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<document id="B19162A5174FDAA2CFB7186D20A1D67B" ID-DOI="10.11646/zootaxa.5020.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5222931" ID-ZooBank="882BBB9D-6E5B-4CE5-99DF-E91AC7971EB5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629386356657" checkinUser="plazi" docAuthor="Kataev, Boris M." docDate="2021" docId="785C87DD333A002EC8F7FF0EC9B947B3" docLanguage="en" docName="zootaxa.5020.1.2.pdf" docOrigin="Zootaxa 5020 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Harpalus pseudoasemus Kataev 2021, sp. n." docType="treatment" docVersion="3" lastPageNumber="41" masterDocId="8465FFA533330024C860FF99CD16402C" masterDocTitle="On some Afrotropical Harpalus, with description of two new species, and remarks on Hypharpax australis (Coleoptera: Carabidae): misidentification, mislabeling and introduction to the Australian region" masterLastPageNumber="56" masterPageNumber="31" pageNumber="40" updateTime="1699070990175" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="F1A1CC402F2A6E63875EA0ED18220D57" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AE47D4853E9E4C21BFE5CB8B464F417F">
<mods:title id="5B910C40C1FAB48F64E52EF4DB54EA99">On some Afrotropical Harpalus, with description of two new species, and remarks on Hypharpax australis (Coleoptera: Carabidae): misidentification, mislabeling and introduction to the Australian region</mods:title>
</mods:titleInfo>
<mods:name id="432CB8DE5F6E5566514492BDDE9CAD2F" type="personal">
<mods:role id="40E761983A33592D4C0028B472A40FE1">
<mods:roleTerm id="70B3DE8E46CC6CF6DD8CBD6106DEC206">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D12C30F043958A09E7741926F250D0E6">Kataev, Boris M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="74F851D481D741EDB335D59ACEF5DD01">text</mods:typeOfResource>
<mods:relatedItem id="1C7D93721D1D033C206AFCFF33115D1B" type="host">
<mods:titleInfo id="F8C126BE978A4E3AB5C0E3F812F640F3">
<mods:title id="E77ED5B5135A18249B59CFE62AF1D1E5">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="AFA908C98ADEAC9BF07F26804E6283DB">
<mods:date id="A505BC45799534AE81E415945E2A8550">2021</mods:date>
<mods:detail id="A08120966238AB64A0C6DC16F5F94141" type="pubDate">
<mods:number id="6A703ECE7ECF6FD1BEAC1FCD9527B254">2021-08-11</mods:number>
</mods:detail>
<mods:detail id="8AE720CFB134D5017CA00555251F5A96" type="volume">
<mods:number id="F3BEAAF37B91085C10151247448EEF90">5020</mods:number>
</mods:detail>
<mods:detail id="1F138B2761BE67433AE43BBBABD14E96" type="issue">
<mods:number id="0C4A5FC09592E50296FDE56F54611991">1</mods:number>
</mods:detail>
<mods:extent id="FCD37E873982726DA1CE004BCA3070AF" unit="page">
<mods:start id="F14C92C8D62D70AAC66DE916333699A1">31</mods:start>
<mods:end id="AEBC9F56D8AA190AA108228426A60970">56</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="82FECFDD10B16C00D93787FE6282320A">journal article</mods:classification>
<mods:identifier id="1694EAEA2EFEA97F3C2020C4CB93A8F7" type="DOI">10.11646/zootaxa.5020.1.2</mods:identifier>
<mods:identifier id="5982247B35818D57328F4D4EE8F1B8F4" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="FFE9871CA4448A684A7AA30D399C3461" type="Zenodo-Dep">5222931</mods:identifier>
<mods:identifier id="127327C5EF0807B94E8E6CEB57E06AAF" type="ZooBank">882BBB9D-6E5B-4CE5-99DF-E91AC7971EB5</mods:identifier>
</mods:mods>
<treatment id="785C87DD333A002EC8F7FF0EC9B947B3" LSID="urn:lsid:plazi:treatment:785C87DD333A002EC8F7FF0EC9B947B3" httpUri="http://treatment.plazi.org/id/785C87DD333A002EC8F7FF0EC9B947B3" lastPageId="10" lastPageNumber="41" pageId="9" pageNumber="40">
<subSubSection id="B8EF6540333A002DC8F7FF0ECF1F409D" box="[151,521,151,178]" pageId="9" pageNumber="40" type="nomenclature">
<paragraph id="F04A36CB333A002DC8F7FF0ECF1F409D" blockId="9.[151,521,151,214]" box="[151,521,151,178]" pageId="9" pageNumber="40">
<heading id="AB0281A7333A002DC8F7FF0ECF1F409D" bold="true" box="[151,521,151,178]" fontSize="11" level="1" pageId="9" pageNumber="40" reason="1">
<emphasis id="C281EAD9333A002DC8F7FF0ECF1F409D" bold="true" box="[151,521,151,178]" pageId="9" pageNumber="40">
<taxonomicName id="37F54D48333A002DC8F7FF0ECCA9409E" authority="Kataev, 2021" authorityName="Kataev" authorityYear="2021" box="[151,447,151,178]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="pseudoasemus" status="sp. nov.">
