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<document id="C728BF7666B2FF5ADCA8AA07CCFFB9B5" ID-CLB-Dataset="302074" ID-DOI="10.3897/phytokeys.246.129476" ID-GBIF-Dataset="b381880a-e0bb-4b02-ad0f-2f90a0b6c995" ID-publisher-id="129476" URI-arpha="7631F16E-4CD8-5A76-9F28-45E5F398CE20" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1725298343909" checkinUser="pensoft" docAuthor="Meerts, Pierre J. &amp; Paton, Alan J." docDate="2024" docId="787486148B305397B7C861C438456D7B" docLanguage="en" docName="PhytoKeys 246: 71-178" docOrigin="PhytoKeys 246" docSource="https://phytokeys.pensoft.net/article/129476/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.phytokeys" docStyleName="PensoftTaxPub.0000.journal_article.phytokeys" docTitle="Coleus welwitschii Briq." docType="treatment" docVersion="5" dtd-version="3.0" lastPageNumber="178" masterDocId="7631F16E4CD85A769F2845E5F398CE20" masterDocTitle="The genus Coleus (Lamiaceae) in Central Africa (Democratic Republic of the Congo, Rwanda, Burundi), with the description of 15 new species" masterLastPageNumber="178" masterPageNumber="71" pageNumber="71" updateTime="1725322260938" updateUser="ExternalLinkService">
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<mods:title id="394F642E815472D41105BC453ED2E96D">The genus Coleus (Lamiaceae) in Central Africa (Democratic Republic of the Congo, Rwanda, Burundi), with the description of 15 new species</mods:title>
</mods:titleInfo>
<mods:name id="FB035F46E53254E9C4FB089CDAC4293F" type="personal">
<mods:role id="72A13A964041D5A29603F7E21266E6E4">
<mods:roleTerm id="727632B197A6174D75C3E8928D571933">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D0C8B1A5025BB266E06322D1788262F4">Meerts, Pierre J.</mods:namePart>
<mods:nameIdentifier id="A79586318A5214DF7678B055FC6AEB50" type="ORCID">0000-0003-4215-027X</mods:nameIdentifier>
<mods:affiliation id="8531C57260E6223270E0945B9C0169AB">Meise Botanic Garden, Nieuwelaan 38, BE- 1860 Meise, Belgium &amp; Université Libre de Bruxelles, Av. F. D. Roosevelt 50 CP 244, BE- 1050 Brussels, Belgium &amp; Fédération Wallonie-Bruxelles, Service général de l’Enseignement supérieur et de la Recherche scientifique, Brussels, Belgium</mods:affiliation>
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</mods:role>
<mods:namePart id="F39D649E6DE5D5E37CE9A8013339ABBF">Paton, Alan J.</mods:namePart>
<mods:nameIdentifier id="70E5485CEEAA2C2CE00A2FD185715BF1" type="ORCID">0000-0002-6052-6675</mods:nameIdentifier>
<mods:affiliation id="9AEE692065C503E6B02539533B7BAD6A">The Herbarium, Royal Botanic Gardens, Kew, Richmond TW 9 3 AB, UK</mods:affiliation>
</mods:name>
<mods:typeOfResource id="113A8D0037C4760308AF2ABED53B76C5">text</mods:typeOfResource>
<mods:relatedItem id="F6AA640EFAAC23C1D5078013A65BE377" type="host">
<mods:titleInfo id="9BE0A90A787B822AB79864006324574D">
<mods:title id="2F5642B0D7EC7036D90E9F582406C97E">PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part id="645F6AE4737A334322EEDBDA971183AA">
<mods:date id="289DAA00170700D2B338544B0AD2B03A">2024</mods:date>
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<mods:number id="430B5D8A61B385C7C3F2BDDD4564D60F">2024-09-02</mods:number>
</mods:detail>
<mods:detail id="3A66EB9774349BE5B3A479E71006AC2B" type="volume">
<mods:number id="56FD5297EBB0FA445C7AC21550DAE940">246</mods:number>
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</mods:extent>
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<mods:classification id="5D56207807E26895A5A86CCC98389963">journal article</mods:classification>
<mods:identifier id="7448003BA85060CFF00E2C2C838356D7" type="CLB-Dataset">302074</mods:identifier>
<mods:identifier id="5D9E79D27CACB2F2EA5AB482EE44820F" type="DOI">10.3897/phytokeys.246.129476</mods:identifier>
<mods:identifier id="51022224AFEC95E0CA67ED427A46D247" type="GBIF-Dataset">b381880a-e0bb-4b02-ad0f-2f90a0b6c995</mods:identifier>
</mods:mods>
<treatment id="787486148B305397B7C861C438456D7B" ID-DOI="http://doi.org/10.5281/zenodo.13629924" ID-GBIF-Taxon="240061296" ID-Zenodo-Dep="13629924" ID-arpha="78748614-8B30-5397-B7C8-61C438456D7B" LSID="urn:lsid:plazi:treatment:787486148B305397B7C861C438456D7B" httpUri="http://treatment.plazi.org/id/787486148B305397B7C861C438456D7B">
<subSubSection id="34AC0C893806C19E5053A276FA88D4A1" type="nomenclature">
<paragraph id="13A0D5BC040010DC63A779AC831CE3C7">
<treatmentCitation id="865A20877C8B8AA9BC47070D9D72BE15" author="Briq." journal="Bot. Jahrb. Syst." page="185" score="144.29617767960897" volume="19" year="1895">
<taxonomicName id="FB9D8E2DCD2B333EAAAE0D4DA7438A81" ID-CoL="X2BV" authority="Briq." authorityName="Briq." class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="welwitschii">
<emphasis id="DD86B24791B6D1E1AB7B6E6BE06FF634" italics="true">Coleus welwitschii</emphasis>
Briq.
