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<document id="94182D6BCC959691657345E75C7FCCBD" ID-CLB-Dataset="7077" ID-DOI="10.11646/zootaxa.4964.1.11" ID-GBIF-Dataset="5fc30ab2-a55d-49fc-8e7f-2c0abaee239e" ID-ISSN="1175-5326" ID-Zenodo-Dep="4706654" ID-ZooBank="014333E3-4B5F-4F44-899B-515F8211D3CD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618986106579" checkinUser="plazi" docAuthor="Bouyer, Thierry, Drumont, Alain, Roland, Lily-Arison Rene De, Slootmaekers, Dan, Pirkl, Jiri &amp; Jocque, Merlijn" docDate="2021" docId="783F8796FB38FF97FF765A18FA6E3286" docLanguage="en" docName="zootaxa.4964.1.11.pdf" docOrigin="Zootaxa 4964 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Schizodontus mahimborondroensis Bouyer &amp; Drumont &amp; Roland &amp; Slootmaekers &amp; Pirkl &amp; Jocque 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="193" masterDocId="8406FFEEFB3BFF92FFE15C05FFF33114" masterDocTitle="Notes on a small collection of Prioninae from North-East Madagascar with the description of a new Schizodontus Quentin &amp; Villiers, 1974 (Cerambycidae Closterini)" masterLastPageNumber="194" masterPageNumber="188" pageNumber="191" updateTime="1698937292149" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="75668E3CD7F5883CDDB16228DF38C32B">Notes on a small collection of Prioninae from North-East Madagascar with the description of a new Schizodontus Quentin &amp; Villiers, 1974 (Cerambycidae Closterini)</mods:title>
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<mods:affiliation id="48997CB81113824C56E9A756F0A25833">Rue Genot 57, B- 4032, Chênée, Belgium.</mods:affiliation>
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<mods:namePart id="BAAC76C0B5175F3552A5B609E940CF00">Drumont, Alain</mods:namePart>
<mods:affiliation id="DC75E406F362D0D0EE7C4BF57D717697">O. D. Taxonomy and Phylogeny-Entomology, Royal Belgian Institute of Natural Sciences (RBINS), Vautierstreet 29, B- 1000 Brussels, Belgium.</mods:affiliation>
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<mods:namePart id="43B8289A86E9CD665CF940727AA5469A">Roland, Lily-Arison Rene De</mods:namePart>
<mods:affiliation id="9E4245EFEEC7D0198DF59CCDD2A0E13E">The Peregrine Fund Madagascar, PO Box 4113, Antananarivo, Madagascar.</mods:affiliation>
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<mods:namePart id="51755DCEE16DBC96AF0EC5AEC2FF8888">Slootmaekers, Dan</mods:namePart>
<mods:affiliation id="9FFFA63649785BB5883E71B414F7E67E">Biodiversity Inventory for Conservation (BINCO) NPO, Walmersumstraat 44, B- 3380 Glabbeek, Belgium. danslootmaekers @ gmail. com</mods:affiliation>
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<mods:namePart id="00820E7CA1F61F305B6876C9B36FA4D7">Pirkl, Jiri</mods:namePart>
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<mods:affiliation id="758CA2EBAD830088EB67456E7A36DA83">Komenskeho 4440, CZ- 430 03 Chomutov, Czech Republic. &amp; School of Geography and the Environment, University of Oxford, Oxford, UK. &amp; BirdLife South Africa, Centre for Biodiversity Conservation, Kirstenbosch Botanical Gardens, Cape Town, South Africa. dalewr @ gmail. com; https: // orcid. org / 0000 - 0003 - 3946 - 3582</mods:affiliation>
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<mods:namePart id="3FB3A6DB80FF4F5868AFD5EE1C91CE66">Jocque, Merlijn</mods:namePart>
<mods:affiliation id="B0C1D3B009E5F835497D97B7F035C18F">O. D. Taxonomy and Phylogeny-Entomology, Royal Belgian Institute of Natural Sciences (RBINS), Vautierstreet 29, B- 1000 Brussels, Belgium. &amp; Aquatic and terrestrial Ecology (ATECO), Royal Belgian Institute of Natural Sciences (RBINS), Vautierstreet 29, B- 1000 Brussels Belgium.</mods:affiliation>
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<taxonomicName id="37964D03FB38FF91FF765A18FDB1372C" ID-CoL="BLLQD" authority="Bouyer &amp; Drumont &amp; Roland &amp; Slootmaekers &amp; Pirkl &amp; Jocque, 2021" authorityName="Bouyer &amp; Drumont &amp; Roland &amp; Slootmaekers &amp; Pirkl &amp; Jocque" authorityYear="2021" box="[151,578,1565,1592]" class="Insecta" family="Cerambycidae" genus="Schizodontus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="191" phylum="Arthropoda" rank="species" species="mahimborondroensis" status="sp. nov.">
<emphasis id="C2E2EA92FB38FF91FF765A18FDB1372C" bold="true" box="[151,578,1565,1592]" italics="true" pageId="3" pageNumber="191">Schizodontus mahimborondroensis</emphasis>
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Bouyer &amp; Drumont
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<paragraph id="F0293680FB38FF91FF765A44FE973748" blockId="3.[151,932,1565,1628]" box="[151,356,1601,1628]" pageId="3" pageNumber="191">
(
