266 lines
33 KiB
XML
266 lines
33 KiB
XML
<document ID-CLB-Dataset="209209" ID-DOI="http://dx.doi.org/10.3897/asp.80.e90570" ID-GBIF-Dataset="bac17de6-baf9-4fee-9ded-268adbb26a61" ID-Pensoft-Pub="1864-8312-80-649" ID-Pensoft-UUID="FAE6DFEF69735531A4CBA275E078320D" ID-ZooBank="AD0B1CC262414460B6B888E6B95F20D7" ModsDocID="1864-8312-80-649" checkinTime="1671114540108" checkinUser="pensoft" docAuthor="Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S. & Harms, Danilo" docDate="2022" docId="78D6D54E21FC5C548F4EDE39CB81241B" docLanguage="en" docName="ArthropodSystPhyl 80: 649-691" docOrigin="Arthropod Systematics & amp; Phylogeny 80" docPubDate="2022-12-14" docSource="http://dx.doi.org/10.3897/asp.80.e90570" docTitle="Toliaranella Lorenz & Loria & Harvey & Harms 2022, gen. nov." docType="treatment" docVersion="5" id="FAE6DFEF69735531A4CBA275E078320D" lastPageNumber="649" masterDocId="FAE6DFEF69735531A4CBA275E078320D" masterDocTitle="The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots" masterLastPageNumber="691" masterPageNumber="649" pageNumber="649" updateTime="1732733992417" updateUser="ExternalLinkService">
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<mods:titleInfo id="7CF05EC048C2911FAF6A10422ADBF116">
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<mods:title id="EE069F334CEDA7BB0F591E51485D16FA">The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots</mods:title>
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<mods:name id="C90F0DABABFD8DC8FAA03CF36C6267BA" type="personal">
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<mods:namePart id="E7D0CB062C3E09577B070EBE2C773394">Lorenz, Michelle</mods:namePart>
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<mods:affiliation id="E30B8C75BC8D6FF69916E89EF6AD04B2">Museum of Nature Hamburg - Zoology, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany & University of Hamburg, Biology Department, Ohnhorststrasse 18, 22609 Hamburg, Germany</mods:affiliation>
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<mods:nameIdentifier id="721654620CE62E52B91C4F96ABC266A2" type="email">michellelorenz@gmx.net</mods:nameIdentifier>
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<mods:roleTerm id="AEF15748307212A73F2CBFA4C0AA2811">Author</mods:roleTerm>
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<mods:namePart id="207D314C8D5E1219670C13B27D0E6975">Loria, Stephanie F.</mods:namePart>
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<mods:affiliation id="FA765CDA97F69118E8EE5C89F6E0379E">Museum of Nature Hamburg - Zoology, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
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</mods:name>
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<mods:name id="705794EDBF032446395493988C5688EA" type="personal">
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<mods:roleTerm id="281140C2F7A75320C54756B757FE3E26">Author</mods:roleTerm>
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<mods:namePart id="6E38004927595D4C199745E131C9C727">Harvey, Mark S.</mods:namePart>
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<mods:nameIdentifier id="040BB00FE3C23033877CB622ADB08D81" type="ORCID">https://orcid.org/0000-0003-1482-0109</mods:nameIdentifier>
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<mods:affiliation id="8C7DF33573984051B6B82E36BA65EF54">Western Australian Museum, Collections & Research, 49 Kew Street, Welshpool, WA 6106, Australia & University of Western Australia, School of Biological Sciences, Crawley WA 6009, Australia</mods:affiliation>
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<mods:namePart id="3A9396D94227D03EBEA4BCEBA203A3F9">Harms, Danilo</mods:namePart>
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<mods:nameIdentifier id="F20709EA3B29269B1ACA8F0383A2ADF2" type="ORCID">https://orcid.org/0000-0002-7189-5345</mods:nameIdentifier>
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<mods:affiliation id="DFBE7118891D4326C557466CB9E9DADE">Museum of Nature Hamburg - Zoology, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
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<mods:typeOfResource id="514440700804DF269F7C1BF32C58433E">text</mods:typeOfResource>
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<mods:title id="8F14641BE20CEF8AD7E6E3112CEA367E">Arthropod Systematics & amp; Phylogeny</mods:title>
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<mods:part id="C2A15E8192766D3B5F1F16A05EDE76A6">
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<mods:date id="09109E010400D8D1DA9F7032ACE8C2F7">2022</mods:date>
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<mods:number id="FF3AEEC561F557E8B38317DEF87CBEC4">2022-12-14</mods:number>
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<mods:number id="154EEF4F899DF6549D5C415F0800201A">80</mods:number>
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<mods:end id="167A493201B118F89DCF43E9ABE261D3">691</mods:end>
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<mods:url id="F647106F90897CA03BF39EAFEC4D8506">http://dx.doi.org/10.3897/asp.80.e90570</mods:url>
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<mods:classification id="E93882B9AEEF92F5D3FDCD5592F0CEA5">journal article</mods:classification>
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<mods:identifier id="14B3E11B4DB3F1FB4A2B8525941FB355" type="DOI">http://dx.doi.org/10.3897/asp.80.e90570</mods:identifier>