</taxonomicName>
, Bot. Jahrb. Syst. 19: 185. 1895
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BA402E10C7D23DAF32F949EC0AE64DAC" type="reference_group">
<paragraph id="8CE6DFCB3A821C5D4B4C45B236EAB60E">
<treatmentCitationGroup id="D56980E2A05D59C0561F016A4661779B">
=
<treatmentCitation id="8700037DF7FE2ECD4EC5CAAB3C7E7749" author="Briq." journal="Bull. Soc. Roy. Bot. Belgique" page="70" score="3.0378308617219716" volume="37" year="1899">
<taxonomicName id="F46D7E2B0A5BFFB8CC782EF7ED0B3F03" ID-CoL="X24S" authority="Briq." authorityName="Briq." class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="dupuisii">
<emphasis id="1F13234C6B60E993EBFA9B0D54D1980A" italics="true">Coleus dupuisii</emphasis>
Briq.
</taxonomicName>
, Bull. Soc. Roy. Bot.
<collectingCountry id="9B4BBA4DA1407211B120D8B3DE246935" name="Belgium">Belgique</collectingCountry>
37: 70. 1899
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="4F5E1E61ED389ABFED1368B41BA9428E">
<treatmentCitationGroup id="DBA6F767E06134CFE2A105A72769073A">
≡
<treatmentCitation id="F99873BABDB33A0AB36A5F68D03F3D8F" author="E. Afr." page="329" volumeTitle="Lamiac." year="2009">
<taxonomicName id="76353BA9F06823A4C48FCB09C1BA8E99" ID-CoL="4K47W" authority="(Briq.) A. J. Paton" authorityName="A. J. Paton" baseAuthorityName="Briq." class="Magnoliopsida" family="Lamiaceae" genus="Plectranthus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="dupuisii">
<emphasis id="94229B1A8D255F1CC0F6CFB64F5A8925" italics="true">Plectranthus dupuisii</emphasis>
(Briq.) A. J. Paton
</taxonomicName>
, Fl. Trop. E. Afr., Lamiac.: 329. 2009
</treatmentCitation>
. Type:
<collectingCountry id="218DE996119990BF46906896C1A01F98">DR. Congo</collectingCountry>
, Kasai, Mayumbe,
<date id="4CAAA31249259151CF9A0F5B686466D0" value="1893-07">Jul 1893</date>
,
<emphasis id="D572C38C168F94067CC45CDD6AE0942B" italics="true">Dupuis s. n.</emphasis>
(
<typeStatus id="B745CC20365B80D091070252221F1409" type="holotype">holotype</typeStatus>
<collectionCode id="BCBAF8497AC529F3ABDA341B2E36456C" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
[
<specimenCode id="512A43B92E4A4DCB14282F2C9CE78FA4">
<collectionCode id="6C6E4E73317D410BF2461AACD6A9974D" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
0000008109132
</specimenCode>
]).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A4DF22E03DFFEA1260F14EBDE4C7EEEC">
<treatmentCitationGroup id="F5BB4C3D6D589112D721CF7A0D71A0D3">
=
<treatmentCitation id="5303A645F07B1AF6E8093B4D72A79313" author="Vollesen" journal="Opera Bot." page="85" score="153.25706264698434" volume="59" year="1980">
<taxonomicName id="8DCC8BA7B6AA7B74D207F5474C82C06F" authority="(Lebrun &amp; L. Touss.) Vollesen" authorityName="Vollesen" baseAuthorityName="Lebrun &amp; L. Touss." class="Magnoliopsida" family="Lamiaceae" genus="Solenostemon" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="thyrsiflorum">
<emphasis id="E5F616AA52793358EDEE1526803031D3" italics="true">Solenostemon thyrsiflorum</emphasis>
(Lebrun &amp; L. Touss.) Vollesen
</taxonomicName>
, Opera Bot. 59: 85. 1980
</treatmentCitation>
., nom. superfl. Type:
<collectingCountry id="D7223FA2A3E8499655C6F614FC6A6207" name="Rwanda">Rwanda</collectingCountry>
, Kagera, Nyakayaga,
<date id="540E887AB005C48C3AD55057056225FD" value="1938-01">Jan 1938</date>
,
<emphasis id="18CF8B610278D544B36D8224C2FFA618" italics="true">J. Lebrun 9447</emphasis>
(
<typeStatus id="08356F7A046C2768418F9E8FCCA9DC0B" type="holotype">holotype</typeStatus>
<collectionCode id="EB12ACBC3F247C3A16F558FE8F965500" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
[
<specimenCode id="9A9137B4EF288555A255C120F7DD404D">
<collectionCode id="A79D2B885E1C7D59830C3EBD3F20F28D" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
0000006263003
</specimenCode>
];
<typeStatus id="40D2569EC18755ECD0BBA13AC64FFA64" type="isotype">isotype</typeStatus>
<collectionCode id="648A947CB264F7DA8FB625C0BDF2297A" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
, P).