<figureCitation id="68AD2A05FB38FF91FF7E5A44FF1B3748" box="[159,232,1601,1628]" captionStart="FIGURE 2" captionStartId="2.[151,250,1668,1692]" captionTargetBox="[154,1424,885,1623]" captionTargetId="figure-244@2.[151,1436,880,1644]" captionTargetPageId="2" captionText="FIGURE 2. Schizodontus mahimborondroensis sp. nov. Holotype male, 34 mm, dorsal view (RBINS). Figure 3. Schizodontus angustus Quentin &amp; Villiers, 1974. Holotype male, 34 mm, dorsal view (MNHN). Figure 4. Schizodontus latus Quentin &amp; Villiers, 1974. Holotype male, 37 mm, dorsal view (MNHN). (Photos Thierry Bouyer (Fig. 2) &amp; Jiri Pirkl (figs 34))." figureDoi="http://doi.org/10.5281/zenodo.4706658" httpUri="https://zenodo.org/record/4706658/files/figure.png" pageId="3" pageNumber="191">Figs 2</figureCitation>
,
<figureCitation id="68AD2A05FB38FF91FF155A44FED33748" box="[244,288,1601,1628]" captionStart="FIGURES 57" captionStartId="3.[151,264,1171,1195]" captionTargetBox="[160,1421,285,1142]" captionTargetId="figure-50@3.[151,1436,273,1148]" captionTargetPageId="3" captionText="FIGURES 57. Schizodontus mahimborondroensis sp. nov. Male holotype (34 mm). Fig. 5: holotype, head and pronotum, dorsal view. Fig. 6: elytra, lateral view from the left side. Fig. 7: left antenna with numbered articles (Photos Thierry Bouyer)." figureDoi="http://doi.org/10.5281/zenodo.4706660" httpUri="https://zenodo.org/record/4706660/files/figure.png" pageId="3" pageNumber="191">57</figureCitation>
,
<figureCitation id="68AD2A05FB38FF91FECD5A47FEC93748" box="[300,314,1602,1628]" captionStart="FIGURE 8" captionStartId="4.[152,235,1632,1656]" captionTargetBox="[306,1246,192,1563]" captionTargetId="figure-18@4.[305,1281,181,1612]" captionTargetPageId="4" captionText="FIGURE 8. First antennal segments of 1) Schizodontus mahimborondroensis sp. nov. (male), 2) S. latus (male), 3) S. angustus (male) and 4) S. latus (female). 24 after Quentin &amp; Villiers, 1975, figs. 227229." figureDoi="http://doi.org/10.5281/zenodo.4706662" httpUri="https://zenodo.org/record/4706662/files/figure.png" pageId="3" pageNumber="191">8</figureCitation>
(1))
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<paragraph id="F0293680FB38FF91FF765A8FFCE437F8" blockId="3.[151,1437,1673,2024]" pageId="3" pageNumber="191">
<emphasis id="C2E2EA92FB38FF91FF765A8FFEB437B7" bold="true" box="[151,327,1673,1700]" pageId="3" pageNumber="191">Type material.</emphasis>
<materialsCitation id="40FE3CDDFB38FF91FEB15A8CFCE437F8" ID-GBIF-Occurrence="3107296301" collectingDate="2019-02-08" collectionCode="RIDGE, BINCO, MAD, RBINS" collectorName="Merlijn Jocque &amp; Dan Slootmaekers &amp; Th. Bouyer" country="Madagascar" elevation="1717" latitude="-14.2939" location="Bemanevika" longLatPrecision="7" longitude="48.7569" pageId="3" pageNumber="191" specimenCode="MAD19" specimenCount="2" typeStatus="holotype">
<typeStatus id="2F2D8822FB38FF91FEB15A8CFE4B37B7" box="[336,440,1673,1699]" pageId="3" pageNumber="191" type="holotype">Holotype</typeStatus>
male and
<specimenCount id="E690FD09FB38FF91FDD35A8FFD4037B0" box="[562,691,1674,1700]" pageId="3" pageNumber="191" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
male:
<collectingCountry id="88817610FB38FF91FCE55A8CFC7E37B7" box="[772,909,1673,1699]" name="Madagascar" pageId="3" pageNumber="191">Madagascar</collectingCountry>
,
<location id="F549605BFB38FF91FC7B5A8CFBD537B7" LSID="urn:lsid:plazi:treatment:783F8796FB38FF97FF765A18FA6E3286:F549605BFB38FF91FC7B5A8CFBD537B7" box="[922,1062,1673,1699]" country="Madagascar" latitude="-14.2939" longLatPrecision="7" longitude="48.7569" name="Bemanevika" pageId="3" pageNumber="191">Bemanevika</location>
,
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<elevation id="7BBBD1B3FB38FF91FBD75A8FFB7C37B0" box="[1078,1167,1674,1700]" metricMagnitude="3" metricUnit="m" metricValue="1.7169999999999999" pageId="3" pageNumber="191" unit="m" value="1717.0">1717 m</elevation>
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,
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<geoCoordinate id="95A25047FB38FF91FAC15A8CFA6437B0" box="[1312,1431,1673,1700]" degrees="48.7569" direction="east" orientation="longitude" pageId="3" pageNumber="191" precision="5" value="48.7569">48.7569 E</geoCoordinate>
,
<date id="84281040FB38FF91FF765AABFEF337DC" box="[151,256,1710,1736]" pageId="3" pageNumber="191" value="2019-02-08">
<collectingDate id="946CE9A8FB38FF91FF765AABFEF337DC" box="[151,256,1710,1736]" pageId="3" pageNumber="191" value="2019-02-08">8.II.2019</collectingDate>
</date>
,
<collectingMethod id="29D74E97FB38FF91FEED5AA8FE8737DC" box="[268,372,1709,1736]" pageId="3" pageNumber="191">light trap</collectingMethod>
, [
<specimenCode id="A0309EFBFB38FF91FE675AABFE1537DC" box="[390,486,1710,1736]" collectionCode="MAD" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="3" pageNumber="191">MAD19</specimenCode>
<collectionCode id="9687AE45FB38FF91FE0C5AABFDB037DC" box="[493,579,1710,1736]" pageId="3" pageNumber="191">RIDGE</collectionCode>
LT].