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<mods:identifier id="09A095427D33B5CBB9B021C271FDFBA3" type="Pensoft-Pub">1864-8312-80-649</mods:identifier>
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<treatment id="78D6D54E21FC5C548F4EDE39CB81241B" ID-GBIF-Taxon="206273797" LSID="urn:lsid:zoobank.org:act:NomEnClAturAlACts/F9EDCC8D-D7A5-41F4-B101-DA7320782EBA" httpUri="http://treatment.plazi.org/id/78D6D54E21FC5C548F4EDE39CB81241B" lastPageNumber="649" pageId="0" pageNumber="649" scope_family="Feaellidae" scope_order="Pseudoscorpiones" scope_subFamily="Feaellinae" scope_superFamily="Feaelloidea">
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<subSubSection id="B742B13DF366538271B0276E6DE97A6A" pageId="0" pageNumber="649" type="nomenclature">
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<paragraph id="2C3DC858D50069A912FF88D60D766618" pageId="0" pageNumber="649">
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<taxonomicName id="E65D0C38F4FFC6EB44046D98BFD5CECE" LSID="https://zoobank.org/NomenclaturalActs/F9EDCC8D-D7A5-41F4-B101-DA7320782EBA" authority="Lorenz & Loria & Harvey & Harms, 2022" authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" class="Arachnida" family="Feaellidae" genus="Toliaranella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toliaranella" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="genus" status="gen. nov.">Toliaranella</taxonomicName>
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<taxonomicNameLabel id="E2E30BC4B9C70B8FD0BD3E34B1E4FEC0" pageId="0" pageNumber="649">gen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="7C30B2DF1D5FB5BCCDCD1A1555F5F83D" pageId="0" pageNumber="649" type="type species">
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<paragraph id="270A1A80BB030D0FBD3F671B44643D5C" pageId="0" pageNumber="649">Type species.</paragraph>
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<paragraph id="23F7B17005E4F30518706FAF926F215B" pageId="0" pageNumber="649">
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<taxonomicName id="2DE507DAF4922DA91E89EC6E608BEBE5" authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" class="Arachnida" family="Feaellidae" genus="Toliaranella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toliaranella pumila" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="species" species="pumila">
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<emphasis id="6482382B0361ADF5452E71DDFB6F1332" italics="true" pageId="0" pageNumber="649">Toliaranella pumila</emphasis>
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</taxonomicName>
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<emphasis id="809504558A4D8D3AFF64BB8C4E8BF8AB" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="B8E2D0437AC196D17C8B0C52DF17CA0F" pageId="0" pageNumber="649" type="diagnosis">
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<paragraph id="4C07082FB4B173C7E086EAB0EE110949" pageId="0" pageNumber="649">Diagnosis.</paragraph>
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<paragraph id="966C8FC6ABFDED8D01574C079352A3EF" pageId="0" pageNumber="649">
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<taxonomicName id="499B07A8E4BA8A77D2C3B655680CD244" authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" class="Arachnida" family="Feaellidae" genus="Toliaranella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toliaranella" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="genus">
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<emphasis id="7E2C8AE796D07E99B21865ED30263B75" italics="true" pageId="0" pageNumber="649">Toliaranella</emphasis>
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</taxonomicName>
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<emphasis id="192955F230779A1768DDCBABB02EF6E6" bold="true" pageId="0" pageNumber="649">gen. nov.</emphasis>
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differs from all other Malagasy feaellids by the presence of a perforated organ above the coxal spines and smallest body size of any Malagasy groups. Like
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<taxonomicName id="39F08420A9F90BFDC5D58C473B1672FD" authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" class="Arachnida" family="Feaellidae" genus="Antsirananaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Antsirananaella" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="genus">
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<emphasis id="B22A16F9DE23320BD7383A08678A4954" italics="true" pageId="0" pageNumber="649">Antsirananaella</emphasis>
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</taxonomicName>
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<emphasis id="617C7CB0F7E9D16B15560F13134FA6A8" bold="true" pageId="0" pageNumber="649">gen. nov.</emphasis>
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and
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<taxonomicName id="4EA2BF387FB5682D0F30BB3DA2C69AFE" authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" class="Arachnida" family="Feaellidae" genus="Mahajanganella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mahajanganella" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="genus">
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<emphasis id="AF7FC8C58E42B34369C17D13BCBFC0AB" italics="true" pageId="0" pageNumber="649">Mahajanganella</emphasis>