</treatmentCitationGroup>
</paragraph>
<paragraph id="F1ACC25112DCA54299F944453EC81732">
<treatmentCitationGroup id="81CAFF09915D8456572CAABA4B939072">
<treatmentCitation id="DCDDB5E49C5A25B5E53BFF7AB677E08A">
=?
<taxonomicName id="5951DD0BBEE4BA5C51C60D37237AF1E5" ID-CoL="X278" authority="De Wild. (1920)" authorityName="De Wild." authorityYear="1920" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="laurentii">
<emphasis id="CCD39A5040A9C5F93190BCF91317FF1C" italics="true">Coleus laurentii</emphasis>
De Wild. (1920)
</taxonomicName>
121.
<bibCitation id="F30950D0D42DD5F4348DD1775DB4A45D" journal="Bol. Soc. Iber. Ci. Nat." page="121" score="282.3037265619079" volume="19" year="1920">Bol. Soc. Ibér. Ci. Nat. 19: 121. 1920</bibCitation>
. Type:
<collectingCountry id="4E2BA01BE18B96E55F69A4FB8223B5B6">DR. Congo</collectingCountry>
, Gombe,
<date id="24495E0651379CB3EC05F48FB179F95F" value="1903-12-14">14 Dec 1903</date>
,
<emphasis id="895F1AD47C9A8461071D7DD7E00D8E7F" italics="true">
<collectionCode id="68524B34C3B620C176A175BE98A42D7D" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" type="Herbarium">E</collectionCode>
. &amp;
<collectionCode id="925F34EE0228C382BD67D54419EEF515" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" type="Herbarium">M</collectionCode>
. Laurent s. n.
</emphasis>
(
<typeStatus id="8290F8DF72EEB6CB8E5F2AA2A8927DB1" type="holotype">holotype</typeStatus>
<collectionCode id="D139BC43F72A4B88245BCDE55541EAFD" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
[
<specimenCode id="EFAB18879F8C8287819E1395DA46268B">
<collectionCode id="8E74024B3CDE9EEF04E5A41C2CFC044E" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
0000006258580
</specimenCode>
] &amp; [
<specimenCode id="A7DF6E1BCD9CE73E913F79ED2B3BDA40">
<collectionCode id="CE718DDB9E7CF3B515AF96E73695DAAB" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
0000008109798
</specimenCode>
]).
