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<document id="B59D8AF67E507F9FED6284DAD9AE88F2" ID-CLB-Dataset="3142" ID-DOI="10.11646/zootaxa.5072.5.2" ID-GBIF-Dataset="cdde4106-d3d5-4294-9f6c-1e82afd8d580" ID-ISSN="1175-5326" ID-Zenodo-Dep="5748997" ID-ZooBank="B1D30BB4-8CAB-419E-98DE-02D97726137C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638425477373" checkinUser="plazi" docAuthor="Hutchinson, Paul M. &amp; Allsopp, Peter G." docDate="2021" docId="79268507FFC8FFA2CBE5D0B3FBAFFEBD" docLanguage="en" docName="zootaxa.5072.5.2.pdf" docOrigin="Zootaxa 5072 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Enracius richardsi Hutchinson &amp; Allsopp 2021, new species" docType="treatment" docUuid="FDCD40FD-4723-4AB2-B8BD-76E959807F55" docUuidSource="ZooBank" docVersion="8" lastPageNumber="446" masterDocId="851FFD7FFFCDFFA5CB72D024FF89FFA0" masterDocTitle="Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina)" masterLastPageNumber="462" masterPageNumber="439" pageNumber="444" updateTime="1699471722501" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="73A18593AFF92D0FEAD17523DF991390" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="239ED9FC026C026A00E5D5AFB306AE38">
<mods:title id="6C867C21B67902279E25EA707C255FCF">Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina)</mods:title>
</mods:titleInfo>
<mods:name id="0E04C6FB468B95518E88B0E0BC417D3F" type="personal">
<mods:role id="8568D44476696315940FE730075CCFC4">
<mods:roleTerm id="0BA761A690636F4BCEE1C8663CDD69C5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="27E665AD4C341ADC269F2B69EEB349B8">Hutchinson, Paul M.</mods:namePart>
<mods:affiliation id="4060A81B543B5432B8B9B83562B291D7">Department of Primary Industries and Regional Development (Quarantine WA), Locked Bag 69, Welshpool DC, Western Australia 6986, Australia.</mods:affiliation>
</mods:name>
<mods:name id="49A2CA759DD1212029FE53AA9E0A6AC0" type="personal">
<mods:role id="F25D2EEBD1D3AEDF9BB5B7BC37169E33">
<mods:roleTerm id="705014EAD2B3401CEEEB6AF33DF29693">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="611B13F50599A433F2DFEF7A6886B6D9">Allsopp, Peter G.</mods:namePart>
<mods:affiliation id="07DDD6AF64795E0715960F39D2FDEE18">Queensland Museum, PO Box 3300, South Brisbane, Queensland 4101, Australia.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="8F07990D8717441E30189FAA11D700D1">text</mods:typeOfResource>
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<mods:titleInfo id="EEBF03822DB33D8E816984FEE7E25A26">
<mods:title id="C4740F8DF99A540E8203DA57259753ED">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="24F69D6F94A5BA60D13C287376C04913">
<mods:date id="DA045D7570142D82B89A2C6D141F02BA">2021</mods:date>
<mods:detail id="5000F5E8D73D483482720734577D5369" type="pubDate">
<mods:number id="BCD267DB0872E3911B0CFD0A4F702A37">2021-12-02</mods:number>
</mods:detail>
<mods:detail id="85FFBCF31E64F347A96ABF99875D3320" type="volume">
<mods:number id="C572F4A08A756C4A35610499370B5807">5072</mods:number>
</mods:detail>
<mods:detail id="7AB8D85AD866F833C2DDB02F49959D26" type="issue">
<mods:number id="2CB1F7AF345392ADA473C79E2F8BA30C">5</mods:number>
</mods:detail>
<mods:extent id="7E038E51B3FEB1A929A9B5DE3C998F5A" unit="page">
<mods:start id="7CB97F6DB4BD4435378B293E65083CDC">439</mods:start>
<mods:end id="43ECE2EDB7A6A721CC7BF61D8BDE5ABF">462</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:classification id="DC61DA7F05B8A9E9EA58F860368691EF">journal article</mods:classification>
<mods:identifier id="276C85740E2EE179E45CE23B672FDEE6" type="CLB-Dataset">3142</mods:identifier>
<mods:identifier id="0599F3DDE130D1CF9858445F93630D7D" type="DOI">10.11646/zootaxa.5072.5.2</mods:identifier>
<mods:identifier id="D8CEEA786192EA8AAD29E0BE2877AB81" type="GBIF-Dataset">cdde4106-d3d5-4294-9f6c-1e82afd8d580</mods:identifier>