<collectionCode id="9687AE45FB38FF91FD615AABFD2A37DC" box="[640,729,1710,1736]" pageId="3" pageNumber="191">BINCO</collectionCode>
_
<collectionCode id="9687AE45FB38FF91FD095AABFCD837DC" box="[744,811,1710,1736]" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="3" pageNumber="191">MAD</collectionCode>
_19_0008,
<collectorName id="5D635356FB38FF91FC445AABFB7437DC" box="[933,1159,1709,1736]" pageId="3" pageNumber="191">Leg. Merlijn Jocque</collectorName>
&amp;
<collectorName id="5D635356FB38FF91FB4A5AABFA8837D3" box="[1195,1403,1709,1736]" pageId="3" pageNumber="191">Dan Slootmaekers</collectorName>
,
<emphasis id="C2E2EA92FB38FF91FA675AABFA6F37D3" box="[1414,1436,1710,1735]" italics="true" pageId="3" pageNumber="191">in</emphasis>
coll.
<collectionCode id="9687AE45FB38FF91FF2E5AD7FED237F8" box="[207,289,1746,1772]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="3" pageNumber="191" type="Museum">RBINS</collectionCode>
(HT and 1 PT), and coll.
<collectorName id="5D635356FB38FF91FDA05AD4FD3337F8" box="[577,704,1745,1772]" pageId="3" pageNumber="191">Th. Bouyer</collectorName>
(1 PT).
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<emphasis id="C2E2EA92FB38FF91FF265AF0FEAA361B" bold="true" box="[199,345,1781,1807]" pageId="3" pageNumber="191">Description.</emphasis>
</paragraph>
<paragraph id="F0293680FB38FF91FF265B1CFD3D36FC" blockId="3.[151,1437,1673,2024]" pageId="3" pageNumber="191">
<typeStatus id="2F2D8822FB38FF91FF265B1CFEDC3627" box="[199,303,1817,1843]" pageId="3" pageNumber="191" type="holotype">Holotype</typeStatus>
male (
<figureCitation id="68AD2A05FB38FF91FE625B1CFE3F3620" box="[387,460,1817,1844]" captionStart="FIGURE 2" captionStartId="2.[151,250,1668,1692]" captionTargetBox="[154,1424,885,1623]" captionTargetId="figure-244@2.[151,1436,880,1644]" captionTargetPageId="2" captionText="FIGURE 2. Schizodontus mahimborondroensis sp. nov. Holotype male, 34 mm, dorsal view (RBINS). Figure 3. Schizodontus angustus Quentin &amp; Villiers, 1974. Holotype male, 34 mm, dorsal view (MNHN). Figure 4. Schizodontus latus Quentin &amp; Villiers, 1974. Holotype male, 37 mm, dorsal view (MNHN). (Photos Thierry Bouyer (Fig. 2) &amp; Jiri Pirkl (figs 34))." figureDoi="http://doi.org/10.5281/zenodo.4706658" httpUri="https://zenodo.org/record/4706658/files/figure.png" pageId="3" pageNumber="191">Fig. 2</figureCitation>
).
<emphasis id="C2E2EA92FB38FF91FE055B1FFD4F3620" bold="true" box="[484,700,1818,1844]" pageId="3" pageNumber="191">Head and thorax.</emphasis>
Head, pronotum and scutellum (
<figureCitation id="68AD2A05FB38FF91FBDC5B1CFB713620" box="[1085,1154,1817,1844]" captionStart="FIGURES 57" captionStartId="3.[151,264,1171,1195]" captionTargetBox="[160,1421,285,1142]" captionTargetId="figure-50@3.[151,1436,273,1148]" captionTargetPageId="3" captionText="FIGURES 57. Schizodontus mahimborondroensis sp. nov. Male holotype (34 mm). Fig. 5: holotype, head and pronotum, dorsal view. Fig. 6: elytra, lateral view from the left side. Fig. 7: left antenna with numbered articles (Photos Thierry Bouyer)." figureDoi="http://doi.org/10.5281/zenodo.4706660" httpUri="https://zenodo.org/record/4706660/files/figure.png" pageId="3" pageNumber="191">Fig. 5</figureCitation>
) strongly punctuate and covered with a dense mat of small hairs. Interocular space flat, with a well-defined median groove between base of antenna. A shallow transversal supramandibular groove clearly present. Antennae long, reaching just before or just about end of elytra.Antennomeres relatively wide, with an external apical tooth clearly visible from antennomere III onwards, antennomere VII with an internal apical tooth but always much smaller than external apical tooth, antennomeres XXI almost completely merged (
<figureCitation id="68AD2A05FB38FF91FD9D5BC8FD3336FC" box="[636,704,1997,2024]" captionStart="FIGURES 57" captionStartId="3.[151,264,1171,1195]" captionTargetBox="[160,1421,285,1142]" captionTargetId="figure-50@3.[151,1436,273,1148]" captionTargetPageId="3" captionText="FIGURES 57. Schizodontus mahimborondroensis sp. nov. Male holotype (34 mm). Fig. 5: holotype, head and pronotum, dorsal view. Fig. 6: elytra, lateral view from the left side. Fig. 7: left antenna with numbered articles (Photos Thierry Bouyer)." figureDoi="http://doi.org/10.5281/zenodo.4706660" httpUri="https://zenodo.org/record/4706660/files/figure.png" pageId="3" pageNumber="191">Fig. 7</figureCitation>
).
</paragraph>
<caption id="A4E96608FB3FFF96FF795A65FC26378E" ID-DOI="http://doi.org/10.5281/zenodo.4706662" ID-Zenodo-Dep="4706662" httpUri="https://zenodo.org/record/4706662/files/figure.png" pageId="4" pageNumber="192" startId="4.[152,235,1632,1656]" targetBox="[306,1246,192,1563]" targetPageId="4">
<paragraph id="F0293680FB3FFF96FF795A65FC26378E" blockId="4.[151,1419,1629,1690]" pageId="4" pageNumber="192">
<emphasis id="C2E2EA92FB3FFF96FF795A65FEF6376C" bold="true" box="[152,261,1632,1656]" pageId="4" pageNumber="192">FIGURE 8.</emphasis>
First antennal segments of 1)
<taxonomicName id="37964D03FB3FFF96FDD75A58FC643761" authority="Bouyer &amp; Drumont &amp; Roland &amp; Slootmaekers &amp; Pirkl &amp; Jocque, 2021" authorityName="Bouyer &amp; Drumont &amp; Roland &amp; Slootmaekers &amp; Pirkl &amp; Jocque" authorityYear="2021" box="[566,919,1629,1653]" class="Insecta" family="Cerambycidae" genus="Schizodontus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="mahimborondroensis" status="sp. nov.">
<emphasis id="C2E2EA92FB3FFF96FDD75A58FC643761" box="[566,919,1629,1653]" italics="true" pageId="4" pageNumber="192">Schizodontus mahimborondroensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D9D157E9FB3FFF96FC7F5A5BFC1E3761" box="[926,1005,1630,1653]" pageId="4" pageNumber="192" rank="species">sp. nov.</taxonomicNameLabel>
(male), 2)
<taxonomicName id="37964D03FB3FFF96FBBC5A5BFB543761" authorityName="Quentin &amp; Villiers" authorityYear="1974" box="[1117,1191,1629,1653]" class="Insecta" family="Cerambycidae" genus="Schizodontus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="C2E2EA92FB3FFF96FBBC5A5BFB543761" box="[1117,1191,1629,1653]" italics="true" pageId="4" pageNumber="192">S. latus</emphasis>
</taxonomicName>
(male), 3)
<taxonomicName id="37964D03FB3FFF96FAF95A5BFA783761" authorityName="Quentin &amp; Villiers" authorityYear="1974" box="[1304,1419,1630,1653]" class="Insecta" family="Cerambycidae" genus="Schizodontus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="angustus">
<emphasis id="C2E2EA92FB3FFF96FAF95A5BFA783761" box="[1304,1419,1630,1653]" italics="true" pageId="4" pageNumber="192">S. angustus</emphasis>
</taxonomicName>
(male) and 4)
<taxonomicName id="37964D03FB3FFF96FEC75A87FE83378D" authorityName="Quentin &amp; Villiers" authorityYear="1974" box="[294,368,1665,1689]" class="Insecta" family="Cerambycidae" genus="Schizodontus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="C2E2EA92FB3FFF96FEC75A87FE83378D" box="[294,368,1665,1689]" italics="true" pageId="4" pageNumber="192">S. latus</emphasis>
</taxonomicName>
(female). 24 after
<bibRefCitation id="94074B71FB3FFF96FDDB5A84FCCB378D" author="Quentin, R. M. &amp; Villiers, A." box="[570,824,1665,1690]" pageId="4" pageNumber="192" refId="ref3475" refString="Quentin, R. M. &amp; Villiers, A. (1975) Faune de Madagascar, volume 40: Insectes Coleopteres Cerambycidae - Parandrinae et Prioninae. ORSTOM &amp; CNRS, Paris. pp. 251." type="book" year="1975">Quentin &amp; Villiers, 1975</bibRefCitation>
, figs. 227229.
</paragraph>
</caption>
<paragraph id="F0293680FB3FFF96FF265ABCFE54360F" blockId="4.[151,1436,1721,2036]" pageId="4" pageNumber="192">Lateral pronotal spines medium to small, central spines almost equal in size to anterior spines. Prosternum clearly and densely setose. Pronotum with a well-defined medial notch in posterior ridge behind head. No local depression in pronotum.</paragraph>
<paragraph id="F0293680FB3FFF96FF265B20FECE369C" blockId="4.[151,1436,1721,2036]" pageId="4" pageNumber="192">
<emphasis id="C2E2EA92FB3FFF96FF265B20FE37362B" bold="true" box="[199,452,1829,1855]" pageId="4" pageNumber="192">Abdomen and elytra.</emphasis>
General color dark brown. Elytra covered with a short, sparse and barely visible mat of fine hairs, providing shiny elytra, especially at level of disc. Elytra finely punctuated with well-defined longitudinal ridges (
<figureCitation id="68AD2A05FB3FFF96FF0A5B68FEDC369C" box="[235,303,1901,1928]" captionStart="FIGURES 57" captionStartId="3.[151,264,1171,1195]" captionTargetBox="[160,1421,285,1142]" captionTargetId="figure-50@3.[151,1436,273,1148]" captionTargetPageId="3" captionText="FIGURES 57. Schizodontus mahimborondroensis sp. nov. Male holotype (34 mm). Fig. 5: holotype, head and pronotum, dorsal view. Fig. 6: elytra, lateral view from the left side. Fig. 7: left antenna with numbered articles (Photos Thierry Bouyer)." figureDoi="http://doi.org/10.5281/zenodo.4706660" httpUri="https://zenodo.org/record/4706660/files/figure.png" pageId="4" pageNumber="192">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="F0293680FB3FFF96FF265B94FE6936BF" blockId="4.[151,1436,1721,2036]" box="[199,410,1937,1963]" pageId="4" pageNumber="192">
<emphasis id="C2E2EA92FB3FFF96FF265B94FED636BF" bold="true" box="[199,293,1937,1963]" pageId="4" pageNumber="192">Female.</emphasis>
Unknown
</paragraph>
<paragraph id="F0293680FB3FFF96FF265BB0FBC736E7" blockId="4.[151,1436,1721,2036]" pageId="4" pageNumber="192">
<emphasis id="C2E2EA92FB3FFF96FF265BB0FEAA36DB" bold="true" box="[199,345,1973,1999]" pageId="4" pageNumber="192">Dimensions.</emphasis>
<typeStatus id="2F2D8822FB3FFF96FE815BB0FE3B36DB" box="[352,456,1973,1999]" pageId="4" pageNumber="192" type="holotype">Holotype</typeStatus>
is
<quantity id="376E9B65FB3FFF96FE085BB0FDCA36DB" box="[489,569,1973,1999]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="4" pageNumber="192" unit="mm" value="34.0">34 mm</quantity>
(
<figureCitation id="68AD2A05FB3FFF96FDA95BB0FD7E36DB" box="[584,653,1973,1999]" captionStart="FIGURE 2" captionStartId="2.[151,250,1668,1692]" captionTargetBox="[154,1424,885,1623]" captionTargetId="figure-244@2.[151,1436,880,1644]" captionTargetPageId="2" captionText="FIGURE 2. Schizodontus mahimborondroensis sp. nov. Holotype male, 34 mm, dorsal view (RBINS). Figure 3. Schizodontus angustus Quentin &amp; Villiers, 1974. Holotype male, 34 mm, dorsal view (MNHN). Figure 4. Schizodontus latus Quentin &amp; Villiers, 1974. Holotype male, 37 mm, dorsal view (MNHN). (Photos Thierry Bouyer (Fig. 2) &amp; Jiri Pirkl (figs 34))." figureDoi="http://doi.org/10.5281/zenodo.4706658" httpUri="https://zenodo.org/record/4706658/files/figure.png" pageId="4" pageNumber="192">Fig. 2</figureCitation>
). Range length for
<specimenCount id="E690FD09FB3FFF96FC875BB0FC2836DB" box="[870,987,1973,1999]" pageId="4" pageNumber="192" type="male">three male</specimenCount>
specimens collected is
<quantity id="376E9B65FB3FFF96FB045BB0FA9236DB" box="[1253,1377,1973,1999]" metricMagnitude="-2" metricUnit="m" metricValue="3.15" metricValueMax="3.4" metricValueMin="2.9" pageId="4" pageNumber="192" unit="mm" value="31.5" valueMax="34.0" valueMin="29.0">2934 mm</quantity>
. Pronotum length:
<quantity id="376E9B65FB3FFF96FEDA5BDCFE3636E7" box="[315,453,2009,2036]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="4" pageNumber="192" unit="mm" value="4.0" valueMax="4.5" valueMin="3.5">3.54.5 mm</quantity>
. Pronotum maximal width without spines:
<quantity id="376E9B65FB3FFF96FC475BDCFBC336E7" box="[934,1072,2009,2036]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="7.5" metricValueMin="6.5" pageId="4" pageNumber="192" unit="mm" value="7.0" valueMax="7.5" valueMin="6.5">6.57.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="B88C650BFB3EFF97FF265C93FC1B31C0" pageId="5" pageNumber="193" type="etymology">
<paragraph id="F0293680FB3EFF97FF265C93FC1B31C0" blockId="5.[151,1437,150,573]" pageId="5" pageNumber="193">
<emphasis id="C2E2EA92FB3EFF97FF265C93FEBD31A4" bold="true" box="[199,334,150,176]" pageId="5" pageNumber="193">Etymology.</emphasis>
The species is named after the Malagasy name for the
<typeStatus id="2F2D8822FB3EFF97FC5A5C92FC1831A5" box="[955,1003,151,177]" pageId="5" pageNumber="193">type</typeStatus>
locality: Mahimborondro, meaning “a place to feel the clouds” and refers to the cloud forest habitat of the species.
</paragraph>
</subSubSection>
<subSubSection id="B88C650BFB3EFF97FF265CDBFB5D3390" pageId="5" pageNumber="193" type="biology_ecology">
<paragraph id="F0293680FB3EFF97FF265CDBFCD33008" blockId="5.[151,1437,150,573]" pageId="5" pageNumber="193">
<emphasis id="C2E2EA92FB3EFF97FF265CDBFEDF31EC" bold="true" box="[199,300,222,248]" pageId="5" pageNumber="193">Ecology.</emphasis>
Nothing is known about the ecology of the species at this point, but the specimens were collected in dense high elevation ridge clad in montane forest (
<figureCitation id="68AD2A05FB3EFF97FD2F5D07FCE13008" box="[718,786,258,284]" captionStart="FIGURE 9" captionStartId="5.[151,250,1841,1865]" captionTargetBox="[151,1436,968,1824]" captionTargetId="figure-594@5.[151,1436,968,1824]" captionTargetPageId="5" captionText="FIGURE 9. Montane ridge forest where Schizodontus mahimborondroensis sp. nov. was collected (Photo Merlijn Jocque)." figureDoi="http://doi.org/10.5281/zenodo.4706664" httpUri="https://zenodo.org/record/4706664/files/figure.png" pageId="5" pageNumber="193">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="F0293680FB3EFF97FF265D23FD9E3328" blockId="5.[151,1437,150,573]" pageId="5" pageNumber="193">
<emphasis id="C2E2EA92FB3EFF97FF265D23FE733054" bold="true" box="[199,384,294,320]" pageId="5" pageNumber="193">Similar species.</emphasis>
There are only three described species in the genus.
<typeStatus id="2F2D8822FB3EFF97FC305D23FBB73054" box="[977,1092,294,320]" pageId="5" pageNumber="193" type="holotype">Holotypes</typeStatus>
of
<taxonomicName id="37964D03FB3EFF97FB8B5D22FB4F3054" authorityName="Quentin &amp; Villiers" authorityYear="1974" box="[1130,1212,294,321]" class="Insecta" family="Cerambycidae" genus="Schizodontus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="C2E2EA92FB3EFF97FB8B5D22FB4F3054" box="[1130,1212,294,321]" italics="true" pageId="5" pageNumber="193">S. latus</emphasis>
</taxonomicName>
and
<taxonomicName id="37964D03FB3EFF97FB125D22FA803054" authorityName="Quentin &amp; Villiers" authorityYear="1974" box="[1267,1395,295,321]" class="Insecta" family="Cerambycidae" genus="Schizodontus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="angustus">
<emphasis id="C2E2EA92FB3EFF97FB125D22FA803054" box="[1267,1395,295,321]" italics="true" pageId="5" pageNumber="193">S. angustus</emphasis>
</taxonomicName>
are preserved in MNHN (Figs 34) and high resolution images of these two species are available through the website of the
<taxonomicName id="37964D03FB3EFF97FF005D6BFEBF309C" authorityName="Latreille" authorityYear="1802" box="[225,332,366,392]" class="Insecta" family="Cerambycidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="subFamily" subFamily="Prioninae">Prioninae</taxonomicName>
(http://www.
<taxonomicName id="37964D03FB3EFF97FE015D6BFDBC309C" authorityName="Latreille" authorityYear="1802" box="[480,591,366,392]" class="Insecta" family="Cerambycidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="subFamily" subFamily="Prioninae">prioninae</taxonomicName>
.eu/) managed by one of the authors (Pirkl). Males of the three species can most easily be separated based on the structure of the antennae (
<figureCitation id="68AD2A05FB3EFF97FC8C5D97FC4030B8" box="[877,947,402,428]" captionStart="FIGURE 8" captionStartId="4.[152,235,1632,1656]" captionTargetBox="[306,1246,192,1563]" captionTargetId="figure-18@4.[305,1281,181,1612]" captionTargetPageId="4" captionText="FIGURE 8. First antennal segments of 1) Schizodontus mahimborondroensis sp. nov. (male), 2) S. latus (male), 3) S. angustus (male) and 4) S. latus (female). 24 after Quentin &amp; Villiers, 1975, figs. 227229." figureDoi="http://doi.org/10.5281/zenodo.4706662" httpUri="https://zenodo.org/record/4706662/files/figure.png" pageId="5" pageNumber="193">Fig. 8</figureCitation>
). Additionally,
<taxonomicName id="37964D03FB3EFF97FB805D96FB4730B8" authorityName="Quentin &amp; Villiers" authorityYear="1974" box="[1121,1204,402,429]" class="Insecta" family="Cerambycidae" genus="Schizodontus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="C2E2EA92FB3EFF97FB805D96FB4730B8" box="[1121,1204,402,429]" italics="true" pageId="5" pageNumber="193">S. latus</emphasis>
</taxonomicName>
has a smooth posterior pronotum, while both
<taxonomicName id="37964D03FB3EFF97FE215DB2FDB330C4" authorityName="Quentin &amp; Villiers" authorityYear="1974" box="[448,576,439,465]" class="Insecta" family="Cerambycidae" genus="Schizodontus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="angustus">
<emphasis id="C2E2EA92FB3EFF97FE215DB2FDB330C4" box="[448,576,439,465]" italics="true" pageId="5" pageNumber="193">S. angustus</emphasis>
</taxonomicName>
and
<taxonomicName id="37964D03FB3EFF97FD975DB2FC8D30C4" authority="Bouyer &amp; Drumont &amp; Roland &amp; Slootmaekers &amp; Pirkl &amp; Jocque, 2021" authorityName="Bouyer &amp; Drumont &amp; Roland &amp; Slootmaekers &amp; Pirkl &amp; Jocque" authorityYear="2021" box="[630,894,438,465]" class="Insecta" family="Cerambycidae" genus="Schizodontus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="mahimborondroensis" status="sp. nov.">
<emphasis id="C2E2EA92FB3EFF97FD975DB2FC8D30C4" box="[630,894,438,465]" italics="true" pageId="5" pageNumber="193">S. mahimborondroensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D9D157E9FB3EFF97FC645DB2FC2F30C5" box="[901,988,439,465]" pageId="5" pageNumber="193" rank="species">sp. nov.</taxonomicNameLabel>
have a small well-defined medial notch in the posterior pronotum edge. Also, the colouration and brightness of the elytra differs between the species
<emphasis id="C2E2EA92FB3EFF97FA8D5DDFFE53330C" italics="true" pageId="5" pageNumber="193">
with
<taxonomicName id="37964D03FB3EFF97FF765DFAFE53330C" authority="Bouyer &amp; Drumont &amp; Roland &amp; Slootmaekers &amp; Pirkl &amp; Jocque, 2021" authorityName="Bouyer &amp; Drumont &amp; Roland &amp; Slootmaekers &amp; Pirkl &amp; Jocque" authorityYear="2021" box="[151,416,510,537]" class="Insecta" family="Cerambycidae" genus="Schizodontus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="mahimborondroensis" status="sp. nov.">S. mahimborondroensis</taxonomicName>
</emphasis>
<taxonomicNameLabel id="D9D157E9FB3EFF97FE465DFAFE0C330D" box="[423,511,511,537]" pageId="5" pageNumber="193" rank="species">sp. nov.</taxonomicNameLabel>
dark brown compared to reddish brown in
<taxonomicName id="37964D03FB3EFF97FC095DFAFBC8330C" authorityName="Quentin &amp; Villiers" authorityYear="1974" box="[1000,1083,510,537]" class="Insecta" family="Cerambycidae" genus="Schizodontus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="C2E2EA92FB3EFF97FC095DFAFBC8330C" box="[1000,1083,510,537]" italics="true" pageId="5" pageNumber="193">S. latus</emphasis>
</taxonomicName>
(Fig. 4) and its elytra are more reflecting compared to both other species.
</paragraph>
<paragraph id="F0293680FB3EFF97FF765E6FFB5D3390" blockId="5.[151,1198,618,644]" box="[151,1198,618,644]" pageId="5" pageNumber="193">
<emphasis id="C2E2EA92FB3EFF97FF765E6FFEE33390" bold="true" box="[151,272,618,644]" pageId="5" pageNumber="193">The three</emphasis>
<taxonomicName id="37964D03FB3EFF97FEF65E6FFDE33390" box="[279,528,618,644]" class="Insecta" family="Cerambycidae" genus="Schizodontus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="C2E2EA92FB3EFF97FEF65E6FFE413390" box="[279,434,618,644]" italics="true" pageId="5" pageNumber="193">Schizodontus</emphasis>
<emphasis id="C2E2EA92FB3EFF97FE5B5E6FFDE33390" bold="true" box="[442,528,618,644]" pageId="5" pageNumber="193">species</emphasis>
</taxonomicName>
<emphasis id="C2E2EA92FB3EFF97FDF65E6FFB5D3390" bold="true" box="[535,1198,618,644]" pageId="5" pageNumber="193">can be identified with the following dichotomous key.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="B88C650BFB3EFF97FF765EAAFA6E3286" pageId="5" pageNumber="193" type="key">
<key id="840761B1FB3EFF97FF765EAAFA6E3286" pageId="5" pageNumber="193">
<keyStep id="4B622E25FB3EFF97FF765EAAFA6F3208" pageId="5" pageNumber="193">
<paragraph id="F0293680FB3EFF97FF765EAAFA6F33F6" blockId="5.[151,1437,687,914]" pageId="5" pageNumber="193">
<keyLead id="4B6795B5FB3EFF97FF765EAAFA6F33F6" pageId="5" pageNumber="193">
1 - Antennae with narrow articles (ratio length / width for antennomere III: 2.9); antennomere III with subparallel sides and without tooth at apical angles (
<figureCitation id="68AD2A05FB3EFF97FE3E5EC9FDEA33F7" box="[479,537,716,739]" captionStart="FIGURE 8" captionStartId="4.[152,235,1632,1656]" captionTargetBox="[306,1246,192,1563]" captionTargetId="figure-18@4.[305,1281,181,1612]" captionTargetPageId="4" captionText="FIGURE 8. First antennal segments of 1) Schizodontus mahimborondroensis sp. nov. (male), 2) S. latus (male), 3) S. angustus (male) and 4) S. latus (female). 24 after Quentin &amp; Villiers, 1975, figs. 227229." figureDoi="http://doi.org/10.5281/zenodo.4706662" httpUri="https://zenodo.org/record/4706662/files/figure.png" pageId="5" pageNumber="193">Fig. 8</figureCitation>
).............................................
<taxonomicName id="37964D03FB3EFF97FBDA5EC8FA6F33F6" ID-CoL="6Y8VZ" authority="Quentin &amp; Villiers, 1974" authorityName="Quentin &amp; Villiers" authorityYear="1974" box="[1083,1436,716,738]" class="Insecta" family="Cerambycidae" genus="Schizodontus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="angustus">
<emphasis id="C2E2EA92FB3EFF97FBDA5EC8FB5B33F6" box="[1083,1192,717,738]" italics="true" pageId="5" pageNumber="193">S. angustus</emphasis>
<bibRefCitation id="94074B71FB3EFF97FB4F5EC9FA6F33F6" author="Quentin, R. M. &amp; Villiers, A." box="[1198,1436,716,738]" pageId="5" pageNumber="193" pagination="249 - 341" refId="ref3434" refString="Quentin, R. M. &amp; Villiers, A. (1974) Revision des Closterini de Madagascar (Col., Cerambycidae Prioninae). Annales de la Societe Entomologique de France, 10, 249 - 341." type="journal article" year="1974">Quentin &amp; Villiers, 1974</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="F0293680FB3EFF97FF765EEFFA6F3208" blockId="5.[151,1437,687,914]" pageId="5" pageNumber="193">
<keyLead id="4B6795B5FB3EFF97FF765EEFFA6F3208" pageId="5" pageNumber="193">
- Antennae with wide articles (ratio length / width for antennomere 3: 2.12.4), antennomere III widened anteriorly with a small well defined apical external tooth (
<figureCitation id="68AD2A05FB3EFF97FDD15F02FD993209" box="[560,618,775,797]" captionStart="FIGURE 8" captionStartId="4.[152,235,1632,1656]" captionTargetBox="[306,1246,192,1563]" captionTargetId="figure-18@4.[305,1281,181,1612]" captionTargetPageId="4" captionText="FIGURE 8. First antennal segments of 1) Schizodontus mahimborondroensis sp. nov. (male), 2) S. latus (male), 3) S. angustus (male) and 4) S. latus (female). 24 after Quentin &amp; Villiers, 1975, figs. 227229." figureDoi="http://doi.org/10.5281/zenodo.4706662" httpUri="https://zenodo.org/record/4706662/files/figure.png" pageId="5" pageNumber="193">Fig. 8</figureCitation>
).................................................................. 2
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<paragraph id="F0293680FB3EFF97FF765F20FA6F3243" blockId="5.[151,1437,687,914]" pageId="5" pageNumber="193">
<keyLead id="4B6795B5FB3EFF97FF765F20FA6F3243" pageId="5" pageNumber="193">
2 - Prominent external apical tooth present only on antennomeres V or VI, anterior pronotum ridge smooth and uninterrupted, reddish brown elytra.............................................................
<taxonomicName id="37964D03FB3EFF97FB805F47FA6F3243" ID-CoL="6Y97Z" authority="Quentin &amp; Villiers, 1974" authorityName="Quentin &amp; Villiers" authorityYear="1974" box="[1121,1436,834,856]" class="Insecta" family="Cerambycidae" genus="Schizodontus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="C2E2EA92FB3EFF97FB805F47FB5B3243" box="[1121,1192,834,855]" italics="true" pageId="5" pageNumber="193">S. latus</emphasis>
<bibRefCitation id="94074B71FB3EFF97FB4F5F47FA6F3243" author="Quentin, R. M. &amp; Villiers, A." box="[1198,1436,834,856]" pageId="5" pageNumber="193" pagination="249 - 341" refId="ref3434" refString="Quentin, R. M. &amp; Villiers, A. (1974) Revision des Closterini de Madagascar (Col., Cerambycidae Prioninae). Annales de la Societe Entomologique de France, 10, 249 - 341." type="journal article" year="1974">Quentin &amp; Villiers, 1974</bibRefCitation>
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<keyLead id="4B6795B5FB3EFF97FF765F65FA6E3286" pageId="5" pageNumber="193">
- Prominent external apical tooth on antennomere III, anterior pronotum ridge centrally with a small notch, dark brown elytra...........................................................................
<taxonomicName id="37964D03FB3EFF97FB815F79FABE3286" ID-CoL="BLLQD" authority="Bouyer &amp; Drumont &amp; Roland &amp; Slootmaekers &amp; Pirkl &amp; Jocque, 2021" authorityName="Bouyer &amp; Drumont &amp; Roland &amp; Slootmaekers &amp; Pirkl &amp; Jocque" authorityYear="2021" box="[1120,1357,892,914]" class="Insecta" family="Cerambycidae" genus="Schizodontus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="mahimborondroensis" status="sp. nov.">
<emphasis id="C2E2EA92FB3EFF97FB815F79FABE3286" bold="true" box="[1120,1357,892,914]" italics="true" pageId="5" pageNumber="193">S. mahimborondroensis</emphasis>
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