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<document id="B806709187EAC4D834DABE4FCF3F7A1A" ID-DOI="10.11646/zootaxa.5352.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="8407084" ID-ZooBank="1FCBB820-6683-44D0-A98D-8ADA327BEA43" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1696428007364" checkinUser="plazi" docAuthor="Volynkin, Anton V. &amp; Černý, Karel" docDate="2023" docId="78771043CB74FFAA02E1FAACFA9A6BB3" docLanguage="en" docName="zootaxa.5352.2.1.pdf" docOrigin="Zootaxa 5352 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5352.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Philippinithosia Volynkin &amp; Černý 2023, gen. n." docType="treatment" docVersion="2" lastPageNumber="159" masterDocId="844E683BCB73FFA20276FFBAFFA1685F" masterDocTitle="On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini)" masterLastPageNumber="176" masterPageNumber="151" pageNumber="158" updateTime="1696428496350" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="BBAF01DBAE84C0E38FC936527D371467">On the taxonomy of “ Tigrioides ” leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini)</mods:title>
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<mods:namePart id="9FE59CB18ED11013D179A0C1A6CA3C4A">Volynkin, Anton V.</mods:namePart>
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<mods:namePart id="FD65884FC6A461118AE1E6EDFD3AC68C">Černý, Karel</mods:namePart>
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<taxonomicName id="37DEDAD6CB74FFA502E1FAACFEF86D6E" authority="Volynkin &amp; Černý, 2023" authorityName="Volynkin &amp; Černý" authorityYear="2023" box="[151,345,1302,1329]" class="Insecta" family="Erebidae" genus="Philippinithosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis id="C2AA7D47CB74FFA502E1FAACFEF86D6E" bold="true" box="[151,345,1302,1329]" italics="true" pageId="7" pageNumber="158">Philippinithosia</emphasis>
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species:
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<taxonomicName id="37DEDAD6CB74FFA5035AFAC5FD896DC8" authority="Volynkin &amp; Černý, 2023" authorityName="Volynkin &amp; Černý" authorityYear="2023" box="[300,552,1407,1431]" class="Insecta" family="Erebidae" genus="Philippinithosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="lourensi" status="sp. n.">
<emphasis id="C2AA7D47CB74FFA5035AFAC5FD896DC8" box="[300,552,1407,1431]" italics="true" pageId="7" pageNumber="158">Philippinithosia lourensi</emphasis>
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, by present designation.
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<emphasis id="C2AA7D47CB74FFA502E1FA7FFEAB6D80" bold="true" box="[151,266,1477,1503]" pageId="7" pageNumber="158">Diagnosis</emphasis>
. Species of the new genus (
<figureCitation id="68E5BDD0CB74FFA5003EFA7FFD6B6DBF" box="[584,714,1477,1504]" captionStart-0="FIGURES 2431" captionStart-1="FIGURES 3239" captionStartId-0="14.[151,264,1943,1967]" captionStartId-1="15.[151,264,1943,1967]" captionTargetBox-0="[198,1390,184,1908]" captionTargetBox-1="[222,1366,184,1908]" captionTargetId-0="figure-25@14.[185,1402,181,1919]" captionTargetId-1="figure-21@15.[209,1378,181,1919]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURES 2431. Leucanithosia spp. (2427) and Philippinithosia lourensi (2831): adults. Depositories of the specimens: 24, 26, and 2831 in MWM/ZSM (24 and 26 ex CKC); 25 and 27 in CKC." captionText-1="FIGURES 3239. Philippinithosia spp. (3235), Tigrioides alterna (36, 37), and Dubatoloviana trimacula (38, 39): adults. Depositories of the specimens: 3235, 38 and 39 in MWM/ZSM; 36 in OUMNH; 37 in NHMUK (©The Trustees of NHMUK)." figureDoi-0="http://doi.org/10.5281/zenodo.8407097" figureDoi-1="http://doi.org/10.5281/zenodo.8407102" httpUri-0="https://zenodo.org/record/8407097/files/figure.png" httpUri-1="https://zenodo.org/record/8407102/files/figure.png" pageId="7" pageNumber="158">Figs 2835</figureCitation>
) are externally very similar to
<taxonomicName id="37DEDAD6CB74FFA5065FFA7FFB696D80" authorityName="Volynkin &amp; Černý" authorityYear="2023" box="[1065,1224,1477,1503]" class="Insecta" family="Erebidae" genus="Leucanithosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="C2AA7D47CB74FFA5065FFA7FFB696D80" box="[1065,1224,1477,1503]" italics="true" pageId="7" pageNumber="158">Leucanithosia</emphasis>
</taxonomicName>
(
<figureCitation id="68E5BDD0CB74FFA506AEFA7FFAEB6D80" box="[1240,1354,1477,1503]" captionStart-0="FIGURES 17" captionStart-1="FIGURES 815" captionStart-2="FIGURES 1623" captionStart-3="FIGURES 2431" captionStartId-0="11.[151,264,1943,1967]" captionStartId-1="12.[151,264,1943,1967]" captionStartId-2="13.[151,264,1943,1967]" captionStartId-3="14.[151,264,1943,1967]" captionTargetBox-0="[198,1389,190,1909]" captionTargetBox-1="[177,1421,182,1908]" captionTargetBox-2="[226,1388,182,1875]" captionTargetBox-3="[198,1390,184,1908]" captionTargetId-0="figure-21@11.[186,1402,181,1919]" captionTargetId-1="figure-25@12.[164,1424,181,1919]" captionTargetId-2="figure-21@13.[164,1424,181,1886]" captionTargetId-3="figure-25@14.[185,1402,181,1919]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionText-0="FIGURES 17. Leucanithosia spp.: adults. Depositories of the specimens: 1 in OUMNH; 24, 6 and 8 in MWM/ZSM; 5 in CKC." captionText-1="FIGURES 815. Leucanithosia spp.: adults. Depositories of the specimens: 8, 11, 12 and 14 in MWM/ZSM (11 and 14 ex CKC); 9 and 10 in CBP; 13 in GMF-B; 15 in CKC." captionText-2="FIGURES 1623. Leucanithosia spp.: adults. The specimens are deposited in MWM/ZSM." captionText-3="FIGURES 2431. Leucanithosia spp. (2427) and Philippinithosia lourensi (2831): adults. Depositories of the specimens: 24, 26, and 2831 in MWM/ZSM (24 and 26 ex CKC); 25 and 27 in CKC." figureDoi-0="http://doi.org/10.5281/zenodo.8407088" figureDoi-1="http://doi.org/10.5281/zenodo.8407090" figureDoi-2="http://doi.org/10.5281/zenodo.8407094" figureDoi-3="http://doi.org/10.5281/zenodo.8407097" httpUri-0="https://zenodo.org/record/8407088/files/figure.png" httpUri-1="https://zenodo.org/record/8407090/files/figure.png" httpUri-2="https://zenodo.org/record/8407094/files/figure.png" httpUri-3="https://zenodo.org/record/8407097/files/figure.png" pageId="7" pageNumber="158">Figs 127</figureCitation>
) but in males, the androconial tuft on the foreleg consists of ochreous yellow and shorter scales, which are elongate and grey in
<taxonomicName id="37DEDAD6CB74FFA50287F9B7FE316E78" authorityName="Volynkin &amp; Černý" authorityYear="2023" box="[241,400,1549,1575]" class="Insecta" family="Erebidae" genus="Leucanithosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="C2AA7D47CB74FFA50287F9B7FE316E78" box="[241,400,1549,1575]" italics="true" pageId="7" pageNumber="158">Leucanithosia</emphasis>
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. The male genital capsule of
<taxonomicName id="37DEDAD6CB74FFA50091F9B7FC3A6E78" authorityName="Volynkin &amp; Černý" authorityYear="2023" box="[743,923,1549,1575]" class="Insecta" family="Erebidae" genus="Philippinithosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="C2AA7D47CB74FFA50091F9B7FC3A6E78" box="[743,923,1549,1575]" italics="true" pageId="7" pageNumber="158">Philippinithosia</emphasis>
</taxonomicName>
(
<figureCitation id="68E5BDD0CB74FFA501DBF9B7FB916E77" box="[941,1072,1549,1576]" captionStart-0="FIGURES 5860" captionStart-1="FIGURES 6164" captionStartId-0="21.[151,264,1898,1922]" captionStartId-1="22.[151,264,1936,1960]" captionTargetBox-0="[190,1398,194,1861]" captionTargetBox-1="[174,1414,194,1899]" captionTargetId-0="figure-21@21.[175,1411,181,1874]" captionTargetId-1="figure-25@22.[161,1426,181,1912]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="FIGURES 5860. Philippinithosia spp.: male genitalia. The specimens dissected are deposited in MWM/ZSM." captionText-1="FIGURES 6164. Philippinithosia, Tigrioides and Dubatoloviana spp.: male genitalia. Depositories of the specimens dissected: 61, 62 and 64 in MWM/ZSM; 63 in OUMNH." figureDoi-0="http://doi.org/10.5281/zenodo.8407119" figureDoi-1="http://doi.org/10.5281/zenodo.8407123" httpUri-0="https://zenodo.org/record/8407119/files/figure.png" httpUri-1="https://zenodo.org/record/8407123/files/figure.png" pageId="7" pageNumber="158">Figs 5862</figureCitation>
) is characterised by the distally dilated and swollen dorsal part of the valva and the distal saccular process, which both are densely covered with minute setae, the feature characteristic of the genus. Additionally, compared to
<taxonomicName id="37DEDAD6CB74FFA5067DF9EFFB0B6E30" authorityName="Volynkin &amp; Černý" authorityYear="2023" box="[1035,1194,1621,1647]" class="Insecta" family="Erebidae" genus="Leucanithosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="C2AA7D47CB74FFA5067DF9EFFB0B6E30" box="[1035,1194,1621,1647]" italics="true" pageId="7" pageNumber="158">Leucanithosia</emphasis>
</taxonomicName>
(
<figureCitation id="68E5BDD0CB74FFA506CFF9EFFA9A6E2F" box="[1209,1339,1621,1648]" captionStart-0="FIGURES 4043" captionStart-1="FIGURES 4447" captionStart-2="FIGURES 4850" captionStart-3="FIGURES 5154" captionStart-4="FIGURES 5557" captionStartId-0="16.[151,264,1939,1963]" captionStartId-1="17.[151,264,1939,1963]" captionStartId-2="18.[151,264,1939,1963]" captionStartId-3="19.[151,264,1939,1963]" captionStartId-4="20.[151,264,1939,1963]" captionTargetBox-0="[171,1417,193,1902]" captionTargetBox-1="[161,1429,195,1903]" captionTargetBox-2="[204,1384,181,1902]" captionTargetBox-3="[152,1428,184,1903]" captionTargetBox-4="[192,1394,193,1902]" captionTargetId-0="figure-25@16.[158,1430,181,1915]" captionTargetId-1="figure-21@17.[151,1435,181,1915]" captionTargetId-2="figure-25@18.[191,1397,181,1915]" captionTargetId-3="figure-21@19.[151,1435,181,1915]" captionTargetId-4="figure-25@20.[180,1407,181,1915]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionTargetPageId-3="19" captionTargetPageId-4="20" captionText-0="FIGURES 4043. Leucanithosia leucanioides: male genitalia. Depositories of the specimens dissected: 40 and 41 in MWM/ ZSM; 42 and 43 in CKC." captionText-1="FIGURES 4447. Leucanithosia spp.: male genitalia. Depositories of the specimens dissected: 44, 45 and 47 in MWM/ZSM (47 ex CKC); 46 in CBP." captionText-2="FIGURES 4850. Leucanithosia spp.: male genitalia. Depositories of the specimens dissected: 48 and 50 in MWM/ZSM (50 ex CKC); 49 in GMF-B." captionText-3="FIGURES 5154. Leucanithosia witti: male genitalia. Depositories of the specimens dissected: 5153 in MWM/ZSM; 54 in CKC." captionText-4="FIGURES 5557. Leucanithosia spp.: male genitalia. The specimens dissected are deposited in MWM/ZSM (56 and 57 ex CKC)." figureDoi-0="http://doi.org/10.5281/zenodo.8407104" figureDoi-1="http://doi.org/10.5281/zenodo.8407108" figureDoi-2="http://doi.org/10.5281/zenodo.8407110" figureDoi-3="http://doi.org/10.5281/zenodo.8407114" figureDoi-4="http://doi.org/10.5281/zenodo.8407116" httpUri-0="https://zenodo.org/record/8407104/files/figure.png" httpUri-1="https://zenodo.org/record/8407108/files/figure.png" httpUri-2="https://zenodo.org/record/8407110/files/figure.png" httpUri-3="https://zenodo.org/record/8407114/files/figure.png" httpUri-4="https://zenodo.org/record/8407116/files/figure.png" pageId="7" pageNumber="158">Figs 4057</figureCitation>
),