<mods:identifier id="F8032240C855EDE38A978656D3CDD616" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A8B46FD2DD181CB260E649D6D11DFD3A" type="Zenodo-Dep">5748997</mods:identifier>
<mods:identifier id="B8A8E7DED6B66D7140DBEBF7D53417E7" type="ZooBank">B1D30BB4-8CAB-419E-98DE-02D97726137C</mods:identifier>
</mods:mods>
<treatment id="79268507FFC8FFA2CBE5D0B3FBAFFEBD" ID-DOI="http://doi.org/10.5281/zenodo.5750344" ID-GBIF-Taxon="190874938" ID-Zenodo-Dep="5750344" LSID="urn:lsid:zoobank.org:act:FDCD40FD-4723-4AB2-B8BD-76E959807F55" httpUri="http://treatment.plazi.org/id/79268507FFC8FFA2CBE5D0B3FBAFFEBD" lastPageId="7" lastPageNumber="446" pageId="5" pageNumber="444">
<subSubSection id="B995679AFFC8FFA0CBE5D0B3FC47FF76" pageId="5" pageNumber="444" type="nomenclature">
<paragraph id="F1303411FFC8FFA0CBE5D0B3FC47FF76" blockId="5.[151,974,151,214]" pageId="5" pageNumber="444">
<heading id="AA78837DFFC8FFA0CBE5D0B3FCBEFF11" bold="true" box="[151,823,151,178]" fontSize="11" level="1" pageId="5" pageNumber="444" reason="1">
<emphasis id="C3FBE803FFC8FFA0CBE5D0B3FCBEFF11" bold="true" box="[151,823,151,178]" pageId="5" pageNumber="444">
<taxonomicName id="368F4F92FFC8FFA0CBE5D0B3FD13FF11" authority="Hutchinson &amp; Allsopp" authorityName="Hutchinson &amp; Allsopp" authorityYear="2021" box="[151,666,151,178]" class="Insecta" family="Dynastidae" genus="Enracius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="richardsi" status="sp. nov.">
<emphasis id="C3FBE803FFC8FFA0CBE5D0B3FEF3FF12" bold="true" box="[151,378,151,178]" italics="true" pageId="5" pageNumber="444">Enracius richardsi</emphasis>
Hutchinson &amp; Allsopp
</taxonomicName>
,
<taxonomicNameLabel id="D8C85578FFC8FFA0C9D5D0B3FCBEFF11" box="[679,823,151,177]" pageId="5" pageNumber="444" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
<uri id="851E3813FFC8FFA0CBE5D09FFC47FF76" box="[151,974,187,214]" pageId="5" pageNumber="444">
urn:lsid:zoobank.org:act:
<uuid id="85290EC4FFC8FFA0CAD5D09FFC47FF76" box="[423,974,187,214]" pageId="5" pageNumber="444">FDCD40FD-4723-4AB2-B8BD-76E959807F55</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="B995679AFFC8FFA0CBE5D127FCB1FE71" pageId="5" pageNumber="444" type="materials_examined">
<paragraph id="F1303411FFC8FFA0CBE5D127FAFCFEC5" blockId="5.[151,1437,259,1942]" pageId="5" pageNumber="444">
<emphasis id="C3FBE803FFC8FFA0CBE5D127FEA0FEBD" bold="true" box="[151,297,259,285]" pageId="5" pageNumber="444">Type series.</emphasis>
<materialsCitation id="41E73E4CFFC8FFA0CA44D127FAFCFEC5" ID-GBIF-Occurrence="3413046301" collectingDate="1970-12-18" collectionCode="WADA" collectorName="W. Aust. &amp; K. T. Richards" country="Australia" determinerName="P. B. Carne" latitude="-34.0" location="Western Australia" longLatPrecision="0" longitude="119.7" municipality="Fitzgerald River National Park" pageId="5" pageNumber="444" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<emphasis id="C3FBE803FFC8FFA0CA44D127FD2DFEBD" bold="true" box="[310,676,259,285]" pageId="5" pageNumber="444">
<typeStatus id="2E348AB3FFC8FFA0CA44D127FE2AFEBD" box="[310,419,259,285]" pageId="5" pageNumber="444" type="holotype">Holotype</typeStatus>
female (
<figureCitation id="69B42894FFC8FFA0C966D127FD1EFEBD" box="[532,663,259,285]" captionStart="FIGURES 8–13" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[173,1415,219,1920]" captionTargetId="figure-26@6.[173,1415,181,1920]" captionTargetPageId="6" captionText="FIGURES 8–13. Enracius richardsi Hutchinson &amp; Allsopp new species, holotype female. 8, Dorsal view; 9, ventral view; 10, lateral view; 11, head dorsal view; 12, head ventral view; 13, labels and left antennomeres 2–10." figureDoi="http://doi.org/10.5281/zenodo.5749007" httpUri="https://zenodo.org/record/5749007/files/figure.png" pageId="5" pageNumber="444">Figs. 8–13</figureCitation>
).