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEWJM" type="material">
<paragraph id="C5E8C34A23292107BC6F7BD100A422D9">
<heading id="8CA613C28FB250B83D57BFD0A0A2ABD3" reason="title">Type.</heading>
</paragraph>
<paragraph id="96CEEE22B9D7824C208DF5C7C42CFDAB">
<collectingCountry id="3C636B91159A64274326D0DB4426B97E" name="Angola">Angola</collectingCountry>
, Pungo Andongo,
<emphasis id="27F64DABA5FB1EA3D9AA7E7972506302" italics="true">F. Welwitsch 5589</emphasis>
(
<typeStatus id="9D5BFFDF6915F5297C4CDC2E4A95BC07" type="syntype">syntypes</typeStatus>
<collectionCode id="7203881B05814B6447D9542D8806077B" country="United Kingdom" name="Bristol Museum">BM</collectionCode>
[
<specimenCode id="61426065A2402CE84B74D90C3A47654F">
<collectionCode id="0CFFB563430DB62D171A774CB301C0E2" country="United Kingdom" name="Bristol Museum">BM</collectionCode>
000564040
</specimenCode>
], [
<specimenCode id="87A3B19BE9A1C82749C6F5246142D07F">
<collectionCode id="55A50A182CB1064D73D57FBD1344A1C2" country="United Kingdom" name="Bristol Museum">BM</collectionCode>
000999976
</specimenCode>
],
<collectionCode id="21384C695645EE01780535867408FA07" country="Denmark" name="University of Copenhagen" type="Herbarium">C</collectionCode>
[
<collectionCode id="DBF880DA74334F1BE2B2BA50DB4FE029" country="Denmark" name="University of Copenhagen" type="Herbarium">C</collectionCode>
10001537],
<collectionCode id="486ECAE7A6E61CAA8A51B8168D639782" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" type="Herbarium">G</collectionCode>
,
<collectionCode id="E9B9A82F2A188798AEFC8FD286898F11" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
,
<collectionCode id="D2EC173B99633CCC6F22807A9571B5E1" country="Portugal" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15046" lsid="urn:lsid:biocol.org:col:15046" name="Museu Nacional de História Natural" type="Herbarium">LISU</collectionCode>
[
<collectionCode id="52F0C8F6B9F8156802F93C55865D30A5" country="Portugal" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15046" lsid="urn:lsid:biocol.org:col:15046" name="Museu Nacional de História Natural" type="Herbarium">LISU</collectionCode>
220997], [
<collectionCode id="0186C8D3FB692BCC40CC7FBA4E9B8BB7" country="Portugal" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15046" lsid="urn:lsid:biocol.org:col:15046" name="Museu Nacional de História Natural" type="Herbarium">LISU</collectionCode>
220998],
<collectionCode id="BA0CF15BCF6312D9361C66E73255A5AD" country="Nicaragua" httpUri="http://grbio.org/cool/hd8z-yxxc" name="Museo Entomologico de Leon" type="Museum or Herbarium: Private non-profit">MEL</collectionCode>
, P), &amp;
<emphasis id="D5B96A38AF0699576B3415BB8B1EDC94" italics="true">A. von Mechow 75</emphasis>
(
<typeStatus id="253557CA90A7AFC52D70ABC25B0CC8A8" type="syntype">syntype</typeStatus>
, not seen).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EOWJM" type="description">
<paragraph id="F1AF18620728313E016D39827C8558B1">
<heading id="415E2D8A295A87046555B3789B7946C4" reason="title">Description.</heading>
</paragraph>
<paragraph id="4262400DA40C8060AF612BD46849105D">
<bibRefCitation id="591CC8F80727D1719BD30E64CFB69229" author="Paton AJ &amp; Bramley G &amp; Ryding O &amp; Polhill RM &amp; Harvey YB &amp; Iwarsson M &amp; Willis F &amp; Phillipson PB &amp; Balkwill K &amp; Lukhoba CW &amp; Otieno D &amp; Harley RM" editor="Beentje HJ &amp; Ghazanfar SA &amp; Polhill RM" etAl="et al." firstAuthor="Paton" journalOrPublisher="Royal Botanic Gardens, Kew, Richmond" page="329" pagination="1-430" refId="B20" refString="Paton AJ, Bramley G, Ryding O, Polhill RM, Harvey YB, Iwarsson M, Willis F, Phillipson PB, Balkwill K, Lukhoba CW, Otieno D, Harley RM (2009) Lamiaceae (Labiatae). In: Beentje HJ, Ghazanfar SA, Polhill RM (Eds) Flora of Tropical East Africa. Royal Botanic Gardens, Kew, Richmond, 1–430." title="Lamiaceae (Labiatae)." volumeTitle="Flora of Tropical East Africa." year="2009">Paton et al. (2009: 329)</bibRefCitation>
,