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</taxonomicName>
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<emphasis id="F5387BF6439B028E9A545D9381697778" bold="true" pageId="0" pageNumber="649">gen. nov.</emphasis>
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, it differs from
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<taxonomicName id="39D3FFECA0EBDD776D2C92AA733E63BF" authorityName="Judson" authorityYear="2017" class="Arachnida" family="Feaellidae" genus="Cybella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cybella" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="genus">
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<emphasis id="1C637B5F06AE93B494983F4233FFF89E" italics="true" pageId="0" pageNumber="649">Cybella</emphasis>
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</taxonomicName>
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by having platelets on the pleural membrane (absent in
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<taxonomicName id="AC8E6B63BE3E3D8FD6657B9846154232" authorityName="Judson" authorityYear="2017" class="Arachnida" family="Feaellidae" genus="Cybella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cybella" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="genus">
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<emphasis id="6AAA9A7E637E95ADA83C3EC15AF3B6DC" italics="true" pageId="0" pageNumber="649">Cybella</emphasis>
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</taxonomicName>
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), from
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<taxonomicName id="8293993A21B5B1DD8BE12B0BF6555B7B" authorityName="Harvey, Andrade & Pinto-da-Rocha" authorityYear="2016" class="Arachnida" family="Feaellidae" genus="Iporangella" kingdom="Animalia" lsidName="Iporangella" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="genus">
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<emphasis id="E376DAE821FC23B6592B12C41E989653" italics="true" pageId="0" pageNumber="649">Iporangella</emphasis>
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</taxonomicName>
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by the presence of specialized setae on the movable chelal finger (absent in
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<taxonomicName id="C9E6ED18684CB09924F1641D9613675E" authorityName="Harvey, Andrade & Pinto-da-Rocha" authorityYear="2016" class="Arachnida" family="Feaellidae" genus="Iporangella" kingdom="Animalia" lsidName="Iporangella" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="genus">
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<emphasis id="F0E353D319BB35E92C86C5E911D18022" italics="true" pageId="0" pageNumber="649">Iporangella</emphasis>
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</taxonomicName>
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), from
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<taxonomicName id="29810CEDD884E31D466A5AD2A26EC596" authority="" class="Arachnida" family="Feaellidae" genus="Feaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feaella (Difeaella)" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="subGenus" subGenus="Difeaella">Feaella (Difeaella)</taxonomicName>
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and
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<taxonomicName id="2FFA89C9E4EA3DE8666F3544C1DC6B10" authority="(Feaella)" class="Arachnida" family="Feaellidae" genus="Feaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feaella (Feaella)" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="subGenus" subGenus="Feaella">Feaella (Feaella)</taxonomicName>
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by the presence of four anterior carapaceal lobes (two and six, respectively), from
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<taxonomicName id="E0EFBB6B972D8004848854729A944F14" authority="" class="Arachnida" family="Feaellidae" genus="Feaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feaella (Tetrafeaella)" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="subGenus" subGenus="Tetrafeaella">Feaella (Tetrafeaella)</taxonomicName>
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in continental Africa by having a less pronounced depression on the base of coxa I and on top of coxa II (distinctly more pronounced in
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<taxonomicName id="7753AE32F5D9EFE43C7FF54D689FC0C5" authorityName="Tullgren" authorityYear="1907" class="Arachnida" family="Feaellidae" genus="Feaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feaella (T.) mucronata" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="species" species="mucronata" subGenus="T.">Feaella (T.) cf. mucronata</taxonomicName>
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) and from the Australian species presently attributed to
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<taxonomicName id="D2FC45ACF7FDFD7529447C30CE706DEF" authority="" class="Arachnida" family="Feaellidae" genus="Feaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feaella (Tetrafeaella)" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="subGenus" subGenus="Tetrafeaella">Feaella (Tetrafeaella)</taxonomicName>
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by having fewer coxal spines (one pair versus three or four in the Australian taxa).