</emphasis>
<collectingCountry id="89987481FFC8FFA0C9C2D127FCDBFEBD" box="[688,850,259,285]" name="Australia" pageId="5" pageNumber="444">AUSTRALIA</collectingCountry>
,
<collectingRegion id="334BFAF3FFC8FFA0C810D127FB1CFEBD" box="[866,1173,259,285]" country="Australia" name="Western Australia" pageId="5" pageNumber="444">WESTERN AUSTRALIA</collectingRegion>
: “Fitzgerald Riv. Res. [
<collectingMunicipality id="1154AE6BFFC8FFA0CBEFD103FD81FEE1" box="[157,520,295,321]" pageId="5" pageNumber="444">Fitzgerald River National Park</collectingMunicipality>
,
<geoCoordinate id="94BB52D6FFC8FFA0C965D103FDD9FEE1" box="[535,592,295,321]" degrees="34" direction="south" orientation="latitude" pageId="5" pageNumber="444" precision="55555" value="-34.0">34°S</geoCoordinate>
,
<geoCoordinate id="94BB52D6FFC8FFA0C912D102FD34FEE1" box="[608,701,294,321]" degrees="119.7" direction="east" orientation="longitude" pageId="5" pageNumber="444" precision="5555" value="119.7">119.7°E</geoCoordinate>
]
<collectorName id="5C7A51C7FFC8FFA0C9BCD10CFCBAFEE1" box="[718,819,295,322]" pageId="5" pageNumber="444">W. Aust.</collectorName>
<date id="853112D1FFC8FFA0C84CD103FC37FEE1" box="[830,958,295,321]" pageId="5" pageNumber="444" value="1970-12-18">
<collectingDate id="9575EB39FFC8FFA0C84CD103FC37FEE1" box="[830,958,295,321]" pageId="5" pageNumber="444" value="1970-12-18">18.xii.1970</collectingDate>
</date>
<collectorName id="5C7A51C7FFC8FFA0C8BAD10CFBE0FEE1" box="[968,1129,295,322]" pageId="5" pageNumber="444">K.T. Richards</collectorName>
| 119 | Agriculture (Dept)
<location id="F45062CAFFC8FFA0CBE5D168FED6FEC5" LSID="urn:lsid:plazi:treatment:79268507FFC8FFA2CBE5D0B3FBAFFEBD:F45062CAFFC8FFA0CBE5D168FED6FEC5" box="[151,351,331,358]" country="Australia" latitude="-34.0" longLatPrecision="0" longitude="119.7" municipality="Fitzgerald River National Park" name="Western Australia" pageId="5" pageNumber="444" stateProvince="Western Australia">Western Australia</location>
37851 | Dynastine genus?
<determinerName id="04ECD8D5FFC8FFA0C9FED168FC8AFEC5" box="[652,771,331,358]" pageId="5" pageNumber="444">P.B. Carne</determinerName>
det. 1980”; left antennomeres 2–10 on card; in
<collectionCode id="979EACD4FFC8FFA0CE68D16FFAFBFEC5" box="[1306,1394,331,357]" country="Australia" httpUri="http://grbio.org/cool/mnrx-zeat" name="Western Australia Department of Agriculture" pageId="5" pageNumber="444">WADA</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="F1303411FFC8FFA0CBB5D14BFCB1FE71" blockId="5.[151,1437,259,1942]" pageId="5" pageNumber="444">
<materialsCitation id="41E73E4CFFC8FFA0CBB5D14BFCBDFE71" ID-GBIF-Occurrence="3413046323" collectingDate="1974-12-10" collectionCode="WADA" collectorName="W. Aust. &amp; K. T. Richards" country="Australia" county="Mt. Ragged" determinerName="P. B. Carne" latitude="-33.8" location="Western Australia" longLatPrecision="7223" longitude="123.5" municipality="Cape Arid National Park" pageId="5" pageNumber="444" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia" typeStatus="paratype">
<emphasis id="C3FBE803FFC8FFA0CBB5D14BFEB2FE29" bold="true" box="[199,315,367,393]" pageId="5" pageNumber="444">
<typeStatus id="2E348AB3FFC8FFA0CBB5D14BFEBFFE29" box="[199,310,367,393]" pageId="5" pageNumber="444" type="paratype">Paratype</typeStatus>
.
</emphasis>
<collectingCountry id="89987481FFC8FFA0CA34D14BFE61FE29" box="[326,488,367,393]" name="Australia" pageId="5" pageNumber="444">AUSTRALIA</collectingCountry>
,
<collectingRegion id="334BFAF3FFC8FFA0CA85D14BFCA0FE29" box="[503,809,367,393]" country="Australia" name="Western Australia" pageId="5" pageNumber="444">WESTERN AUSTRALIA</collectingRegion>
:
<specimenCount id="E789FF98FFC8FFA0C84BD14BFCD7FE29" box="[825,862,367,393]" pageId="5" pageNumber="444" type="female">1♀</specimenCount>
: “
<quantity id="367799F4FFC8FFA0C808D14BFC6BFE29" box="[890,994,367,394]" metricMagnitude="4" metricUnit="m" metricValue="4.02336" pageId="5" pageNumber="444" unit="mi" value="25.0">25 miles</quantity>
[
<quantity id="367799F4FFC8FFA0C886D14BFBCFFE29" box="[1012,1094,366,393]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="444" unit="km" value="40.0">40 km</quantity>
] sth
<collectingCounty id="18514C9DFFC8FFA0CFF4D154FA86FE29" box="[1158,1295,367,394]" pageId="5" pageNumber="444">Mt. Ragged</collectingCounty>
[
<collectingMunicipality id="1154AE6BFFC8FFA0CE50D14BFEBEFE0D" pageId="5" pageNumber="444">Cape Arid National Park</collectingMunicipality>
,
<geoCoordinate id="94BB52D6FFC8FFA0CA36D1B7FE1BFE0D" box="[324,402,403,429]" degrees="33.8" direction="south" orientation="latitude" pageId="5" pageNumber="444" precision="5555" value="-33.8">33.8°S</geoCoordinate>
,
<geoCoordinate id="94BB52D6FFC8FFA0CAD2D1B6FE77FE0D" box="[416,510,402,429]" degrees="123.5" direction="east" orientation="longitude" pageId="5" pageNumber="444" precision="5555" value="123.5">123.5°E</geoCoordinate>
]
<collectorName id="5C7A51C7FFC8FFA0C97FD1B0FDF9FE0D" box="[525,624,403,430]" pageId="5" pageNumber="444">W. Aust.</collectorName>
<date id="853112D1FFC8FFA0C90AD1B7FD7EFE0D" box="[632,759,403,429]" pageId="5" pageNumber="444" value="1974-12-10">
<collectingDate id="9575EB39FFC8FFA0C90AD1B7FD7EFE0D" box="[632,759,403,429]" pageId="5" pageNumber="444" value="1974-12-10">10.xii.1974</collectingDate>
</date>
<collectorName id="5C7A51C7FFC8FFA0C872D1B0FC29FE0D" box="[768,928,403,430]" pageId="5" pageNumber="444">K.T.Richards</collectorName>
| 165 | Agriculture (Dept)
<location id="F45062CAFFC8FFA0CFA1D1B0FA14FE0D" LSID="urn:lsid:plazi:treatment:79268507FFC8FFA2CBE5D0B3FBAFFEBD:F45062CAFFC8FFA0CFA1D1B0FA14FE0D" box="[1235,1437,403,430]" country="Australia" county="Mt. Ragged" latitude="-33.8" longLatPrecision="7223" longitude="123.5" municipality="Cape Arid National Park" name="Western Australia" pageId="5" pageNumber="444" stateProvince="Western Australia">Western Australia</location>
37850 | Dynastine genus?
<determinerName id="04ECD8D5FFC8FFA0CACCD19CFDBCFE71" box="[446,565,439,466]" pageId="5" pageNumber="444">P.B. Carne</determinerName>
det. 1980”; in
<collectionCode id="979EACD4FFC8FFA0C9AED193FCBDFE71" box="[732,820,439,465]" country="Australia" httpUri="http://grbio.org/cool/mnrx-zeat" name="Western Australia Department of Agriculture" pageId="5" pageNumber="444">WADA</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B995679AFFC8FFA0CBB5D1FFFB1AFA3D" pageId="5" pageNumber="444" type="description">
<paragraph id="F1303411FFC8FFA0CBB5D1FFFA12FA91" blockId="5.[151,1437,259,1942]" pageId="5" pageNumber="444">
<emphasis id="C3FBE803FFC8FFA0CBB5D1FFFD35FE55" bold="true" box="[199,700,475,501]" pageId="5" pageNumber="444">
Description of
<typeStatus id="2E348AB3FFC8FFA0CA03D1FFFE51FE55" box="[369,472,475,501]" pageId="5" pageNumber="444" type="holotype">holotype</typeStatus>
female (
<figureCitation id="69B42894FFC8FFA0C944D1FFFD26FE55" box="[566,687,475,501]" captionStart="FIGURES 8–13" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[173,1415,219,1920]" captionTargetId="figure-26@6.[173,1415,181,1920]" captionTargetPageId="6" captionText="FIGURES 8–13. Enracius richardsi Hutchinson &amp; Allsopp new species, holotype female. 8, Dorsal view; 9, ventral view; 10, lateral view; 11, head dorsal view; 12, head ventral view; 13, labels and left antennomeres 2–10." figureDoi="http://doi.org/10.5281/zenodo.5749007" httpUri="https://zenodo.org/record/5749007/files/figure.png" pageId="5" pageNumber="444">Figs. 8–12</figureCitation>
).
</emphasis>
Protarsomeres 2–5,left mesotarsomeres 3–5,right mesotarsomeres 4–5, right metatarsomeres 4–5 missing. Body
<quantity id="367799F4FFC8FFA0C9E7D1DBFD71FDBA" box="[661,760,511,538]" metricMagnitude="-2" metricUnit="m" metricValue="2.12" pageId="5" pageNumber="444" unit="mm" value="21.2">21.2 mm</quantity>
long, pronotum 9.0 mm wide at widest point, elytra
<quantity id="367799F4FFC8FFA0CE49D1DBFA14FDBA" box="[1339,1437,511,538]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="5" pageNumber="444" unit="mm" value="11.3">11.3 mm</quantity>
wide;
<taxonomicName id="368F4F92FFC8FFA0CBACD207FEE4FD9E" box="[222,365,547,574]" form="convex" pageId="5" pageNumber="444" rank="form">form convex</taxonomicName>
depressed, robust; dorsal surface matt, head, pronotum, elytra and pygidium black; scutellum, elytral suture, sternum, abdominal ventrites and legs dark brown to black. Maxillary palp with palpomeres 1 and 3 subequal in length, palpomere 2 longer but shorter than palpomere 4, palpomere 4 about 3 times longer than greatest width and bearing small, elongate, shallow sensorium on basal half; mandibles with lateral margin bearing long setae; mentum wide, bearing very long setae laterally, glabrous medially. Antenna with 10 antennomeres, club subequal in length to antennomeres 2–7 combined, scape bearing long setae, antennomeres 2–7 glabrous. Clypeus transverse, almost trapezoidal, anterior angles rounded, anteriorly emarginate, truncate face vertical and glabrous, surface rugulose; clypeofrontal ridge posteriorly arcuate, weakly raised.Frons rugulose; ocular canthi sub-continuous with line of clypeus, dorsal surface rugulose, anterior margin bearing row of stout setae. Pronotum widest basally, subparallel on basal half, angulate medially, linear and tapering to acute anterior angles, recurved to linear transverse anterior margin, anterior margin membranous, basal, lateral and anterior ridges complete; disc evenly convex bearing sparse micropunctures becoming larger and denser adjacent to margins. Scutellum sparsely punctate on disc. Elytral striae linear-punctate, intervals almost impunctate, apical declivity rugulose, suture punctate-striate, epipleura nonemarginate, continuous to suture, lateral setae restricted to base adjacent to mesepimeron. Pygidium moderately convex, glabrous, disc sparsely micropunctate, denser at lateral margins, apical ridge glabrous. Prosternal process long, slender, apex angled posteriorly and setose. Metasternum and metepisternum densely punctate, laterally rugulose and bearing long, light-brown setae. Abdominal sternites with a line of sparse setae laterally adjacent to posterior margins, suture between sternites 6 and 7 wide and glossy yellow, sternite 7 bearing narrow sulcus laterally, apical margin bearing row of long setae. Protibia tridentate, denticles equidistant; meso- and metatibia unicarinate, apical cilia short, blunt and widely set; metafemur stout, anterior margin very convex; metatibia stout, flat on inner surface with dorsal margin concave post medial carina, inner spur 1.5 times length of outer spur, both broad, straight and widest pre-apically and apex at oblique angle; metatarsomeres combined longer than metatibia.
</paragraph>
<paragraph id="F1303411FFC8FFA0CBB5D51FFE01FAF5" blockId="5.[151,1437,259,1942]" box="[199,392,1339,1365]" pageId="5" pageNumber="444">
<emphasis id="C3FBE803FFC8FFA0CBB5D51FFE82FAF5" bold="true" box="[199,267,1339,1365]" pageId="5" pageNumber="444">Male.</emphasis>
Unknown.
</paragraph>
<paragraph id="F1303411FFC8FFA0CBB5D57BFB1AFA3D" blockId="5.[151,1437,259,1942]" pageId="5" pageNumber="444">
<emphasis id="C3FBE803FFC8FFA0CBB5D57BFEB6FAD9" bold="true" box="[199,319,1375,1401]" pageId="5" pageNumber="444">Variation.</emphasis>
<typeStatus id="2E348AB3FFC8FFA0CA39D544FE24FADA" box="[331,429,1376,1402]" pageId="5" pageNumber="444" type="paratype">Paratype</typeStatus>
length
<quantity id="367799F4FFC8FFA0C979D57BFDFFFADA" box="[523,630,1375,1402]" metricMagnitude="-2" metricUnit="m" metricValue="1.91" pageId="5" pageNumber="444" unit="mm" value="19.1">19.1 mm</quantity>
, pronotal width
<quantity id="367799F4FFC8FFA0C849D57BFC11FADA" box="[827,920,1375,1402]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="5" pageNumber="444" unit="mm" value="8.4">8.4 mm</quantity>
, elytral width
<quantity id="367799F4FFC8FFA0CF3AD57BFB3DFADA" box="[1096,1204,1375,1402]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="5" pageNumber="444" unit="mm" value="10.3">10.3 mm</quantity>
; elytra dark brown, pronotum laterally suffused brown; left meso- and metatibia present and longer than tibiae.
</paragraph>
</subSubSection>
<subSubSection id="B995679AFFC8FFA0CBB5D583FE41F8A6" pageId="5" pageNumber="444" type="diagnosis">
<paragraph id="F1303411FFC8FFA0CBB5D583FE41F8A6" blockId="5.[151,1437,259,1942]" pageId="5" pageNumber="444">
<emphasis id="C3FBE803FFC8FFA0CBB5D583FE59FA61" bold="true" box="[199,464,1447,1473]" pageId="5" pageNumber="444">Differential diagnosis.</emphasis>
Females of
<taxonomicName id="368F4F92FFC8FFA0C92DD58CFCBAFA61" authorityName="Hutchinson &amp; Allsopp" authorityYear="2021" box="[607,819,1447,1473]" class="Insecta" family="Dynastidae" genus="Enracius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="richardsi">
<emphasis id="C3FBE803FFC8FFA0C92DD58CFCBAFA61" box="[607,819,1447,1473]" italics="true" pageId="5" pageNumber="444">Enracius richardsi</emphasis>
</taxonomicName>
can be easily differentiated from females of
<taxonomicName id="368F4F92FFC8FFA0CE45D58CFF7FFA45" authorityName="Dechambre" authorityYear="1999" class="Insecta" family="Dynastidae" genus="Enracius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="C3FBE803FFC8FFA0CE45D58CFF7FFA45" italics="true" pageId="5" pageNumber="444">Enracius longipes</emphasis>
</taxonomicName>
by the following characters (
<taxonomicName id="368F4F92FFC8FFA0C93FD5E8FD47FA45" authorityName="Dechambre" authorityYear="1999" box="[589,718,1483,1509]" class="Insecta" family="Dynastidae" genus="Enracius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="C3FBE803FFC8FFA0C93FD5E8FD47FA45" box="[589,718,1483,1509]" italics="true" pageId="5" pageNumber="444">E. longipes</emphasis>
</taxonomicName>
characters in brackets): body&gt;
<quantity id="367799F4FFC8FFA0CF34D5EFFB11FA46" box="[1094,1176,1483,1510]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="444" unit="mm" value="19.0">19 mm</quantity>
long (
<quantity id="367799F4FFC8FFA0CF96D5EFFA06FA46" box="[1252,1423,1483,1510]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.75" metricValueMin="1.35" pageId="5" pageNumber="444" unit="mm" value="15.5" valueMax="17.5" valueMin="13.5">13.5–17.5 mm</quantity>
); clypeofrontal ridge posteriorly arcuate (linear); ocular canthi subcontinuous with line of clypeus (angular to clypeus); maxillary palpomeres 1–3 of unequal length (subequal); mentum broader; pronotum micropunctate and evenly arcuate (bearing fine punctures becoming rugulose along lateral and anterior margins, anterior margin weakly raised medially, preceding small flat area and narrow impunctate midline); elytra stria deep (weakly impressed), intervals almost impunctate (abundantly micropunctate to rugulose), epipleural lateral setae basal (lateral setae to apical calli); pygidium moderately convex (almost flat); abdominal ventrite 7 bearing pre-apical sulcus (absent); meso-and metatibial cilia short and blunt (apical cilia long, sharp); apical spurs short and stout (metatibial inner spur 1.5 times length of outer spur).
</paragraph>
</subSubSection>
<subSubSection id="B995679AFFC8FFA0CBB5D72BFEFAF835" pageId="5" pageNumber="444" type="etymology">
<paragraph id="F1303411FFC8FFA0CBB5D72BFEFAF835" blockId="5.[151,1437,259,1942]" pageId="5" pageNumber="444">
<emphasis id="C3FBE803FFC8FFA0CBB5D72BFEC7F889" bold="true" box="[199,334,1807,1833]" pageId="5" pageNumber="444">Etymology.</emphasis>
Named for WADA entomologist Kevin T. Richards (1932–2009) who completed a Master’s degree on potential pests in the Ord Irrigation Scheme in the Kimberley Region of
<collectingRegion id="334BFAF3FFC8FFA0C8AAD710FB2BF8ED" box="[984,1186,1843,1870]" country="Australia" name="Western Australia" pageId="5" pageNumber="444">Western Australia</collectingRegion>
. Later in Perth, he was curator of the WADA Entomology collection of which he made many contributions. He collected both specimens of this new species.
</paragraph>
</subSubSection>
<caption id="A5F06499FFCBFFA3CBE5D7B3FBE1F874" ID-DOI="http://doi.org/10.5281/zenodo.5749007" ID-Zenodo-Dep="5749007" httpUri="https://zenodo.org/record/5749007/files/figure.png" pageId="6" pageNumber="445" startId="6.[151,264,1943,1967]" targetBox="[173,1415,219,1920]" targetPageId="6">
<paragraph id="F1303411FFCBFFA3CBE5D7B3FBE1F874" blockId="6.[151,1436,1943,2004]" pageId="6" pageNumber="445">
<emphasis id="C3FBE803FFCBFFA3CBE5D7B3FECEF80F" bold="true" box="[151,327,1943,1967]" pageId="6" pageNumber="445">FIGURES 8–13.</emphasis>
<taxonomicName id="368F4F92FFCBFFA3CA3CD7BCFD7FF810" authority="Hutchinson &amp; Allsopp" authorityName="Hutchinson &amp; Allsopp" authorityYear="2021" box="[334,758,1944,1968]" class="Insecta" family="Dynastidae" genus="Enracius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="richardsi" status="sp. nov.">
<emphasis id="C3FBE803FFCBFFA3CA3CD7BCFD82F810" box="[334,523,1944,1968]" italics="true" pageId="6" pageNumber="445">Enracius richardsi</emphasis>
Hutchinson &amp; Allsopp
</taxonomicName>
<emphasis id="C3FBE803FFCBFFA3C98ED7BCFCF0F80F" bold="true" box="[764,889,1943,1968]" pageId="6" pageNumber="445">
<taxonomicNameLabel id="D8C85578FFCBFFA3C98ED7BCFCF0F80F" box="[764,889,1943,1968]" pageId="6" pageNumber="445" rank="species">new species</taxonomicNameLabel>
</emphasis>
, holotype female.
<emphasis id="C3FBE803FFCBFFA3CF44D7B3FBC0F80F" bold="true" box="[1078,1097,1943,1967]" pageId="6" pageNumber="445">8,</emphasis>
Dorsal view;
<emphasis id="C3FBE803FFCBFFA3CFAAD7B3FB62F80F" bold="true" box="[1240,1259,1943,1967]" pageId="6" pageNumber="445">9,</emphasis>
ventral view;
<emphasis id="C3FBE803FFCBFFA3CE0FD7B3FA15F80F" bold="true" box="[1405,1436,1943,1967]" pageId="6" pageNumber="445">10,</emphasis>
lateral view;
<emphasis id="C3FBE803FFCBFFA3CA69D79FFEB0F873" bold="true" box="[283,313,1979,2003]" pageId="6" pageNumber="445">11,</emphasis>
head dorsal view;
<emphasis id="C3FBE803FFCBFFA3CA8BD79FFD91F873" bold="true" box="[505,536,1979,2003]" pageId="6" pageNumber="445">12,</emphasis>
head ventral view;
<emphasis id="C3FBE803FFCBFFA3C992D79FFC89F873" bold="true" box="[736,768,1979,2003]" pageId="6" pageNumber="445">13,</emphasis>
labels and left antennomeres 2–10.
</paragraph>
</caption>
<subSubSection id="B995679AFFCAFFA2CBB5D0B2FBAFFEBD" pageId="7" pageNumber="446" type="distribution">
<paragraph id="F1303411FFCAFFA2CBB5D0B2FBAFFEBD" blockId="7.[151,1436,150,285]" pageId="7" pageNumber="446">
<emphasis id="C3FBE803FFCAFFA2CBB5D0B2FE79FF10" bold="true" box="[199,496,150,177]" pageId="7" pageNumber="446">Distribution and habitat.</emphasis>
Known only from the southern coast of
<collectingRegion id="334BFAF3FFCAFFA2C8C2D0B3FBFEFF11" box="[944,1143,151,177]" country="Australia" name="Western Australia" pageId="7" pageNumber="446">Western Australia</collectingRegion>
(
<figureCitation id="69B42894FFCAFFA2CFF7D0B3FB5FFF11" box="[1157,1238,151,177]" captionStart="FIGURE 14" captionStartId="7.[151,250,1114,1138]" captionTargetBox="[151,1435,309,1090]" captionTargetId="figure-82@7.[151,1435,309,1090]" captionTargetPageId="7" captionText="FIGURE 14. Known distribution of Enracius longipes Dechambre, 1999 and E. richardsi Hutchinson &amp; Allsopp new species in Western Australia." figureDoi="http://doi.org/10.5281/zenodo.5749009" httpUri="https://zenodo.org/record/5749009/files/figure.png" pageId="7" pageNumber="446">Fig. 14</figureCitation>
) (Köppen-Geiger climate of Csb, warm-summer Mediterranean). Both specimens have incomplete tarsomeres on most legs, and the
<typeStatus id="2E348AB3FFCAFFA2CBE5D0FBFF70FF59" box="[151,249,223,249]" pageId="7" pageNumber="446" type="holotype">holotype</typeStatus>
was covered in a dense layer of fine undetermined matter and moth scales along margins. The collection method is not noted on their labels, although the moth scales indicate a light trap.
</paragraph>
</subSubSection>
</treatment>
</document>
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