<bibRefCitation id="04D11D4F9F00D36556CF1ED86CEC6FC8" author="Paton AJ &amp; Bramley G &amp; Ryding O &amp; Polhill RM &amp; Harvey YB &amp; Iwarsson M &amp; Willis F &amp; Phillipson PB &amp; Balkwill K &amp; Lukhoba CW &amp; Otieno D &amp; Harley RM" editor="Timberlake J" etAl="et al." firstAuthor="Paton" journalOrPublisher="Royal Botanic Gardens, Kew, Richmond" page="268" pagination="1-346" refId="B21" refString="Paton AJ, Bramley G, Ryding O, Polhill RM, Harvey YB, Iwarsson M, Willis F, Phillipson PB, Balkwill K, Lukhoba CW, Otieno D, Harley RM (2013) Lamiaceae. In: Timberlake J (Ed.) Flora Zambesiaca 8 (8). Royal Botanic Gardens, Kew, Richmond, 1–346." title="Lamiaceae." volumeTitle="Flora Zambesiaca 8 (8)." year="2013">Paton et al. (2013: 268)</bibRefCitation>
, as
<taxonomicName id="CF5FD5F9140CC7ABE2BC4AA29BE56EB3" authority="(Briq.) A. J. Paton." authorityName="A. J. Paton." baseAuthorityName="Briq." class="Magnoliopsida" family="Lamiaceae" genus="Plectranthus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="dupuisii">
<emphasis id="AD65194DC3694F773010841E56D1AD52" italics="true">Plectranthus dupuisii</emphasis>
(Briq.) A. J. Paton.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EHXJM" type="distribution">
<paragraph id="775B5C5F0DC1C433B1352AEBBFBDEA1B">
<heading id="91D04EE868BF0058192231A3FD090032" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="F8EE6F1F94A0A21F66082C69CC028D28">
<collectingCountry id="8B47D9503E1B7A74D0109DBF52484886" name="Ethiopia">Ethiopia</collectingCountry>
to S. Tropical Africa.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EMXJM" type="habitat">
<paragraph id="21AED226BAE7120C2BBA4ACC0395170C">
<heading id="759C45D49AF260AABDD287BB25BA21CA" reason="title">Habitat and ecology.</heading>
</paragraph>
<paragraph id="33B03CA5FA04588B4018982B45CCD3BF">
Savannah, steppic savannah, often on shallow rocky soil, xerophilous scrub, more rarely forest and woodland (termite mounds);
<quantity id="4B30ED84236627C71C08BD41D19D36A5" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="2.2" metricValueMin="0.1" unit="m" value="1150.0" valueMax="2200.0" valueMin="100.0">100–2200 m</quantity>
elev.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERXJM" type="Additional specimens">
<paragraph id="BAAF29B09CFFF8428320D4273857C636">
<heading id="87926C62FCF8A9E616251D814865B34F" reason="title">Additional specimens.</heading>
</paragraph>
<paragraph id="7D31FC9B9FB25C3735B079D35A1D2E52">
<collectingCountry id="3B5C70F16C78F8AB5780B443C8A20BE2">
DR.
<collectingRegion id="3F37334F6813CEC33C5D2502521B5516" country="Republic of the Congo" name="Bas-Congo">Congo</collectingRegion>
</collectingCountry>
,
<emphasis id="8CB6791184EC02C5D21629DF0DAFE6DB" bold="true" italics="true">Côtier</emphasis>
, Banana,
<date id="49FE1FDC954053362935EEB39321853D" value="1948-03">
<collectingRegion id="001A6F25A70385715977F93218BC48B1" country="South Sudan" name="Lakes">Mar</collectingRegion>
1948
</date>
,
<emphasis id="1C4C54C02598FAD7F3766A4F51A4A6AC" italics="true">
<collectionCode id="32F79CD4231247716E73FD52E8F9AE23" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" type="Herbarium">H</collectionCode>
. Callens 1066
</emphasis>
(
<collectingRegion id="8D80AA2182855C3A413CC03156546295" country="Serbia" name="Bor">BR</collectingRegion>
);
<emphasis id="A287874FAE776D5D644032D79AA43A83" bold="true" italics="true">
<collectingRegion id="9FEA310F68E07E3979D313CBA4C09A83" country="Republic of the Congo" name="Bas-Congo">Bas-Congo</collectingRegion>
</emphasis>
, Léopoldvile [
<collectingRegion id="529AD3DDC137A7DE1697FF56F5B93AAF" country="Republic of the Congo" name="Kinshasa">Kinshasa</collectingRegion>
], Matete,
<date id="CC5BBD7846D91189D129CA821B7E57AA" value="1956-08-05">5 Aug 1956</date>
,
<emphasis id="F01175AFAFC7B1C964E9CC69842153B6" italics="true">A. Carlier 334</emphasis>
(
<collectingRegion id="75087A0BA24519CDA2C30575664AD2A9" country="Serbia" name="Bor">BR</collectingRegion>
);
<emphasis id="108AE7219E453068B48FDD7E85A0272D" bold="true" italics="true">Ubangi-Uele</emphasis>
, Garamba,
<date id="01A69B934811E537D39003BBE64EB9CD" value="1950-05-14">14 May 1950</date>
,
<emphasis id="0FE22B6050AB78086C9E5F9C0B19EC32" italics="true">
<collectionCode id="0ED7E67088FDEE22232AEE240266E3CD" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" type="Herbarium">H</collectionCode>
. De Saeger 511
</emphasis>
(
<collectingRegion id="8460AE43A90DBAD931039C7DF59155F6" country="Serbia" name="Bor">BR</collectingRegion>
); Garamba, route Dungu-Bagbele, km 17 de Bagbele,
<date id="EF9169F4E7B97D4C8C58D6E29486C217" value="1952-08-27">27 Aug 1952</date>
,
<emphasis id="430421A1C262510429BCBB54D4723D0A" italics="true">
<collectionCode id="3BB46AB4CE51E9424B184960126EF1BA" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" type="Herbarium">G</collectionCode>
. Troupin 2028
</emphasis>
(
<collectingRegion id="279A27C445BF7460604938C6D90EDC9A" country="Serbia" name="Bor">BR</collectingRegion>
);
<emphasis id="78C04B5BD18323DE73803604EC9CF36C" bold="true" italics="true">
<collectingRegion id="9348E99B4BA2764B5E7D84352E93F5EE" country="Ivory Coast" name="Lacs">Lacs</collectingRegion>
Edouard et
<collectingRegion id="B4CD2569CAFE91FCEE3029246A50A3D4" country="Republic of the Congo" name="Sud-Kivu">Kivu</collectingRegion>
</emphasis>
, Plaine de la Ruzizi, sommet Tsamate,
<date id="5A39936ACC3853F4EDFC271D6785BDA5" value="1950-04">Apr 1950</date>
,
<emphasis id="F6C54638B5545283F53D7F691E48C251" italics="true">
<collectionCode id="B720EFA637BCA9132317ACF45EFFEED6" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
. Germain 6826
</emphasis>
(
<collectingRegion id="CD36A493E98CF6865EA6BDA64CA562E7" country="Serbia" name="Bor">BR</collectingRegion>
).
</paragraph>
<paragraph id="8DCEB419F188EAEEB901CAA58CAA3C6D">
<collectingCountry id="D73A137C9B29DC2D1CA43C0AC2D7C2CD" name="Rwanda">Rwanda</collectingCountry>
, Bugesera,
<date id="132038445CBB0DEBA00FC97FE8F5C93B" value="1970-03-27">27 Mar 1970</date>
,
<emphasis id="13F4931A01C9EDA387168567ADE8CB19" italics="true">
<collectionCode id="AC329BF7B4B0C31084AE514DBD3743AA" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" type="Herbarium">G</collectionCode>
. Bouxin &amp;
<collectionCode id="51401931629D254E0AE28C18999154DE" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" type="Herbarium">M</collectionCode>
. Radoux 1631
</emphasis>
(
<collectionCode id="5135C3B436E449F33D9C7FBF7C8D7938" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
); Rusumo, route vers Nyarubuye,
<date id="651512E9CCD6B4369687A7BC9945CDCC" value="1980-01-25">25 Jan 1980</date>
,
<emphasis id="6337129567EEB1E77892F51A96CE1993" italics="true">D. Bridson 299</emphasis>
(
<collectionCode id="8E8D69A3E3A3997600FE93F29AD9D3CB" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
,
<collectionCode id="745AB94929C163C4CB6DC9AE576A006A" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15578" lsid="urn:lsid:biocol.org:col:15578" name="Wageningen University" type="Herbarium">WAG</collectionCode>
); Kibungo, Rusumo,
<date id="C29588A74BA7D77FDAA77B83700ED20B" value="1974-10-16">16 Oct 1974</date>
,
<emphasis id="E401C28D1E4272414D90CCFFB2A5BEAA" italics="true">J. Lambinon 74 / 1574</emphasis>
(
<collectionCode id="168C2E51B7303A784FAC6BAB42F55F44" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
,
<collectionCode id="D1951B9F92E5F9D7056FA33D7038EAD2" country="Belgium" name="Université de Liège" type="Herbarium">LG</collectionCode>
); Route Nyamasheke-Kibuye,
<date id="AEBDCD3249ABDDDAF41A3858666F1EE7" value="1972-03-29">29 Mar 1972</date>
,
<emphasis id="DA82445E96FD42D75A7D69BE2CA538DD" italics="true">
<collectionCode id="C9EEC46182D3614753E90DB3755C8FC0" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" type="Herbarium">G</collectionCode>
. Troupin 6510
</emphasis>
(
<collectionCode id="DB9F6508CFA62DE19C131A3D659CBCCC" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
,
<collectionCode id="31554C7D418E9FAB2C081275E7EFECBD" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15578" lsid="urn:lsid:biocol.org:col:15578" name="Wageningen University" type="Herbarium">WAG</collectionCode>
); Kitega, Bweru, Muhweza,
<date id="853B3BDAA4E593A9E2F5E8E1FD3D2603" value="1957-11-27">27 Nov 1957</date>
,
<emphasis id="D70E30F4E11B5527B061DD990090AC9E" italics="true">van der Ben 1727</emphasis>
(
<collectionCode id="E8199F5DA8C90E665F8E726937CF1B0F" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
).
</paragraph>
<paragraph id="9968FC0EA14FEC5299D13BF85DCB8C60">
<collectingCountry id="20DEE8F4D7FFCCD16D2F2BCD7B501090" name="Burundi">Burundi</collectingCountry>
, Buterama, Gasorwe,
<date id="900D812188FDCF5E8EAE43D209B1628D" value="1959-05-22">22 May 1959</date>
,
<emphasis id="455731ACF1D15441EAE4191B8356D0DC" italics="true">van der Ben 2558</emphasis>
(
<collectionCode id="40812E94871E05B8DFB67E91E5D02CA4" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
); Kameramagambo,
<date id="3596A0D791EAA179F78DA33DFB291E1F" value="1965-12-27">27 Dec 1965</date>
,
<emphasis id="EC77B109E408CE2064B5A67A286E3CFC" italics="true">J. Lewalle 155</emphasis>
(
<collectionCode id="03D3DB291F146BE101315FDDF5EEE837" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
,
<collectionCode id="12FFC979ECF7755F701C7DED1DCFB63C" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" type="Herbarium">MO</collectionCode>
); Gitwenge,
<date id="076067F466BDF162A9CBE51DD2D6EA36" value="1979-01-02">2 Jan 1979</date>
,
<emphasis id="A34EDC78C1D6805335C954850FBE1E9E" italics="true">
<collectionCode id="90A61889930D3E3D544310A3C22E2CC0" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" type="Herbarium">M</collectionCode>
. Reekmans 7448
</emphasis>
(
<collectionCode id="566519EBC977DACD7362E2A26E47171E" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
); Kagoma,
<collectionCode id="844E1877C7AEF46A0E9B3207FEEEB9EA" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" type="Herbarium">E</collectionCode>
de Gitwenge,
<date id="1F809E400E8C042F851E82B49B5146D8" value="1979-02-07">7 Feb 1979</date>
,
<emphasis id="7F4EE369F73F13BF139C58684D3E30C9" italics="true">
<collectionCode id="1600A48F01EA2E5F10388575ADADD907" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" type="Herbarium">M</collectionCode>
. Reekmans 7639
</emphasis>
(
<collectionCode id="BC5B1DE32585A6CEDCC427ED2204BFE8" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
,
<collectionCode id="DA4F4FCCF1E775E50667A47512682BCA" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15578" lsid="urn:lsid:biocol.org:col:15578" name="Wageningen University" type="Herbarium">WAG</collectionCode>
);
<collectingRegion id="8E0741841F8730BFC9C884DBA1BCBB51" country="Burundi" name="Muramvya">Muramvya</collectingRegion>
, Komwe,
<date id="98C0B05A729278F85E90C4C2FCC6366C" value="1981-05-24">24 May 1981</date>
,
<emphasis id="AFF300EC0F4C60F5C57E7FA42E47B56B" italics="true">
<collectionCode id="CD681E970A260AF842969BE2A18E1AF0" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" type="Herbarium">M</collectionCode>
. Reekmans 10355
</emphasis>
(
<collectionCode id="DF743044E67813B181736C209E230D3A" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
,
<collectionCode id="23B239596DB6AE81C0C83A6A8644EB73" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" type="Herbarium">MO</collectionCode>
,
<collectionCode id="2A5314F881A9C93A796658ECCD2DB3B7" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15578" lsid="urn:lsid:biocol.org:col:15578" name="Wageningen University" type="Herbarium">WAG</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVZJM" type="notes">
<paragraph id="0BC44103CD131F687864D61FDC71A9A6">
<heading id="DBA455B56771A90AE6DE2F4E572B26D9" reason="title">Notes.</heading>
</paragraph>
<paragraph id="4AEBF84C794B075E6C513F5CE6B91136">
1.
<taxonomicName id="6F16D766FD6910FEF6B4B2667588D13E" authority="De Wild." authorityName="De Wild." class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="laurentii">
<emphasis id="E8A8084DEA314281D1572896D0CFB2F5" italics="true">Coleus laurentii</emphasis>
De Wild.
</taxonomicName>
is considered as a synonym of
<taxonomicName id="01B04BD5843FC4B19D1026A05B412D6C" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="welwitschii">
<emphasis id="8F2AD7EBBA72538CAF397E8AEB010309" italics="true">C. welwitschii</emphasis>
</taxonomicName>
by
<bibRefCitation id="2F1CC7084F84699E4C3170C51D276EBD" DOI="10.3897/phytokeys.129.34988" author="Paton AJ &amp; Mwanyambo M &amp; Govaerts RH &amp; Smith AK &amp; Suddee S &amp; Phillipson PB &amp; Wilson TC &amp; Forster PI &amp; Culham A" etAl="et al." firstAuthor="Paton" journalOrPublisher="PhytoKeys" pagination="1-158" refId="B23" refString="Paton AJ, Mwanyambo M, Govaerts RH, Smith AK, Suddee S, Phillipson PB, Wilson TC, Forster PI, Culham A (2019) Nomenclatural changes in Coleus and Plectranthus (Lamiaceae): A tale of more than two genera. PhytoKeys 129: 1–158. https://doi.org/10.3897/phytokeys.129.34988" title="Nomenclatural changes in Coleus and Plectranthus (Lamiaceae): A tale of more than two genera." volume="129" year="2019">Paton et al. (2019)</bibRefCitation>
. However, the type material (
<emphasis id="C62932E2546F43915361E736FCAC2A6D" italics="true">
<collectionCode id="C540D4793D1D4E68E60B3B203EA7F4E2" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" type="Herbarium">E</collectionCode>
. &amp;
<collectionCode id="7ADC93919A7E09F1BF21ED5E2D3F3865" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" type="Herbarium">M</collectionCode>
. Laurent s. n.
</emphasis>
[
<specimenCode id="AFF6E941427FDCFDF6DF8085AD6C54DA">
<collectingRegion id="8D08C65B96AB69D96F3D2E34FEF2A037" country="Serbia" name="Bor">BR</collectingRegion>
0000008109798
</specimenCode>
], [
<specimenCode id="07D084458FD7651AF38042AE02DE1BA5">
<collectingRegion id="6FD24171CC4712DE4CF400A0299D108F" country="Serbia" name="Bor">BR</collectingRegion>
0000006258580
</specimenCode>
]) is remarkable in being almost glabrous in all vegetative parts; collecting notes on the label indicate the presence of white tubers (not collected); it could represent a different taxon (
<taxonomicName id="32849950403A06AD52A630F9FC6AF2BD" class="Magnoliopsida" family="Lamiaceae" genus="Plectranthus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="rotundifolius">
<emphasis id="5AF0E5E23C51916CA8E64B01A441A2B7" italics="true">C. rotundifolius</emphasis>
</taxonomicName>
?).
</paragraph>
<paragraph id="B6A6001062BC4CB157B66E98666BBD3F">
2.
<emphasis id="BC38C365E957B4B19046980C3DC15BA5" italics="true">L. Liben 2855</emphasis>
is somewhat intermediate between
<taxonomicName id="94192BA9A69E1EBEC45C6D4FD7A34C1D" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="bojeri">
<emphasis id="E427EE4A8C7BFE941E60CC0F08DAE842" italics="true">C. bojeri</emphasis>
</taxonomicName>
and
<taxonomicName id="C744752C53925096C3B7E0A2B2CE531A" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="welwitschii">
<emphasis id="869B498A12FB09DAF4CED3C5A6ADDEAF" italics="true">C. welwitschii</emphasis>
</taxonomicName>
, having robust lignified shoots, but lacking a true rhizome.
</paragraph>
<paragraph id="B4907A831D1D963A7BC61A784B8846A8">
3. Many specimens from
<collectingCountry id="766712B69827D2B4481688C348A24934" name="Rwanda">Rwanda</collectingCountry>
,
<collectingCountry id="01E7F29EDCD900B888811AC760AB3338" name="Burundi">Burundi</collectingCountry>
and
<collectionCode id="C97103884DC36F770FAB47A60309707C" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" type="Herbarium">W</collectionCode>
<collectingCountry id="5FDAD369F6D8B0A8B3A13F3B1D487CC4" name="Tanzania">Tanzania</collectingCountry>
have greyish-tomentose lower leaf surface and thicker leaves, corresponding to the type specimen of
<taxonomicName id="905F8FF3E68051EE267BC60557E4272F" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="thyrsiflorus">
<emphasis id="02B698DB7C5F4DC7770AA7CA1B24EF18" italics="true">Coleus thyrsiflorus</emphasis>
</taxonomicName>
, but there is no real discontinuity with less pubescent forms.
</paragraph>
<paragraph id="F52C723F9B282EA914DDA5DC47C4601B">
4.
<collectingRegion id="69264A4F7278B6FF1A6609D46DD948A0" country="South Sudan" name="Lakes">See</collectingRegion>
also note under
<taxonomicName id="949C58B1A9CD07EF70849C60312F9C09" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="brazzavillensis">
<emphasis id="4247DF2D4B3E1C7FAED2C52FBE079F5A" italics="true">C. brazzavillensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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