<taxonomicName id="37DEDAD6CB74FFA50738F9EFFEA66ECC" authorityName="Volynkin &amp; Černý" authorityYear="2023" class="Insecta" family="Erebidae" genus="Philippinithosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="C2AA7D47CB74FFA50738F9EFFEA66ECC" italics="true" pageId="7" pageNumber="158">Philippinithosia</emphasis>
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lacks the intervalval commissure, and its juxta is weakly sclerotised whereas in
<taxonomicName id="37DEDAD6CB74FFA50609F9C3FABF6ECC" authorityName="Volynkin &amp; Černý" authorityYear="2023" box="[1151,1310,1657,1683]" class="Insecta" family="Erebidae" genus="Leucanithosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="C2AA7D47CB74FFA50609F9C3FABF6ECC" box="[1151,1310,1657,1683]" italics="true" pageId="7" pageNumber="158">Leucanithosia</emphasis>
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, the valvae are proximally connected by a commissure, and the posterior section of the juxta is heavily sclerotised and laterally fused with transtillae and ventrally connected to the intervalval commissure. The uncus of the new genus is proximally narrow and distally dilated and swollen whereas it is evenly swollen and basally dilated in
<taxonomicName id="37DEDAD6CB74FFA506B0F95FFAC46EA0" authorityName="Volynkin &amp; Černý" authorityYear="2023" box="[1222,1381,1765,1791]" class="Insecta" family="Erebidae" genus="Leucanithosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="C2AA7D47CB74FFA506B0F95FFAC46EA0" box="[1222,1381,1765,1791]" italics="true" pageId="7" pageNumber="158">Leucanithosia</emphasis>
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. The phallus of
<taxonomicName id="37DEDAD6CB74FFA50379F8B3FE626F7C" authorityName="Volynkin &amp; Černý" authorityYear="2023" box="[271,451,1801,1827]" class="Insecta" family="Erebidae" genus="Philippinithosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="C2AA7D47CB74FFA50379F8B3FE626F7C" box="[271,451,1801,1827]" italics="true" pageId="7" pageNumber="158">Philippinithosia</emphasis>
</taxonomicName>
is broader than in
<taxonomicName id="37DEDAD6CB74FFA500EFF8B3FC996F7C" authorityName="Volynkin &amp; Černý" authorityYear="2023" box="[665,824,1801,1827]" class="Insecta" family="Erebidae" genus="Leucanithosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="C2AA7D47CB74FFA500EFF8B3FC996F7C" box="[665,824,1801,1827]" italics="true" pageId="7" pageNumber="158">Leucanithosia</emphasis>
</taxonomicName>
and lacks the distal serrulate plate. The vesicae of the two genera display no fundamental differences. The female genitalia ground plan of
<taxonomicName id="37DEDAD6CB74FFA50626F897FAA56F18" authorityName="Volynkin &amp; Černý" authorityYear="2023" box="[1104,1284,1837,1863]" class="Insecta" family="Erebidae" genus="Philippinithosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="C2AA7D47CB74FFA50626F897FAA56F18" box="[1104,1284,1837,1863]" italics="true" pageId="7" pageNumber="158">Philippinithosia</emphasis>
</taxonomicName>
(
<figureCitation id="68E5BDD0CB74FFA50762F897FA356F18" box="[1300,1428,1837,1863]" captionStart-0="FIGURES 6972" captionStart-1="FIGURES 7376" captionStartId-0="24.[151,264,1938,1962]" captionStartId-1="25.[151,264,1943,1967]" captionTargetBox-0="[200,1387,194,1905]" captionTargetBox-1="[173,1419,185,1910]" captionTargetId-0="figure-25@24.[187,1399,181,1914]" captionTargetId-1="figure-20@25.[160,1427,181,1919]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionText-0="FIGURES 6972. Leucanithosia (6971) and Philippinithosia (72) spp.: female genitalia. Depositories of the specimens dissected: 69, 70, and 72 in MWM/ZSM; 71 in CKC." captionText-1="FIGURES 7376. Philippinithosia, Tigrioides and Dubatoloviana spp.: female genitalia. Depositories of the specimens dissected: 73, 74 and 76 in MWM/ZSM; 75 in NHMUK (©The Trustees of NHMUK)." figureDoi-0="http://doi.org/10.5281/zenodo.8407129" figureDoi-1="http://doi.org/10.5281/zenodo.8407131" httpUri-0="https://zenodo.org/record/8407129/files/figure.png" httpUri-1="https://zenodo.org/record/8407131/files/figure.png" pageId="7" pageNumber="158">Figs 7274</figureCitation>
) is similar to
<taxonomicName id="37DEDAD6CB74FFA5035FF8EBFE696F34" authorityName="Volynkin &amp; Černý" authorityYear="2023" box="[297,456,1873,1899]" class="Insecta" family="Erebidae" genus="Leucanithosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="C2AA7D47CB74FFA5035FF8EBFE696F34" box="[297,456,1873,1899]" italics="true" pageId="7" pageNumber="158">Leucanithosia</emphasis>
</taxonomicName>
(
<figureCitation id="68E5BDD0CB74FFA503AFF8EBFDC06F33" box="[473,609,1873,1900]" captionStart-0="FIGURES 6568" captionStart-1="FIGURES 6972" captionStartId-0="23.[151,264,1943,1967]" captionStartId-1="24.[151,264,1938,1962]" captionTargetBox-0="[237,1350,187,1910]" captionTargetBox-1="[200,1387,194,1905]" captionTargetId-0="figure-21@23.[224,1363,181,1919]" captionTargetId-1="figure-25@24.[187,1399,181,1914]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionText-0="FIGURES 6568. Leucanithosia spp.: female genitalia. Depositories of the specimens dissected: 65 in OUMNH; 6668 in CKC." captionText-1="FIGURES 6972. Leucanithosia (6971) and Philippinithosia (72) spp.: female genitalia. Depositories of the specimens dissected: 69, 70, and 72 in MWM/ZSM; 71 in CKC." figureDoi-0="http://doi.org/10.5281/zenodo.8407125" figureDoi-1="http://doi.org/10.5281/zenodo.8407129" httpUri-0="https://zenodo.org/record/8407125/files/figure.png" httpUri-1="https://zenodo.org/record/8407129/files/figure.png" pageId="7" pageNumber="158">Figs. 6571</figureCitation>
) but in the current new genus, the appendix bursae is weakly expressed whereas the bursa copulatrix of
<taxonomicName id="37DEDAD6CB74FFA50074F8CFFD006FD0" authorityName="Volynkin &amp; Černý" authorityYear="2023" box="[514,673,1909,1935]" class="Insecta" family="Erebidae" genus="Leucanithosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="C2AA7D47CB74FFA50074F8CFFD006FD0" box="[514,673,1909,1935]" italics="true" pageId="7" pageNumber="158">Leucanithosia</emphasis>
</taxonomicName>
is clearly subdivided into a corpus bursae and a sack-like appendix bursae.
</paragraph>
</subSubSection>
<subSubSection id="B8C4F2DECB74FFAA02B1F807FF666BDF" lastPageId="8" lastPageNumber="159" pageId="7" pageNumber="158" type="description">
<paragraph id="F061A155CB74FFAA02B1F807FF666BDF" blockId="7.[151,1437,1477,2008]" lastBlockId="8.[151,1437,150,1005]" lastPageId="8" lastPageNumber="159" pageId="7" pageNumber="158">
<emphasis id="C2AA7D47CB74FFA502B1F807FEF06F88" bold="true" box="[199,337,1981,2007]" pageId="7" pageNumber="158">Description</emphasis>
.
<emphasis id="C2AA7D47CB74FFA50316F807FE0E6F88" bold="true" box="[352,431,1981,2007]" pageId="7" pageNumber="158">Adults</emphasis>
. Sexual dimorphism limited: female somewhat larger than male and with slightly broader forewing. Antenna dark grey, ciliate in both sexes. Head creamy or ochreous creamy. Thorax creamy. Pro- and metathorax with black spot medio-dorsally. Patagia ochreous creamy. Male foreleg with androconial tuft consisting of ochreous yellow scales. Tegula creamy with black spot medially. Forewing elongate and narrow, with almost parallel margins; costal margin straight proximally and medially and downcurved distally; anal margin convex. Forewing ground colour creamy or ochreous creamy. Pattern represented by dark grey or brown suffusion along veins, reduced in certain species. Forewing cilia creamy or ochreous creamy. Hindwing unicolorous creamy or ochreous creamy, paler than forewing. Abdomen creamy with intense admixture of ochreous scales laterally and distally.
<emphasis id="C2AA7D47CB7BFFAA0283FE28FE3E69F3" bold="true" box="[245,415,402,428]" pageId="8" pageNumber="159">Male genitalia</emphasis>
. Uncus proximally narow and distally dilated and swollen, with tiny claw-shaped tip. Tegumen weakly sclerotised, with posteriorly dilated arms. Tuba analis membranous, as broad as half of tegumen length. Vinculum shorter than tegumen, with narrowly U-shaped saccus. Valva lobular, somewhat dilated medially. Valva apex dilated, swollen and densely covered with short setae. Sacculus narrow and elongate, distal saccular process dilated and densely setose apically. Juxta moderately sclerotised, trapezoidal with medial anterior incision. Anellus membranous. Phallus short but broad, cylindrical. Proximal section of vesica broadly tubular. Distal section of vesica narrower than proximal one, tubular, terminating with one or two tubular diverticula bearing one or two short but basally broad cornuti apically. Vesica ejaculatorius originates medially, with stick-shaped basal plate.
<emphasis id="C2AA7D47CB7BFFAA0733FD34FF5F6A93" bold="true" pageId="8" pageNumber="159">Female genitalia</emphasis>
. Papilla analis trapezoidal with rounded corners, weakly setose. Apophyses elongate and thin, apophysis anterioris shorter or equal in length to apophysis posterioris. Postvaginal area membranous. Ductus bursae elongate, tubular, rugose. Corpus bursae broad sack-like with two small and rounded signa, rugose posteriorly, in certain species with short and semiglobular postero-lateral rugose appendix at junction with ductus bursae. Appendix bursae not separated from corpus bursae, represented as broad lateral protrusion with ductus seminalis originating laterally.
</paragraph>
</subSubSection>
<subSubSection id="B8C4F2DECB7BFFAA02B1FC30FBDC6BFB" box="[199,1149,906,933]" pageId="8" pageNumber="159" type="distribution">
<paragraph id="F061A155CB7BFFAA02B1FC30FBDC6BFB" blockId="8.[151,1437,150,1005]" box="[199,1149,906,933]" pageId="8" pageNumber="159">
<emphasis id="C2AA7D47CB7BFFAA02B1FC30FEF86BFB" bold="true" box="[199,345,906,932]" pageId="8" pageNumber="159">Distribution</emphasis>
. The genus is endemic to the
<collectingCountry id="88C9E1C5CB7BFFAA00D1FC30FC846BFB" box="[679,805,906,932]" name="Philippines" pageId="8" pageNumber="159">Philippines</collectingCountry>
(absent from
<collectingRegion id="321A6FB7CB7BFFAA01B4FC30FB806BFB" box="[962,1057,906,932]" country="Philippines" name="Palawan" pageId="8" pageNumber="159">Palawan</collectingRegion>
Island).
</paragraph>
</subSubSection>
<subSubSection id="B8C4F2DECB7BFFAA02B1FC14FA9A6BB3" pageId="8" pageNumber="159" type="etymology">
<paragraph id="F061A155CB7BFFAA02B1FC14FA9A6BB3" blockId="8.[151,1437,150,1005]" pageId="8" pageNumber="159">
<emphasis id="C2AA7D47CB7BFFAA02B1FC14FEE96B97" bold="true" box="[199,328,942,968]" pageId="8" pageNumber="159">Etymology</emphasis>
. The new genus name is an aggregate of the toponym The Philippines and the genus group name
<taxonomicName id="37DEDAD6CB7BFFAA02E1FC68FF526BB3" authorityName="Fabricius" authorityYear="1798" box="[151,243,978,1004]" class="Insecta" family="Erebidae" genus="Lithosia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="genus">
<emphasis id="C2AA7D47CB7BFFAA02E1FC68FF526BB3" box="[151,243,978,1004]" italics="true" pageId="8" pageNumber="159">Lithosia</emphasis>
</taxonomicName>
, and refers to the Philippine Archipelago to which the new genus is endemic. Gender is feminine.
</paragraph>
</subSubSection>
</treatment>
</document>