<emphasis id="C281EAD9333A002DC8F7FF0ECCA9409E" bold="true" box="[151,447,151,178]" italics="true" pageId="9" pageNumber="40">Harpalus pseudoasemus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D9B257A2333A002DC9A6FF0ECF1F409D" box="[454,521,151,177]" pageId="9" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B8EF6540333A002DC8F7FF22CC4740FA" box="[151,337,187,214]" pageId="9" pageNumber="40" type="description">
<paragraph id="F04A36CB333A002DC8F7FF22CC4740FA" blockId="9.[151,521,151,214]" box="[151,337,187,214]" pageId="9" pageNumber="40">
(
<figureCitation id="68CE2A4E333A002DC8FFFF22CDE140F9" box="[159,247,187,213]" captionStart="FIGURES 20, 21" captionStartId="10.[151,264,1278,1302]" captionTargetBox="[163,1420,193,1250]" captionTargetId="figure-23@10.[152,1435,181,1254]" captionTargetPageId="10" captionText="FIGURES 20, 21. Harpalus spp., habitus. 20, H. pseudoasemus sp. n. (paratype from Naivasha); 21, H. merkli sp. n. (holotype). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5222943" httpUri="https://zenodo.org/record/5222943/files/figure.png" pageId="9" pageNumber="40">Figs 20</figureCitation>
, 22–26)
</paragraph>
</subSubSection>
<subSubSection id="B8EF6540333A002DC8F7FE99CF5B4134" box="[151,589,255,280]" pageId="9" pageNumber="40" type="reference_group">
<paragraph id="F04A36CB333A002DC8F7FE99CF5B4134" blockId="9.[151,589,255,280]" box="[151,589,255,280]" pageId="9" pageNumber="40">
<taxonomicName id="37F54D48333A002DC8F7FE99CF1E413B" ID-CoL="3JSHC" authority=": Basilewsky, 1951" authorityName="Basilewsky" authorityYear="1951" box="[151,520,255,280]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="asemus">
<emphasis id="C281EAD9333A002DC8F7FE99CC5E413B" box="[151,328,256,280]" italics="true" pageId="9" pageNumber="40">Harpalus asemus</emphasis>
:
<bibRefCitation id="94644B3A333A002DC935FE99CF1E413B" author="Basilewsky, P." box="[341,520,255,280]" pageId="9" pageNumber="40" pagination="1 - 333" refId="ref14741" refString="Basilewsky, P. (1951) Revision generale des Harpalinae d'Afrique et de Madagascar (Coleoptera Carabidae). Deuxieme partie. Annales du Musee Royal du Congo Belge, Serie in 8 °, Sciences Zoologiques, 9, 1 - 333, pls. I - VI." type="journal article" year="1951">Basilewsky, 1951</bibRefCitation>
</taxonomicName>
(part).
</paragraph>
</subSubSection>
<subSubSection id="B8EF6540333A002DC8F7FEDFCC114270" pageId="9" pageNumber="40" type="materials_examined">
<paragraph id="F04A36CB333A002DC8F7FEDFCC114270" blockId="9.[151,1437,325,1972]" pageId="9" pageNumber="40">
<emphasis id="C281EAD9333A002DC8F7FEDFCC534173" bold="true" box="[151,325,325,352]" pageId="9" pageNumber="40">Type material.</emphasis>
<materialsCitation id="409D3C96333A002DC92CFEDFCEF14185" collectingDate="1957-04-22" collectionCode="ZIN" collectorName="Coll. Mus. &amp; P. Basilewsy &amp; N. Leleup &amp; P. Basilewsky" country="Tanzania" elevation="1850" location="Oldonyo Dambu" pageId="9" pageNumber="40" specimenCount="1" specimenCount-male="1" stateProvince="Arusha" typeStatus="holotype">
<typeStatus id="2F4E8869333A002DC92CFEDFCCAE414C" box="[332,440,326,352]" pageId="9" pageNumber="40" type="holotype">Holotype</typeStatus>
.
<specimenCount id="E6F3FD42333A002DC9A3FEDFCCCC414C" box="[451,474,326,352]" pageId="9" pageNumber="40" type="male">♂</specimenCount>
,
<emphasis id="C281EAD9333A002DC986FEDCCF454173" bold="true" box="[486,595,325,351]" pageId="9" pageNumber="40">
<collectingCountry id="88E2765B333A002DC986FEDCCF454173" box="[486,595,325,351]" name="Tanzania" pageId="9" pageNumber="40">Tanzania</collectingCountry>
</emphasis>
: “
<collectorName id="5D00531D333A002DCA0DFEDFCFF1414C" box="[621,743,326,352]" pageId="9" pageNumber="40">Coll. Mus.</collectorName>
Congo Tanganyika Terr.:
<location id="F52A6010333A002DCC6BFEDFC9DF414C" LSID="urn:lsid:plazi:treatment:785C87DD333A002EC8F7FF0EC9B947B3:F52A6010333A002DCC6BFEDFC9DF414C" box="[1035,1225,326,352]" country="Tanzania" name="Oldonyo Dambu" pageId="9" pageNumber="40" stateProvince="Arusha">Oldonyo Dambu</location>
,
<quantity id="370D9B2E333A002DCCB5FEDFC80A414C" box="[1237,1308,326,353]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="40" unit="km" value="15.0">15 km</quantity>
N.
<collectingRegion id="3231F829333A002DCD24FEDFC88E414C" box="[1348,1432,326,352]" country="Tanzania" name="Arusha" pageId="9" pageNumber="40">Arusha</collectingRegion>
, cratère,
<quantity id="370D9B2E333A002DC892FEF3CC5141A8" box="[242,327,362,389]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="9" pageNumber="40" unit="m" value="1850.0">
<elevation id="7BD8D1F8333A002DC892FEF3CC5141A8" box="[242,327,362,389]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="9" pageNumber="40" unit="m" value="1850.0">1850 m</elevation>
</quantity>
<date id="844B100B333A002DC92EFEF3CCF641A9" box="[334,480,362,389]" pageId="9" pageNumber="40" value="1957-04-22">
<collectingDate id="940FE9E3333A002DC92EFEF3CCF641A9" box="[334,480,362,389]" pageId="9" pageNumber="40" value="1957-04-22">22–IV–1957</collectingDate>
</date>
”, “Mission Zoolog. I.R.S.A.C. en Afrique orientale (
<collectorName id="5D00531D333A002DCC4FFEF3C9A841A8" box="[1071,1214,362,388]" pageId="9" pageNumber="40">P. Basilewsy</collectorName>
et
<collectorName id="5D00531D333A002DCC80FEF3C84141A9" box="[1248,1367,362,389]" pageId="9" pageNumber="40">N. Leleup</collectorName>
)”, “
<taxonomicName id="37F54D48333A002DCDE0FEF3CCEE4184" authority="Basilew. P. Basilewsky" authorityName="Basilew. P. Basilewsky" authorityYear="1947" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="asemus">H. asemus Basilew. P. Basilewsky</taxonomicName>
det. 19” (
<collectionCode id="96E4AE0E333A002DCA0CFE17CF894184" box="[620,671,398,424]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="9" pageNumber="40" type="Museum">ZIN</collectionCode>
; donation by
<collectorName id="5D00531D333A002DCB5DFE17CEC94185" box="[829,991,398,425]" pageId="9" pageNumber="40">P. Basilewsky</collectorName>
)
</materialsCitation>
.
<materialsCitation id="409D3C96333A002DCB93FE17CC7541E0" collectingDate="1957-04-22" collectionCode="ZIN" collectorName="Coll. Mus. &amp; P. Basilewsy &amp; N. Leleup &amp; P. Basilewsky" country="Tanzania" elevation="1850" location="Oldonyo Dambu" pageId="9" pageNumber="40" specimenCount="1" specimenCount-female="1" stateProvince="Arusha" typeStatus="paratype">
<typeStatus id="2F4E8869333A002DCB93FE17C9724184" box="[1011,1124,398,424]" pageId="9" pageNumber="40" type="paratype">Paratypes</typeStatus>
.
<emphasis id="C281EAD9333A002DCC10FE14C9F1418B" bold="true" box="[1136,1255,397,423]" pageId="9" pageNumber="40">
<collectingCountry id="88E2765B333A002DCC10FE14C9F4418B" box="[1136,1250,397,423]" name="Tanzania" pageId="9" pageNumber="40">Tanzania</collectingCountry>
:
</emphasis>
<specimenCount id="E6F3FD42333A002DCC90FE17C80D4184" box="[1264,1307,398,424]" pageId="9" pageNumber="40" type="female">1 ♀</specimenCount>
, same data as holotype (
<collectionCode id="96E4AE0E333A002DC947FE2BCC4D41E0" box="[295,347,434,460]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="9" pageNumber="40" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="409D3C96333A002DC90FFE2BC9F541DC" collectingDate="1981-03-24" collectingDateMax="1982-03-06" collectingDateMin="1981-03-24" collectionCode="MFNB, ZIN" collectorName="H. J. Bremer" country="Tanzania" county="Arusha" elevation="1200" location="Ngare Sero" municipality="Usa river" pageId="9" pageNumber="40" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Arusha" typeStatus="paratype">
<specimenCount id="E6F3FD42333A002DC90FFE2BCC8A41E0" box="[367,412,434,460]" pageId="9" pageNumber="40" type="male">1 ♂</specimenCount>
, “
<collectingCountry id="88E2765B333A002DC9D1FE2BCF0F41E0" box="[433,537,434,460]" name="Tanzania" pageId="9" pageNumber="40">Tanzania</collectingCountry>
sept.
<location id="F52A6010333A002DCA3BFE2BCFCB41E0" LSID="urn:lsid:plazi:treatment:785C87DD333A002EC8F7FF0EC9B947B3:F52A6010333A002DCA3BFE2BCFCB41E0" box="[603,733,434,460]" country="Tanzania" county="Arusha" municipality="Usa river" name="Ngare Sero" pageId="9" pageNumber="40">Ngare Sero</location>
,
<quantity id="370D9B2E333A002DCA89FE2BCE2841E0" box="[745,830,434,460]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="40" unit="m" value="1200.0">
<elevation id="7BD8D1F8333A002DCA89FE2BCE2841E0" box="[745,830,434,460]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="40" unit="m" value="1200.0">1200 m</elevation>
</quantity>
near
<collectingMunicipality id="102EACB1333A002DCB1CFE2BCEF241E0" box="[892,996,434,460]" pageId="9" pageNumber="40">Usa river</collectingMunicipality>
<quantity id="370D9B2E333A002DCB8BFE2BC92541E0" box="[1003,1075,434,460]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="40" unit="km" value="20.0">20 km</quantity>
east of
<collectingCounty id="192B4E47333A002DCCE9FE2BC9CC41E0" box="[1161,1242,434,460]" pageId="9" pageNumber="40">Arusha</collectingCounty>
<date id="844B100B333A002DCC82FE2BC88B41E0" box="[1250,1437,434,460]" pageId="9" pageNumber="40" value="1982-02-17" valueMax="1982-03-06" valueMin="1982-02-17">
<collectingDate id="940FE9E3333A002DCC82FE2BC88B41E0" box="[1250,1437,434,460]" pageId="9" pageNumber="40" value="1982-02-17" valueMax="1982-03-06" valueMin="1982-02-17">17.II.–6.III.1982</collectingDate>
</date>
<collectorName id="5D00531D333A002DC8F7FE4FCC3741DC" box="[151,289,470,496]" pageId="9" pageNumber="40">H.J. Bremer</collectorName>
leg. (
<collectionCode id="96E4AE0E333A002DC903FE4FCCAF41DC" box="[355,441,470,496]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="40">MFNB</collectionCode>
);
<specimenCount id="E6F3FD42333A002DC9AEFE4FCF1941DC" box="[462,527,470,496]" pageId="9" pageNumber="40" type="female">3 ♀♀</specimenCount>
, same data but also “sub lapide
<date id="844B100B333A002DCBE3FE4FC93B41DC" box="[899,1069,470,496]" pageId="9" pageNumber="40" value="1981-03-24" valueMax="1981-03-31" valueMin="1981-03-24">
<collectingDate id="940FE9E3333A002DCBE3FE4FC93B41DC" box="[899,1069,470,496]" pageId="9" pageNumber="40" value="1981-03-24" valueMax="1981-03-31" valueMin="1981-03-24">24–31.III.1981</collectingDate>
</date>
” (
<collectionCode id="96E4AE0E333A002DCC27FE4FC98D41DC" box="[1095,1179,470,496]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="40">MFNB</collectionCode>
,
<collectionCode id="96E4AE0E333A002DCCC8FE4FC9CD41DC" box="[1192,1243,470,496]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="9" pageNumber="40" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="409D3C96333A002DCC90FE4FCF7D4238" collectingDate="1981-03-24" collectingDateMax="1982-03-06" collectingDateMin="1981-03-24" collectionCode="MFNB" collectorName="H. J. Bremer" country="Tanzania" county="Arusha" elevation="1200" location="Gebusch" municipality="Usa river" pageId="9" pageNumber="40" specimenCount="1" specimenCount-female="1" stateProvince="Arusha" typeStatus="paratype">
<specimenCount id="E6F3FD42333A002DCC90FE4FC80A41DC" box="[1264,1308,470,496]" pageId="9" pageNumber="40" type="female">1 ♀</specimenCount>
, same data but also “aus
<location id="F52A6010333A002DC94DFE63CC994238" LSID="urn:lsid:plazi:treatment:785C87DD333A002EC8F7FF0EC9B947B3:F52A6010333A002DC94DFE63CC994238" box="[301,399,506,532]" country="Tanzania" name="Gebusch" pageId="9" pageNumber="40">Gebüsch</location>
geklopft” (
<collectionCode id="96E4AE0E333A002DCA6DFE63CF754238" box="[525,611,506,532]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="40">MFNB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="409D3C96333A002DCA15FE63CC154270" collectionCode="MNHN" collectorName="P. - A. Chappuis" country="Kenya" county="Jeannel" determinerName="P. Basilewsky" elevation="2" location="Naivasha" municipality="Arambourg" pageId="9" pageNumber="40" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="C281EAD9333A002DCA15FE63CFDA4238" bold="true" box="[629,716,506,532]" pageId="9" pageNumber="40">
<collectingCountry id="88E2765B333A002DCA15FE63CFD14238" box="[629,711,506,532]" name="Kenya" pageId="9" pageNumber="40">Kenya</collectingCountry>
:
</emphasis>
<specimenCount id="E6F3FD42333A002DCAB2FE63CFE84238" box="[722,766,506,532]" pageId="9" pageNumber="40" type="male">1 ♂</specimenCount>
, “
<collectingCountry id="88E2765B333A002DCB73FE63CE494238" box="[787,863,506,532]" name="Kenya" pageId="9" pageNumber="40">Kenya</collectingCountry>
<location id="F52A6010333A002DCB05FE63CEC64238" LSID="urn:lsid:plazi:treatment:785C87DD333A002EC8F7FF0EC9B947B3:F52A6010333A002DCB05FE63CEC64238" box="[869,976,506,532]" country="Kenya" county="Jeannel" municipality="Arambourg" name="Naivasha" pageId="9" pageNumber="40">Naivasha</location>
,
<location id="F52A6010333A002DCBBAFE63C9424238" LSID="urn:lsid:plazi:treatment:785C87DD333A002EC8F7FF0EC9B947B3:F52A6010333A002DCBBAFE63C9424238" box="[986,1108,506,532]" country="Kenya" county="Jeannel" municipality="Arambourg" name="Rift Valley" pageId="9" pageNumber="40">Rift Valley</location>
<quantity id="370D9B2E333A002DCC3AFE63C9AF4238" box="[1114,1209,506,532]" metricMagnitude="0" metricUnit="m" metricValue="1.9" pageId="9" pageNumber="40" unit="m" value="1.9">
<elevation id="7BD8D1F8333A002DCC3AFE63C9AF4238" box="[1114,1209,506,532]" metricMagnitude="0" metricUnit="m" metricValue="1.9" pageId="9" pageNumber="40" unit="m" value="1.9">1.900 m</elevation>
</quantity>
”, “
<location id="F52A6010333A002DCCB8FE63CC744214" LSID="urn:lsid:plazi:treatment:785C87DD333A002EC8F7FF0EC9B947B3:F52A6010333A002DCCB8FE63CC744214" country="Kenya" county="Jeannel" municipality="Arambourg" name="Museum de Paris Mission de l'Omo" pageId="9" pageNumber="40">Muséum de Paris Mission de l’Omo</location>
C.
<collectingMunicipality id="102EACB1333A002DC9E7FD87CF1E4214" box="[391,520,542,568]" pageId="9" pageNumber="40">Arambourg</collectingMunicipality>
<collectorName id="5D00531D333A002DCA6EFD87CFA14214" box="[526,695,542,568]" pageId="9" pageNumber="40">P.-A. Chappuis</collectorName>
&amp; R.
<collectingCounty id="192B4E47333A002DCA9AFD87CE584214" box="[762,846,542,568]" pageId="9" pageNumber="40">Jeannel</collectingCounty>
1932–33”, “
<taxonomicName id="37F54D48333A002DCBBAFD87C98C4214" authorityName="Basilew. P. Basilewsky" authorityYear="1947" box="[986,1178,542,568]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="asemus">Harpalus asemus</taxonomicName>
m.
<determinerName id="0596DA0F333A002DCCA5FD87C8764214" box="[1221,1376,542,568]" pageId="9" pageNumber="40">P. Basilewsky</determinerName>
det.” (
<collectionCode id="96E4AE0E333A002DC8FFFDDBCDED4270" box="[159,251,578,604]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="40">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B8EF6540333A002EC8A7FDFCCF3E4677" lastPageId="10" lastPageNumber="41" pageId="9" pageNumber="40" type="description">
<paragraph id="F04A36CB333A002DC8A7FDFCCCB5433C" blockId="9.[151,1437,325,1972]" pageId="9" pageNumber="40">
<emphasis id="C281EAD9333A002DC8A7FDFCCC474253" bold="true" box="[199,337,613,639]" pageId="9" pageNumber="40">Description</emphasis>
(
<specimenCount id="E6F3FD42333A002DC901FDFFCC8942AC" box="[353,415,614,640]" pageId="9" pageNumber="40" type="male">3 ♂♂</specimenCount>
and
<specimenCount id="E6F3FD42333A002DC9B4FDFFCF0542AC" box="[468,531,614,641]" pageId="9" pageNumber="40" type="female">5 ♀♀</specimenCount>
measured). Body length
<quantity id="370D9B2E333A002DCB4AFDFFCEA542AC" box="[810,947,614,641]" metricMagnitude="-3" metricUnit="m" metricValue="5.55" metricValueMax="5.8" metricValueMin="5.3" pageId="9" pageNumber="40" unit="mm" value="5.55" valueMax="5.8" valueMin="5.3">5.3–5.8 mm</quantity>
. Habitus as in
<figureCitation id="68CE2A4E333A002DCC34FDFFC9B042AC" box="[1108,1190,614,640]" captionStart="FIGURES 20, 21" captionStartId="10.[151,264,1278,1302]" captionTargetBox="[163,1420,193,1250]" captionTargetId="figure-23@10.[152,1435,181,1254]" captionTargetPageId="10" captionText="FIGURES 20, 21. Harpalus spp., habitus. 20, H. pseudoasemus sp. n. (paratype from Naivasha); 21, H. merkli sp. n. (holotype). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5222943" httpUri="https://zenodo.org/record/5222943/files/figure.png" pageId="9" pageNumber="40">Fig. 20</figureCitation>
. Dorsum black, shiny, occasionally with light greenish or copper luster; ventral side black; palpomeres at least apically and two (occasionally one) basal antennomeres brownish yellow; palpomeres partly and antennomeres 3–11 or 2–11 distinctly infuscate, blackish brown. Legs brown, with femora, apical portion of tibiae and often tarsi more or less infuscate, usually blackish brown.
</paragraph>
<paragraph id="F04A36CB333A002DC8A7FC83C89743E8" blockId="9.[151,1437,325,1972]" pageId="9" pageNumber="40">
Head moderately sized (HWmax/PWmax 0.64–0.70, HWmin/PWmax 0.53–0.56), impunctate, with large and convex eyes (HWmax/HWmin 1.18–1.28). Tempora oblique, short and flat. Frontal foveae punctiform. Supraorbital seta situated near supraorbital furrow at level of posterior margin of eye or slightly before it. Genae about as wide as antennomere 1. Labrum almost straight or with very shallow emargination anteriorly. Labium similar to that of
<taxonomicName id="37F54D48333A002DCDE1FC1ECC1543E8" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="impressus">
<emphasis id="C281EAD9333A002DCDE1FC1ECC1543E8" italics="true" pageId="9" pageNumber="40">H. impressus</emphasis>
</taxonomicName>
; paraglossae each with one marginal seta. Dorsal microsculpture fine, consisting of isodiametric meshes.
</paragraph>
<paragraph id="F04A36CB333A002DC8A7FC57CC6E4500" blockId="9.[151,1437,325,1972]" pageId="9" pageNumber="40">Pronotum transverse (PWmax/PL 1.36–1.44), almost equally narrowed apically and basally, widest slightly before middle (PWmax/PWmin 1.14–1.21). Sides in apical half evenly rounded, in basal half more widely rounded or almost straight. One pair of lateral setae in widest point of pronotum. Apical margin shallowly emarginate, almost straight or arcuate in middle portion, bordered only laterally. Apical angles rounded, slightly prominent. Basal angles obtuse, blunted or narrowly rounded at apices. Basal edge completely bordered, glabrous, more or less straight in middle portion, oblique or very widely rounded laterally, slightly longer than apical margin and slightly shorter than elytral base. Surface convex, without lateral depressions, with basal foveae small, elongate, shallow, very finely and sparsely punctate; several very fine punctures occasionally also present just at basal angles. Microsculpture fine, consisting of distinct transverse meshes on disc and of more or less isodiametric meshes within small area at basal angles.</paragraph>
<paragraph id="F04A36CB333A002DC8A7FAAFCE9446B8" blockId="9.[151,1437,325,1972]" pageId="9" pageNumber="40">Elytra comparatively wide (in male, EL/EW 1.41–1.45, EL/PL 2.33–2.46, EW/PWmax 1.16–1.23; in female, these indices 1.39–1.45, 2.38–2.57, and 1.19–1.30, respectively), widest about the middle; sides almost rectilinearly diverging in basal half and rounded in apical half; preapical sinuation very shallow. Shoulders slightly prominent, angularly rounded, each with very small and indistinct denticle recognizable from behind. Basal border glabrous, forming a very obtuse angle with lateral elytral margin. Sutural angle not projecting posteriorly, slightly less than 90°, blunted at tip. Striae impunctate, superficial almost along entire length, only slightly impressed apically. Para- scutellar setigerous pore present. Parascutellar (abbreviate) striole more or less strongly reduced. Intervals impunctate and glabrous, flat on disc, only slightly convex at apex. Interval 3 with or without discal setigerous pore; intervals 5 and 7 without discal pores. Marginal umbilicate series more or less widely interrupted at middle. Microsculpture very distinct, consisting of moderately transverse meshes.</paragraph>
<paragraph id="F04A36CB333A002DC8A7F907CF114694" blockId="9.[151,1437,325,1972]" box="[199,519,1694,1720]" pageId="9" pageNumber="40">Hind wings fully developed.</paragraph>
<paragraph id="F04A36CB333A002DC8A7F95BCF70472C" blockId="9.[151,1437,325,1972]" pageId="9" pageNumber="40">Prosternum with very fine and short setae along anterior margin and laterally. Metepisternum noticeably longer than wide, strongly narrowed posteriorly.</paragraph>
<paragraph id="F04A36CB333A002DC8A7F893CE574764" blockId="9.[151,1437,325,1972]" pageId="9" pageNumber="40">
Metacoxa, in addition to the two obligatory setigerous pores, with a posteromedial setigerous pore (Fig. 22). Femora, tibiae and tarsi as described above for
<taxonomicName id="37F54D48333A002DCAC9F8B6CE2C4764" box="[681,826,1839,1864]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="impressus">
<emphasis id="C281EAD9333A002DCAC9F8B6CE2C4764" box="[681,826,1839,1864]" italics="true" pageId="9" pageNumber="40">H. impressus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="F04A36CB333A002DC8A7F8CBCC574798" blockId="9.[151,1437,325,1972]" pageId="9" pageNumber="40">Abdominal sternites without additional long setae, only medial portions of sternites III and IV finely pubescent. Last visible abdominal sternite (VII) in male subtruncate, in female rounded at apex, in both sexes with two pairs of marginal setae.</paragraph>
<caption id="A48A66433339002EC8F7FB67CCB34516" ID-DOI="http://doi.org/10.5281/zenodo.5222943" ID-Zenodo-Dep="5222943" httpUri="https://zenodo.org/record/5222943/files/figure.png" pageId="10" pageNumber="41" startId="10.[151,264,1278,1302]" targetBox="[163,1420,193,1250]" targetPageId="10">
<paragraph id="F04A36CB3339002EC8F7FB67CCB34516" blockId="10.[151,1436,1278,1339]" pageId="10" pageNumber="41">
<emphasis id="C281EAD93339002EC8F7FB67CC40453A" bold="true" box="[151,342,1278,1302]" pageId="10" pageNumber="41">FIGURES 20, 21.</emphasis>
<taxonomicName id="37F54D483339002EC93EFB67CCE7453A" box="[350,497,1278,1302]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="C281EAD93339002EC93EFB67CCA8453A" box="[350,446,1278,1302]" italics="true" pageId="10" pageNumber="41">Harpalus</emphasis>
spp.
</taxonomicName>
, habitus. 20,
<taxonomicName id="37F54D483339002ECA1BFB66CE3B453A" authority="Kataev, 2021" authorityName="Kataev" authorityYear="2021" box="[635,813,1278,1302]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="pseudoasemus" status="sp. nov.">
<emphasis id="C281EAD93339002ECA1BFB66CE3B453A" box="[635,813,1278,1302]" italics="true" pageId="10" pageNumber="41">H. pseudoasemus</emphasis>
</taxonomicName>
<emphasis id="C281EAD93339002ECB55FB66CE66453B" bold="true" box="[821,880,1279,1303]" pageId="10" pageNumber="41">
<taxonomicNameLabel id="D9B257A23339002ECB55FB66CE66453B" box="[821,880,1279,1303]" pageId="10" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratype from Naivasha); 21,
<taxonomicName id="37F54D483339002ECCD4FB66C803453A" authority="Kataev, 2021" authorityName="Kataev" authorityYear="2021" box="[1204,1301,1278,1302]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="merkli" status="sp. nov.">
<emphasis id="C281EAD93339002ECCD4FB66C803453A" box="[1204,1301,1278,1302]" italics="true" pageId="10" pageNumber="41">H. merkli</emphasis>
</taxonomicName>
<emphasis id="C281EAD93339002ECD7CFB66C841453B" bold="true" box="[1308,1367,1279,1303]" pageId="10" pageNumber="41">
<taxonomicNameLabel id="D9B257A23339002ECD7CFB66C841453B" box="[1308,1367,1279,1303]" pageId="10" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(holotype). Scale bar = 1.0 mm.
</paragraph>
</caption>
<paragraph id="F04A36CB3339002EC8A7FAF0CC1A458B" blockId="10.[151,1437,1384,1951]" pageId="10" pageNumber="41">Female genitalia (Figs 25, 26): gonocoxite moderately wide, with long setae on dorsal edge and in scrobe of outer side.</paragraph>
<paragraph id="F04A36CB3339002EC8A7FA28CF3E4677" blockId="10.[151,1437,1384,1951]" pageId="10" pageNumber="41">Median lobe of aedeagus (Figs 23, 24) bent ventrally about the middle, with apical portion rather strongly curved to the right; ventral margin in lateral view markedly convex in apical half. Terminal lamella in dorsal view slightly longer than wide, slightly narrowed posteriorly and rounded apically, in lateral aspect its apex protruding ventrally. Apical orifice shifted to the left. Internal sac with two spines in apical portion of median lobe: distal spine larger and wider than proximal one.</paragraph>
</subSubSection>
<subSubSection id="B8EF65403339002EC8A7F9FCC9BB4652" box="[199,1197,1637,1663]" pageId="10" pageNumber="41" type="etymology">
<paragraph id="F04A36CB3339002EC8A7F9FCC9BB4652" blockId="10.[151,1437,1384,1951]" box="[199,1197,1637,1663]" pageId="10" pageNumber="41">
<emphasis id="C281EAD93339002EC8A7F9FCCC5E4653" bold="true" box="[199,328,1637,1663]" pageId="10" pageNumber="41">Etymology</emphasis>
. The species name refers to the similarity of the new species with
<emphasis id="C281EAD93339002ECC51F9FCC9BB4652" box="[1073,1197,1637,1662]" italics="true" pageId="10" pageNumber="41">
<taxonomicName id="37F54D483339002ECC51F9FCC9BF4652" authorityName="Basilew. P. Basilewsky" authorityYear="1947" box="[1073,1193,1637,1662]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="asemus">H. asemus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="B8EF65403339002EC8A7F911C9B947B3" pageId="10" pageNumber="41" type="distribution">
<paragraph id="F04A36CB3339002EC8A7F911CE0046EB" blockId="10.[151,1437,1384,1951]" pageId="10" pageNumber="41">
<emphasis id="C281EAD93339002EC8A7F911CC77468E" bold="true" box="[199,353,1672,1698]" pageId="10" pageNumber="41">Distribution.</emphasis>
Known from
<collectingCountry id="88E2765B3339002EC999F910CF48468F" box="[505,606,1673,1699]" name="Tanzania" pageId="10" pageNumber="41">Tanzania</collectingCountry>
and
<collectingCountry id="88E2765B3339002ECAF1F910CFC8468F" box="[657,734,1673,1699]" name="Kenya" pageId="10" pageNumber="41">Kenya</collectingCountry>
.At least some of the published records of
<taxonomicName id="37F54D483339002ECCCCF910C809468E" authorityName="Basilew. P. Basilewsky" authorityYear="1947" box="[1196,1311,1673,1698]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="asemus">
<emphasis id="C281EAD93339002ECCCCF910C809468E" box="[1196,1311,1673,1698]" italics="true" pageId="10" pageNumber="41">H. asemus</emphasis>
</taxonomicName>
(
<bibRefCitation id="94644B3A3339002ECD4CF910CDE246EB" author="Basilewsky, P." pageId="10" pageNumber="41" pagination="1 - 333" refId="ref14741" refString="Basilewsky, P. (1951) Revision generale des Harpalinae d'Afrique et de Madagascar (Coleoptera Carabidae). Deuxieme partie. Annales du Musee Royal du Congo Belge, Serie in 8 °, Sciences Zoologiques, 9, 1 - 333, pls. I - VI." type="journal article" year="1951">Basilewsky 1951</bibRefCitation>
,
<bibRefCitation id="94644B3A3339002EC960F934CC2F46E4" author="Basilewsky, P." box="[256,313,1709,1736]" pageId="10" pageNumber="41" pagination="48 - 337" refId="ref14947" refString="Basilewsky, P. (1962) Mission zoologique de l`I. R. S. A. C. en Afrique orientale (P. Basilewsky et N. Leleup, 1957). LX: Coleop- tera Carabidae. Annales du Musee royal de l'Afrique centrale, Serie in- 8 °, Sciences Zoologiques, 107, 48 - 337." type="journal article" year="1962">1962</bibRefCitation>
) are apparently based on this new species.
</paragraph>
<paragraph id="F04A36CB3339002EC8A7F949C9B947B3" blockId="10.[151,1437,1384,1951]" pageId="10" pageNumber="41">
<emphasis id="C281EAD93339002EC8A7F949CC7446C6" bold="true" box="[199,354,1744,1770]" pageId="10" pageNumber="41">Comparison.</emphasis>
This new species is very similar in habitus to
<taxonomicName id="37F54D483339002ECB0AF948CEC946C6" authorityName="Basilew. P. Basilewsky" authorityYear="1947" box="[874,991,1745,1770]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="asemus">
<emphasis id="C281EAD93339002ECB0AF948CEC946C6" box="[874,991,1745,1770]" italics="true" pageId="10" pageNumber="41">H. asemus</emphasis>
</taxonomicName>
, but, on average, smaller and stouter; it easily differs from the latter, as well as from
<taxonomicName id="37F54D483339002ECAE4F96CCE024722" box="[644,788,1781,1806]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="impressus">
<emphasis id="C281EAD93339002ECAE4F96CCE024722" box="[644,788,1781,1806]" italics="true" pageId="10" pageNumber="41">H. impressus</emphasis>
</taxonomicName>
, in having elytral microsculpture consisting of moderately transverse meshes, metacoxa with posteromedial pore and last visible abdominal sternite (VII) of male subtruncate. In addition, unlike
<taxonomicName id="37F54D483339002EC9C8F8A4CF0A477A" authorityName="Basilew. P. Basilewsky" authorityYear="1947" box="[424,540,1853,1878]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="asemus">
<emphasis id="C281EAD93339002EC9C8F8A4CF0A477A" box="[424,540,1853,1878]" italics="true" pageId="10" pageNumber="41">H. asemus</emphasis>
</taxonomicName>
, median lobe of aedeagus of
<taxonomicName id="37F54D483339002ECB07F8A4C93D477B" authority="Kataev, 2021" authorityName="Kataev" authorityYear="2021" box="[871,1067,1853,1879]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="pseudoasemus" status="sp. nov.">
<emphasis id="C281EAD93339002ECB07F8A4C93D477B" box="[871,1067,1853,1879]" italics="true" pageId="10" pageNumber="41">H. pseudoasemus</emphasis>
</taxonomicName>
<emphasis id="C281EAD93339002ECC52F8A4C964477B" bold="true" box="[1074,1138,1853,1879]" pageId="10" pageNumber="41">
<taxonomicNameLabel id="D9B257A23339002ECC52F8A4C964477B" box="[1074,1138,1853,1879]" pageId="10" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is bent ventrally about the middle, with apical portion more strongly curved to the right and with ventral margin in lateral view markedly more convex in apical half; its internal sac has two spines situated in apical portion of median lobe.
</paragraph>
</subSubSection>
</treatment>
</document>
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