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</paragraph>
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</subSubSection>
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<subSubSection id="CBC6FCAECDB9AC24D5E8A2C851728F60" pageId="0" pageNumber="649" type="etymology">
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<paragraph id="25EBB2A58EFD280232A98031E7C5A01E" pageId="0" pageNumber="649">Etymology.</paragraph>
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<paragraph id="0DE8A37442A9065D3820BD4B0A503A46" pageId="0" pageNumber="649">This genus is named after the former Toliara Province, where all specimens were collected. The gender is feminine.</paragraph>
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</subSubSection>
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<subSubSection id="A6FD097AEA945A3D6A0C4467F6C40C69" pageId="0" pageNumber="649" type="description">
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<paragraph id="64B9D9AB5EBFB55063125A20B70FF046" pageId="0" pageNumber="649">Description.</paragraph>
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<paragraph id="EBD48042880245E9C07C38F77C26AD56" pageId="0" pageNumber="649">
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The following description is based on holotype and allotype. -
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<emphasis id="EFB9294337034456C1716F78E46B633C" italics="true" pageId="0" pageNumber="649">Carapace</emphasis>
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(Figs
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<figureCitation id="F2C91C39AEF6B14CA55F4D5EF3D5B80A" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Toliaranella gen. nov., scanning electron micrographs. Specimens used for imaging: A T. griswoldi sp. nov. male paratype (ZMH-A 0016740); B, C T. meridionalis sp. nov. male paratype (ZMH-A 0016753). A carapace, dorsal view; B sternum ventral overview; C coxal spines and perforated organ in detail (highlighted in red). Scale bars: 0.2 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/asp.80.e90570.figure18" httpUri="https://binary.pensoft.net/fig/782010" pageId="0" pageNumber="649">18A</figureCitation>
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,
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<figureCitation id="D4706EE5A2F41D67EE53ECA55F5AC06C" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Drawings of Toliaranella pumila sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure20" httpUri="https://binary.pensoft.net/fig/782012" pageId="0" pageNumber="649">20A</figureCitation>
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,
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<figureCitation id="85AF56CCB3F37001AD6C6B88D02C7A31" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Drawings of Toliaranella fisheri sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure21" httpUri="https://binary.pensoft.net/fig/782013" pageId="0" pageNumber="649">21A</figureCitation>
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,
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<figureCitation id="11B0ADDD74A969E6E8C1BCC6EEC09BB9" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Drawings of Toliaranella griswoldi sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure22" httpUri="https://binary.pensoft.net/fig/782014" pageId="0" pageNumber="649">22A</figureCitation>
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,
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<figureCitation id="6579D29D97E5EEA9FB450BC058CA6D48" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Drawings of Toliaranella mahnerti sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure23" httpUri="https://binary.pensoft.net/fig/782015" pageId="0" pageNumber="649">23A</figureCitation>
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,
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<figureCitation id="E6EAE8843522F58C889C123B20935B42" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Drawings of Toliaranella meridionalis sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure24" httpUri="https://binary.pensoft.net/fig/782016" pageId="0" pageNumber="649">24A</figureCitation>
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|
): Strongly granulate, all four anterior lobes with same distance to each other and rounded at tips. -
|
|
<emphasis id="F87FA18FFE6C61533FE3E5E31752F2CD" italics="true" pageId="0" pageNumber="649">Pedipalps</emphasis>
|
|
(Figs
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|
<figureCitation id="FC9B2628D9CDE47631D3B3D9EF790465" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Toliaranella gen. nov.. Specimen used for imaging: T. griswoldi sp. nov. holotype (CAS 9071500). A dorsal overview; B ventral overview; C lateral overview. Scale bar: 0.2 mm." figureDoi="10.3897/asp.80.e90570.figure17" httpUri="https://binary.pensoft.net/fig/782009" pageId="0" pageNumber="649">17A, B</figureCitation>
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,
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|
<figureCitation id="484C64D88FE08577EFD698DB371BBA52" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Toliaranella, gen. nov., scanning electron micrographs. Specimen used for imaging: T. pumila sp. nov. male paratype (ZMH-A 0016722). A right chela from ventral with specialized setae in a group in detail (highlighted in red); B left chela with double trichobothrium dt in detail (highlighted in red); C left chela from medial with terminal teeth in a compact group in detail. Scale bar: 0.2 mm." figureDoi="10.3897/asp.80.e90570.figure19" httpUri="https://binary.pensoft.net/fig/782011" pageId="0" pageNumber="649">19A-C</figureCitation>
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,
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<figureCitation id="BAC50F8328434D5E02EA44E745D68B20" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Drawings of Toliaranella pumila sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure20" httpUri="https://binary.pensoft.net/fig/782012" pageId="0" pageNumber="649">20C, D</figureCitation>
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,
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|
<figureCitation id="5C09D00F1C02289F8A5D61587CE41272" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Drawings of Toliaranella fisheri sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure21" httpUri="https://binary.pensoft.net/fig/782013" pageId="0" pageNumber="649">21C, D</figureCitation>
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,
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|
<figureCitation id="7A83CAAFBD3408EDFBDDC0DCA35004DF" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Drawings of Toliaranella griswoldi sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure22" httpUri="https://binary.pensoft.net/fig/782014" pageId="0" pageNumber="649">22C, D</figureCitation>
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,
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|
<figureCitation id="3C7B584A1267E4B10F4B38D4BB64A5B0" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Drawings of Toliaranella mahnerti sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure23" httpUri="https://binary.pensoft.net/fig/782015" pageId="0" pageNumber="649">23C, D</figureCitation>
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,
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|
<figureCitation id="7FB1FFA0EB8D473CDA52A3E269C65FA6" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Drawings of Toliaranella meridionalis sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure24" httpUri="https://binary.pensoft.net/fig/782016" pageId="0" pageNumber="649">24C, D</figureCitation>
|
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): Terminal teeth with 7-8 teeth on both the fixed and the movable fingers (including 1 large tooth on fixed finger). -
|
|
<emphasis id="2E8E18342592C2DD00C5F5D4E9E86431" italics="true" pageId="0" pageNumber="649">Coxal region</emphasis>
|
|
(Figs
|
|
<figureCitation id="E7A32E1EE8C8BC72A820791A96BB7EA3" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Toliaranella gen. nov.. Specimen used for imaging: T. griswoldi sp. nov. holotype (CAS 9071500). A dorsal overview; B ventral overview; C lateral overview. Scale bar: 0.2 mm." figureDoi="10.3897/asp.80.e90570.figure17" httpUri="https://binary.pensoft.net/fig/782009" pageId="0" pageNumber="649">17B</figureCitation>
|
|
,
|
|
<figureCitation id="F17C32332DC6B62B8DD5467641CE9B63" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Toliaranella gen. nov., scanning electron micrographs. Specimens used for imaging: A T. griswoldi sp. nov. male paratype (ZMH-A 0016740); B, C T. meridionalis sp. nov. male paratype (ZMH-A 0016753). A carapace, dorsal view; B sternum ventral overview; C coxal spines and perforated organ in detail (highlighted in red). Scale bars: 0.2 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/asp.80.e90570.figure18" httpUri="https://binary.pensoft.net/fig/782010" pageId="0" pageNumber="649">18B, C</figureCitation>
|
|
): Pedipalpal coxae each with one spike laterally near base; coxae I with basal depression, each side with one small coxal spine; slightly more cranial of coxal spine one novel organ on each side: rose-shaped, not granulate but perforated organ (Fig.
|
|
<figureCitation id="F9F3BE4CFB7D75863928C7AAEE38DD22" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Toliaranella gen. nov., scanning electron micrographs. Specimens used for imaging: A T. griswoldi sp. nov. male paratype (ZMH-A 0016740); B, C T. meridionalis sp. nov. male paratype (ZMH-A 0016753). A carapace, dorsal view; B sternum ventral overview; C coxal spines and perforated organ in detail (highlighted in red). Scale bars: 0.2 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/asp.80.e90570.figure18" httpUri="https://binary.pensoft.net/fig/782010" pageId="0" pageNumber="649">18B, C</figureCitation>
|
|
) filling the base of the depression; coxae II with irregular shaped spines framing depression of coxae I.
|
|
</paragraph>
|
|
<caption id="886374ACA11BA62D15859789DC1918E7" doi="10.3897/asp.80.e90570.figure17" httpUri="https://binary.pensoft.net/fig/782009" pageId="0" pageNumber="649" start="Figure 17" startId="F17">
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|
<paragraph id="0D853984A08A1EBBC2CC9E3808077C85" pageId="0" pageNumber="649">
|
|
<emphasis id="80386B6831E0C693DC7C700BE77D930C" bold="true" pageId="0" pageNumber="649">Figure 17.</emphasis>
|
|
<taxonomicName id="5355C14D743209261B464DA80179EBF8" authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" class="Arachnida" family="Feaellidae" genus="Toliaranella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toliaranella" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="4AD532A3DFDF00DF8D10071490DB04EB" italics="true" pageId="0" pageNumber="649">Toliaranella</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="EB4325008BF51671B2F7C52718F7EF06" bold="true" pageId="0" pageNumber="649">gen. nov.</emphasis>
|
|
. Specimen used for imaging:
|
|
<taxonomicName id="1A114FDD4BF782C6B48215DE7DED2AC5" lsidName="T. griswoldi" pageId="0" pageNumber="649" rank="species" species="griswoldi">
|
|
<emphasis id="7E29FE58F6DA176D321BAD609CE6B6CC" italics="true" pageId="0" pageNumber="649">T. griswoldi</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="5CBAB7A88CF7251FB2E2A14939BBEB5C" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
|
holotype (CAS 9071500).
|
|
<emphasis id="232F5EB1E6B2FE8401A4DE7E5C73FD75" bold="true" pageId="0" pageNumber="649">A</emphasis>
|
|
dorsal overview;
|
|
<emphasis id="8095B4F0FEA79F4565E9EEE1456DBE61" bold="true" pageId="0" pageNumber="649">B</emphasis>
|
|
ventral overview;
|
|
<emphasis id="F55C6A3FEBABFF16A5C320B89FCBBCEC" bold="true" pageId="0" pageNumber="649">C</emphasis>
|
|
lateral overview. Scale bar: 0.2 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="244307A8D2F8090A5D012D62AA68D288" doi="10.3897/asp.80.e90570.figure18" httpUri="https://binary.pensoft.net/fig/782010" pageId="0" pageNumber="649" start="Figure 18" startId="F18">
|
|
<paragraph id="03D00450C23054ED1DA0FD165699C769" pageId="0" pageNumber="649">
|
|
<emphasis id="98921DDF0CF0D71EDBE9255C3FEDAD5D" bold="true" pageId="0" pageNumber="649">Figure 18.</emphasis>
|
|
<taxonomicName id="C1A1157BBBF4DB17EE4917B0B3400855" authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" class="Arachnida" family="Feaellidae" genus="Toliaranella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toliaranella" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B20269B08EC673505193173932D8FF7E" italics="true" pageId="0" pageNumber="649">Toliaranella</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="F40640F1C4BB2A9DEE5C8C0E166BCCD6" bold="true" pageId="0" pageNumber="649">gen. nov.</emphasis>
|
|
, scanning electron micrographs. Specimens used for imaging:
|
|
<emphasis id="899B817DB5621EEDEAE9EC01D4EA14F2" bold="true" pageId="0" pageNumber="649">A</emphasis>
|
|
<taxonomicName id="3893638F4DC549E5D4C5B57B26059061" lsidName="T. griswoldi" pageId="0" pageNumber="649" rank="species" species="griswoldi">
|
|
<emphasis id="B6D004B5A435886F0DA67EAF859E4978" italics="true" pageId="0" pageNumber="649">T. griswoldi</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="BCE891DDF5722690657261713C78A286" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
|
male paratype (ZMH-A0016740);
|
|
<emphasis id="B41A7AE89CA8BBE820ED0A6D5A1A9098" bold="true" pageId="0" pageNumber="649">B</emphasis>
|
|
,
|
|
<emphasis id="AF2CAECCB7E13F52452B87EA013E1ECA" bold="true" pageId="0" pageNumber="649">C</emphasis>
|
|
<taxonomicName id="11CFC21993BE1D2A896EED27DF141EEA" lsidName="T. meridionalis" pageId="0" pageNumber="649" rank="species" species="meridionalis">
|
|
<emphasis id="13274315A2098B11B0816B2C4FA0E35D" italics="true" pageId="0" pageNumber="649">T. meridionalis</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="6F2DDCA06E199C3FC1C716D5A79DAE41" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
|
male paratype (ZMH-A0016753).
|
|
<emphasis id="484ADFC532CEE03FF58019EC89E38E30" bold="true" pageId="0" pageNumber="649">A</emphasis>
|
|
carapace, dorsal view;
|
|
<emphasis id="E23B9D59986E7569F612B145C5270B64" bold="true" pageId="0" pageNumber="649">B</emphasis>
|
|
sternum ventral overview;
|
|
<emphasis id="AABDFF7D22D8FDA17AB70B3C2E1498CE" bold="true" pageId="0" pageNumber="649">C</emphasis>
|
|
coxal spines and perforated organ in detail (highlighted in red). Scale bars: 0.2 mm (
|
|
<emphasis id="65BE266C24470F22794DFDF3FD240520" bold="true" pageId="0" pageNumber="649">A</emphasis>
|
|
,
|
|
<emphasis id="FD6E5B6688FFA76560C968D2882D2D60" bold="true" pageId="0" pageNumber="649">B</emphasis>
|
|
); 0.1 mm (
|
|
<emphasis id="2E721E6A88CCD2D2BE6B11EB7C015E37" bold="true" pageId="0" pageNumber="649">C</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="43D470F9B99A83B9C310D2913C0044A7" pageId="0" pageNumber="649" type="included species">
|
|
<paragraph id="13B9D4475DB2A15EAF3E3EBB20D60108" pageId="0" pageNumber="649">Included species.</paragraph>
|
|
<paragraph id="D3200D9DCF64EE09A5C412FA8AB7A966" pageId="0" pageNumber="649">
|
|
<taxonomicName id="F6DB550630AD3B02B85E03D513224254" authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" class="Arachnida" family="Feaellidae" genus="Toliaranella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toliaranella fisheri" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="species" species="fisheri">
|
|
<emphasis id="2409A7D09B318175C6099D1D1EDE4AB2" italics="true" pageId="0" pageNumber="649">Toliaranella fisheri</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="3BAFD1806E5AA250BD63EE117E6271B9" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
|
;
|
|
<taxonomicName id="7B60122860CF4F2104E56B6DA744839B" authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" class="Arachnida" family="Feaellidae" genus="Toliaranella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toliaranella griswoldi" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="species" species="griswoldi">
|
|
<emphasis id="B882F345D4B457D7223E3EAC496DA8BA" italics="true" pageId="0" pageNumber="649">Toliaranella griswoldi</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="0A5530ACEDFE8A6F43C406CF49DD91F1" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
|
;
|
|
<taxonomicName id="CA88007D13D70C357FD5714E2DF887E0" authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" class="Arachnida" family="Feaellidae" genus="Toliaranella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toliaranella mahnerti" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="species" species="mahnerti">
|
|
<emphasis id="4B68B0AD8C38BFAB8A57A48E828BA341" italics="true" pageId="0" pageNumber="649">Toliaranella mahnerti</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="1D0A92A19ADB9C935A5ED6676EE3E92B" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
|
;
|
|
<taxonomicName id="F608FC23E07AA0FDC9F30E114479F681" authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" class="Arachnida" family="Feaellidae" genus="Toliaranella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toliaranella meridionalis" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="species" species="meridionalis">
|
|
<emphasis id="2C206188E52AF0EACAAF4C08E86BC165" italics="true" pageId="0" pageNumber="649">Toliaranella meridionalis</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="3C7AD7FCFBF336DDBF40BDC074AF6FEB" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
|
; and
|
|
<taxonomicName id="1FE5A709F2FE72C265B806B1FBCCB244" authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" class="Arachnida" family="Feaellidae" genus="Toliaranella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toliaranella pumila" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="species" species="pumila">
|
|
<emphasis id="70CDF94730F38FB8333E99150F110A0F" italics="true" pageId="0" pageNumber="649">Toliaranella pumila</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="4D71097E7A898DA52CEB9AEDF197CCFA" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="FF7C913BD8D15045BB17D22C8206B2ED" pageId="0" pageNumber="649" type="distribution">
|
|
<paragraph id="77AB75E4643C7987FCD18A635CB665A8" pageId="0" pageNumber="649">Distribution.</paragraph>
|
|
<paragraph id="5E797F4DF150E7A9B0323B29A995417B" pageId="0" pageNumber="649">Members of this genus are localized to the southern part of Madagascar in the Androy, Anosy and Atsimo-Andrefana Regions (formerly Toliara Province) where the main landscape is covered by arid spiny bush vegetation.</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |