240 lines
32 KiB
XML
240 lines
32 KiB
XML
<document id="2B26D5BB5FBEC0F34A9488EEDC60A786" ID-DOI="http://dx.doi.org/10.3897/zookeys.542.5975" ID-GBIF-Dataset="b53aa92a-b9ac-43bd-a9b9-f468d62f0bdc" ID-PMC="PMC4714360" ID-Pensoft-Pub="1313-2970-542-1" ID-PubMed="26798285" ID-ZooBank="026407877355425BAB10BF1674510F12" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-542-1" ModsDocOrigin="ZooKeys 542" ModsDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" checkinTime="1451243652205" checkinUser="pensoft" docAuthor="Bistroem, Olof, Nilsson, Anders N. & Bergsten, Johannes" docDate="2015" docId="78541A217044B285DD8E3FEDAA60031A" docLanguage="en" docName="ZooKeys 542: 1-379" docOrigin="ZooKeys 542" docSource="http://dx.doi.org/10.3897/zookeys.542.5975" docTitle="Laccophilus pictipennis Sharp 1882" docType="treatment" docVersion="5" lastPageNumber="45" masterDocId="E3118410B56DFF839B32FF89FFCCFFB0" masterDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" masterLastPageNumber="379" masterPageNumber="1" pageNumber="44" updateTime="1701314941532" updateUser="plazi">
|
|
<mods:mods id="6381A5E4537FF3B1DF6B2C499F81C28F" xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo id="14789B4D83C42E4836CCD90B60CA9E0E">
|
|
<mods:title id="750A9D686D64240970CD3CEAB6965DE4">Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name id="C46A9FB20EA27A55FA3A3A8ADADFFAB5" type="personal">
|
|
<mods:role id="635AF32FB550253DC48699001D1457D3">
|
|
<mods:roleTerm id="43B99F10DDAD0D8750ADD7D373484FAE">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="21B053FFEFABE7BA857329AFD27C3E07">Bistroem, Olof</mods:namePart>
|
|
</mods:name>
|
|
<mods:name id="3621B4E9DEB8BCFF09F0F322504A7C6E" type="personal">
|
|
<mods:role id="966D7F416E3927FD6569AE1D72379AAB">
|
|
<mods:roleTerm id="419E9418233FB10175D1BE1B795EDA30">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="EF7B027CBA3B060EE683941BEB4A98A1">Nilsson, Anders N.</mods:namePart>
|
|
</mods:name>
|
|
<mods:name id="123AFB064DB004B630FB7CD31331E31B" type="personal">
|
|
<mods:role id="1FB403803926ACCA59B996CAB8DD60CC">
|
|
<mods:roleTerm id="2359D90F29DD1C450B86B946D50020F3">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="CE552E47BD08A607EBB9882D9CB29D6F">Bergsten, Johannes</mods:namePart>
|
|
</mods:name>
|
|
<mods:typeOfResource id="B78CDA1B818EEE3B02568C4ED8637061">text</mods:typeOfResource>
|
|
<mods:relatedItem id="41721B893190B886D903EAB61C7BD09C" type="host">
|
|
<mods:titleInfo id="A7E24D6F7E3DBD4D843A41002470E5CF">
|
|
<mods:title id="0C6BC502D6B46CBF22463FC143146ABC">ZooKeys</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part id="F3C89CAE28EEFF687BCBD7869550E0D9">
|
|
<mods:date id="8B6CDC74BA01957B5A0A300466A6CE2A">2015</mods:date>
|
|
<mods:detail id="3A62C80DD92F6F19D7142C6901251B4C" type="volume">
|
|
<mods:number id="74E61A525F8E79446496F2F8F4A4ACDD">542</mods:number>
|
|
</mods:detail>
|
|
<mods:extent id="866E9CC1A441844685FFC954BB6BC0C7" unit="page">
|
|
<mods:start id="0A18D925CB9279AF431D9A3084B86D2D">1</mods:start>
|
|
<mods:end id="15AA9A9D1D5BD143287FA442D7A8DBDE">379</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location id="AF9692A85D9B5570432D25A6B38982A7">
|
|
<mods:url id="04BDBDE1F818012BE5875227CC8EA5DD">http://dx.doi.org/10.3897/zookeys.542.5975</mods:url>
|
|
</mods:location>
|
|
<mods:classification id="F6D213902E7162462B6685FE866FE7F8">journal article</mods:classification>
|
|
<mods:identifier id="08A4A2445E0774A023BE720F5879861D" type="DOI">http://dx.doi.org/10.3897/zookeys.542.5975</mods:identifier>
|
|
<mods:identifier id="61AC7B0FE1945BC23E70F1B9590226EA" type="Pensoft-Pub">1313-2970-542-1</mods:identifier>
|
|
<mods:identifier id="9EA4B42A085FB9D01CFB55A9C857CDEB" type="ZooBank">026407877355425BAB10BF1674510F12</mods:identifier>
|
|
</mods:mods>
|
|
<treatment id="78541A217044B285DD8E3FEDAA60031A" ID-GBIF-Taxon="127862740" LSID="urn:lsid:plazi:treatment:78541A217044B285DD8E3FEDAA60031A" httpUri="http://treatment.plazi.org/id/78541A217044B285DD8E3FEDAA60031A" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44">
|
|
<subSubSection id="92A3FCEFF8E4CAC2ABE8304F6C4EF7FB" pageId="43" pageNumber="44" type="multiple">
|
|
<paragraph id="1DEABF5AFE9E70E2014C3BB5A051AAA0" pageId="43" pageNumber="44">
|
|
<pageBreakToken id="8D4D5AFB94C1C677B0B38F208745EE6E" pageId="43" pageNumber="44" start="start">Taxon</pageBreakToken>
|
|
classification Animalia Coleoptera Dytiscidae
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="AE2BEE0CD3AD3EBF06321E5675ED446F" pageId="43" pageNumber="44" type="nomenclature">
|
|
<paragraph id="462A2E05F616FCBA09330FBF6EFA4C41" pageId="43" pageNumber="44">
|
|
<taxonomicName id="A3DE7B33824E3138AB26438E0B76E62D" ID-CoL="6NRLS" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pictipennis" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="pictipennis">Laccophilus pictipennis Sharp, 1882</taxonomicName>
|
|
Figs 45-46, 247-248, 398-399, 534
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="F5673BECD895F762A52959418FDB0A20" pageId="44" pageNumber="45" type="reference_group">
|
|
<paragraph id="59D5D98700BBA6A6DE2B899D7C0460C9" pageId="44" pageNumber="45">
|
|
<taxonomicName id="9D526DE949E5072E317F4F4CB4A368F1" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pictipennis" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="pictipennis">
|
|
<pageBreakToken id="473B93476A8A2A48C28B526FFF19C49C" pageId="44" pageNumber="45" start="start">Laccophilus</pageBreakToken>
|
|
pictipennis
|
|
</taxonomicName>
|
|
<bibRefCitation id="A3C9AFE7662C4A1904296A93A574D35B" author="Sharp, D" journalOrPublisher="The Scientific Transactions of the Royal Dublin Society" pageId="263" pageNumber="264" pagination="179 - 1003" title="On aquatic carnivorous Coleoptera or Dytiscidae." volume="2" year="1882">Sharp 1882</bibRefCitation>
|
|
: 305 (original description, faunistics); v. d.
|
|
<bibRefCitation id="8ED652698D9675B406F4201B33D29505" author="Branden, C van den" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="255" pageNumber="256" pagination="1 - 118" title="Catalogue des coleopteres carnassiers aquatiques (Haliplidae, Amphizoidae, Pelobiidae et Dytiscidae)." volume="28" year="1885">Branden 1885</bibRefCitation>
|
|
: 23 (catalogue, faunistics);
|
|
<bibRefCitation id="EA18030F506895F5FA834D074E9F1873" author="Regimbart, M" journalOrPublisher="Memoires de la Societe entomologique de Belgique" pageId="262" pageNumber="263" pagination="1 - 244" title="Revision des Dytiscidae et Gyrinidae d'Afrique, Madagascar et iles voisines. En contribution a la faune entomologique du Congo." volume="4" year="1895">
|
|
<normalizedToken id="CB975BFB3A3348E4D28768F517D226A2" originalValue="Régimbart">Regimbart</normalizedToken>
|
|
1895
|
|
</bibRefCitation>
|
|
: 131, 132 (description, faunistics, discussion);
|
|
<bibRefCitation id="23586483436D411CC675B513DC23B387" author="Zimmermann, A" journalOrPublisher="Junk, Berlin" pageId="263" pageNumber="264" title="Dytiscidae, Haliplidae, Hygrobiidae, Amphizoidae. In: Schenkling S (Ed.) Coleopterorum Catalogus. Vol. 4 pars 71. W." year="1920 a">Zimmermann 1920a</bibRefCitation>
|
|
: 24 (catalogue);
|
|
<bibRefCitation id="00F4343667AF728758879885165696E0" author="Zimmermann, A" journalOrPublisher="Troppau" pageId="263" pageNumber="264" pagination="35 - 118" title="Monographie der palaearktischen Dytiscidae. I. Noterinae, Laccophilinae, Hydroporinae (1. Teil)." volumeTitle="Koleopterologische Rundschau" year="1930">Zimmermann 1930</bibRefCitation>
|
|
: 21, 23 (description, faunistics);
|
|
<bibRefCitation id="912F90396CF6B00B20061B4BB230AE04" author="Balfour-Browne, J" journalOrPublisher="16. Birtish Museum (Natural History) Expedition to South-West Arabia 1937 - 1938" pageId="252" pageNumber="253" pagination="179 - 220" title="Coleoptera: Haliplidae, Dytiscidae, Gyrinidae, Hydraenidae, Hydrophilidae." volume="1" year="1951">Balfour-Browne 1951</bibRefCitation>
|
|
: 193 (discussion, faunistics);
|
|
<bibRefCitation id="CD13ACD0CD1866359B63DE3DA63C2100" author="Guignot, F" journalOrPublisher="Bulletin Institut royal des Sciences Naturelles de Belgique" pageId="257" pageNumber="258" pagination="1 - 7" title="Description de nouveaux dytiscides africains." volume="28" year="1952 d">Guignot 1952d</bibRefCitation>
|
|
: 4, 5 (discussion);
|
|
<bibRefCitation id="FBEA4EB79D1F2082A4D3BFA20D8BBC7C" author="Guignot, F" journalOrPublisher="Annales du Musee Royal du Congo Belge (Serie 8 vo) Sciences Zoologiques" pageId="258" pageNumber="259" pagination="323 - 648" title="Revision des hydrocanthares d'Afrique (ColeopteraDytiscoidea). 2." volume="78" year="1959 a">Guignot 1959a</bibRefCitation>
|
|
: 533, 536 description, faunistics);
|
|
<bibRefCitation id="244C9F16314182906B01F7DFE3F7C53E" author="Brancucci, M" journalOrPublisher="Fauna of Saudi Arabia" pageId="255" pageNumber="256" pagination="156 - 161" title="Insects of Saudi Arabia Coleoptera: Fam. Haliplidae, Dytiscidae, Gyrinidae." volume="1" year="1979">Brancucci 1979</bibRefCitation>
|
|
: 159 (faunistics, description, discussion);
|
|
<bibRefCitation id="17BA2E34F0AC273D42DF075C98B8FC2C" author="Brancucci, M" journalOrPublisher="Entomologische Arbeiten aus dem Museum G. Frey" pageId="255" pageNumber="256" pagination="241 - 426" title="Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae)." volume="31 / 32" year="1983 b">Brancucci 1983b</bibRefCitation>
|
|
:274, 394, 416 (description, faunistics; lectotype designation);
|
|
<bibRefCitation id="E5D639D0F628CABBBF7D2AA3B44DD7B7" author="Rocchi, S" journalOrPublisher="Fauna of Saudi Arabia" pageId="262" pageNumber="263" pagination="444 - 450" title="Insects of the Yemen Arab Republic. Coleoptera: Fam. Haliplidae, Dytiscidae, Gyrinidae." volume="6" year="1984">Rocchi 1984</bibRefCitation>
|
|
: 447 (faunistics.);
|
|
<bibRefCitation id="C4298C3C5A8A649FBCFE49AB0EC237BE" author="Nilsson, AN" journalOrPublisher="Entomologica Fennica" pageId="260" pageNumber="261" pagination="57 - 94" title="Taxonomy, distribution and habitats of the Dytiscidae (Coleoptera) of Ethiopia." volume="4" year="1993">Nilsson and Persson 1993</bibRefCitation>
|
|
: 79, 94 (faunistics, discussion);
|
|
<bibRefCitation id="B77F8921460016D74BA09759533918E8" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
|
|
: 248 (catalogue, faunistics);
|
|
<bibRefCitation id="8BD53CFA252C16A0A7D0563ACDD90C6E" author="Nilsson, AN" editor="Loebl, I" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" pagination="35 - 78" title="Dytiscidae." volumeTitle="Catalogue of Palaearctic Coleoptera, Vol. 1. Archostemata-Myxophaga-Adephaga" year="2003">Nilsson 2003</bibRefCitation>
|
|
: 77 (faunistics, list);
|
|
<bibRefCitation id="E7E3844C9F12F469AC5C776BC1067959" author="Angus, R" journalOrPublisher="Latissimus, newsletter" pageId="252" pageNumber="253" pagination="15 - 17" title="Egyptian Dytiscidae and Noteridae." volume="2003" year="2003">Angus 2003</bibRefCitation>
|
|
: 16 (synonymy, discussion); Hajek and Reiter 2014: (faunistics, biology);
|
|
<bibRefCitation id="892C9F8028E3E7AF6BB608B8BEEDEC38" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
|
|
: 168 (catalogue, faunistics).
|
|
</paragraph>
|
|
<paragraph id="F5AFC885549103C9DD46A6EDC46BFA44" pageId="44" pageNumber="45">
|
|
<taxonomicName id="30239F14373D7AA6CBD5695F12437CFF" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus wehnckei" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="wehnckei">Laccophilus wehnckei</taxonomicName>
|
|
<bibRefCitation id="42B4FA4A026332DF14F6E913A692C4EB" author="Sharp, D" journalOrPublisher="The Scientific Transactions of the Royal Dublin Society" pageId="263" pageNumber="264" pagination="179 - 1003" title="On aquatic carnivorous Coleoptera or Dytiscidae." volume="2" year="1882">Sharp 1882</bibRefCitation>
|
|
: 306 (original description, faunistics); v. d.
|
|
<bibRefCitation id="07C9622F8F45BE1FFFDBD31644281296" author="Branden, C van den" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="255" pageNumber="256" pagination="1 - 118" title="Catalogue des coleopteres carnassiers aquatiques (Haliplidae, Amphizoidae, Pelobiidae et Dytiscidae)." volume="28" year="1885">Branden 1885</bibRefCitation>
|
|
: 24 (catalogue, faunistics);
|
|
<bibRefCitation id="368566F8B7671012CA56F1802BC3E095" author="Regimbart, M" journalOrPublisher="Memoires de la Societe entomologique de Belgique" pageId="262" pageNumber="263" pagination="1 - 244" title="Revision des Dytiscidae et Gyrinidae d'Afrique, Madagascar et iles voisines. En contribution a la faune entomologique du Congo." volume="4" year="1895">
|
|
<normalizedToken id="0F5F9D47961D40609ECAA7A176E461BE" originalValue="Régimbart">Regimbart</normalizedToken>
|
|
1895
|
|
</bibRefCitation>
|
|
: 131 (description, faunistics, discussion);
|
|
<bibRefCitation id="7D98BAD70655CF86248AA6372F65D531" author="Zimmermann, A" journalOrPublisher="Junk, Berlin" pageId="263" pageNumber="264" title="Dytiscidae, Haliplidae, Hygrobiidae, Amphizoidae. In: Schenkling S (Ed.) Coleopterorum Catalogus. Vol. 4 pars 71. W." year="1920 a">Zimmermann1920a</bibRefCitation>
|
|
: 28 (catalogue);
|
|
<bibRefCitation id="025C4B8EF6EFBFDE790D6403AAA14CE9" author="Guignot, F" journalOrPublisher="Revue Francaise d'Entomologie" pageId="256" pageNumber="257" pagination="81 - 103" title="Dytiscidae et Gyrinidae de la Cote d'Ivoire et du Soudan francais." volume="9" year="1943">Guignot 1943</bibRefCitation>
|
|
: 99 (description, faunistics);
|
|
<bibRefCitation id="FDB658612C7FA77337F7D7B8E2780DAF" author="Guignot, F" journalOrPublisher="Memoires du Museum National d'Histoire Naturelle Paris" pageId="256" pageNumber="257" pagination="215 - 322" title="Mission scientifique de l'Omo 6 (58). ColeopteraDytiscidae (2 e partie)." volume="19" year="1946 c">Guignot 1946c</bibRefCitation>
|
|
: 270, 273, 277, 312, 315 (
|
|
<taxonomicName id="90841BB64893D0E637FEE78D48817F96" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus wehnchei" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="wehnchei">Laccophilus wehnchei</taxonomicName>
|
|
Sharp, misspelling: description, faunistics, discussion);
|
|
<bibRefCitation id="4BDBF169404185E61AB12EEDD029BFC8" author="Guignot, F" journalOrPublisher="Bulletin Institut royal des Sciences Naturelles de Belgique" pageId="257" pageNumber="258" pagination="1 - 7" title="Description de nouveaux dytiscides africains." volume="28" year="1952 d">Guignot 1952d</bibRefCitation>
|
|
: 4 (discussion: misidentification by
|
|
<bibRefCitation id="1D22D987339AAF12660651FAC7D446D5" author="Guignot, F" journalOrPublisher="Memoires du Museum National d'Histoire Naturelle Paris" pageId="256" pageNumber="257" pagination="215 - 322" title="Mission scientifique de l'Omo 6 (58). ColeopteraDytiscidae (2 e partie)." volume="19" year="1946 c">Guignot 1946c</bibRefCitation>
|
|
);
|
|
<bibRefCitation id="6652BD2A0118930F9DCDFEABA0463AFD" author="Legros, C" journalOrPublisher="Memoires de l'Institut Francais d'Afrique Noire" pageId="259" pageNumber="260" pagination="265 - 280" title="XXI. Coleopteres Hydrocanthares (Mission M. Lamotte, 1942)." volume="40" year="1954">Legros 1954</bibRefCitation>
|
|
: 268 (faunistics);
|
|
<bibRefCitation id="341648FF5FC2CDA07240575B197E1EDA" author="Legros, C" journalOrPublisher="Memoires de l'Institut Francais d'Afrique Noire" pageId="259" pageNumber="260" pagination="209 - 218" title="IX. Coleopteres Hydrocanthares (Mission M. Lamotte et Roy, juillet-decembre 1951)." volume="53" year="1958">Legros 1958</bibRefCitation>
|
|
: 211 (faunistics);
|
|
<bibRefCitation id="226F53E9093B9ED7503904EF576E60A1" author="Guignot, F" journalOrPublisher="Annales du Musee Royal du Congo Belge (Serie 8 vo) Sciences Zoologiques" pageId="258" pageNumber="259" pagination="323 - 648" title="Revision des hydrocanthares d'Afrique (ColeopteraDytiscoidea). 2." volume="78" year="1959 a">Guignot 1959a</bibRefCitation>
|
|
: 536, 567 (listed as synonym of
|
|
<taxonomicName id="C1C91A3D8A44780EC6EE5BD4D3C67275" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pictipennis" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="pictipennis">Laccophilus pictipennis</taxonomicName>
|
|
Sharp, discussion);
|
|
<bibRefCitation id="8F2EB6B0EEB5DB0311B03BC36CFE2381" author="Nilsson, AN" journalOrPublisher="Entomologica Fennica" pageId="260" pageNumber="261" pagination="57 - 94" title="Taxonomy, distribution and habitats of the Dytiscidae (Coleoptera) of Ethiopia." volume="4" year="1993">Nilsson and Persson 1993</bibRefCitation>
|
|
: 79 (list, synonymy);
|
|
<bibRefCitation id="AD39C01EDFC229FF73F3F849E3AD216E" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
|
|
: 249 (catalogue, faunistics, list, synonymy);
|
|
<bibRefCitation id="7AD5E75E60686506C6D137942CF71B69" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
|
|
: 168 (catalogue, faunistics, list, synonymy). Confirmed synonym.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="5BEF0B1D5F071090FA9C0D221AFA00FA" pageId="44" pageNumber="45" type="type localities">
|
|
<paragraph id="A49BF9EEDBD1B3BEF862EC8033508C6F" pageId="44" pageNumber="45">Type localities.</paragraph>
|
|
<paragraph id="CEC87F2AF3EDB0F34C4F33E48F747ACB" pageId="44" pageNumber="45">
|
|
<taxonomicName id="C482A8833668AE5A6317D1FBED86DE2D" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pictipennis" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="pictipennis">Laccophilus pictipennis</taxonomicName>
|
|
: Saudi Arabia: Hedjaz.
|
|
</paragraph>
|
|
<paragraph id="8D9F86569E89E425547AC5D2A2D612EB" pageId="44" pageNumber="45">
|
|
<taxonomicName id="84EDC6CB4F1097DE97BD5D6532F15676" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus wehnckei" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="wehnckei">Laccophilus wehnckei</taxonomicName>
|
|
: Tanzania: Zanzibar.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="BFFD68D1FA81C3395568E11D709E90BD" pageId="44" pageNumber="45" type="type material">
|
|
<paragraph id="1A3D9C6262580664ED39A607C44A9B2A" pageId="44" pageNumber="45">Type material studied</paragraph>
|
|
<paragraph id="F5DC28C74F9D07EB0C0F754621E39664" pageId="44" pageNumber="45">
|
|
(6 exs.):
|
|
<taxonomicName id="11D347C5363996052BC5A936760D8D8C" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pictipennis" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="pictipennis">Laccophilus pictipennis</taxonomicName>
|
|
: Lectotype (designated by
|
|
<bibRefCitation id="C6F4C5A87BB6B390E4E7D47D11191556" author="Brancucci, M" journalOrPublisher="Entomologische Arbeiten aus dem Museum G. Frey" pageId="255" pageNumber="256" pagination="241 - 426" title="Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae)." volume="31 / 32" year="1983 b">Brancucci (1983b)</bibRefCitation>
|
|
): Male: "Lectotype / Lectotypus / male-label / 566 / Hedjaz Millingen / Sharp Coll. 1905-313 /
|
|
<taxonomicName id="402FFEB337E55A3E7DAF356ADCEAAE03" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pictipennis" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="pictipennis">Laccophilus pictipennis</taxonomicName>
|
|
Shp M.E. Bacchus det. 1977 Syntype / Lectotype
|
|
<taxonomicName id="0743E26E2DA5287EF51D8B50BA73948F" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pictipennis" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="pictipennis">Laccophilus pictipennis</taxonomicName>
|
|
Sharp des. Brancucci 81" (BMNH; habitus in Fig. 398). - Paralectotypes: same data as lectotype but labeled as
|
|
<normalizedToken id="BC6A0DD7DB188FBE83F9920320B3456A" originalValue="“Paralectotype”">"Paralectotype"</normalizedToken>
|
|
(1 ex. BMNH); "Paralectotype / Paralectotypus / Abyssinia / Sharp Coll. 1905-313 / Type 566
|
|
<taxonomicName id="ECBC48F46785F873AE0F1451DB12B772" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pictipennis" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="pictipennis">Laccophilus pictipennis</taxonomicName>
|
|
/
|
|
<taxonomicName id="0FA6BCE01726DBFA76FE5E47150D3C05" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pictipennis" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="pictipennis">Laccophilus pictipennis</taxonomicName>
|
|
Shp M.E. Bacchus det. 1977 Syntype" (1 ex. BMNH); same data but add:
|
|
<normalizedToken id="363C03F93E1B26088334C1B3B6852B29" originalValue="“Raffray”">"Raffray"</normalizedToken>
|
|
(1 ex. BMNH).
|
|
</paragraph>
|
|
<paragraph id="7F6B76B82A0259B59199BE91A9F300C0" pageId="44" pageNumber="45">
|
|
<taxonomicName id="5CCC8296FFEFD75602B2CD091BA8381E" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus wehnckei" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="wehnckei">Laccophilus wehnckei</taxonomicName>
|
|
: Lectotype (by present designation): female: "Type / E. Africa / Sharp Coll. 1905-313 / Type 620
|
|
<taxonomicName id="497639CBD33CC4516EAB31667232EB44" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus wehnckei" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="wehnckei">Laccophilus wehnckei</taxonomicName>
|
|
sp. n. Zanzibar" (BMNH; habitus in Fig. 399). - Paralectotype: female: principally same data as lectotype but labelled as
|
|
<normalizedToken id="4F8D1749E887F41D93CE26CC4BD468AA" originalValue="“cotype”">"cotype"</normalizedToken>
|
|
(1 ex. BMNH).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="9272CB1C959B3A5C7B1EF9700CB49EB4" pageId="44" pageNumber="45" type="materials_examined">
|
|
<paragraph id="45AB0C8785B7250E10E1BBDD849E04AF" pageId="44" pageNumber="45">Additional material studied</paragraph>
|
|
<paragraph id="65380866DF6E3D5215CDA47DA8EE5E97" pageId="44" pageNumber="45">
|
|
(15 exs.). Ethiopia: "Saati Levander" (1 ex. MZH);
|
|
<normalizedToken id="418E8EFEAE549FCE38C3E341D6CF2059" originalValue="“Abyssinia”">"Abyssinia"</normalizedToken>
|
|
(1 ex. ZMHB). - Somalia: "Daragodleh 25.6. 1963 Linnavuori" (6 exs. MZH); "Lasgori /
|
|
<taxonomicName id="CABA9778D870B265C289821B1E0CFA04" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pictipennis" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="pictipennis">Laccophilus pictipennis</taxonomicName>
|
|
Sharp det. Brancucci 1982" (5 exx. ZMHB, 1 ex. NHMB). Non-African record: - Yemen: "W. Aden Prot. nr Lahej 9-15.7. 1963 Linnavuori" (1 ex. MZH).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="1F3B1E16CB675584827305598F353D7A" pageId="44" pageNumber="45" type="comments on synonymy">
|
|
<paragraph id="989AE1CA7D0C9AD3EBFECE88FF8101C4" pageId="44" pageNumber="45">Comments on synonymy.</paragraph>
|
|
<paragraph id="A1AA27309F488BB68FD58B6BEB1E7E7D" pageId="44" pageNumber="45">
|
|
Earlier established synonymy of
|
|
<taxonomicName id="CCEC4C12A7816BE0F3E96E39548A3065" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pictipennis" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="pictipennis">Laccophilus pictipennis</taxonomicName>
|
|
and
|
|
<taxonomicName id="735ABF8BC9D5E4621ADF7F1F354BEEBF" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus wehnckei" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="wehnckei">Laccophilus wehnckei</taxonomicName>
|
|
is confirmed by study of external characters; no diagnostically important differences detected. As no males are available of
|
|
<taxonomicName id="12A1E5683D084C45BB91110CA44DCD25" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus wehnckei" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="wehnckei">Laccophilus wehnckei</taxonomicName>
|
|
we could not in this case undertake comparison of male genitalia.
|
|
<taxonomicName id="C3E618A1B071CD7064D272FF88E4CA26" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus discretus" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="discretus">Laccophilus discretus</taxonomicName>
|
|
Sharp, 1882, described from Saudi Arabia, has earlier been synonymized with
|
|
<taxonomicName id="7B372A261BFC0AADBE4F306216FE43E2" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pictipennis" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="pictipennis">Laccophilus pictipennis</taxonomicName>
|
|
. It has never been recorded from Africa by its own name and accordingly, it is outside the scope of this study.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="61E9F85A8FBAF0AD89CCC596156FE900" pageId="44" pageNumber="45" type="diagnosis">
|
|
<paragraph id="F48B779CA569ED9470B6F9D0C2EED303" pageId="44" pageNumber="45">Diagnosis.</paragraph>
|
|
<paragraph id="6513032374CE714C43396E3ADCA9C7F3" pageId="44" pageNumber="45">
|
|
<taxonomicName id="9864FD17EE25D2E40649098731791BF7" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pictipennis" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="pictipennis">Laccophilus pictipennis</taxonomicName>
|
|
is characterized by peculiar elytral colour pattern in combination with penis, which is slightly and evenly curved, tapering gradually towards its apex. Note also that male apical ventrite lacks asymmetrically located knob, although excavated on each side of midline and slight medial extension (Fig. 45).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="6E69F95690186B3AA42B535F3E823FAB" pageId="44" pageNumber="45" type="description">
|
|
<paragraph id="6C9D3C67F376DA60AFD25324A41E63E8" pageId="44" pageNumber="45">Description.</paragraph>
|
|
<paragraph id="2458233539B7C090E9C68F62C087F2BF" pageId="44" pageNumber="45">Body length 4.4-4.7 mm, width 2.5-2.8 mm. Dorsal, aspect of body with fairly distinct colour pattern. African specimens seem to have vaguer dorsal colour pattern (Figs 398-399).</paragraph>
|
|
<paragraph id="60C5255E1183938A3949A34480FDA45E" pageId="44" pageNumber="45">Head: Pale ferrugineous. Punctation indistinct, almost absent; close to eyes with two minute deptressions with irregular, fine punctures. Shiny, although irregularly and rather finely reticulated. In part double reticulation weakly discernible (delimitation in two distinct size classes of meshes vague).</paragraph>
|
|
<paragraph id="4CF6F6654DF1B002E0EDA228A6CD0C41" pageId="44" pageNumber="45">Pronotum: Pale ferrugineous. At frontal margin with a vague, ferrugineous, almost bilobed marking. Punctation fine, sparse to rather sparse and irregularly distributed. Punctures frontally densest. Rather shiny although microsculptured. Reticulation indistinctly divided into two kinds; smaller meshes sometimes discernible within large meshes. In part, only large meshes well-developed.</paragraph>
|
|
<paragraph id="5777CD1219D110F62F502934305C4465" pageId="44" pageNumber="45">Elytra: Pale ferrugineous, with extensive, distinct, dark ferrugineous to brownish markings (Fig. 398). Sometimes elytral colour pattern rather vague (Fig. 399). Reticulation double; large meshes contain generally 3-6 small meshes. Small meshes fine, sometimes weakly developed and indistinct. Irregular, discal, dorsolateral and lateral rows of punctures are discernible. All rows of punctures rather sparse, not forming straight rows.</paragraph>
|
|
<paragraph id="6B7E8C8EC5C4CFFB23FA5531638E0BF3" pageId="44" pageNumber="45">Ventral aspect: Pale ferrugineous to ferrugineous. Rather shiny, although extensively, finely microsculptured. Besides microsculpture metacoxal plates with shallow, transverse furrows and abdomen especially basally with distinct striae. Apical ventrite lacks knob (Fig. 45). Apex of prosternal process rather narrow and pointed.</paragraph>
|
|
<paragraph id="0CA7A42E8E28937E3541118D76771E59" pageId="44" pageNumber="45">Legs: Pro- and mesotarsus slightly enlarged, with fine suckers.</paragraph>
|
|
<paragraph id="224DBAF1F5D84FFDA4467593136AC7D8" pageId="44" pageNumber="45">Male genitalia: Penis in lateral aspect quite broad, from base slightly and evenly curved to apex (Figs 247-248).</paragraph>
|
|
<paragraph id="7F6D7BB892C3012C0C0D24DF3559FC8A" pageId="44" pageNumber="45">Female: Pro- and mesotarsus slender. Apical ventrite (Fig. 46).</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="7423172E69534012189EE7FB198276D7" pageId="44" pageNumber="45" type="distribution">
|
|
<paragraph id="F5BC506033ABDB20933DF23A437BA263" pageId="44" pageNumber="45">Distribution.</paragraph>
|
|
<paragraph id="2A57291331B7D53030BE89BFD08C586B" pageId="44" pageNumber="45">Somalia, Ethiopia, Tanzania (Zanzibar) (Fig. 534). Material examined also from Arabian Peninsula (Saudi Arabia and Yemen). Only personally examined specimens accepted.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="651F82C35F117082CEE106BC975CB297" pageId="44" pageNumber="45" type="collecting circumstances">
|
|
<paragraph id="E01449612BBA2AA80DCD5EE4D0F47946" pageId="44" pageNumber="45">Collecting circumstances.</paragraph>
|
|
<paragraph id="C06FFCBA14A7CE667294B214481AF246" pageId="44" pageNumber="45">
|
|
Almost unknown in Africa.
|
|
<bibRefCitation id="3292AD9BB922008B7FD68D2A4BF41504" author="Hajek, J" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae" pageId="258" pageNumber="259" pagination="63 - 99" title="Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island." volume="54" year="2014">Hajek and Reiter (2014)</bibRefCitation>
|
|
report the species from Oman being mostly associated with running water, especially in relatively permanent side pools of streams and river at lower and middle altitudes.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |