plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/78/30/06/783006583E355573943A2BDCD6E08DEF.xml

628 lines
52 KiB
XML
Raw Permalink Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="3D06E5691CD243F4287A429A69BED373" ID-CLB-Dataset="304926" ID-DOI="10.3897/aiep.54.135448" ID-GBIF-Dataset="22888b0d-a602-4918-9cc1-db34e6998697" ID-ZooBank="1F507CB1-0C06-4381-9CBF-CB0A9FFC8263" ID-publisher-id="135448" URI-arpha="470541C0-0A45-53EE-8F14-A86CE57F3B99" URI-zoobank="http://zoobank.org/1F507CB1-0C06-4381-9CBF-CB0A9FFC8263" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" article-type="research-article" checkinTime="1730398432430" checkinUser="pensoft" docAuthor="Dewa, Yuna &amp; Motomura, Hiroyuki" docDate="2024" docId="783006583E355573943A2BDCD6E08DEF" docLanguage="en" docName="Acta Ichthyologica et Piscatoria 54: 261-268" docOrigin="Acta Ichthyologica et Piscatoria 54" docSource="https://aiep.pensoft.net/article/135448/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Enneapterygius olivaceus Dewa, Tashiro, et Motomura 2023" docType="treatment" docVersion="4" dtd-version="3.0" lastPageNumber="268" masterDocId="470541C00A4553EE8F14A86CE57F3B99" masterDocTitle="First records of two triplefins, Enneapterygius rhothion and Enneapterygius olivaceus (Actinopterygii: Blenniiformes: Tripterygiidae), from Australia and Vanuatu" masterLastPageNumber="268" masterPageNumber="261" pageNumber="261" updateTime="1730408107366" updateUser="admin">
<mods:mods id="BCE426AFEA54983ECBC58D3088485AC5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DD3F87D69990F493FD3D2B19E1810B0A">
<mods:title id="878B9419B54F5F9F7178D162C5D34AB8">First records of two triplefins, Enneapterygius rhothion and Enneapterygius olivaceus (Actinopterygii: Blenniiformes: Tripterygiidae), from Australia and Vanuatu</mods:title>
</mods:titleInfo>
<mods:name id="4A5B9A6F2DCF0ABFB87AE6068C00B966" type="personal">
<mods:role id="7D088B867C5A764F237949676173EBF0">
<mods:roleTerm id="FD1B49BC5F100A952DA1AEC2BEBC3954">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8EC0DB8DE0445E867BB63A2FCF4CD4AB">Dewa, Yuna</mods:namePart>
<mods:nameIdentifier id="08D0BF44ADFE7E0959D88D1B409DCD09" type="ORCID">0000-0002-8512-6782</mods:nameIdentifier>
<mods:affiliation id="3C3BF4E0FF84618299B4E4557A3C3016">United Graduate School of Agricultural Sciences, Kagoshima University, Kagoshima, Japan</mods:affiliation>
</mods:name>
<mods:name id="F5D2318867FC70B7D6DF6D1A357E78B2" type="personal">
<mods:role id="90D015F6FE8D5B5F724A80B8C8CDA6FE">
<mods:roleTerm id="4F982FAA3587C9FB08E008B43FCADB86">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="177B02E5937D69B1C4D6E78786C31382">Motomura, Hiroyuki</mods:namePart>
<mods:nameIdentifier id="3B9159AC42679E54AC3DE466852EB057" type="ORCID">0000-0002-7448-2482</mods:nameIdentifier>
<mods:affiliation id="B9EC9A561218B5E0FAD842A60031C8CD">Kagoshima University Museum, Kagoshima, Japan</mods:affiliation>
</mods:name>
<mods:typeOfResource id="ADADF6621CBBD0FBA849DFAB835EDEB1">text</mods:typeOfResource>
<mods:relatedItem id="BFCBD8A3B51BD44F77116C7C5B2E77CF" type="host">
<mods:titleInfo id="0F9517F9A7D79ED14BFF1A5EE1EE4576">
<mods:title id="632D1A1821E671411909DC24FD16F9FE">Acta Ichthyologica et Piscatoria</mods:title>
</mods:titleInfo>
<mods:part id="6F9F402DBFF0EB8059A22FFDAE44A7FD">
<mods:date id="AF3EAD5363724480FA10400308618B1A">2024</mods:date>
<mods:detail id="8C71B5206BA569EC865809CEAE9566BB" type="pubDate">
<mods:number id="9E2006D1D9A584D2A8B327C040143AF3">2024-10-31</mods:number>
</mods:detail>
<mods:detail id="525AF7CFB856F8E452B9C8EBB8FCEC6A" type="volume">
<mods:number id="AAFC2696D9F2542567A15EA3DDB9A034">54</mods:number>
</mods:detail>
<mods:extent id="9F5D0D379E52CD11614F89353BA7F8BA" unit="page">
<mods:start id="4D4D0101FE4FDD8C3507D75226F91538">261</mods:start>
<mods:end id="62F38B1089D54137B009FE1E6310317D">268</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="1252FBBC5F811DF39F6049FACFC307D5">journal article</mods:classification>
<mods:identifier id="6E7D993865E265A959AA2A6C8B53A361" type="CLB-Dataset">304926</mods:identifier>
<mods:identifier id="49C4CFD737BC90A2ACB6E5D57604253B" type="DOI">10.3897/aiep.54.135448</mods:identifier>
<mods:identifier id="8B6E58BC45E5A80FC420B3C7E9911B29" type="GBIF-Dataset">22888b0d-a602-4918-9cc1-db34e6998697</mods:identifier>
<mods:identifier id="7AB66202AA93387F595DB71DC3FA6ACA" type="ZooBank">1F507CB1-0C06-4381-9CBF-CB0A9FFC8263</mods:identifier>
</mods:mods>
<treatment id="783006583E355573943A2BDCD6E08DEF" ID-DOI="http://doi.org/10.5281/zenodo.14019629" ID-GBIF-Taxon="245302290" ID-Zenodo-Dep="14019629" ID-arpha="78300658-3E35-5573-943A-2BDCD6E08DEF" LSID="urn:lsid:plazi:treatment:783006583E355573943A2BDCD6E08DEF" httpUri="http://treatment.plazi.org/id/783006583E355573943A2BDCD6E08DEF">
<subSubSection id="181EE76E105FAC823401921FB9118DB8" type="nomenclature">
<paragraph id="A203708C3598E30DA059DC3030215DC0">
<taxonomicName id="FF0088179D0C03F3A223AB3C07C1C7FA" authority="Dewa, Tashiro, et Motomura, 2023" authorityName="Dewa, Tashiro, et Motomura" authorityYear="2023" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Blenniiformes" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="DA544D59F0EA88305B4A3DAE853D360C" italics="true">Enneapterygius olivaceus</emphasis>
Dewa, Tashiro, et Motomura, 2023
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="86A8346E63D8A70B0012E267A091CABB" type="description">
<paragraph id="6D0D13410ED9BBE290E371E97D4D778B">
<figureCitation id="08ED5B920C4859348C6DCA0CE51ECD60" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distributional map of Enneapterygius rhothion (red circles) and E. olivaceus (blue triangles). Closed and open symbols indicate localities determined during the presently reported study and previous records, respectively. Black arrows indicate type localities of the two species." figureDoi="10.3897/aiep.54.135448.figure3" httpUri="https://binary.pensoft.net/fig/1165074">English name: olive green triplefin Figs 3</figureCitation>
,
<figureCitation id="05CC5F455A2D3B0983EC22F7CF33E253" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Photographs of preserved specimens of Enneapterygius olivaceus (A: AMS I. 22732-008, male, 22.1 mm SL, Australia; B: AMS I. 6317, male, 19.7 mm SL, Vanuatu; C: AMS I. 30000-001, female, 18.0 mm SL, Australia)." figureDoi="10.3897/aiep.54.135448.figure4" httpUri="https://binary.pensoft.net/fig/1165075">4</figureCitation>
;
<tableCitation id="DD9A00F23D3430EDC1E379106CC57566" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Measurements and counts of Enneapterygius olivaceus collected from Australia and Vanuatu. Means in parentheses." httpUri="http://table.plazi.org/id/9974914A90050809A67863B109EC1861" tableUuid="9974914A90050809A67863B109EC1861">Table 2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUYAE" type="material">
<paragraph id="1CC1BA4B251C60F6D00EEBA12C81426D">
<heading id="A8F75517B1B3754EA6C83C936F33E935" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="346BFA28150588BB5D234DC7D1706C5B">
<specimenCount id="9E50E74594A91121FB3A9427560CDE97" count="20" type="generic">20 specimens</specimenCount>
(
<quantity id="023B5F1239D6D77B8B62CC3D4100F06C" metricMagnitude="-2" metricUnit="m" metricValue="1.71" metricValueMax="2.21" metricValueMin="1.21" unit="mm" value="17.1" valueMax="22.1" valueMin="12.1">12.1–22.1 mm</quantity>
SL).
<materialsCitation id="E80D1BE54D0BCD583619066A11F1297E" collectionCode="AMS" collectorName="D. Hoese" country="Australia" latitude="-15.3" location="Bird Islet" longLatPrecision="21" longitude="145.44972" municipality="Lizard Island" specimenCode="AMS I. 22732-008" specimenCount="1" specimenCount-male="1" stateProvince="Queensland">
<emphasis id="168A5C8929DA126CCD909B3B1731DA68" bold="true">
<collectingCountry id="665F5BB4B14D4B3AD79381775824319B" name="Australia">AUSTRALIA</collectingCountry>
</emphasis>
:
<specimenCode id="7F0FC464F1C41B79197910855AEBBF93">AMS I. 22732-008</specimenCode>
,
<specimenCount id="F007E538181E40DF355334A560712206" type="male">male</specimenCount>
,
<quantity id="7F8F838F69B6F51BD865087A3A2F2FBA" metricMagnitude="-2" metricUnit="m" metricValue="2.21" unit="mm" value="22.1">22.1 mm</quantity>
SL,
<location id="EBB04615AC8407B82FAFC6B46262C2C8" LSID="urn:lsid:plazi:treatment:783006583E355573943A2BDCD6E08DEF:EBB04615AC8407B82FAFC6B46262C2C8" country="Australia" latitude="-15.3" longLatPrecision="21" longitude="145.44972" municipality="Lizard Island" name="Bird Islet" stateProvince="Queensland">Bird Islet</location>
,
<collectingMunicipality id="C99BC7E651DFF85D63029B10BACF2057">Lizard Island</collectingMunicipality>
,
<collectingRegion id="976B67566A30CFBC283ACB5FBAA0CEBD" country="Australia" name="Queensland">Queensland</collectingRegion>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E6YAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[145.449722,-15.300000]}">
<geoCoordinate id="8AB917D10C6CF69ADADDB23151385368" degrees="15" direction="south" minutes="18" orientation="latitude" precision="15" seconds="00" value="-15.3">15°18′00″S</geoCoordinate>
,
<geoCoordinate id="B81DAA243241C073568780A2BC6B9439" degrees="145" direction="east" minutes="26" orientation="longitude" precision="15" seconds="59" value="145.44972">145°26′59″E</geoCoordinate>
</named-content>
, 1981,
<collectorName id="65DB26C794D0A81AA8D0EB47061F6FA8">D. Hoese</collectorName>
et al.
</materialsCitation>
;
<materialsCitation id="0C50DB57E48A49BF77C2254D53D64FF2" collectionCode="AMS" latitude="-15.33" location="Lizard Island" longLatPrecision="21" longitude="145.46971" specimenCode="AMS I. 30000-001" specimenCount="17" specimenCount-female="8" specimenCount-male="9">
<specimenCode id="6191E54F4D37B262099112E9AA60D988">AMS I. 30000-001</specimenCode>
,
<specimenCount id="DCE29C65207E69CF9751C45B6B7A4272" count="9" type="male">9 males</specimenCount>
and
<specimenCount id="59A9154B4A26DBE6F8FBCB3906CBE74F" count="8" type="female">8 females</specimenCount>
,
<quantity id="08E46B660938286BD744AC4BACD704BA" metricMagnitude="-2" metricUnit="m" metricValue="1.575" metricValueMax="1.94" metricValueMin="1.21" unit="mm" value="15.75" valueMax="19.4" valueMin="12.1">12.1–19.4 mm</quantity>
SL,
<location id="047E54AFA54D0720839505A6A0CE7633" LSID="urn:lsid:plazi:treatment:783006583E355573943A2BDCD6E08DEF:047E54AFA54D0720839505A6A0CE7633" latitude="-15.33" longLatPrecision="21" longitude="145.46971" name="Lizard Island">Lizard Island</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EHZAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[145.469722,-15.330000]}">
<geoCoordinate id="16E76361815EFE23D9B4DED9E9086CFA" degrees="15" direction="south" minutes="19" orientation="latitude" precision="15" seconds="48" value="-15.33">15°19′48″S</geoCoordinate>
,
<geoCoordinate id="ECE4D301551B0AA1B2CFD489340E46C2" degrees="145" direction="east" minutes="28" orientation="longitude" precision="15" seconds="11" value="145.46971">145°28′11″E</geoCoordinate>
</named-content>
, 1989
</materialsCitation>
.
<materialsCitation id="065520A511DC5BD7C292867CF89E9BC6" collectionCode="AMS" collectorName="W. A. Haswell" country="Vanuatu" location="Vila" municipality="Efate" specimenCode="AMS I. 6317" specimenCount="4" specimenCount-male="1">
<emphasis id="A44F4FADBB2047381701A9A1F0BB8D02" bold="true">
<collectingCountry id="91132BF0AE21979536A831144A4087F4" name="Vanuatu">VANUATU</collectingCountry>
</emphasis>
:
<specimenCode id="B38B995561E344FD6A2416DA752DEAF8">AMS I. 6317</specimenCode>
, 1 of
<specimenCount id="F29F45129161B7532F613C0A2F8AE9FB" count="3" type="generic">3 specimens</specimenCount>
,
<specimenCount id="7175191079B0EF40D7D9E343575D0E02" type="male">male</specimenCount>
,
<quantity id="A3D62FAA3A9B91A5EFB7202161895D3E" metricMagnitude="-2" metricUnit="m" metricValue="1.97" unit="mm" value="19.7">19.7 mm</quantity>
SL,
<location id="F44CCFBEA23301BD50A525A06BA79FA5" LSID="urn:lsid:plazi:treatment:783006583E355573943A2BDCD6E08DEF:F44CCFBEA23301BD50A525A06BA79FA5" country="Vanuatu" municipality="Efate" name="Vila">Vila</location>
,
<collectingMunicipality id="1F73C6A35D18CFF3D3F76536962C31FF">Efate</collectingMunicipality>
, 1903,
<collectorName id="763C2E0B9575803F5581BEA633B1D426">W. A. Haswell</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="9632F904B135D135D6AF079AC1FE47D3" collectionCode="AMS" collectorName="W. A. Haswell" location="Vanua Lava" municipality="Banks Islands" specimenCode="AMS I. 6451" specimenCount="4" specimenCount-female="1">
<specimenCode id="65A81B6C740511B5645ED82F79201C9F">AMS I. 6451</specimenCode>
, 1 of
<specimenCount id="7324DFC68019F6A6A708D6B50B55E6F2" count="3" type="generic">3 specimens</specimenCount>
,
<specimenCount id="80E00BE1E325430399A9B94222C4C3DF" type="female">female</specimenCount>
,
<quantity id="8EE8C1F7BA813301550DBAD27D366FDE" metricMagnitude="-2" metricUnit="m" metricValue="1.59" unit="mm" value="15.9">15.9 mm</quantity>
SL,
<location id="6DE8F4363BF937EBDEB6B90E713EA844" LSID="urn:lsid:plazi:treatment:783006583E355573943A2BDCD6E08DEF:6DE8F4363BF937EBDEB6B90E713EA844" municipality="Banks Islands" name="Vanua Lava">Vanua Lava</location>
,
<collectingMunicipality id="80F0E82A3E29C1974456DDEF2CC9E0D8">Banks Islands</collectingMunicipality>
, 1903,
<collectorName id="54433CEA1CB686CF60F4517BDF3D76FF">W. A. Haswell</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EOZAE" type="description">
<paragraph id="9C9BC0A0D68179C21ADE1806AC63FECD">
<heading id="AF1694CD5B11E0E44A1127F6F119461D" reason="title">Description.</heading>
</paragraph>
<paragraph id="17C90B8989BF63F556449DD7B17953D7">
Counts and measurements given in Table
<tableCitation id="AEA0355DBC787954A6A61484D4082210" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Measurements and counts of Enneapterygius olivaceus collected from Australia and Vanuatu. Means in parentheses." httpUri="http://table.plazi.org/id/9974914A90050809A67863B109EC1861" tableUuid="9974914A90050809A67863B109EC1861">2</tableCitation>
. Body moderately elongate, slightly compressed anteriorly, progressively more compressed posteriorly. Dorsal profile of snout straight, moderately steep. Mouth slightly oblique; posterior margin of maxilla reaching to or extending slightly beyond anterior margin of pupil; anterior tip of upper jaw slightly above level of lower margin of orbit (lateral view). Anterior nostril forming membranous tube with short unbranched tentacle, base at level of middle of eye, slightly closer to eye than to upper lip; posterior nostril opening circular, without membranous tube. Eye oriented dorsolaterally; broad leaf-like tentacle on posterodorsal margin, its length longer than nasal tentacle. Interorbital space narrow, its width less than pupil diameter. Opercular margin slightly pointed, reaching to below base of 2 nd or 3 rd spine of first dorsal fin.
</paragraph>
<caption id="9974914A90050809A67863B109EC1861" ID-Table-UUID="9974914A90050809A67863B109EC1861" httpUri="http://table.plazi.org/id/9974914A90050809A67863B109EC1861" startId="T2" targetIsTable="true">
<paragraph id="5D55D354BE56E705D3E96CDC1643D429">
<label id="DFFD8A5BD784AAACB729F0A20B597870">Table 2.</label>
</paragraph>
<paragraph id="1D74B215ACA8895FFAC161E31026788D">
Measurements and counts of
<taxonomicName id="0FD7979D68700D158C4F3351DE30ACE1" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="6034B2340F7964C2A4F2DE21A22CE63A" italics="true">Enneapterygius olivaceus</emphasis>
</taxonomicName>
collected from Australia and Vanuatu. Means in parentheses.
</paragraph>
</caption>
<paragraph id="5802D35B67BB4255358A09ADBB8C992A">
<table id="TID0EQKAG" rules="all">
<tbody id="BB8120237A587D2490C0A87AA0BEA36E">
<tr id="5003000B6A8E21B8AF945DDD2A1B3904">
<th id="6789E6BD1C910C984F4837B6AF7F71ED" colspan="1" rowspan="3">Character</th>
<th id="BC254C9F668EEDD9B8447F1D29527982" colspan="1" rowspan="1">Australia</th>
<th id="E6C7CDC196EE17893F67395AFAEC016A" colspan="2" rowspan="1">Vanuatu</th>
</tr>
<tr id="99BEF110D8695724FEFACAE9FC047109">
<th id="2C6438BCB23BA27E891AF3698F8F6890" colspan="1" rowspan="1">
<emphasis id="885F6E3DA3520AF44FBBF338BC39AAFA" italics="true">n</emphasis>
= 18
</th>
<th id="F94E1047CCD48370F97EE5EDB3ACEAE1" colspan="1" rowspan="1">AMS I. 6317</th>
<th id="60072B78C720135C4680D47B57B4282B" colspan="1" rowspan="1">AMS I. 6451</th>
</tr>
<tr id="B9EDB56591E443FDD7B8394992E9665E">
<th id="4B8D89DB7664D489AAEC02BD8996A440" colspan="1" rowspan="1">Males and females</th>
<th id="F08BEC3D33FF374C8E4A35F3E53E7F89" colspan="1" rowspan="1">Male</th>
<th id="986C74888D723AA89D958EB9861FE6F3" colspan="1" rowspan="1">Female</th>
</tr>
<tr id="244CD3ACFAF6DBACCF1FD06B9D04604B">
<td id="974A7FB07228FA188BACF1DC0EE85551" colspan="4" rowspan="1">
<emphasis id="1DD5D26E68AD6862C120C1B30D8A2942" bold="true">Morphometric data; absolute value [mm</emphasis>
]
</td>
</tr>
<tr id="0F220131A2443AD0CD7B49943A88C1AC">
<td id="B43C1ACF940C79DE43CD50D29EC28F18" colspan="1" rowspan="1">Standard length (SL)</td>
<td id="E72C0221901345634771FC0077E675C2" colspan="1" rowspan="1">12.1–19.4</td>
<td id="547B78DD50D1D61B10ABE5CEA3711A38" colspan="1" rowspan="1">19.7</td>
<td id="39C9146CB14C054607F90DE18DB80EAD" colspan="1" rowspan="1">15.9</td>
</tr>
<tr id="7C87A9D82B16BB632304FB4CE52464FF">
<td id="6CB4E2E20E2F2995774EE9D112BD8DE0" colspan="4" rowspan="1">
<emphasis id="10F1A3577010B8EEB71C9A2F1F22E2ED" bold="true">Morphometric data; relative values [% SL</emphasis>
]
</td>
</tr>
<tr id="193697C917EA2E02990796137BEDACBA">
<td id="EAEF898D0C2553F56691C2DDDE3CFB10" colspan="1" rowspan="1">Body depth</td>
<td id="48CDACBDE833420EC870257D1B76B58C" colspan="1" rowspan="1">20.0–22.1 (21.2)</td>
<td id="DE69285D75AEF09D6DE1415DD363CB3A" colspan="1" rowspan="1">23.5</td>
<td id="91DD48D4AF18E9A00EBE78F6B11B53F4" colspan="1" rowspan="1">20.8</td>
</tr>
<tr id="5F0455D6A9E5FF611072F2F52ECF9251">
<td id="8B3EB333CA71D1DF0F7A3E7BB0493CFC" colspan="1" rowspan="1">Body width</td>
<td id="CCD24A9132841B3902364A4095531535" colspan="1" rowspan="1">17.4–20.5 (19.2)</td>
<td id="58E013C97163226AA4A5C284858FEB1F" colspan="1" rowspan="1">21.4</td>
<td id="FC9AA98942D3DDF424EE4628398BFA13" colspan="1" rowspan="1">19.4</td>
</tr>
<tr id="971277E6618A95983493F0776A116F96">
<td id="FB0D5FAF47E142A3FF57986CEBBBF2B7" colspan="1" rowspan="1">Head length</td>
<td id="10AA6D6104FA2766678A0CCEB9505160" colspan="1" rowspan="1">28.9–31.6 (30.4)</td>
<td id="67BC94D72B4FA33718EE2136E572B29B" colspan="1" rowspan="1">32.3</td>
<td id="F6339BD4BEF3FDCD7BB9333ED12D8CBB" colspan="1" rowspan="1">31.9</td>
</tr>
<tr id="6D83EAB67BC34F628EA8E447A5A46DC5">
<td id="A3C28A505C25B97D6B2708780B55A6B6" colspan="1" rowspan="1">Snout length</td>
<td id="44DB724721809B5133E16B79F320D97A" colspan="1" rowspan="1">7.8–12.3 (10.4)</td>
<td id="B4BF798EDC02D4F3FAAE9296F6002413" colspan="1" rowspan="1">12.3</td>
<td id="2739D25DDBF2773B04CE4BE38B43F948" colspan="1" rowspan="1">13.6</td>
</tr>
<tr id="2C2D232FEFAD70EF0AFB05A4C4AAAA19">
<td id="EBDAA48A1B4FBBD81DEACFFEC9D7F094" colspan="1" rowspan="1">Orbit diameter</td>
<td id="281FD55E85FB889C42297E790096A93B" colspan="1" rowspan="1">8.2–11.0 (9.6)</td>
<td id="4B3B2C4C152FE241130FC71EC3415887" colspan="1" rowspan="1">9.1</td>
<td id="96322C593D13E8BD25971FD758A4FA1D" colspan="1" rowspan="1">10.4</td>
</tr>
<tr id="674DA5C92DDDB53CE2E12869E1353A53">
<td id="3BC99D32CAD2F4358350C75F29B77C00" colspan="1" rowspan="1">Interorbital width</td>
<td id="40F01B8D264F99918D9ED89C003DD76C" colspan="1" rowspan="1">1.2–2.4 (1.9)</td>
<td id="9892837333A95665EA4129643288A8F5" colspan="1" rowspan="1">2.0</td>
<td id="EE9570B874DD3598D049BC2B2462FD76" colspan="1" rowspan="1">1.9</td>
</tr>
<tr id="BEC313B89AB6D5CFCF3ED3DBB7130BE5">
<td id="12C21C16E406A14A18D346E87794F218" colspan="1" rowspan="1">Upper-jaw length</td>
<td id="7C424504B9E246E1D17D271C7E58DB71" colspan="1" rowspan="1">9.3–12.0 (10.5)</td>
<td id="D70479888AD6BB464CBEAB81D6D83E17" colspan="1" rowspan="1">11.8</td>
<td id="7034A3D2FA048176E0929289718578A7" colspan="1" rowspan="1">11.4</td>
</tr>
<tr id="D0C21D38763FD02F4CD3C1F48C203757">
<td id="BB5AE49DD33063929B202A4458E66028" colspan="1" rowspan="1">Postorbital length</td>
<td id="F9FEC233BADBADDBB2AEBB16763351D7" colspan="1" rowspan="1">12.0–14.6 (13.0)</td>
<td id="ED07E315DEF11C7BC9AD953CE6561815" colspan="1" rowspan="1">12.7</td>
<td id="C0F2625F4699DE873C692992488046D2" colspan="1" rowspan="1">11.5</td>
</tr>
<tr id="ECA44F4A36B103154C487559983B4DF2">
<td id="618CD7C5086D7284B20722906F0C66D2" colspan="1" rowspan="1">Pre- 1 st-dorsal-fin length</td>
<td id="96C0F6C298C492EF64ED95BD348B1D52" colspan="1" rowspan="1">23.7–27.3 (25.3)</td>
<td id="559D4605A9FB7320BB089222440D01B9" colspan="1" rowspan="1">26.2</td>
<td id="7DA1A79180FB2FDCEC505CAE907680C7" colspan="1" rowspan="1">27.5</td>
</tr>
<tr id="C1A1A90DE15E57931C9DB18FC5ACF843">
<td id="98B44ABF70F107858EAE0E27310D3535" colspan="1" rowspan="1">Pre- 2 nd-dorsal-fin length</td>
<td id="89A131AA14DA978733180B0EB8296655" colspan="1" rowspan="1">33.2–38.3 (36.5)</td>
<td id="1211FFF2C5B10780E004275194838200" colspan="1" rowspan="1">37.9</td>
<td id="0223242986EF08EB93A0D81068830F6F" colspan="1" rowspan="1">38.2</td>
</tr>
<tr id="5D54A6BB89A1826993209F9D810ED6FA">
<td id="A100DAD3BABE5B390213C8A7BF1F45BF" colspan="1" rowspan="1">Pre- 3 rd-dorsal-fin length</td>
<td id="490583E4035B46285A4B25686E4328E1" colspan="1" rowspan="1">68.5–73.3 (70.9)</td>
<td id="0AB7EDF9542C196DEDF05E86C93BE628" colspan="1" rowspan="1">73.8</td>
<td id="FD222DFE39BAF84CCEA913CEC61A44BF" colspan="1" rowspan="1">74.4</td>
</tr>
<tr id="65ABD5A5AB6CFED54659572AA2519CEC">
<td id="C0A99178987403E2BA48AEA9D81A71C9" colspan="1" rowspan="1">Pre-anal-fin length</td>
<td id="9B7BF6A414EFCB9251D2727B5B2B44F4" colspan="1" rowspan="1">45.7–53.9 (51.3)</td>
<td id="97BD6EAFE7C74DD21C40EA14E4094112" colspan="1" rowspan="1">54.1</td>
<td id="FACDA524AC6EF791E74E1D6D0489C214" colspan="1" rowspan="1">51.0</td>
</tr>
<tr id="FE074FA08E72C14494F381E049FD1CB5">
<td id="B8C588F14678A50F414182D310F3E539" colspan="1" rowspan="1">Anal-fin base length</td>
<td id="BEB4479AC5F83A174BBA7C2C14FE2B18" colspan="1" rowspan="1">38.2–44.0 (41.2)</td>
<td id="6D0795511D3CB9F7FBB163B8F28B46C4" colspan="1" rowspan="1">40.4</td>
<td id="85307B32091DECF6436DC9EC2861DC6C" colspan="1" rowspan="1">38.8</td>
</tr>
<tr id="B9D6159206CBA7460FB27E4D87DB8464">
<td id="6009CFC46364DEAE1724EC133F22183F" colspan="1" rowspan="1">Pre-pectoral-fin length</td>
<td id="9EC9795FE5D1533BA67E0A652208035B" colspan="1" rowspan="1">28.9–35.5 (33.4)</td>
<td id="51CCF43F632577F8244305C6E16B2FE4" colspan="1" rowspan="1">35.6</td>
<td id="68A476136F243FE3EA20D9A6EA3DFCB3" colspan="1" rowspan="1">34.4</td>
</tr>
<tr id="14823B788976097E7D2F8DAF1EDF0785">
<td id="33D37EEE8FBDDC07D5E40154B8F83311" colspan="1" rowspan="1">Pre-pelvic-fin length</td>
<td id="8F7C151210C4590D979ADE4839ACC2D0" colspan="1" rowspan="1">23.6–27.2 (25.6)</td>
<td id="F5A76B6FF010DFE252B45D8FAEB601C6" colspan="1" rowspan="1">28.4</td>
<td id="7ADC767A46C2CF1D3984209BC536CE3F" colspan="1" rowspan="1">25.1</td>
</tr>
<tr id="CC3D743CF9678040E61DA81810BFE27E">
<td id="889E978A7A42FBCE88244A404E6178EE" colspan="1" rowspan="1">Caudal peduncle length</td>
<td id="D36C96E13B2AF60276BAD425B94CE749" colspan="1" rowspan="1">9.1–13.7 (11.8)</td>
<td id="BCEFEB344EAAF16FBF197BCBEFA848E0" colspan="1" rowspan="1">13.5</td>
<td id="8D575AD5DD61AD0CDA99A72C7C384F70" colspan="1" rowspan="1">11.0</td>
</tr>
<tr id="AE6811BAFC2D5DF12AD739F5624BF400">
<td id="7F0EF15C11D1D3989F68DA75E8637E3C" colspan="1" rowspan="1">Caudal peduncle depth</td>
<td id="58DAD89E7F75C0FF5BAAF79B743C0249" colspan="1" rowspan="1">7.1–9.2 (8.5)</td>
<td id="2A46E02723D31D0747FA758F815D1385" colspan="1" rowspan="1">8.6</td>
<td id="8E23A8C1061D3CA85DF8C57B6F5136BF" colspan="1" rowspan="1">8.3</td>
</tr>
<tr id="91488F7E6C1C276A62547B7516B936D6">
<td id="52A8F2F08AA2662CF65F5EF1D3EE8210" colspan="1" rowspan="1">1 st spine length of 1 st dorsal fin</td>
<td id="F9488168E002CD65EA32E738B3F7D9C3" colspan="1" rowspan="1">11.0–15.6 (13.2)</td>
<td id="5F630C128BDC4415FB6587251024AED1" colspan="1" rowspan="1">18.2</td>
<td id="62FA27CEBE3130A7DCC21046FE1B6BFF" colspan="1" rowspan="1">12.6</td>
</tr>
<tr id="A815350B1A5DF34D466B5B3A7787DCDC">
<td id="80AA47231A3C833FF25EA9840796844B" colspan="1" rowspan="1">2 nd spine length of 1 st dorsal fin</td>
<td id="8C3B0BF380410D123D239F2A125AC3DE" colspan="1" rowspan="1">8.6–11.8 (10.6)</td>
<td id="8ACAEEE3CBD095CB2705CB6B80DD8CE3" colspan="1" rowspan="1">14.3</td>
<td id="6B494E8CFCBDAEEEAD9942603CDBF943" colspan="1" rowspan="1">10.7</td>
</tr>
<tr id="28441D471BA3DED28635ECD934BFEA5B">
<td id="D449B28116C96DA197D6ED17644D95F4" colspan="1" rowspan="1">3 rd spine length of 1 st dorsal fin</td>
<td id="F0774F737F4B501C92B9A2E6E28A3268" colspan="1" rowspan="1">6.8–9.9 (8.8)</td>
<td id="CF33F2015DE4E80CBD1B26E8A24F8A21" colspan="1" rowspan="1">11.2</td>
<td id="94AFC980E474C49F7C29114F2EEC619B" colspan="1" rowspan="1">Damaged</td>
</tr>
<tr id="F43E5F1304A52168461C7A812258A0EC">
<td id="C5F4FF6624E79C8FB443583017AF81DF" colspan="1" rowspan="1">1 st dorsal-fin base length</td>
<td id="1956C4FC27F9B8FF748E2D0F32F690B2" colspan="1" rowspan="1">4.3–6.8 (5.2)</td>
<td id="D68466C136E997BFBC28A487643BB4A4" colspan="1" rowspan="1">5.6</td>
<td id="21370C7C29287F0F4EA91851359438BE" colspan="1" rowspan="1">5.0</td>
</tr>
<tr id="57223185976236E8CC908921BC817622">
<td id="B84C8A844A307EB862E3824F3FBBCAD9" colspan="1" rowspan="1">1 st spine length of 2 nd dorsal fin</td>
<td id="C2328573BE6CE85C72D5C359AE284706" colspan="1" rowspan="1">11.9–18.5 (13.5)</td>
<td id="F48EEC0C6247555707C35386349A05E2" colspan="1" rowspan="1">14.8</td>
<td id="77B6C8AB25AC5E32BAA872CC28DAFFD4" colspan="1" rowspan="1">Damaged</td>
</tr>
<tr id="D8D69C77E510C0450D4A8D2BDA5FAA86">
<td id="CB3FDA5172C1D89816CC1F3260200B8B" colspan="1" rowspan="1">2 nd spine length of 2 nd dorsal fin</td>
<td id="512C4EB8010CAF5F2EB2CEEE1D2A401B" colspan="1" rowspan="1">12.6–16.6 (14.5)</td>
<td id="1D605B14F2320BE48B7DB2C8C9F229D7" colspan="1" rowspan="1">16.6</td>
<td id="DAF0B19D9EC22B3C0C6C421588C1EB40" colspan="1" rowspan="1">Damaged</td>
</tr>
<tr id="1CD9BD8AD22BDCCB3C446769C4BFAC52">
<td id="A3DBB9E393BC9224C62D47ED8A29ABA9" colspan="1" rowspan="1">3 rd spine length of 2 nd dorsal fin</td>
<td id="F99278B07FF6BE7367E23002F826ADC4" colspan="1" rowspan="1">10.3–18.6 (14.6)</td>
<td id="1B1FC178D017E7FE5F9D569F951515B0" colspan="1" rowspan="1">14.9</td>
<td id="D7CA5944B61F90D7E68E3C3F03C53364" colspan="1" rowspan="1">Damaged</td>
</tr>
<tr id="C3A8769439080E57012336F77788D6EF">
<td id="54500FBBB758256C2B3ADD74DC65171C" colspan="1" rowspan="1">2 nd dorsal-fin base length</td>
<td id="CCD24D4AD7C1506142F57DE6B31FAE77" colspan="1" rowspan="1">25.5–30.9 (28.8)</td>
<td id="44EBEEF208C9D860F1B044089F802FC6" colspan="1" rowspan="1">29.4</td>
<td id="3088A4A4B8387E34F86EA4F52F2AAF61" colspan="1" rowspan="1">30.8</td>
</tr>
<tr id="9C2D7CCDA793AB94225CA2E742BF2278">
<td id="FD999A8B94BC60C335BC12008705CBFC" colspan="1" rowspan="1">1 st ray length of 3 rd dorsal fin</td>
<td id="2221D42D84F69879EAD88DFE85E319B0" colspan="1" rowspan="1">13.1–19.6 (16.1)</td>
<td id="A97B2B9C1BC31FE6499DB6ECCA4C9499" colspan="1" rowspan="1">18.6</td>
<td id="FF78E1E84F4DC7A7C2EE09A64591EC51" colspan="1" rowspan="1">15.8</td>
</tr>
<tr id="D586852F66E4F949DAB5C7CD99159CDF">
<td id="E7727830997E311A1B01460EB6D1C66A" colspan="1" rowspan="1">2 nd ray length of 3 rd dorsal fin</td>
<td id="93F8B696C9E007C9DDC90F71BA7DA64A" colspan="1" rowspan="1">13.4–18.2 (15.9)</td>
<td id="58BFA61E1F4909652F5C58B5C03E5D80" colspan="1" rowspan="1">17.6</td>
<td id="1772B665F2C339DC5A44816153B34783" colspan="1" rowspan="1">12.6</td>
</tr>
<tr id="6F11BB3FCC54ECAE86D779CFF5941AA0">
<td id="7EFE9D6B0B7508BACB4548003A9B285B" colspan="1" rowspan="1">3 rd ray length of 3 rd dorsal fin</td>
<td id="8C75331370970AB74BC3367258AF6291" colspan="1" rowspan="1">13.6–17.2 (15.4)</td>
<td id="61C15DA49DB7E31F259DA8663065A5D3" colspan="1" rowspan="1">15.5</td>
<td id="8D9A510DCD825DDFB5B87629DC4D1AC0" colspan="1" rowspan="1">Damaged</td>
</tr>
<tr id="11EC233BD6E2C104DADFD8F466788392">
<td id="ABDD820B2D8E5B0AE1D41A02A7C10BD7" colspan="1" rowspan="1">3 rd dorsal fin base length</td>
<td id="5056065B7063907B1177AC52B27FF6F4" colspan="1" rowspan="1">13.9–19.5 (17.0)</td>
<td id="C3776E08F2C062342237D832AC6C7F69" colspan="1" rowspan="1">17.3</td>
<td id="0C193A8D70C2107532BEB3C6363D4485" colspan="1" rowspan="1">15.1</td>
</tr>
<tr id="EA7C0191865EDDAC1038773110DE5C36">
<td id="1290E466E2FF1F815D27F24E00F53376" colspan="1" rowspan="1">Pectoral-fin length</td>
<td id="6911958A109C47B360DBCDEB73A2F357" colspan="1" rowspan="1">27.6–34.9 (32.2)</td>
<td id="7FF80C3A07BC0DB112CA5EEF6A43D34B" colspan="1" rowspan="1">30.9</td>
<td id="5A041BC41B450172541F975D84974216" colspan="1" rowspan="1">31.7</td>
</tr>
<tr id="A2C8B19DCAE38D5125E671B048F5FA4B">
<td id="5F68B3724E1BBEF7C7BA9A852E92DB36" colspan="1" rowspan="1">1 st ray length of pelvic fin</td>
<td id="123E3FEAA4C4A16AF2690FE940BFF1BF" colspan="1" rowspan="1">13.4–18.3 (15.6)</td>
<td id="6B29FED5FBE6F17FC4A9218A742C1DB4" colspan="1" rowspan="1">19.1</td>
<td id="A46D3CC34CE9C2036E3D1C6BC0D2E8D4" colspan="1" rowspan="1">16.3</td>
</tr>
<tr id="36BB05D729527DB4D73861F736DBFE30">
<td id="AD417521C0B14F7C47003FD39BC52DF4" colspan="1" rowspan="1">2 nd ray length of pelvic fin</td>
<td id="574A8474C7335CE68D7D4D03A2381D67" colspan="1" rowspan="1">20.6–26.5 (24.0)</td>
<td id="A7F8AA7D9569846D8FD5954743DB3F90" colspan="1" rowspan="1">30.4</td>
<td id="F0E74F78576124C7A86F2EE0817566C3" colspan="1" rowspan="1">23.9</td>
</tr>
<tr id="328B622038C984B02FD59FED3C8F3C54">
<td id="9B5A836B57087E44BEE481F814E3C245" colspan="4" rowspan="1">
<emphasis id="5DB8DD6869D01DD71B3EB4A7CC34BABD" bold="true">Meristic data (counts)</emphasis>
</td>
</tr>
<tr id="884C0884EDEF4C73A86883FF7AEDC044">
<td id="F98359D616993850CD1BE8E260B32031" colspan="1" rowspan="1">Dorsal-fin rays</td>
<td id="04784808665104055BC79587170E6016" colspan="1" rowspan="1">III, XI – XIII, 8–10</td>
<td id="C9BAC2438EF0B5B92143E41C99804B43" colspan="1" rowspan="1">III, XIII, 9</td>
<td id="C02139824B139997AB11483355DFBEAD" colspan="1" rowspan="1">III, XIII, 8</td>
</tr>
<tr id="DF4A5442845E25C7DC712A353FB9EDAF">
<td id="B55587D30242C6C5573DF0B2DC075D25" colspan="1" rowspan="1">Anal-fin rays</td>
<td id="2C0928DE33C38D52E0ADBBFE44B71308" colspan="1" rowspan="1">I, 16 or 17</td>
<td id="43B40B7FFF86D9858EB5CB12BDF7860D" colspan="1" rowspan="1">I, 16</td>
<td id="934384D01655C094F14111B4FB3E1C8D" colspan="1" rowspan="1">I, 16</td>
</tr>
<tr id="9BDE73AEE2A47E6B42445291E411FD2A">
<td id="D8053BE5A9E0AA0B98D47F1DDCB88219" colspan="1" rowspan="1">Pectoral-fin rays</td>
<td id="38352835E744937991DFD52B2ADB59C5" colspan="1" rowspan="1">iii – v + 3–5 + vi – viii = 14–16</td>
<td id="328A79C2B92A63B28B1F9DDE68F92546" colspan="1" rowspan="1">3 + damaged = 15</td>
<td id="5C5E5C73BE29B11831F479FFA63C731F" colspan="1" rowspan="1">iv + 4 + vii = 15</td>
</tr>
<tr id="314CD4C41CB167D5402CE68676EDA6F8">
<td id="5611DF5EBC0F96ECFDFDCF89DB3ECE8B" colspan="1" rowspan="1">Scale rows in longitudinal series</td>
<td id="82ECC5697766B224BE52B03A46F6154E" colspan="1" rowspan="1">29–32</td>
<td id="780679A54D722685603E629672EB653C" colspan="1" rowspan="1">29</td>
<td id="449CFAE631B8AE31432181770D5EA0ED" colspan="1" rowspan="1">30</td>
</tr>
<tr id="FBF406B741EF382F8CADDE098C3236C9">
<td id="4F9A2EA552A0387651EE3C3A91D0EFFE" colspan="1" rowspan="1">Pored lateral-line scales</td>
<td id="1D4C7AFAF1389EE4CB622AFE2031ADB4" colspan="1" rowspan="1">12–14</td>
<td id="5BE3F74BA32C9BA31870EA6B94B5F671" colspan="1" rowspan="1">13</td>
<td id="70E3AD09E6D9511DDB3FFAEC14B01101" colspan="1" rowspan="1">12</td>
</tr>
<tr id="90E0D6B1E39D588A5173B7A6A45AD2A6">
<td id="E68C1555AC0D86C5F0BC0C71A6B4888D" colspan="1" rowspan="1">Notched lateral-line scales</td>
<td id="B1A03ABAAE3CA0229DDE5580EB968049" colspan="1" rowspan="1">18–21</td>
<td id="B2DB7BC5F8FDA4500B3582410CEB5371" colspan="1" rowspan="1">20</td>
<td id="DE4264839D4ADB425BC5F2686CB2EC17" colspan="1" rowspan="1">19</td>
</tr>
<tr id="04AC1D803DAF2611FFC75BD78D11D203">
<td id="4C8EA15C132E1D85D8C5704A95975FD6" colspan="1" rowspan="1">Scales above of 1 st PLL</td>
<td id="AADF34DF1776FE5BD55AD4F3CABCF7C0" colspan="1" rowspan="1">2 (rarely 1)</td>
<td id="30AFE3D4A5C39F3243C6E5A036433A6D" colspan="1" rowspan="1">2</td>
<td id="DDBC5722C50C85264F9FE3352C3B8183" colspan="1" rowspan="1">2</td>
</tr>
<tr id="8F13B8D8868F70FD17712341F29A91BD">
<td id="0C9D252020ACBB071AF4936E9D807219" colspan="1" rowspan="1">Scales below 2 nd dorsal fin</td>
<td id="0ED34AF4890AE13257610BAA83876B49" colspan="1" rowspan="1">2–3</td>
<td id="A58F701D002EC057F4B48DEA73769A84" colspan="1" rowspan="1">2 ½</td>
<td id="4A3E2BA07B63DD94313AF063AE04FA68" colspan="1" rowspan="1">2 ½</td>
</tr>
<tr id="1D0E07D939A0F6739808C859E169A19B">
<td id="300B7B9C89D1C234836720EC8783AB05" colspan="1" rowspan="1">Scales below of 1 st NLL</td>
<td id="0876A8A6120E68A67D3D95FEC6A9728D" colspan="1" rowspan="1">2 or 3</td>
<td id="E420573E9586093339BB5109EE810608" colspan="1" rowspan="1">3</td>
<td id="CC0A31C6C55CBE8FCF8847667B4E47FE" colspan="1" rowspan="1">2</td>
</tr>
<tr id="F90A7895CC3499EA645D43B2CBA28077">
<td id="3AC3A6D33532E057A68864FFDEB828C4" colspan="1" rowspan="1">Caudal peduncle scales</td>
<td id="6DB03E9E9CFC1C6C538E94B4E324D76D" colspan="1" rowspan="1">8</td>
<td id="EF4DE745215B222E5E8ED0B1187A9275" colspan="1" rowspan="1">8</td>
<td id="DDD630FD7AC78779DB3C664ACEC16361" colspan="1" rowspan="1">8</td>
</tr>
<tr id="DBCD0E76F727C76C7F8D4B7A90CD0966">
<td id="FB916F042A44A9BF8AD516E54147B9CB" colspan="1" rowspan="1">Mandibular pore formula</td>
<td id="3574E18D00FB7BC5B0586533C3187B30" colspan="1" rowspan="1">3 + 1 + 3</td>
<td id="E00D83F88E6E17876404D13568EFE376" colspan="1" rowspan="1">3 + 1 + 3</td>
<td id="61EC4D10616BC225853323FE6AF0BAF2" colspan="1" rowspan="1">3 + 1 + 3</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="EAACF4FA7E5DE1FD059D6DB21F22DF9A">Lateral line discontinuous, with anterior series of pored scales and posterior series of notched scales; pored scale series ending below membrane between 9 th and 10 th spines of second dorsal fin; notched scale series beginning below second scale from last pored scale, ending at caudal-fin base; Body scales ctenoid; scales absent on head, including maxilla, interorbital space, preopercle and opercle, and pectoral-fin base, undersurface of head, abdomen and pre-dorsal-fin region; all fin membranes, except basal part of caudal fin, scaleless.</paragraph>
<paragraph id="39C501660302741080290EB05E8E647D">First dorsal fin triangular to trapezoid, its origin vertically above preopercular margin or midway between preopercular and opercular margins; 1 st spine of first dorsal fin longest, thereafter, becoming shorter posteriorly. Origin of second dorsal fin just above 4 th to 6 th pored lateral-line scales, 2 nd or 3 rd spine longest, thereafter, becoming gradually shorter posteriorly, forming rounded distal margin. Third dorsal fin semicircular to trapezoid, its origin just above 19 th and 20 th longitudinal scales, 1 st or 2 nd ray longest, thereafter becoming gradually shorter posteriorly. Anal-fin membranous margin deeply incised between rays; anal fin origin just below 6 th to 8 th spine base of second dorsal fin, its posteriormost tip close to caudal-fin base. Pectoral fin relatively long, its posterior tip pointed and slightly beyond or reaching vertical through base of last spine of second dorsal fin; upper and lowermost pectoral-fin base level with bases of 1 st spine of second dorsal-fin and 3 rd spine of first dorsal fin, respectively. Pelvic fin origin just below vertical through base of 1 st spine of first dorsal fin, its tip not reaching anus. Caudal fin rounded; its length similar to head length.</paragraph>
</subSubSection>
<subSubSection id="SECID0EFOAG" type="Nuptial male coloration">
<paragraph id="E0F956F1FED5D28C40DC234979775894">
<heading id="82A7F4C59E4975EF9DBD7C87847BC159" reason="title">Nuptial male coloration</heading>
</paragraph>
<paragraph id="8FA193CA946D991A64D08515F973B67C">
<emphasis id="2A82EB3AD173AE4E7730D77FB18140F7" bold="true">(in preservative).</emphasis>
Based on Fig.
<figureCitation id="B9BF91F04C3A29FD69081BC61EAB935D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Photographs of preserved specimens of Enneapterygius olivaceus (A: AMS I. 22732-008, male, 22.1 mm SL, Australia; B: AMS I. 6317, male, 19.7 mm SL, Vanuatu; C: AMS I. 30000-001, female, 18.0 mm SL, Australia)." figureDoi="10.3897/aiep.54.135448.figure4" httpUri="https://binary.pensoft.net/fig/1165075">4</figureCitation>
. Body generally yellowish white or brownish. Head, including eye, snout, lips, cheek, and opercle, and pectoral-fin base black or dark brown with melanophores. Pre-pelvic region and undersurface of abdomen black or dark brown with melanophores. Orbital tentacle translucent white, covered by melanophores. Iris greenish white; pupil silver. First dorsal fin black. Second and third dorsal fins generally transparent, with two brown longitudinal bands on fin margin and base, width of former ca. 1 / 2 length of spines, latter somewhat indistinct. Membranous margin of 1 st to 3 rd rays of third dorsal fin transparent, without brownish pigmentation. Pectoral fin transparent, membranes between lower 6–8 rays brownish. Pelvic fin white, its basal part brownish. Anal fin brown, fin margin and base whitish. Caudal fin white with faint brownish bars.
</paragraph>
<caption id="C0DE96C79E9D1551D1B62F2C3DA5ADC7" ID-DOI="10.3897/aiep.54.135448.figure4" ID-arpha="B0F30AB7-DB1E-5A85-81B9-85C5E30BEA77" httpUri="https://binary.pensoft.net/fig/1165075" startId="F4">
<paragraph id="296B1479C0586CC394753F50825074C1">
<label id="AC5D02F6E35F55D06F6C8B6E2858E70E">Figure 4.</label>
</paragraph>
<paragraph id="FE130108AF72F1BC7CCC802146CCBB4E">
Photographs of preserved specimens of
<taxonomicName id="D7F8001C53F3F484A1AE950EA84E9F4F" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="936C210ABE37E2C777AB93D97026DB49" italics="true">Enneapterygius olivaceus</emphasis>
</taxonomicName>
(
<emphasis id="F9CD4045F07AF565953617E72183F618" bold="true">A</emphasis>
: AMS I. 22732-008, male, 22.1 mm SL, Australia;
<emphasis id="57F57EA1468723325C0F23497F094440" bold="true">B</emphasis>
: AMS I. 6317, male, 19.7 mm SL, Vanuatu;
<emphasis id="705819294A7D6A360010F3A99470835F" bold="true">C</emphasis>
: AMS I. 30000-001, female, 18.0 mm SL, Australia).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EPPAG" type="Female coloration">
<paragraph id="7F3F3E25824BA903B8A41E115AB7DD99">
<heading id="0AEFAE9474728E7842EE1E748DC2A9C1" reason="title">Female coloration</heading>
</paragraph>
<paragraph id="0FAE0A6C8F0E3661BCB7C6B4C27AB601">
<emphasis id="1648259456F69DB2C7FBB394D6A33FFB" bold="true">(in preservative).</emphasis>
Body generally yellowish white; lateral surface with A- or X-shaped brownish bars. Faint brownish stripes extending from tip of snout to anterior margin of eye, width subequal to upper lip width. Prepelvic region and undersurface of abdomen whitish. Orbital tentacle translucent white, covered by melanophores. Iris navy, pupil silver. Caudal-fin base with vertical brown band. First dorsal fin transparent, with scattered melanophores. Second and third dorsal fins generally transparent, with 2 or 3 irregular oblique brownish bands. Anal fin white. Pectoral fin translucent white; fin rays broadly flecked with brown pigmentation forming ca. 4 or 5 vertical bands. Pelvic fins translucent white. Caudal fin translucent white with narrow brownish vertical bars.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXPAG" type="distribution">
<paragraph id="21CC26302B9A5147F7FBCE1B75DEF767">
<heading id="66F095B5348C148531608836953480AD" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="A1D18ECC3D5139669E08B1AD4A88DB01">
Currently known from
<collectingCountry id="493D182C1277FDBCA9129DCB12C62D88" name="Japan">Japan</collectingCountry>
(the southern Ryukyu Islands), the
<collectingCountry id="F642BE6C650F716F702DB2B735745DFD" name="Philippines">Philippines</collectingCountry>
(Talampulan and Negros islands),
<collectingCountry id="BC800775DA7FE6342F560AC99839F07B" name="Australia">Australia</collectingCountry>
(Lizard Island), and
<collectingCountry id="1F77FA04481E4BC21BCE5B20A702DC45" name="Vanuatu">Vanuatu</collectingCountry>
. (
<bibRefCitation id="8CCCD6195A3894F01611C9E768A7BCC1" DOI="10.11646/zootaxa.5374.3.2" author="Dewa" etAl="et al." firstAuthor="Dewa" issue="3" journalOrPublisher="Zootaxa" pagination="333-360" refId="B2" refString="Dewa Y, Tashiro S, Motomura H (2023) A new species of the genus Enneapterygius (Perciformes: Tripterygiidae) from Japan and the Philippines, with a synopsis and synonymy of related species. Zootaxa 5374 (3): 333–360. https://doi.org/10.11646/zootaxa.5374.3.2" title="A new species of the genus Enneapterygius (Perciformes: Tripterygiidae) from Japan and the Philippines, with a synopsis and synonymy of related species." volume="5374" year="2023">Dewa et al. 2023</bibRefCitation>
; this study) (Fig.
<figureCitation id="C81CD0C420418B1F3C1BE55123905AB7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distributional map of Enneapterygius rhothion (red circles) and E. olivaceus (blue triangles). Closed and open symbols indicate localities determined during the presently reported study and previous records, respectively. Black arrows indicate type localities of the two species." figureDoi="10.3897/aiep.54.135448.figure3" httpUri="https://binary.pensoft.net/fig/1165074">3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFQAG" type="remarks">
<paragraph id="20D7CDD69FE9DC889B688D54557A0A2E">
<heading id="EA7D442D3051925D13C7FD75505D9670" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="C306EF0D1556A6AAA01D4C91B7EE1BC2">
The morphological characters of the presently reported South Pacific specimens agreed well with the description of
<taxonomicName id="7DEC17244E5283CF1A83B35834818851" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="C78027F1120F83F736974E71D6F56D16" italics="true">E. olivaceus</emphasis>
</taxonomicName>
provided by
<bibRefCitation id="F68FC4596BAD4FBF5F21519BA3740BDA" DOI="10.11646/zootaxa.5374.3.2" author="Dewa Y &amp; Tashiro S &amp; Motomura H" etAl="et al." firstAuthor="Dewa" issue="3" journalOrPublisher="Zootaxa" pagination="333-360" refId="B2" refString="Dewa Y, Tashiro S, Motomura H (2023) A new species of the genus Enneapterygius (Perciformes: Tripterygiidae) from Japan and the Philippines, with a synopsis and synonymy of related species. Zootaxa 5374 (3): 333–360. https://doi.org/10.11646/zootaxa.5374.3.2" title="A new species of the genus Enneapterygius (Perciformes: Tripterygiidae) from Japan and the Philippines, with a synopsis and synonymy of related species." volume="5374" year="2023">Dewa et al. (2023)</bibRefCitation>
, especially as follows: 11–13 (modally 12) second dorsal-fin spines; 18–21 (modally 20) notched lateral-line scales; mandibular pore formula 3 + 1 + 3; head length 28.9 % – 32.3 % (mean 30.6 %) of SL; upper jaw length 9.3 % – 12.0 % (10.6 %) of SL; 1 st spine of first dorsal fin longer than that of 2 nd dorsal-fin, its length 11.0 % – 18.2 % (13.4 %) of SL; orbital tentacle broad, leaf-shaped; head, pectoral-fin base and lower part of pectoral fin brownish in nuptial males; caudal fin whitish in nuptial males; pectoral fin with bands in females; anal fin without distinct bars or lines in both sexes. Although the presently reported specimens had fewer longitudinal series scale rows and scales below the 1 st notched lateral-line scale compared to the type series of
<taxonomicName id="34B323D474810C0969009B52B2ACB04E" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="D0FE997AF9FAEBD6E9B4DC417E76EA9D" italics="true">E. olivaceus</emphasis>
</taxonomicName>
[29–32 (modally 30) and 2 or 3 (3), respectively, in the former vs. 30–32 (31) and 2–3 (2) in the latter], such differences were considered minor and regarded as intraspecific variations.
</paragraph>
<paragraph id="B0ACD4323AE430C5884218D77D293688">
In Japanese waters,
<taxonomicName id="80AC55F048BD3939D37092D351A81A4A" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="0099BE402B5236930BB171F60A0E0770" italics="true">E. olivaceus</emphasis>
</taxonomicName>
co-occurs with
<taxonomicName id="4CD06FF7F760A293A569E530C7727861" baseAuthorityName="Gunther" baseAuthorityYear="1877" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="minutus">
<emphasis id="0DDD2F89FD269B4179724C9F0BAA7FC9" italics="true">Enneapterygius minutus</emphasis>
</taxonomicName>
in the Ryukyu Islands (
<bibRefCitation id="EAD151458B4C8C9A22F81C5ED13CB532" DOI="10.11646/zootaxa.5374.3.2" author="Dewa" etAl="et al." firstAuthor="Dewa" issue="3" journalOrPublisher="Zootaxa" pagination="333-360" refId="B2" refString="Dewa Y, Tashiro S, Motomura H (2023) A new species of the genus Enneapterygius (Perciformes: Tripterygiidae) from Japan and the Philippines, with a synopsis and synonymy of related species. Zootaxa 5374 (3): 333–360. https://doi.org/10.11646/zootaxa.5374.3.2" title="A new species of the genus Enneapterygius (Perciformes: Tripterygiidae) from Japan and the Philippines, with a synopsis and synonymy of related species." volume="5374" year="2023">Dewa et al. 2023</bibRefCitation>
). Similarly, the two
<collectingCountry id="69C907A7E3B5573EC828686E9BADCE86" name="Vanuatu">Vanuatu</collectingCountry>
specimens of
<taxonomicName id="D3376C831995D62C045BA62A3A87618C" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="74689D50A4C181D13F0A7B8E4AF31797" italics="true">E. olivaceus</emphasis>
</taxonomicName>
examined in this study had been collected with
<taxonomicName id="89B3C12E2DE330CF7C7BD9610933B6DC" baseAuthorityName="Gunther" baseAuthorityYear="1877" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="minutus">
<emphasis id="119A8F82085E83EFC55214654F2EE198" italics="true">E. minutus</emphasis>
</taxonomicName>
specimens (AMS I. 6317, 2 of
<specimenCount id="994EB0D07499D34AEC71969B2E336B64" count="3" type="generic">3 specimens</specimenCount>
,
<quantity id="2E879049B4DA2C7C74EAF4A4951CD129" metricMagnitude="-2" metricUnit="m" metricValue="1.9799999999999998" metricValueMax="2.02" metricValueMin="1.94" unit="mm" value="19.799999999999997" valueMax="20.2" valueMin="19.4">19.4–20.2 mm</quantity>
SL; AMS I. 6451, 2 of
<specimenCount id="79E7E90728BFA8181BB4AE206930E7C9" count="3" type="generic">3 specimens</specimenCount>
,
<quantity id="542F11D80C16AE1A71BF981FAB4F26DC" metricMagnitude="-2" metricUnit="m" metricValue="1.66" metricValueMax="1.67" metricValueMin="1.65" unit="mm" value="16.6" valueMax="16.7" valueMin="16.5">16.5–16.7 mm</quantity>
SL).
<taxonomicName id="7AB737B53BBFB0C61B13678AD6686A3C" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="F9BBC8FFE3E652D7D75BEDB855AED114" italics="true">Enneapterygius olivaceus</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="F1083C9F7BD9401C36D481CC659B41CC" baseAuthorityName="Gunther" baseAuthorityYear="1877" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="minutus">
<emphasis id="8F5D84A8E3D81F7A41C79890CC3FC241" italics="true">E. minutus</emphasis>
</taxonomicName>
by the following features: orbital tentacle large and leaf-shaped (vs. slender and pointed in
<taxonomicName id="A6C48E39F3D9B3A88E2534280131AE9B" baseAuthorityName="Gunther" baseAuthorityYear="1877" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="minutus">
<emphasis id="568F68CB413C9CC387E9B59F9F867C0D" italics="true">E. minutus</emphasis>
</taxonomicName>
); head long, length 28.4 % – 33.0 % of SL (vs. short, 24.7 % – 32.4 % of SL); upper jaw long, length 9.3 % – 13.1 % of SL (vs. short, 7.4 % – 11.0 % of SL); 1 st spine of first dorsal fin relatively long, length 11.0 % – 18.2 % of SL (vs. short, 7.6 % – 11.4 % of SL); black areas in nuptial males restricted to head and all fins, except for caudal fin [body (except nape and all fins) entirely black in nuptial males]; pectoral fin with faint broad bands in females (with distinct narrow bands in females) (
<bibRefCitation id="48BDD93A47F1B6F810B1B7CA89E16613" DOI="10.11646/zootaxa.5374.3.2" author="Dewa" etAl="et al." firstAuthor="Dewa" issue="3" journalOrPublisher="Zootaxa" pagination="333-360" refId="B2" refString="Dewa Y, Tashiro S, Motomura H (2023) A new species of the genus Enneapterygius (Perciformes: Tripterygiidae) from Japan and the Philippines, with a synopsis and synonymy of related species. Zootaxa 5374 (3): 333–360. https://doi.org/10.11646/zootaxa.5374.3.2" title="A new species of the genus Enneapterygius (Perciformes: Tripterygiidae) from Japan and the Philippines, with a synopsis and synonymy of related species." volume="5374" year="2023">Dewa et al. 2023</bibRefCitation>
; this study).
</paragraph>
<paragraph id="C11FF6D962D64975B71BD728F7C8AF3B">
<taxonomicName id="631CFB4F47A3836B8DA562CCCD5B8E4F" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="EFC954294EBA84C904F4DD69FBA5CB67" italics="true">Enneapterygius olivaceus</emphasis>
</taxonomicName>
was originally described based on
<specimenCount id="23D4FCEF58D8CD6EB86D59CD5D57B77A" count="28" type="generic">28 specimens</specimenCount>
from the southern Ryukyu Islands,
<collectingCountry id="59B27D7AF52DC67AAB014F034052E940" name="Japan">Japan</collectingCountry>
and Talampulan and Negros islands, the
<collectingCountry id="909AE0953B5E1EADD446005748E631CE" name="Philippines">Philippines</collectingCountry>
(
<bibRefCitation id="152314B6B6857A7BB22594C027DAA3CF" DOI="10.11646/zootaxa.5374.3.2" author="Dewa" etAl="et al." firstAuthor="Dewa" issue="3" journalOrPublisher="Zootaxa" pagination="333-360" refId="B2" refString="Dewa Y, Tashiro S, Motomura H (2023) A new species of the genus Enneapterygius (Perciformes: Tripterygiidae) from Japan and the Philippines, with a synopsis and synonymy of related species. Zootaxa 5374 (3): 333–360. https://doi.org/10.11646/zootaxa.5374.3.2" title="A new species of the genus Enneapterygius (Perciformes: Tripterygiidae) from Japan and the Philippines, with a synopsis and synonymy of related species." volume="5374" year="2023">Dewa et al. 2023</bibRefCitation>
). Accordingly, the presently reported specimens, collected from
<collectingCountry id="B90AFC906D78CF11A98784D547886A42" name="Australia">Australia</collectingCountry>
and
<collectingCountry id="6616986FE928F2A5AAC07D5C737061CF" name="Vanuatu">Vanuatu</collectingCountry>
, represent the first records of the species from those localities and the Southern Hemisphere.
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink