pulchripes, detail of records in Colombia. C. Distribution of P. calypso, detail of records. P. calypso: yellow circle = records, yellow circle with black border = Type locality. P. pulchripes: red square = records, red square with black border = Type locality." httpUri="https://zenodo.org/record/546848/files/figure.png" pageId="2" pageNumber="359">Fig. 8A</figureCitation>
, C).
</paragraph>
</subSubSection>
<subSubSection id="BB9C652FF621FA58D1D17542FC4BFB9F" pageId="2" pageNumber="359" type="etymology">
<paragraph id="F33936A4F621FA58D1D17542FC4BFB9F" blockId="2.[151,1437,258,1111]" pageId="2" pageNumber="359">
<emphasis id="C1F2EAB6F621FA58D1D17542FEE5FBFB" bold="true" box="[199,334,1050,1075]" pageId="2" pageNumber="359">Etymology.</emphasis>
Species name is dedicated to music rhythm 
<taxonomicName id="34864D27F621FA58D2797542FC65FBFB" box="[879,974,1050,1075]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="372" phylum="Arthropoda" rank="species" species="calypso">Calypso</taxonomicName>
, typical from 
<collectingCountry id="8B917634F621FA58D56D7542FADFFBFB" box="[1147,1396,1050,1075]" name="Trinidad and Tobago" pageId="2" pageNumber="359">
Trinidad and 
<collectingRegion id="3142F846F621FA58D40B7542FADFFBFB" box="[1309,1396,1050,1075]" country="Trinidad and Tobago" name="Tobago" pageId="2" pageNumber="359">Tobago</collectingRegion>
</collectingCountry>
. In commemoration to cultural expression of this country. Name in apposition.
</paragraph>
</subSubSection>
<caption id="A7F9662CF621FA58D1817626FCD9F87B" httpUri="https://zenodo.org/record/546841/files/figure.png" pageId="2" pageNumber="359" targetBox="[240,1349,1167,1893]" targetPageId="2">
<paragraph id="F33936A4F621FA58D1817626FCD9F87B" blockId="2.[151,1436,1918,1971]" pageId="2" pageNumber="359">
<emphasis id="C1F2EAB6F621FA58D1817626FEBCF85C" bold="true" box="[151,279,1918,1940]" pageId="2" pageNumber="359">FIGURE 1.</emphasis>
Compared female genitalia: A. 
<taxonomicName id="34864D27F621FA58D3647626FCB5F85C" box="[626,798,1918,1940]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="359" phylum="Arthropoda" rank="species" species="calypso" status="sp. nov.">
<emphasis id="C1F2EAB6F621FA58D3647626FCB5F85C" box="[626,798,1918,1940]" italics="true" pageId="2" pageNumber="359">Phrynus calypso</emphasis>
</taxonomicName>
<emphasis id="C1F2EAB6F621FA58D23F7627FCD4F85C" bold="true" box="[809,895,1919,1940]" pageId="2" pageNumber="359">
<taxonomicNameLabel id="DAC157CDF621FA58D23F7627FCD2F85C" box="[809,889,1919,1940]" pageId="2" pageNumber="359" rank="species">sp. nov</taxonomicNameLabel>
,
</emphasis>
Holotype. B. 
<emphasis id="C1F2EAB6F621FA58D50D7626FB40F85C" box="[1051,1259,1918,1940]" italics="true" pageId="2" pageNumber="359">
<taxonomicName id="34864D27F621FA58D50D7626FB43F85C" box="[1051,1256,1918,1940]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="359" phylum="Arthropoda" rank="species" species="pulchripes">Phrynus pulchripes</taxonomicName>
,
</emphasis>
ICN-Am 85. C. 
<taxonomicName id="34864D27F621FA58D18176C5FEC5F87B" box="[151,366,1949,1971]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="359" phylum="Arthropoda" rank="species" species="barbadensis">
<emphasis id="C1F2EAB6F621FA58D18176C5FEC5F87B" box="[151,366,1949,1971]" italics="true" pageId="2" pageNumber="359">Phrynus barbadensis</emphasis>
</taxonomicName>
. MUD-AM 403. D. 
<taxonomicName id="34864D27F621FA58D32876C5FD49F87B" box="[574,738,1949,1971]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="359" phylum="Arthropoda" rank="species" species="panche">
<emphasis id="C1F2EAB6F621FA58D32876C5FD49F87B" box="[574,738,1949,1971]" italics="true" pageId="2" pageNumber="359">Phrynus panche</emphasis>
</taxonomicName>
. ICN-AM 64.
</paragraph>
</caption>
<caption id="A7F9662CF620FA59D1817445FA86FA9A" httpUri="https://zenodo.org/record/546842/files/figure.png" pageId="3" pageNumber="360" targetBox="[209,1370,197,1274]" targetPageId="3">
<paragraph id="F33936A4F620FA59D1817445FA86FA9A" blockId="3.[151,1435,1309,1362]" pageId="3" pageNumber="360">
<emphasis id="C1F2EAB6F620FA59D1817445FEB8FAFB" bold="true" box="[151,275,1309,1331]" pageId="3" pageNumber="360">FIGURE 2.</emphasis>
Variation in female genitalia: 
<taxonomicName id="34864D27F620FA59D35C7445FCBAFAFB" box="[586,785,1309,1331]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="pulchripes">
<emphasis id="C1F2EAB6F620FA59D35C7445FCBAFAFB" box="[586,785,1309,1331]" italics="true" pageId="3" pageNumber="360">Phrynus pulchripes</emphasis>
</taxonomicName>
: MZUSP 70924-1: A. Dorsal view. B. posterior view. MZUSP 70924-2: C. Dorsal view. D. posterior view. 
<taxonomicName id="34864D27F620FA59D3417464FD54FA9A" box="[599,767,1340,1362]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="calypso">
<emphasis id="C1F2EAB6F620FA59D3417464FD54FA9A" box="[599,767,1340,1362]" italics="true" pageId="3" pageNumber="360">Phrynus calypso</emphasis>
</taxonomicName>
: CASENT 9060553: E. Dorsal view. F. posterior view.
</paragraph>
</caption>
<subSubSection id="BB9C652FF620FA5CD1D17421FF49F9BC" lastPageId="6" lastPageNumber="363" pageId="3" pageNumber="360" type="diagnosis">
<paragraph id="F33936A4F620FA59D1D17421FB12F83D" blockId="3.[151,1437,1401,2038]" pageId="3" pageNumber="360">
<emphasis id="C1F2EAB6F620FA59D1D17421FEEAFA5A" bold="true" box="[199,321,1401,1426]" pageId="3" pageNumber="360">Diagnosis.</emphasis>
<taxonomicName id="34864D27F620FA59D05F7421FDA8FA5A" box="[329,515,1401,1426]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="calypso" status="sp. nov.">
<emphasis id="C1F2EAB6F620FA59D05F7421FDA8FA5A" box="[329,515,1401,1426]" italics="true" pageId="3" pageNumber="360">Phrynus calypso</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAC157CDF620FA59D31D7421FDCEFA5A" box="[523,613,1401,1426]" pageId="3" pageNumber="360" rank="species">
<emphasis id="C1F2EAB6F620FA59D31D7421FDCEFA5A" bold="true" box="[523,613,1401,1426]" pageId="3" pageNumber="360">sp. nov.</emphasis>
</taxonomicNameLabel>
is distinguished from most 
<taxonomicName id="34864D27F620FA59D2B67421FC55FA5A" box="[928,1022,1401,1426]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="genus">
<emphasis id="C1F2EAB6F620FA59D2B67421FC55FA5A" box="[928,1022,1401,1426]" italics="true" pageId="3" pageNumber="360">Phrynus</emphasis>
</taxonomicName>
species by the presence of two teeth in the external line of the base of chelicerae, both with an acuminated apex, and a small spine on dorso-mesal surface of pedipalp tarsus. These two conditions are present together in just four more species: 
<taxonomicName id="34864D27F620FA59D5E57499FE5BFA36" authority="Colmenares &amp; Villarreal, 2008" authorityName="Colmenares &amp; Villarreal" authorityYear="2008" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="araya">
<emphasis id="C1F2EAB6F620FA59D5E57499FA37FA12" box="[1267,1436,1473,1498]" italics="true" pageId="3" pageNumber="360">Phrynus araya</emphasis>
<bibRefCitation id="97174B55F620FA59D18174BDFE5BFA36" author="Colmenares" box="[151,496,1509,1534]" pageId="3" pageNumber="360" refString="Colmenares, P. &amp; Villarreal, O. (2008) Una nueva especie de Phrynus Lamarck, 1801 (Amblypygi: Phrynidae), de La Sierra de Perija, Venezuela. Boletin Sociedad Entomologica Aragonesa, 43, 89 - 93." type="journal article" year="2008">Colmenares &amp; Villarreal, 2008</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="34864D27F620FA59D0E874BDFC7FFA36" authority="Armas &amp; Angarita, 2008" authorityName="Armas &amp; Angarita" authorityYear="2008" box="[510,980,1509,1534]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="panche">
<emphasis id="C1F2EAB6F620FA59D0E874BDFD1FFA36" box="[510,692,1509,1534]" italics="true" pageId="3" pageNumber="360">Phrynus panche</emphasis>
<bibRefCitation id="97174B55F620FA59D3AD74BEFC7FFA36" author="Armas" box="[699,980,1509,1534]" pageId="3" pageNumber="360" refString="Armas, L. F. &amp; Angarita, A. (2008) Nueva especie de Phrynus Lamark, 1801 (Amblypygi: Phrynidae) de Colombia. Boletin de la Sociedad Entomologica Aragonesa (S. E. A), 43, 25 - 28." type="journal article" year="2008">Armas &amp; Angarita, 2008</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="34864D27F620FA59D2C974BDFAC7FA36" authority="Pocock, 1894" authorityName="Pocock" authorityYear="1894" box="[991,1388,1509,1534]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="pulchripes">
<emphasis id="C1F2EAB6F620FA59D2C974BDFB11FA36" box="[991,1210,1509,1534]" italics="true" pageId="3" pageNumber="360">Phrynus pulchripes</emphasis>
(
<bibRefCitation id="97174B55F620FA59D5DF74BDFACFFA36" author="Pocock" box="[1225,1380,1509,1534]" pageId="3" pageNumber="360" refString="Pocock, R. I. (1894) Notes on the Pedipalpi of the family Tarantulidae contained in the collection of the British Museum. Annals and Magazine of Natural History, 14 (6), 273 - 298." type="journal article" year="1894">Pocock, 1894</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="34864D27F620FA59D1817751FE75F9EA" authority="Harvey, 2002" authorityName="Harvey" authorityYear="2002" box="[151,478,1545,1570]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="exsul">
<emphasis id="C1F2EAB6F620FA59D1817751FE93F9EA" box="[151,312,1545,1570]" italics="true" pageId="3" pageNumber="360">Phrynus exsul</emphasis>
<bibRefCitation id="97174B55F620FA59D0577752FE75F9EA" author="Harvey" box="[321,478,1546,1570]" pageId="3" pageNumber="360" refString="Harvey, M. S. (2002) The first old world species of Phrynidae (Amblypygi): Phrynus exsul from Indonesia. 2002. The Journal of Arachnology, 30, 470 - 474." type="journal article" year="2002">Harvey, 2002</bibRefCitation>
</taxonomicName>
. 
<taxonomicName id="34864D27F620FA59D0FD7753FDF5F9EA" box="[491,606,1545,1570]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="calypso">
<emphasis id="C1F2EAB6F620FA59D0FD7753FDF5F9EA" box="[491,606,1545,1570]" italics="true" pageId="3" pageNumber="360">P. calypso</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="34864D27F620FA59D3ED7753FCFCF9EA" box="[763,855,1547,1570]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="araya">
<emphasis id="C1F2EAB6F620FA59D3ED7753FCFCF9EA" box="[763,855,1547,1570]" italics="true" pageId="3" pageNumber="360">P. araya</emphasis>
</taxonomicName>
on the number o spines between Fv3 and Fv5. 
<taxonomicName id="34864D27F620FA59D4917753FF73F98D" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="araya">
<emphasis id="C1F2EAB6F620FA59D4917753FF73F98D" italics="true" pageId="3" pageNumber="360">P. araya</emphasis>
</taxonomicName>
presents three spines between Fv3 and Fv5; Fv4 and two more, according to nomenclature proposed by 
<bibRefCitation id="97174B55F620FA59D1817709FEF9F9A2" author="Quintero" box="[151,338,1617,1642]" pageId="3" pageNumber="360" refString="Quintero, D. (1981) The Amblypygid genus Phrynus in the Americas (Amblypygi, Phrynidae). Journal of Arachnology, 9, 117 - 166." type="journal article" year="1981">Quintero (1981)</bibRefCitation>
. 
<taxonomicName id="34864D27F620FA59D048770BFE65F9A2" box="[350,462,1617,1642]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="calypso">
<emphasis id="C1F2EAB6F620FA59D048770BFE65F9A2" box="[350,462,1617,1642]" italics="true" pageId="3" pageNumber="360">P. calypso</emphasis>
</taxonomicName>
has usually just one Fv4 spine and at most can present two spines between Fv3 and Fv5. 
<taxonomicName id="34864D27F620FA59D1C7772EFE84F945" box="[209,303,1654,1677]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="araya">
<emphasis id="C1F2EAB6F620FA59D1C7772EFE84F945" box="[209,303,1654,1677]" italics="true" pageId="3" pageNumber="360">P. araya</emphasis>
</taxonomicName>
has Fv3 equal or longer than Fv5, in 
<taxonomicName id="34864D27F620FA59D3CB772EFCDCF946" authority="Fv" authorityName="Fv" box="[733,887,1653,1678]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="calypso">
<emphasis id="C1F2EAB6F620FA59D3CB772EFCE7F946" box="[733,844,1653,1678]" italics="true" pageId="3" pageNumber="360">P. calypso</emphasis>
Fv
</taxonomicName>
3 is shorter than Fv5. The female genitalia of 
<emphasis id="C1F2EAB6F620FA59D491772EFEFAF97A" italics="true" pageId="3" pageNumber="360">
<taxonomicName id="34864D27F620FA59D491772EFF40F97A" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="panche">P. panche</taxonomicName>
, 
<taxonomicName id="34864D27F620FA59D1EF77C3FEFAF97A" box="[249,337,1689,1714]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="exsul">P. exsul</taxonomicName>
</emphasis>
and 
<taxonomicName id="34864D27F620FA59D09977C3FE5BF97A" box="[399,496,1691,1714]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="araya">
<emphasis id="C1F2EAB6F620FA59D09977C3FE5BF97A" box="[399,496,1691,1714]" italics="true" pageId="3" pageNumber="360">P. araya</emphasis>
</taxonomicName>
are very similar, all have the sclerites of gonopods with a rectangular elongated shape, this sticks out from seminal receptacles (
<figureCitation id="6BBD2A21F620FA59D3F477E5FC82F91E" box="[738,809,1725,1750]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[240,1349,1167,1893]" captionTargetId="figure@2.[236,1352,1164,1897]" captionTargetPageId="2" captionText="FIGURE 1. Compared female genitalia: A. Phrynus calypso sp. nov, Holotype. B. Phrynus pulchripes, ICN-Am 85. C. Phrynus barbadensis. MUD-AM 403. D. Phrynus panche. ICN-AM 64." httpUri="https://zenodo.org/record/546841/files/figure.png" pageId="3" pageNumber="360">Fig. 1</figureCitation>
D), 
<taxonomicName id="34864D27F620FA59D24977E6FC7FF91E" box="[863,980,1725,1750]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="calypso">
<emphasis id="C1F2EAB6F620FA59D24977E6FC7FF91E" box="[863,980,1725,1750]" italics="true" pageId="3" pageNumber="360">P. calypso</emphasis>
</taxonomicName>
has short, fusiform sclerites with an acuminated apex (
<figureCitation id="6BBD2A21F620FA59D07177B9FE03F932" box="[359,424,1761,1786]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[240,1349,1167,1893]" captionTargetId="figure@2.[236,1352,1164,1897]" captionTargetPageId="2" captionText="FIGURE 1. Compared female genitalia: A. Phrynus calypso sp. nov, Holotype. B. Phrynus pulchripes, ICN-Am 85. C. Phrynus barbadensis. MUD-AM 403. D. Phrynus panche. ICN-AM 64." httpUri="https://zenodo.org/record/546841/files/figure.png" pageId="3" pageNumber="360">Fig. 1</figureCitation>
A). In 
<taxonomicName id="34864D27F620FA59D0EF77BBFDFBF932" box="[505,592,1761,1786]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="exsul">
<emphasis id="C1F2EAB6F620FA59D0EF77BBFDFBF932" box="[505,592,1761,1786]" italics="true" pageId="3" pageNumber="360">P. exsul</emphasis>
</taxonomicName>
spine Fd4 is absent, and in this position there is a small setal tubercle. 
<emphasis id="C1F2EAB6F620FA59D49177BBFEE7F8D6" italics="true" pageId="3" pageNumber="360">
<taxonomicName id="34864D27F620FA59D49177BBFF78F8D6" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="exsul">P. exsul</taxonomicName>
, 
<taxonomicName id="34864D27F620FA59D1C9765FFEE7F8D6" box="[223,332,1797,1822]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="panche">P. panche</taxonomicName>
</emphasis>
and 
<taxonomicName id="34864D27F620FA59D093765FFDBCF8D6" box="[389,535,1797,1822]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="pulchripes">
<emphasis id="C1F2EAB6F620FA59D093765FFDBCF8D6" box="[389,535,1797,1822]" italics="true" pageId="3" pageNumber="360">P. pulchripes</emphasis>
</taxonomicName>
present three small spines placed over the Td3, 
<taxonomicName id="34864D27F620FA59D52D765FFB02F8D6" box="[1083,1193,1797,1822]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="calypso">
<emphasis id="C1F2EAB6F620FA59D52D765FFB02F8D6" box="[1083,1193,1797,1822]" italics="true" pageId="3" pageNumber="360">P. calypso</emphasis>
</taxonomicName>
presents at most one spine on the base of Td3. The most similar species to 
<taxonomicName id="34864D27F620FA59D3EE7673FCCCF88A" box="[760,871,1833,1858]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="calypso">
<emphasis id="C1F2EAB6F620FA59D3EE7673FCCCF88A" box="[760,871,1833,1858]" italics="true" pageId="3" pageNumber="360">P. calypso</emphasis>
</taxonomicName>
is 
<taxonomicName id="34864D27F620FA59D29A7673FBB5F88A" box="[908,1054,1833,1858]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="pulchripes">
<emphasis id="C1F2EAB6F620FA59D29A7673FBB5F88A" box="[908,1054,1833,1858]" italics="true" pageId="3" pageNumber="360">P. pulchripes</emphasis>
</taxonomicName>
. They differ mainly in the female genitalia: in 
<taxonomicName id="34864D27F620FA59D0307617FE11F8AE" box="[294,442,1869,1894]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="pulchripes">
<emphasis id="C1F2EAB6F620FA59D0307617FE11F8AE" box="[294,442,1869,1894]" italics="true" pageId="3" pageNumber="360">P. pulchripes</emphasis>
</taxonomicName>
sclerites do not have an acuminated apex, and are shorter and thicker, as well as more sclerotized. Seminal receptacles in 
<taxonomicName id="34864D27F620FA59D332762BFD18F842" box="[548,691,1905,1930]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="pulchripes">
<emphasis id="C1F2EAB6F620FA59D332762BFD18F842" box="[548,691,1905,1930]" italics="true" pageId="3" pageNumber="360">P. pulchripes</emphasis>
</taxonomicName>
are completely sclerotized (
<figureCitation id="6BBD2A21F620FA59D2E37629FB9EF842" box="[1013,1077,1905,1930]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[240,1349,1167,1893]" captionTargetId="figure@2.[236,1352,1164,1897]" captionTargetPageId="2" captionText="FIGURE 1. Compared female genitalia: A. Phrynus calypso sp. nov, Holotype. B. Phrynus pulchripes, ICN-Am 85. C. Phrynus barbadensis. MUD-AM 403. D. Phrynus panche. ICN-AM 64." httpUri="https://zenodo.org/record/546841/files/figure.png" pageId="3" pageNumber="360">Fig. 1</figureCitation>
B), in 
<taxonomicName id="34864D27F620FA59D569762BFB44F842" box="[1151,1263,1905,1930]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="calypso">
<emphasis id="C1F2EAB6F620FA59D569762BFB44F842" box="[1151,1263,1905,1930]" italics="true" pageId="3" pageNumber="360">P. calypso</emphasis>
</taxonomicName>
the sclerotized region covers just the basal area. Female genitalia in 
<taxonomicName id="34864D27F620FA59D20676CFFC29F866" box="[784,898,1941,1966]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="calypso">
<emphasis id="C1F2EAB6F620FA59D20676CFFC29F866" box="[784,898,1941,1966]" italics="true" pageId="3" pageNumber="360">P. calypso</emphasis>
</taxonomicName>
can be confused with the genital of 
<taxonomicName id="34864D27F620FA59D42876CDFE7EF81A" authority="Pocock, 1894" authorityName="Pocock" authorityYear="1894" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="barbadensis">
<emphasis id="C1F2EAB6F620FA59D42876CDFE8BF81A" italics="true" pageId="3" pageNumber="360">Phrynus barbadensis</emphasis>
(
<bibRefCitation id="97174B55F620FA59D03976E1FE66F81A" author="Pocock" box="[303,461,1977,2002]" pageId="3" pageNumber="360" refString="Pocock, R. I. (1894) Notes on the Pedipalpi of the family Tarantulidae contained in the collection of the British Museum. Annals and Magazine of Natural History, 14 (6), 273 - 298." type="journal article" year="1894">Pocock, 1894</bibRefCitation>
)
</taxonomicName>
, which inhabit the same localities, however this species has the gonopod apex sclerite at least three times thinner than the base (
<figureCitation id="6BBD2A21F620FA59D3717685FD0DF83E" box="[615,678,2013,2038]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[240,1349,1167,1893]" captionTargetId="figure@2.[236,1352,1164,1897]" captionTargetPageId="2" captionText="FIGURE 1. Compared female genitalia: A. Phrynus calypso sp. nov, Holotype. B. Phrynus pulchripes, ICN-Am 85. C. Phrynus barbadensis. MUD-AM 403. D. Phrynus panche. ICN-AM 64." httpUri="https://zenodo.org/record/546841/files/figure.png" pageId="3" pageNumber="360">Fig. 1</figureCitation>
C), a condition that is not present in 
<taxonomicName id="34864D27F620FA59D5577687FB19F83E" box="[1089,1202,2013,2038]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="calypso">
<emphasis id="C1F2EAB6F620FA59D5577687FB19F83E" box="[1089,1202,2013,2038]" italics="true" pageId="3" pageNumber="360">P. calypso</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="A7F9662CF627FA5ED181779EFD20F932" httpUri="https://zenodo.org/record/546843/files/figure.png" pageId="4" pageNumber="361" targetBox="[178,1412,200,1694]" targetPageId="4">
<paragraph id="F33936A4F627FA5ED181779EFD20F932" blockId="4.[151,1436,1734,1786]" pageId="4" pageNumber="361">
<emphasis id="C1F2EAB6F627FA5ED181779EFEB9F913" bold="true" box="[151,274,1734,1756]" pageId="4" pageNumber="361">FIGURE 3.</emphasis>
<taxonomicName id="34864D27F627FA5ED00E779EFE6BF914" box="[280,448,1734,1756]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="361" phylum="Arthropoda" rank="species" species="calypso" status="sp. nov.">
<emphasis id="C1F2EAB6F627FA5ED00E779EFE6BF914" box="[280,448,1734,1756]" italics="true" pageId="4" pageNumber="361">Phrynus calypso</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAC157CDF627FA5ED0D0779EFDBDF914" box="[454,534,1734,1756]" pageId="4" pageNumber="361" rank="species">
<emphasis id="C1F2EAB6F627FA5ED0D0779EFDBBF913" bold="true" box="[454,528,1734,1755]" pageId="4" pageNumber="361">sp. nov</emphasis>
.
</taxonomicNameLabel>
Female Holotype: A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view.
</paragraph>
</caption>
<paragraph id="F33936A4F627FA5FD1D1767AFCF9FD97" blockId="4.[151,1437,1826,2030]" lastBlockId="5.[151,1436,151,608]" lastPageId="5" lastPageNumber="362" pageId="4" pageNumber="361">
Female genitalia present many characters frequently used for to identify species in 
<taxonomicName id="34864D27F627FA5ED57E767AFA32F8F2" authority="Quintero 1981" authorityName="Quintero" authorityYear="1981" box="[1128,1433,1826,1851]" class="Arachnida" family="Phrynidae" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="361" phylum="Arthropoda" rank="family">
Phrynidae (
<bibRefCitation id="97174B55F627FA5ED5F1767AFA3BF8F2" author="Quintero" box="[1255,1424,1826,1851]" pageId="4" pageNumber="361" refString="Quintero, D. (1981) The Amblypygid genus Phrynus in the Americas (Amblypygi, Phrynidae). Journal of Arachnology, 9, 117 - 166." type="journal article" year="1981">Quintero 1981</bibRefCitation>
)
</taxonomicName>
. However, little is known about ontogenetic variation of female genitalia (
<bibRefCitation id="97174B55F627FA5ED2D0761DFBD3F896" author="Weygoldt" box="[966,1144,1861,1886]" pageId="4" pageNumber="361" refString="Weygoldt, P. (2000) Whip spiders (Chelicerata: Amblypygi). Their biology, morphology and systematics. Apollo Books, Stenstrup, 163 pp." type="book" year="2000">Weygoldt 2000</bibRefCitation>
). One reasonable question is if differences between two putative species could be attributed to variation between smaller and larger individuals of the same species. In this research all available female genitalia of 
<taxonomicName id="34864D27F627FA5ED53576D7FB3AF86E" box="[1059,1169,1933,1958]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="361" phylum="Arthropoda" rank="species" species="calypso">
<emphasis id="C1F2EAB6F627FA5ED53576D7FB3AF86E" box="[1059,1169,1933,1958]" italics="true" pageId="4" pageNumber="361">P. calypso</emphasis>
</taxonomicName>
and 
<taxonomicName id="34864D27F627FA5ED5DA76D7FAF5F86E" box="[1228,1374,1933,1958]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="361" phylum="Arthropoda" rank="species" species="pulchripes">
<emphasis id="C1F2EAB6F627FA5ED5DA76D7FAF5F86E" box="[1228,1374,1933,1958]" italics="true" pageId="4" pageNumber="361">P. pulchripes</emphasis>
</taxonomicName>
were examined, even in young exemplars. In all cases pattern of shape, coloration, proportions and position were constant, with some variations in size. Some characteristics can be accentuated in larger specimens, like curvature in sclerites of 
<taxonomicName id="34864D27F626FA5FD02171C1FE62FF78" box="[311,457,151,176]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="5" pageNumber="362" phylum="Arthropoda" rank="species" species="pulchripes">
<emphasis id="C1F2EAB6F626FA5FD02171C1FE62FF78" box="[311,457,151,176]" italics="true" pageId="5" pageNumber="362">P. pulchripes</emphasis>
</taxonomicName>
(
<figureCitation id="6BBD2A21F626FA5FD0CC71CFFDBEFF78" box="[474,533,151,176]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[240,1349,1167,1893]" captionTargetId="figure@2.[236,1352,1164,1897]" captionTargetPageId="2" captionText="FIGURE 1. Compared female genitalia: A. Phrynus calypso sp. nov, Holotype. B. Phrynus pulchripes, ICN-Am 85. C. Phrynus barbadensis. MUD-AM 403. D. Phrynus panche. ICN-AM 64." httpUri="https://zenodo.org/record/546841/files/figure.png" pageId="5" pageNumber="362">Fig.1</figureCitation>
B, 2A–D), and size of sclerites in both species. Also, in both species, coloration of sclerites and seminal receptacles could be a little lighter in smaller specimens (
<figureCitation id="6BBD2A21F626FA5FD57471E4FB0EFF1C" box="[1122,1189,188,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[240,1349,1167,1893]" captionTargetId="figure@2.[236,1352,1164,1897]" captionTargetPageId="2" captionText="FIGURE 1. Compared female genitalia: A. Phrynus calypso sp. nov, Holotype. B. Phrynus pulchripes, ICN-Am 85. C. Phrynus barbadensis. MUD-AM 403. D. Phrynus panche. ICN-AM 64." httpUri="https://zenodo.org/record/546841/files/figure.png" pageId="5" pageNumber="362">Fig. 1</figureCitation>
A–B, 2). Differences between these two species were constant, a posterior view of female sclerites allowed a clearer identification of the differences in shape, and small details like curvatures in 
<taxonomicName id="34864D27F626FA5FD202705DFC0DFED5" box="[788,934,260,285]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="5" pageNumber="362" phylum="Arthropoda" rank="species" species="pulchripes">
<emphasis id="C1F2EAB6F626FA5FD202705DFC0DFED5" box="[788,934,260,285]" italics="true" pageId="5" pageNumber="362">P. pulchripes</emphasis>
</taxonomicName>
and the less sclerotized ending in 
<taxonomicName id="34864D27F626FA5FD43C705DFA37FED5" box="[1322,1436,260,285]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="5" pageNumber="362" phylum="Arthropoda" rank="species" species="calypso">
<emphasis id="C1F2EAB6F626FA5FD43C705DFA37FED5" box="[1322,1436,260,285]" italics="true" pageId="5" pageNumber="362">P. calypso</emphasis>
</taxonomicName>
(cf. 
<taxonomicName id="34864D27F626FA5FD1D37071FEF3FE88" box="[197,344,295,320]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="5" pageNumber="362" phylum="Arthropoda" rank="species" species="pulchripes">
<emphasis id="C1F2EAB6F626FA5FD1D37071FEF3FE88" box="[197,344,295,320]" italics="true" pageId="5" pageNumber="362">P. pulchripes</emphasis>
</taxonomicName>
<figureCitation id="6BBD2A21F626FA5FD076707FFE0FFE88" box="[352,420,295,320]" captionStart="FIGURE 2" captionStartId="3.[151,250,1309,1331]" captionTargetBox="[209,1370,197,1274]" captionTargetId="figure@3.[206,1380,193,1288]" captionTargetPageId="3" captionText="FIGURE 2. Variation in female genitalia: Phrynus pulchripes: MZUSP 70924 - 1: A. Dorsal view. B. posterior view. MZUSP 70924 - 2: C. Dorsal view. D. posterior view. Phrynus calypso: CASENT 9060553: E. Dorsal view. F. posterior view." httpUri="https://zenodo.org/record/546842/files/figure.png" pageId="5" pageNumber="362">Fig. 2</figureCitation>
B, D, vs 
<taxonomicName id="34864D27F626FA5FD31C7071FDD2FE88" box="[522,633,295,320]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="5" pageNumber="362" phylum="Arthropoda" rank="species" species="calypso">
<emphasis id="C1F2EAB6F626FA5FD31C7071FDD2FE88" box="[522,633,295,320]" italics="true" pageId="5" pageNumber="362">P. calypso</emphasis>
</taxonomicName>
<figureCitation id="6BBD2A21F626FA5FD392707FFD63FE88" box="[644,712,295,320]" captionStart="FIGURE 2" captionStartId="3.[151,250,1309,1331]" captionTargetBox="[209,1370,197,1274]" captionTargetId="figure@3.[206,1380,193,1288]" captionTargetPageId="3" captionText="FIGURE 2. Variation in female genitalia: Phrynus pulchripes: MZUSP 70924 - 1: A. Dorsal view. B. posterior view. MZUSP 70924 - 2: C. Dorsal view. D. posterior view. Phrynus calypso: CASENT 9060553: E. Dorsal view. F. posterior view." httpUri="https://zenodo.org/record/546842/files/figure.png" pageId="5" pageNumber="362">Fig. 2</figureCitation>
F). These differences were also visible when we compared with illustrations of 
<typeStatus id="2C3D8806F626FA5FD0547015FED9FEAD" box="[322,370,333,357]" pageId="5" pageNumber="362">type</typeStatus>
material of 
<emphasis id="C1F2EAB6F626FA5FD0EA7015FD3DFEAD" box="[508,662,332,357]" italics="true" pageId="5" pageNumber="362">
<taxonomicName id="34864D27F626FA5FD0EA7015FD3AFEAD" box="[508,657,332,357]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="5" pageNumber="362" phylum="Arthropoda" rank="species" species="pulchripes">P. pulchripes</taxonomicName>
,
</emphasis>
(See Fig. 
<quantity id="347E9B41F626FA5FD21D7014FCF8FEAD" box="[779,851,332,357]" metricMagnitude="0" metricUnit="m" metricValue="3.2004" pageId="5" pageNumber="362" unit="in" value="126.0">126 in</quantity>
<bibRefCitation id="97174B55F626FA5FD24C7014FBAAFEAC" author="Quintero" box="[858,1025,332,357]" pageId="5" pageNumber="362" refString="Quintero, D. (1981) The Amblypygid genus Phrynus in the Americas (Amblypygi, Phrynidae). Journal of Arachnology, 9, 117 - 166." type="journal article" year="1981">Quintero 1981</bibRefCitation>
). Recently, 
<taxonomicName id="34864D27F626FA5FD5937015FAAFFEAC" box="[1157,1284,333,356]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="5" pageNumber="362" phylum="Arthropoda" rank="species" species="gervaisii">
<emphasis id="C1F2EAB6F626FA5FD5937015FAAFFEAC" box="[1157,1284,333,356]" italics="true" pageId="5" pageNumber="362">P. gervaisii</emphasis>
</taxonomicName>
was reported as a synonym of 
<taxonomicName id="34864D27F626FA5FD0447029FE40FE40" box="[338,491,367,392]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="5" pageNumber="362" phylum="Arthropoda" rank="species" species="barbadensi">
<emphasis id="C1F2EAB6F626FA5FD0447029FE40FE40" box="[338,491,367,392]" italics="true" pageId="5" pageNumber="362">P. barbadensi</emphasis>
</taxonomicName>
s (
<bibRefCitation id="97174B55F626FA5FD3127028FD38FE40" author="Armas" box="[516,659,368,392]" pageId="5" pageNumber="362" refString="Armas, L. F. &amp; Selter, M. (2013) Phrynus gervaisii (Pocock, 1894) is a junior synonym of Phrynus barbadensis (Pocock, 1893) (Amblypygi: Phrynidae). Revista Iberica de Aracnologia, 23, 128 - 132." type="journal article" year="2013">Armas 2013</bibRefCitation>
). Specimens of 
<taxonomicName id="34864D27F626FA5FD2557029FC14FE40" box="[835,959,369,392]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="5" pageNumber="362" phylum="Arthropoda" rank="species" species="gervaisii">
<emphasis id="C1F2EAB6F626FA5FD2557029FC14FE40" box="[835,959,369,392]" italics="true" pageId="5" pageNumber="362">P. gervaisii</emphasis>
</taxonomicName>
were considerably smaller than specimens of 
<emphasis id="C1F2EAB6F626FA5FD1AF70CDFEC3FE65" box="[185,360,404,429]" italics="true" pageId="5" pageNumber="362">
<taxonomicName id="34864D27F626FA5FD1AF70CDFECFFE65" box="[185,356,404,429]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="5" pageNumber="362" phylum="Arthropoda" rank="species" species="barbadensis">P. barbadensis</taxonomicName>
,
</emphasis>
in this case female genital of both are almost equal, with just some differences in size and accentuation of coloration (
<bibRefCitation id="97174B55F626FA5FD0DD70E0FDF7FE18" author="Armas" box="[459,604,440,464]" pageId="5" pageNumber="362" refString="Armas, L. F. &amp; Selter, M. (2013) Phrynus gervaisii (Pocock, 1894) is a junior synonym of Phrynus barbadensis (Pocock, 1893) (Amblypygi: Phrynidae). Revista Iberica de Aracnologia, 23, 128 - 132." type="journal article" year="2013">Armas 2013</bibRefCitation>
). Unlike this case, differences between 
<taxonomicName id="34864D27F626FA5FD50C70E1FB23FE18" box="[1050,1160,439,464]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="5" pageNumber="362" phylum="Arthropoda" rank="species" species="calypso">
<emphasis id="C1F2EAB6F626FA5FD50C70E1FB23FE18" box="[1050,1160,439,464]" italics="true" pageId="5" pageNumber="362">P. calypso</emphasis>
</taxonomicName>
and 
<taxonomicName id="34864D27F626FA5FD5D570E1FAFEFE18" box="[1219,1365,439,464]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="5" pageNumber="362" phylum="Arthropoda" rank="species" species="puclhripes">
<emphasis id="C1F2EAB6F626FA5FD5D570E1FAFEFE18" box="[1219,1365,439,464]" italics="true" pageId="5" pageNumber="362">P. puclhripes</emphasis>
</taxonomicName>
are in relation to shape, patterns of sclerotization and proportions. There are no intermediate character states between larger females of 
<taxonomicName id="34864D27F626FA5FD0777359FE7BFDD0" box="[353,464,511,536]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="5" pageNumber="362" phylum="Arthropoda" rank="species" species="calypso">
<emphasis id="C1F2EAB6F626FA5FD0777359FE7BFDD0" box="[353,464,511,536]" italics="true" pageId="5" pageNumber="362">P. calypso</emphasis>
</taxonomicName>
and smaller females of 
<taxonomicName id="34864D27F626FA5FD3FC7359FCD5FDD0" box="[746,894,511,536]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="5" pageNumber="362" phylum="Arthropoda" rank="species" species="pulchripes">
<emphasis id="C1F2EAB6F626FA5FD3FC7359FCD5FDD0" box="[746,894,511,536]" italics="true" pageId="5" pageNumber="362">P. pulchripes</emphasis>
</taxonomicName>
. In spite of the somatic similarities, the female genitalia of 
<taxonomicName id="34864D27F626FA5FD036737DFE3BFDF5" box="[288,400,548,573]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="5" pageNumber="362" phylum="Arthropoda" rank="species" species="calypso">
<emphasis id="C1F2EAB6F626FA5FD036737DFE3BFDF5" box="[288,400,548,573]" italics="true" pageId="5" pageNumber="362">P. calypso</emphasis>
</taxonomicName>
and 
<taxonomicName id="34864D27F626FA5FD0DA737DFDF4FDF5" box="[460,607,548,573]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="5" pageNumber="362" phylum="Arthropoda" rank="species" species="pulchripes">
<emphasis id="C1F2EAB6F626FA5FD0DA737DFDF4FDF5" box="[460,607,548,573]" italics="true" pageId="5" pageNumber="362">P. pulchripes</emphasis>
</taxonomicName>
present clear differences between them, as interspecific differences with genitals of other species such as 
<taxonomicName id="34864D27F626FA5FD3127311FD02FDA8" box="[516,681,583,608]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="5" pageNumber="362" phylum="Arthropoda" rank="species" species="barbadensis">
<emphasis id="C1F2EAB6F626FA5FD3127311FD02FDA8" box="[516,681,583,608]" italics="true" pageId="5" pageNumber="362">P. barbadensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="34864D27F626FA5FD3C97311FCE7FDA8" box="[735,844,583,608]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="5" pageNumber="362" phylum="Arthropoda" rank="species" species="panche">
<emphasis id="C1F2EAB6F626FA5FD3C97311FCE7FDA8" box="[735,844,583,608]" italics="true" pageId="5" pageNumber="362">P. panche</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="A7F9662CF626FA5FD18176FFFCB6F814" httpUri="https://zenodo.org/record/546844/files/figure.png" pageId="5" pageNumber="362" targetBox="[158,1419,669,1919]" targetPageId="5">
<paragraph id="F33936A4F626FA5FD18176FFFCB6F814" blockId="5.[151,1436,1959,2012]" pageId="5" pageNumber="362">
<emphasis id="C1F2EAB6F626FA5FD18176FFFEB8F874" bold="true" box="[151,275,1959,1981]" pageId="5" pageNumber="362">FIGURE 4.</emphasis>
<taxonomicName id="34864D27F626FA5FD00F76FFFE6AF875" box="[281,449,1959,1981]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="5" pageNumber="362" phylum="Arthropoda" rank="species" species="calypso" status="sp. nov.">
<emphasis id="C1F2EAB6F626FA5FD00F76FFFE6AF875" box="[281,449,1959,1981]" italics="true" pageId="5" pageNumber="362">Phrynus calypso</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAC157CDF626FA5FD0DE76FFFDB2F875" box="[456,537,1959,1981]" pageId="5" pageNumber="362" rank="species">
<emphasis id="C1F2EAB6F626FA5FD0DE76FFFDB9F874" bold="true" box="[456,530,1959,1980]" pageId="5" pageNumber="362">sp. nov</emphasis>
.
</taxonomicNameLabel>
Female Holotype: A. Ventral patella (Pv). B. Dorsal patella (Pd). C. Ventral femur (Fv). D. Dorsal femur (Fd). E. Ventral tibia (Tv). F. Dorsal tibia (Td).
</paragraph>
</caption>
<paragraph id="F33936A4F625FA5CD1937366F9EEFDA3" blockId="6.[131,1862,574,758]" pageId="6" pageNumber="363">
<emphasis id="C1F2EAB6F625FA5CD1937366FF47FD99" bold="true" box="[133,236,574,593]" pageId="6" pageNumber="363">TABLE ·.</emphasis>
Measures of bođy, peđipalp segments anđ femur of legs in 
<taxonomicName id="34864D27F625FA5CD23E7366FC69FD99" box="[808,962,574,593]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="6" pageNumber="363" phylum="Arthropoda" rank="species" species="calypso" status="sp. nov.">
<emphasis id="C1F2EAB6F625FA5CD23E7366FC69FD99" box="[808,962,574,593]" italics="true" pageId="6" pageNumber="363">Phrynus calypso</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAC157CDF625FA5CD2DC7366FBBCFD99" box="[970,1047,574,593]" pageId="6" pageNumber="363" rank="species">
<emphasis id="C1F2EAB6F625FA5CD2DC7366FBB9FD99" bold="true" box="[970,1042,574,593]" pageId="6" pageNumber="363">sp. nov</emphasis>
.
</taxonomicNameLabel>
, measures in mm. When character is missing cell is empty. Paratypes (1-10): 1 anđ 2 corresponđ to CASENT 9060553, 3 anđ 4 to CASENT 9060537, 5 to MZUSP 70892-1, 6 to MZUSP 70892-2, 7 anđ 8 MNRJ 69315, 9 anđ 10 to MNRJ 69312.
</paragraph>
<paragraph id="F33936A4F625FA5CD03D73D8F977FD5B" blockId="6.[131,1862,574,758]" box="[299,1756,640,659]" pageId="6" pageNumber="363">
Holotype 1 2 3 4 5 6 7 
<date id="87381064F625FA5CD4AC73D8F977FD5B" box="[1466,1756,640,659]" pageId="6" pageNumber="363" value="2010-09-08">8 9 10</date>
</paragraph>
<paragraph id="F33936A4F625FA5CD03D73FDFEBCFD3E" blockId="6.[131,1862,574,758]" pageId="6" pageNumber="363">Female Female Male Female Female Male Male Female Male Male Female length 12.7 12.3 14.6 13.0 11.6 20.8 13.6 17 21.8 17.8 18.3 with chelicerae)</paragraph>
<paragraph id="F33936A4F625FA5CD16A7249FF67FCEC" blockId="6.[119,1862,785,1652]" box="[124,204,785,804]" pageId="6" pageNumber="363">Prosoma</paragraph>
<paragraph id="F33936A4F625FA5CD19E726CFF00FCA9" blockId="6.[119,1862,785,1652]" pageId="6" pageNumber="363">Maximum 7.4 7.2 9.3 8.4 6.8 11.7 7.9 9.4 11.2 10.1 9.7 wiđth</paragraph>
<paragraph id="F33936A4F625FA5CD19E7230FF1BFC5D" blockId="6.[119,1862,785,1652]" pageId="6" pageNumber="363">Maximum 4.1 4.1 6.6 6.1 4.7 8.1 5.5 6.6 8.2 6.9 7.2 length</paragraph>
<paragraph id="F33936A4F625FA5CD16172C4FF5BFC67" blockId="6.[119,1862,785,1652]" box="[119,240,924,943]" pageId="6" pageNumber="363">Opisthosoma</paragraph>
<paragraph id="F33936A4F625FA5CD19E7298FF00FC25" blockId="6.[119,1862,785,1652]" pageId="6" pageNumber="363">Maximum 5.5 5.1 5.8 5.8 6.1 8.4 5.5 7.4 7 6.1 6.2 wiđth</paragraph>
<paragraph id="F33936A4F625FA5CD19E72ACFF1BFBE9" blockId="6.[119,1862,785,1652]" pageId="6" pageNumber="363">Maximum 7.7 7.1 8.6 7.4 7.4 11.6 7.6 9.6 11.7 9.7 9.8 length</paragraph>
<paragraph id="F33936A4F625FA5CD03D7572F8EFFBF5" blockId="6.[119,1862,785,1652]" box="[299,1860,1066,1085]" pageId="6" pageNumber="363">Right Left Right Left Right Left Right Left Right Left Right Left Right Left Right Left Right Left Right Left Right Left</paragraph>
<paragraph id="F33936A4F625FA5CD1617517FEA4FBAA" blockId="6.[119,1862,785,1652]" box="[119,271,1103,1122]" pageId="6" pageNumber="363">Peđipalp lengths</paragraph>
<paragraph id="F33936A4F625FA5CD19E752BFF53FAF4" blockId="6.[119,1862,785,1652]" pageId="6" pageNumber="363">Trochanter 1.8 1.8 2 2 2.5 2.5 2.1 2.1 1.5 1.5 3.1 3.3 2.4 2.4 2.4 2.6 3.1 3.1 2.4 2.4 3 3 Femur 3.6 3.4 4.8 4.8 5.3 5.6 4.7 4.7 3.4 3.4 7.7 7.6 4.2 4.4 5.4 5 7.7 7.9 6.1 6.1 5.7 5.7 Patella 3.6 3.6 4.6 4.6 6.2 6.3 5.6 5.6 4.1 4.1 8.0 7.7 4.8 4.9 5.5 5.5 8 8.3 6.3 6.4 6.2 6.2 Tibia 2.1 2.0 2.4 2.4 3 2.9 2.7 2.4 1.9 1.7 4.1 4.0 2.3 2.2 3 3 4.2 4.2 3.2 3.2 3.4 3.4 Tarsus+apotele 2.5 2.4 2.5 2.5 2.7 2.7 3.1 3.1 2.0 2.0 4.2 4.1 2.4 2.4 3.9 3.7 4.2 4.1 3.8 3.6 3.6 3.6 Femur length</paragraph>
<paragraph id="F33936A4F625FA5CD19E7415F8EDFA03" blockId="6.[119,1862,785,1652]" pageId="6" pageNumber="363">Leg I 10.2 10.2 11.8 12.0 16.0 15.5 12.8 12.7 9.7 8.0 21.4 21.4 12.3 12.2 16.3 16.1 23.3 23.2 17.5 17.6 17.8 17.8 Leg II 7.6 7.7 8.1 8.1 11.5 11.4 9.1 9.3 6.9 6.8 17.2 16.0 8.6 8.6 10.9 10.8 16.3 16.3 12.6 12.7 12.9 12.5 Leg III 8.8 8.6 9.0 9.2 13.1 12.8 10.5 10.2 7.9 7.5 17.1 17.1 10.0 9.8 12.4 12.5 8.7 8.6 13.3 13.3 13.4 13.4 Leg IV 7.4 7.0 7.8 7.8 10.4 10.4 8.6 8.6 6.5 6.4 13.7 13.6 6.4 7.8 10.4 10.4 14.5 14.4 11.6 11.6 11.3 11.2</paragraph>
<paragraph id="F33936A4F625FA5CD1807486F89EFA39" blockId="6.[119,1862,785,1652]" box="[150,1845,1502,1521]" pageId="6" pageNumber="363">. segments 33 31 32 32 32 31 31 31 31 31 31 29 31 30 31</paragraph>
<paragraph id="F33936A4F625FA5CD1B374A1FF06F9C4" blockId="6.[119,1862,785,1652]" box="[165,173,1529,1548]" pageId="6" pageNumber="363">I</paragraph>
<paragraph id="F33936A4F625FA5CD180774BFF11F988" blockId="6.[119,1862,785,1652]" pageId="6" pageNumber="363">. segments 67 68 68 67 75 68 68&gt;58 66&gt;50&gt;64 61 68 65 67 tarsus I</paragraph>
<paragraph id="F33936A4F625FA5CD180771FFF49F9BC" blockId="6.[119,1862,785,1652]" pageId="6" pageNumber="363">. segments 3 3 3 3 3 3 3 3 3 3 3 3 3 3 3 3 3 3 3 3 3 basitibia IV</paragraph>
</subSubSection>
<subSubSection id="BB9C652FF624FA52D1D171CFFCE2FD4C" lastPageId="8" lastPageNumber="365" pageId="7" pageNumber="364" type="description">
<paragraph id="F33936A4F624FA5DD1D171CFFE0AFF30" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="364">
<emphasis id="C1F2EAB6F624FA5DD1D171CFFDFFFF78" bold="true" box="[199,596,151,176]" pageId="7" pageNumber="364">
Description. Female (
<typeStatus id="2C3D8806F624FA5DD0C571CFFDECFF78" box="[467,583,151,176]" pageId="7" pageNumber="364" type="holotype">Holotype</typeStatus>
):
</emphasis>
Total length 
<quantity id="347E9B41F624FA5DD3E871C0FCC7FF67" box="[766,876,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="7" pageNumber="364" unit="mm" value="12.7">12.7 mm</quantity>
. Carapace and opisthosoma color dark brown, pedipalps and legs a lighter brown. Dimensions of prosoma, opisthosoma, pedipalpal segments, and all leg femora are provided in Table 1.
</paragraph>
<paragraph id="F33936A4F624FA5DD1D1705BFD0FFEAC" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="364">
<emphasis id="C1F2EAB6F624FA5DD1D1705BFE97FED4" bold="true" box="[199,316,259,284]" italics="true" pageId="7" pageNumber="364">Carapace.</emphasis>
Frontal margin with one lobe on each side, lobes are not joined in the middle area. Carapace presents a group of granules widely spaced, inferior margin with white regions. Lateral and anteromedial eyes clearly visible, ocular tubercle black (
<figureCitation id="6BBD2A21F624FA5DD32B7014FDD4FEAC" box="[573,639,332,357]" captionStart="FIGURE 3" captionStartId="4.[151,250,1734,1756]" captionTargetBox="[178,1412,200,1694]" captionTargetId="figure@4.[174,1412,193,1706]" captionTargetPageId="4" captionText="FIGURE 3. Phrynus calypso sp. nov. Female Holotype: A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." httpUri="https://zenodo.org/record/546843/files/figure.png" pageId="7" pageNumber="364">Fig. 3</figureCitation>
A).
</paragraph>
<paragraph id="F33936A4F624FA5DD1D17037FEF1FE07" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="364">
<emphasis id="C1F2EAB6F624FA5DD1D17037FE84FE40" bold="true" box="[199,303,367,392]" italics="true" pageId="7" pageNumber="364">Sternum.</emphasis>
Tri-segmented; all segments clearly sclerotized, area around segments is sclerotized too. Tritosternum projected anteriorly, elongated, conical, with four setae in basal region, two in medial region and two more in the apex.
</paragraph>
<paragraph id="F33936A4F624FA5DD1D17084FA23FDF4" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="364">
Second segment (tetrasternum) hexagonal with one setae on each side of medial region, rounded shape. Third segment (pentasternum) rhomboidal slightly shorter than the second segment, with one setae on each corner of medial region. Metasternum longitudinally divided with two setae on the posterior margin of each half (
<figureCitation id="6BBD2A21F624FA5DD434737CFACEFDF4" box="[1314,1381,548,573]" captionStart="FIGURE 3" captionStartId="4.[151,250,1734,1756]" captionTargetBox="[178,1412,200,1694]" captionTargetId="figure@4.[174,1412,193,1706]" captionTargetPageId="4" captionText="FIGURE 3. Phrynus calypso sp. nov. Female Holotype: A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." httpUri="https://zenodo.org/record/546843/files/figure.png" pageId="7" pageNumber="364">Fig. 3</figureCitation>
B).
</paragraph>
<paragraph id="F33936A4F624FA5DD1D1731FFBE2FD4C" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="364">
<emphasis id="C1F2EAB6F624FA5DD1D1731FFE97FDA8" bold="true" box="[199,316,583,608]" italics="true" pageId="7" pageNumber="364">Abdomen.</emphasis>
Oblong, color dark brown, with soft punctuations of lighter coloration. Presents patches of light coloration at both sides of each tergite. Carapace wider than the Abdomen (
<figureCitation id="6BBD2A21F624FA5DD2F27334FB8CFD4C" box="[996,1063,620,645]" captionStart="FIGURE 3" captionStartId="4.[151,250,1734,1756]" captionTargetBox="[178,1412,200,1694]" captionTargetId="figure@4.[174,1412,193,1706]" captionTargetPageId="4" captionText="FIGURE 3. Phrynus calypso sp. nov. Female Holotype: A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." httpUri="https://zenodo.org/record/546843/files/figure.png" pageId="7" pageNumber="364">Fig. 3</figureCitation>
C).
</paragraph>
<paragraph id="F33936A4F624FA5DD1D173D7FADFFCDC" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="364">
<emphasis id="C1F2EAB6F624FA5DD1D173D7FE33FD60" bold="true" box="[199,408,655,680]" italics="true" pageId="7" pageNumber="364">Female genitalia.</emphasis>
Genital operculum pentagonal, it extends to second segment of opistosoma, seminal receptacles rounded, dorsal surface lightly sclerotized on basal area, with coloration darker than the rest of the structure which is mainly white. Sclerite of gonopod, with fusiform shape, acuminated apex which is less sclerotized, almost transparent. Sclerite coloration light brown, with apical region translucent (
<figureCitation id="6BBD2A21F624FA5DD5AC73A4FB52FCDC" box="[1210,1273,764,789]" captionStart="FIGURE 1" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[240,1349,1167,1893]" captionTargetId="figure@2.[236,1352,1164,1897]" captionTargetPageId="2" captionText="FIGURE 1. Compared female genitalia: A. Phrynus calypso sp. nov, Holotype. B. Phrynus pulchripes, ICN-Am 85. C. Phrynus barbadensis. MUD-AM 403. D. Phrynus panche. ICN-AM 64." httpUri="https://zenodo.org/record/546841/files/figure.png" pageId="7" pageNumber="364">Fig. 1</figureCitation>
A, 2 E–F).
</paragraph>
<paragraph id="F33936A4F624FA5DD1D17247FD38FC00" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="364">
<emphasis id="C1F2EAB6F624FA5DD1D17247FE96FCF0" bold="true" box="[199,317,799,824]" italics="true" pageId="7" pageNumber="364">Chelicera.</emphasis>
Mesal row of the basal segment of chelicerae with three teeth, the first bilobed, placed in the proximal portion, followed by one tooth shorter in medial portion, and the third, bigger than the others, placed in the distal region (
<figureCitation id="6BBD2A21F624FA5DD048723FFE0AFC48" box="[350,417,871,896]" captionStart="FIGURE 3" captionStartId="4.[151,250,1734,1756]" captionTargetBox="[178,1412,200,1694]" captionTargetId="figure@4.[174,1412,193,1706]" captionTargetPageId="4" captionText="FIGURE 3. Phrynus calypso sp. nov. Female Holotype: A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." httpUri="https://zenodo.org/record/546843/files/figure.png" pageId="7" pageNumber="364">Fig. 3</figureCitation>
D). Ectal row with two teeth of similar sizes, both with an acuminated apex, placed in the more proximal region, and previous to the medial region. There is a small keel on the distal region (
<figureCitation id="6BBD2A21F624FA5DD42072D4FAD7FC6C" box="[1334,1404,908,933]" captionStart="FIGURE 3" captionStartId="4.[151,250,1734,1756]" captionTargetBox="[178,1412,200,1694]" captionTargetId="figure@4.[174,1412,193,1706]" captionTargetPageId="4" captionText="FIGURE 3. Phrynus calypso sp. nov. Female Holotype: A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." httpUri="https://zenodo.org/record/546843/files/figure.png" pageId="7" pageNumber="364">Fig. 3</figureCitation>
F). Mobile segment of chelicerae with five teeth.
</paragraph>
<paragraph id="F33936A4F624FA5DD1D1728BFBE3F9BD" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="364">
<emphasis id="C1F2EAB6F624FA5DD1D1728BFE1EFC25" box="[199,437,979,1005]" italics="true" pageId="7" pageNumber="364">
<emphasis id="C1F2EAB6F624FA5DD1D1728BFE99FC24" bold="true" box="[199,306,979,1004]" italics="true" pageId="7" pageNumber="364">Pedipalp.</emphasis>
Trochanter
</emphasis>
: Prolateral face with four spines; spines Tr1 and Tr3 placed in medial region, Tr2 placed near to the ventral margin. Tr3 and Tr4 with subequal lengths. Spine lengths: Tr2&gt;Tr1&gt;Tr4ŻTr3. Dorsal oblique series of five setal tubercles. Dorsomedial area with one spine with a length slightly longer than one tubercle (
<figureCitation id="6BBD2A21F624FA5DD4797544FF0EFB90" captionStart="FIGURE 3" captionStartId="4.[151,250,1734,1756]" captionTargetBox="[178,1412,200,1694]" captionTargetId="figure@4.[174,1412,193,1706]" captionTargetPageId="4" captionText="FIGURE 3. Phrynus calypso sp. nov. Female Holotype: A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." httpUri="https://zenodo.org/record/546843/files/figure.png" pageId="7" pageNumber="364">Fig. 3</figureCitation>
E). 
<emphasis id="C1F2EAB6F624FA5DD1D87519FEB2FB90" box="[206,281,1089,1112]" italics="true" pageId="7" pageNumber="364">Femur</emphasis>
: Dorsal face with five conspicuous spines, spine Fd4 is reduced, similar in size to a setal tubercle, and there are two little spines more distal than Fd6, in total there are eight spines in the dorsal femur, Fd1 and Fd2 share the same base, Fd3 is adjacent to Fd2, between Fd5 and Fd6 there is a small tubercle. Spine lengths: Fd3&gt;Fd2&gt;Fd5&gt;Fd1&gt;Fd6&gt;Fd7&gt;Fd8&gt;Fd4 (
<figureCitation id="6BBD2A21F624FA5DD36775F4FD1EFB0C" box="[625,693,1196,1221]" captionStart="FIGURE 4" captionStartId="5.[151,250,1959,1981]" captionTargetBox="[158,1419,669,1919]" captionTargetId="figure@5.[151,1435,661,1937]" captionTargetPageId="5" captionText="FIGURE 4. Phrynus calypso sp. nov. Female Holotype: A. Ventral patella (Pv). B. Dorsal patella (Pd). C. Ventral femur (Fv). D. Dorsal femur (Fd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." httpUri="https://zenodo.org/record/546844/files/figure.png" pageId="7" pageNumber="364">Fig. 4</figureCitation>
D). Ventral face with six conspicuous spines, Fv4 very small, more distal than Fv6 are two tubercles present on both pedipalps, and there is also one tubercle between Fv5 and Fv6. Spine lengths: Fv1&gt;Fv2&gt;Fv5&gt;Fv3&gt;Fv6&gt;Fv4 (
<figureCitation id="6BBD2A21F624FA5DD3B475ACFD4CFAC4" box="[674,743,1268,1293]" captionStart="FIGURE 4" captionStartId="5.[151,250,1959,1981]" captionTargetBox="[158,1419,669,1919]" captionTargetId="figure@5.[151,1435,661,1937]" captionTargetPageId="5" captionText="FIGURE 4. Phrynus calypso sp. nov. Female Holotype: A. Ventral patella (Pv). B. Dorsal patella (Pd). C. Ventral femur (Fv). D. Dorsal femur (Fd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." httpUri="https://zenodo.org/record/546844/files/figure.png" pageId="7" pageNumber="364">Fig. 4</figureCitation>
C). 
<emphasis id="C1F2EAB6F624FA5DD21975ACFCCBFAC5" box="[783,864,1268,1293]" italics="true" pageId="7" pageNumber="364">Patella</emphasis>
: Dorsal face with eight major spines, between Pd1 and Pd2 there is a small spine here named Pd1b, to conserve the spines numeration proposed by 
<bibRefCitation id="97174B55F624FA5DD5C8744FFA33FAF8" author="Quintero" box="[1246,1432,1303,1328]" pageId="7" pageNumber="364" refString="Quintero, D. (1981) The Amblypygid genus Phrynus in the Americas (Amblypygi, Phrynidae). Journal of Arachnology, 9, 117 - 166." type="journal article" year="1981">Quintero (1981)</bibRefCitation>
: Pd3&gt;Pd5&gt;Pd2&gt;Pd4&gt;Pd6&gt;Pd1&gt;Pd7&gt;Pd8&gt;Pd1b (
<figureCitation id="6BBD2A21F624FA5DD3AD7464FCAAFA9C" box="[699,769,1340,1365]" captionStart="FIGURE 4" captionStartId="5.[151,250,1959,1981]" captionTargetBox="[158,1419,669,1919]" captionTargetId="figure@5.[151,1435,661,1937]" captionTargetPageId="5" captionText="FIGURE 4. Phrynus calypso sp. nov. Female Holotype: A. Ventral patella (Pv). B. Dorsal patella (Pd). C. Ventral femur (Fv). D. Dorsal femur (Fd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." httpUri="https://zenodo.org/record/546844/files/figure.png" pageId="7" pageNumber="364">Fig. 4</figureCitation>
B). Ventral face with five major spines. Pv3 is small, but it is present on both pedipalps, five very small spines of similar size are present between each major spine: one more proximal than Pv1, between, Pv1 and Pv2, Pv4 and Pv5, Pv5 and Pv6, and one more distal than Pv6. Lengths: Pv2&gt;Pv5&gt;Pv1&gt;Pv4&gt;Pv6&gt;Pv3 (
<figureCitation id="6BBD2A21F624FA5DD0ED74FFFD95FA08" box="[507,574,1447,1472]" captionStart="FIGURE 4" captionStartId="5.[151,250,1959,1981]" captionTargetBox="[158,1419,669,1919]" captionTargetId="figure@5.[151,1435,661,1937]" captionTargetPageId="5" captionText="FIGURE 4. Phrynus calypso sp. nov. Female Holotype: A. Ventral patella (Pv). B. Dorsal patella (Pd). C. Ventral femur (Fv). D. Dorsal femur (Fd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." httpUri="https://zenodo.org/record/546844/files/figure.png" pageId="7" pageNumber="364">Fig. 4</figureCitation>
A). 
<emphasis id="C1F2EAB6F624FA5DD37F74FFFD09FA08" box="[617,674,1447,1472]" italics="true" pageId="7" pageNumber="364">Tibia</emphasis>
: Dorsal face with three major spines, between Td2 and Td3 there is one tubercle and one small spine, more distal than Td3 there are two tubercles, and a small spine smaller than others, here named Td4. Spine lengths: Td2&gt;Td3&gt;Td1&gt;Td4 (
<figureCitation id="6BBD2A21F624FA5DD25E74B7FC27F9C0" box="[840,908,1519,1544]" captionStart="FIGURE 4" captionStartId="5.[151,250,1959,1981]" captionTargetBox="[158,1419,669,1919]" captionTargetId="figure@5.[151,1435,661,1937]" captionTargetPageId="5" captionText="FIGURE 4. Phrynus calypso sp. nov. Female Holotype: A. Ventral patella (Pv). B. Dorsal patella (Pd). C. Ventral femur (Fv). D. Dorsal femur (Fd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." httpUri="https://zenodo.org/record/546844/files/figure.png" pageId="7" pageNumber="364">Fig. 4</figureCitation>
F). Td1 has half the length of Td3. Ventral face with three major spines, and two tubercles between Tv2 and Tv3. Spine lengths: Tv2&gt;Tv3&gt;Tv1 (
<figureCitation id="6BBD2A21F624FA5DD424774CFAD7F9E4" box="[1330,1404,1556,1581]" captionStart="FIGURE 4" captionStartId="5.[151,250,1959,1981]" captionTargetBox="[158,1419,669,1919]" captionTargetId="figure@5.[151,1435,661,1937]" captionTargetPageId="5" captionText="FIGURE 4. Phrynus calypso sp. nov. Female Holotype: A. Ventral patella (Pv). B. Dorsal patella (Pd). C. Ventral femur (Fv). D. Dorsal femur (Fd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." httpUri="https://zenodo.org/record/546844/files/figure.png" pageId="7" pageNumber="364">Fig. 4</figureCitation>
E). 
<emphasis id="C1F2EAB6F624FA5DD181776FFEE2F998" box="[151,329,1591,1616]" italics="true" pageId="7" pageNumber="364">Tarsus+Apotele</emphasis>
: Internal face with two lines of dorsomedial bristles; suture between tarsus and apotele is not visible. Tarsus with an inconspicuous spine on proximal end of dorso-mesal surface.
</paragraph>
<paragraph id="F33936A4F624FA5DD1D177D8FD7FF8CC" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="364">
<emphasis id="C1F2EAB6F624FA5DD1D177D8FEAFF950" bold="true" box="[199,260,1663,1688]" pageId="7" pageNumber="364">
<emphasis id="C1F2EAB6F624FA5DD1D177D8FF56F95F" bold="true" box="[199,253,1664,1687]" italics="true" pageId="7" pageNumber="364">Legs</emphasis>
.
</emphasis>
Femora with lighter coloration rings. Femora lengths: I&gt;III&gt;II&gt;IV (Table 1). Leg I: Tibia with 33 segments and tarsus with 67 segments in the right leg; left leg with 31 segments in the tibia and 68 segments in the tarsus. Leg IV: Basitibia with three segments. Basitibia-distitibia lengths: BT1&gt;DT&gt;BT3&gt;BT2. Basitarsus and Telotarsus with subequal sizes. Tarsus tetramerous.
</paragraph>
<paragraph id="F33936A4F624FA52D1D17657FA37FE64" blockId="7.[151,1437,151,2013]" lastBlockId="8.[151,1437,151,645]" lastPageId="8" lastPageNumber="365" pageId="7" pageNumber="364">
<emphasis id="C1F2EAB6F624FA5DD1D17657FE97F8E0" bold="true" box="[199,316,1807,1832]" pageId="7" pageNumber="364">Variation.</emphasis>
<typeStatus id="2C3D8806F624FA5DD05F7649FE00F8E1" box="[329,427,1809,1833]" pageId="7" pageNumber="364" type="paratype">Paratype</typeStatus>
series measurements and number of segments in basitibia IV, Tibia I and tarsus I, are summarized in Table 1, just adult specimens were measured. 
<emphasis id="C1F2EAB6F624FA5DD253766BFC1AF885" bold="true" box="[837,945,1843,1869]" pageId="7" pageNumber="364">
<emphasis id="C1F2EAB6F624FA5DD253766BFC0CF884" bold="true" box="[837,935,1843,1868]" italics="true" pageId="7" pageNumber="364">Sternum</emphasis>
:
</emphasis>
Setae over all segments of sternum are very variable. 
<emphasis id="C1F2EAB6F624FA5DD1E9760FFE6BF8B8" box="[255,448,1879,1904]" italics="true" pageId="7" pageNumber="364">
<emphasis id="C1F2EAB6F624FA5DD1E9760FFEC5F8B8" bold="true" box="[255,366,1879,1904]" italics="true" pageId="7" pageNumber="364">Pedipalp:</emphasis>
Femur
</emphasis>
: Dorsal face: the size and presence of mentioned tubercles in the 
<typeStatus id="2C3D8806F624FA5DD5B4760FFAAFF8B8" box="[1186,1284,1879,1904]" pageId="7" pageNumber="364" type="holotype">holotype</typeStatus>
is variable in 
<typeStatus id="2C3D8806F624FA5DD1817625FEADF85D" box="[151,262,1917,1941]" pageId="7" pageNumber="364" type="paratype">paratypes</typeStatus>
. Ventral face: the tubercle’s presence is variable, in larger specimens two tubercles can be observed between Fv5 and Fv6, and can be interpreted as small spines. In specimens from 
<collectingCountry id="8B917634F624FA5DD54E76F8FB60F871" box="[1112,1227,1952,1977]" name="Venezuela" pageId="7" pageNumber="364">Venezuela</collectingCountry>
one or two small tubercles may be observed between Fv3 and Fv5. In one male (MNRJ 09315) there were two spines observed between Fv3 and Fv5, and these were present just in the right pedipalp. 
<emphasis id="C1F2EAB6F62BFA52D2DA71CFFBB6FF78" box="[972,1053,151,176]" italics="true" pageId="8" pageNumber="365">Patella</emphasis>
: Dorsal face: in some individuals there is a tubercle anterior to spine Pd1, spine Pd1b is present in many individuals but can be less conspicuous, or even be absent in others. In larger individuals there is one spine smaller than the others that was more distal than Pd8. Ventral face: Tubercles were more conspicuous in larger sized individuals; in some cases, a second tubercle was present more proximal than Pv1. 
<emphasis id="C1F2EAB6F62BFA52D32D707FFDD8FE88" box="[571,627,295,320]" italics="true" pageId="8" pageNumber="365">Tibia</emphasis>
: Dorsal face: presence of tubercles between spines was variable. Ventral face: In some individuals there was just one tubercle between Tv2 and Tv3. 
<emphasis id="C1F2EAB6F62BFA52D2FF7014FB8CFEAB" bold="true" box="[1001,1063,332,355]" italics="true" pageId="8" pageNumber="365">Legs.</emphasis>
Number of segments of tibia and tarsus of leg I was variable among the individuals, number of tibial segments keeps between 31 and 33, and tarsal segments between 61 and 75. In observed specimens, number of segments did not depend on size or sex (Table. 1).
</paragraph>
<paragraph id="F33936A4F62BFA52D1D170EFFBCEFE3D" blockId="8.[151,1437,151,645]" pageId="8" pageNumber="365">
<emphasis id="C1F2EAB6F62BFA52D1D170EFFEA0FE07" bold="true" box="[199,267,439,464]" pageId="8" pageNumber="365">Male.</emphasis>
Observed males were larger in size than females (Table. 1), spination pattern as in females, genital operculum has an inferior margin oval and cover part of third segment of opisthosoma.
</paragraph>
<paragraph id="F33936A4F62BFA52D1D170A7FCE2FD4C" blockId="8.[151,1437,151,645]" pageId="8" pageNumber="365">
<emphasis id="C1F2EAB6F62BFA52D1D170A7FED3FDD0" bold="true" box="[199,376,511,536]" pageId="8" pageNumber="365">
<emphasis id="C1F2EAB6F62BFA52D1D170A7FEDBFDD0" bold="true" box="[199,368,511,536]" italics="true" pageId="8" pageNumber="365">Male genitalia</emphasis>
.
</emphasis>
(MZUSP 70892): LaM longer than Pi. LoL 1 and LoL 2 subequal in length, both densely covered with minute projections, in LoL 1 projections are tubular, thin and elongated, placed close together; in LoL 2 are tubular and short, some of these with ramifications at the apex, spaced over a base clearly visible. In dorsal view the LoD rounded and shorter than LoL1 (
<figureCitation id="6BBD2A21F62BFA52D3B27334FD4DFD4C" box="[676,742,620,645]" captionStart="FIGURE 7" captionStartId="11.[151,250,1801,1823]" captionTargetBox="[162,1417,273,1774]" captionTargetId="figure@11.[157,1429,265,1780]" captionTargetPageId="11" captionText="FIGURE 7. Male genitalia: Phrynus pulchripes ICN-AM 86: A. Dorsal view. B. Ventral view. E. Detail of ornamentation of LoL 1 and LoL 2. Phrynus calypso sp. nov. MZUSP 70892: C. Dorsal view. D. Ventral view. F. Detail of ornamentation of LoL 1 and LoL 2. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1 = lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod." httpUri="https://zenodo.org/record/546847/files/figure.png" pageId="8" pageNumber="365">Fig. 7</figureCitation>
C, D, F).
</paragraph>
</subSubSection>
</treatment>
</document>