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<document id="CEE35CC74463C257EDA947922ABE8EF0" ID-DOI="10.5281/zenodo.4504075" ID-ISSN="2345-7600" ID-Zenodo-Dep="4504075" ID-ZooBank="2300F64E-BD30-4E06-A24F-A226185A4BD3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612465590805" checkinUser="carolina" docAuthor="Siriboon, Thanit, Sutcharit, Chirasak, Naggs, Fred, Rowson, Ben &amp; Panha, Somsak" docDate="2014" docId="781A87B5C44B9C35FF9D3FB9FC1C9DB6" docLanguage="en" docName="RafflesBZool.62.161-174.pdf" docOrigin="Raffles Bulletin of Zoology 62" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Carinartemis vesperus Siriboon &amp; Sutcharit &amp; Naggs &amp; Rowson &amp; Panha 2014, new species" docType="treatment" docUuid="F5241159-8FA4-4052-9A20-FFF21070A5D4" docUuidSource="ZooBank" docVersion="5" lastPageNumber="171" masterDocId="8423FFCDC44C9C3FFFE23E44FFF99F43" masterDocTitle="Revision of the carnivorous snail genus Indoartemon Forcart, 1946 and a new genus Carinartemis from Thailand (Pulmonata: Streptaxidae)" masterLastPageNumber="174" masterPageNumber="161" pageNumber="168" updateTime="1698865986014" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="20E1CFE2E56C8FAD7E81EB03967A3A2D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E3A740DD879F0CBAE383D3DDF823AFA3">
<mods:title id="DCA0DEE0368D287F864AA33908AC1EE9">Revision of the carnivorous snail genus Indoartemon Forcart, 1946 and a new genus Carinartemis from Thailand (Pulmonata: Streptaxidae)</mods:title>
</mods:titleInfo>
<mods:name id="D330997E9425C03582C0651DEB4D5CC6" type="personal">
<mods:role id="FEB232F8DEAE969039C018ABD868D8B0">
<mods:roleTerm id="AE324AB50550CCB78A7EE6546CCCD6DE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B6B3CE096E090FD80208102053C23CF4">Siriboon, Thanit</mods:namePart>
</mods:name>
<mods:name id="44033B87D1277655139E4A8297AC4B72" type="personal">
<mods:role id="5F66AB5737A097EB57BCD64AE21EF256">
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<mods:namePart id="756F242F1E0C5C9D018098139962AAE6">Sutcharit, Chirasak</mods:namePart>
</mods:name>
<mods:name id="51972215B95839ED05C6806A70FDDC66" type="personal">
<mods:role id="034FF3A57AA3C0646710A852BEB4A45C">
<mods:roleTerm id="701211ADEE0CB03F1C519351B2E1C8E7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="378CEA765BF70B8AFDC0D3DB3FD9F762">Naggs, Fred</mods:namePart>
</mods:name>
<mods:name id="4061709F0A556CF0F9848BF14AFA20EE" type="personal">
<mods:role id="1A11A41498A2FD15136924FC721B7ED8">
<mods:roleTerm id="80A2A7DB3F28CC027ABEFBB7DD7C8B67">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="95BDCDCD736AB47C7CD9B757A3CD6020">Rowson, Ben</mods:namePart>
</mods:name>
<mods:name id="52565F6114C5175BBDDC0548BAE9569B" type="personal">
<mods:role id="4B2CD121CDB4AD20ECA97D264D5B7D6F">
<mods:roleTerm id="21A07C85C739647F1A3282D691CF6EDA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E1B3F2302556C482F78C16DDEEF75B1E">Panha, Somsak</mods:namePart>
</mods:name>
<mods:typeOfResource id="1D3B74D804B291898170A7584B9AA4B9">text</mods:typeOfResource>
<mods:relatedItem id="CB893358DAB084450B151E36E7CD4104" type="host">
<mods:titleInfo id="D42066AE2AABBA6F2D869BA0017B2BC5">
<mods:title id="B22F8EEB7C1F28EEDBED859CD3DAA0D0">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="CDE8A84D5EE4BF86C06DD86336607AC3">
<mods:date id="0001581313C81E6C107FECFD3D4F0A6B">2014</mods:date>
<mods:detail id="32D4BEC12137B6463AF43E12C303F672" type="pubDate">
<mods:number id="3C04D3D8B4AFEB652D150CFFAE833757">2014-04-04</mods:number>
</mods:detail>
<mods:detail id="D898ED4F333A5D5F6F732BA9210B0FD0" type="volume">
<mods:number id="2CE65B40C427C440066E99987186F61B">62</mods:number>
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<mods:start id="F82997A6BC953EF83FC43D0D8A3CFEAC">161</mods:start>
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<mods:classification id="B292AA206686E0B8994A8C51B813F8DE">journal article</mods:classification>
<mods:identifier id="155D6A67691075F8535766DE75E751BF" type="DOI">10.5281/zenodo.4504075</mods:identifier>
<mods:identifier id="16BD4F19C51B7E2F141415CC8C4B3DD8" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="4E898BB8626351D1F6D83F9142961D0F" type="Zenodo-Dep">4504075</mods:identifier>
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</mods:mods>
<treatment id="781A87B5C44B9C35FF9D3FB9FC1C9DB6" LSID="urn:lsid:zoobank.org:act:F5241159-8FA4-4052-9A20-FFF21070A5D4" httpUri="http://treatment.plazi.org/id/781A87B5C44B9C35FF9D3FB9FC1C9DB6" lastPageId="10" lastPageNumber="171" pageId="7" pageNumber="168">
<subSubSection id="B8A96528C44B9C38FF9D3FB9FD1A9D57" box="[127,739,509,533]" pageId="7" pageNumber="168" type="nomenclature">
<paragraph id="F00C36A3C44B9C38FF9D3FB9FD1A9D57" blockId="7.[127,739,509,564]" box="[127,739,509,533]" pageId="7" pageNumber="168">
<heading id="AB4481CFC44B9C38FF9D3FB9FD1A9D57" bold="true" box="[127,739,509,533]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="168" reason="2">
<emphasis id="C2C7EAB1C44B9C38FF9D3FB9FD1A9D57" bold="true" box="[127,739,509,533]" pageId="7" pageNumber="168">
<taxonomicName id="37B34D20C44B9C38FF9D3FB9FDA89D57" ID-CoL="RD95" authority="Siriboon &amp; Panha" authorityName="Siriboon &amp; Sutcharit &amp; Naggs &amp; Rowson &amp; Panha" authorityYear="2014" box="[127,593,509,533]" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="168" phylum="Mollusca" rank="species" species="vesperus" status="sp. nov.">
<emphasis id="C2C7EAB1C44B9C38FF9D3FB9FE8E9D57" bold="true" box="[127,375,509,533]" italics="true" pageId="7" pageNumber="168">Carinartemis vesperus</emphasis>
Siriboon &amp; Panha
</taxonomicName>
,
<taxonomicNameLabel id="D9F457CAC44B9C38FDBD3FB9FD1A9D57" box="[607,739,509,532]" pageId="7" pageNumber="168" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B8A96528C44B9C38FF2C3C59FD6D9D77" box="[206,660,541,564]" pageId="7" pageNumber="168" type="description">
<paragraph id="F00C36A3C44B9C38FF2C3C59FD6D9D77" blockId="7.[127,739,509,564]" box="[206,660,541,564]" pageId="7" pageNumber="168">
(
<figureCitation id="68882A26C44B9C38FF343C59FEE39D77" box="[214,282,541,564]" captionStart="Fig" captionStartId="2.[136,171,1832,1853]" captionTargetBox="[138,776,989,1810]" captionTargetId="figure-7@2.[137,776,989,1810]" captionTargetPageId="2" captionText="Fig. 1. Distribution map of Indoartemon spp. and Carinartemis spp. Indoartemon spp (white circle): (1) Indoartemon cingalensis (Benson, 1853); (2) Indoartemon layardianus (Benson, 1853); (3) Indoartemon glacilis (Collett, 1898); (4) Indoartemon eburneus (Pfeiffer, 1861); (5) Indoartemon fuchsianus (Gredler, 1881); (6) Indoartemon bidens (Möllendorff, 1883); (7) Indoartemon tridens (Möllendorff, 1898); (8) Indoartemon laevis (Blanford, 1899); (9) Indoartemon prestoni (Gude, 1903); and (10) Indoartemon medius, new species. Carinartemis spp. (black circle): (1) Carinartemis vesperus, new species; and (2) Carinartemis striatus, new species." figureDoi="http://doi.org/10.5281/zenodo.4504077" httpUri="https://zenodo.org/record/4504077/files/figure.png" pageId="7" pageNumber="168">Figs 1</figureCitation>
,
<figureCitation id="68882A26C44B9C38FECB3C59FEB29D77" box="[297,331,541,564]" captionStart="Fig" captionStartId="2.[836,872,1975,1996]" captionTargetBox="[836,1473,561,1952]" captionTargetId="figure-361@2.[836,1473,561,1952]" captionTargetPageId="2" captionText="Fig. 2. Living snails of: A, Indoartemon medius, new species, paratype CUMZ 5017 (shell width about 8 mm); B, Carinartemis vesperus, new species, paratype CUMZ 6201 (shell width about 10 mm); C, Carinartemis striatus, new species, paratype CUMZ 6205 (shell width about 12 mm)." figureDoi="http://doi.org/10.5281/zenodo.4504079" httpUri="https://zenodo.org/record/4504079/files/figure.png" pageId="7" pageNumber="168">2B</figureCitation>
,
<figureCitation id="68882A26C44B9C38FEB53C59FE9D9D77" box="[343,356,541,564]" captionStart="Fig" captionStartId="7.[113,148,2005,2026]" captionTargetBox="[113,1453,825,1982]" captionTargetId="figure-13@7.[113,1453,825,1982]" captionTargetPageId="7" captionText="Fig. 6. A, Shell of Stemmatopsis poirieri lectotype MNHN 26980; B–E, Shells of Carinartemis vesperus, new species, (B) holotype CUMZ 5019 with apertural dentition, (C, D) paratype CUMZ 6200 from Khao Kling, Khao Yoi District, Phetchaburi, and (E) specimen NMW 1955.158.25233 from Khao Yoi, Petchaburi; F, G, Carinartemis striatus, new species, (F) holotype CUMZ 6203 with apertural dentition, and (G) paratype CUMZ 6117 from Tam Kra Sae, Sai Yok, Kanchanaburi." figureDoi="http://doi.org/10.5281/zenodo.4504087" httpUri="https://zenodo.org/record/4504087/files/figure.png" pageId="7" pageNumber="168">6</figureCitation>
B–E, 7A, B, 8A–F,
<tableCitation id="BD310318C44B9C38FDD93C59FD739D77" box="[571,650,541,564]" captionStart="Table 1" captionStartId="4.[136,189,191,212]" captionText="Table 1. Shell measurements of the three Indoartemon species and two Carinatemis species. Specimen collections and catalogue numbers" pageId="7" pageNumber="168">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B8A96528C44B9C38FF933C18FC7B9E3C" pageId="7" pageNumber="168" type="materials_examined">
<paragraph id="F00C36A3C44B9C38FF933C18FE299DA7" blockId="7.[113,753,604,740]" pageId="7" pageNumber="168">
<emphasis id="C2C7EAB1C44B9C38FF933C18FEF89D32" bold="true" box="[113,257,604,625]" pageId="7" pageNumber="168">Type material.</emphasis>
<materialsCitation id="40DB3CFEC44B9C38FEE73C18FDCD9DCD" collectionCode="CUMZ" pageId="7" pageNumber="168" specimenCode="CUMZ 5019" specimenCount="1" typeStatus="holotype">
<typeStatus id="2F088801C44B9C38FEE73C18FEA59D32" box="[261,348,604,625]" pageId="7" pageNumber="168" type="holotype">Holotype</typeStatus>
<specimenCode id="A0159ED8C44B9C38FE833C18FE219D32" box="[353,472,604,625]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="7" pageNumber="168">CUMZ 5019</specimenCode>
(
<figureCitation id="68882A26C44B9C38FE013C18FDD29D32" box="[483,555,604,625]" captionStart="Fig" captionStartId="7.[113,148,2005,2026]" captionTargetBox="[113,1453,825,1982]" captionTargetId="figure-13@7.[113,1453,825,1982]" captionTargetPageId="7" captionText="Fig. 6. A, Shell of Stemmatopsis poirieri lectotype MNHN 26980; B–E, Shells of Carinartemis vesperus, new species, (B) holotype CUMZ 5019 with apertural dentition, (C, D) paratype CUMZ 6200 from Khao Kling, Khao Yoi District, Phetchaburi, and (E) specimen NMW 1955.158.25233 from Khao Yoi, Petchaburi; F, G, Carinartemis striatus, new species, (F) holotype CUMZ 6203 with apertural dentition, and (G) paratype CUMZ 6117 from Tam Kra Sae, Sai Yok, Kanchanaburi." figureDoi="http://doi.org/10.5281/zenodo.4504087" httpUri="https://zenodo.org/record/4504087/files/figure.png" pageId="7" pageNumber="168">Fig. 6B</figureCitation>
). Measurement: shell height
<quantity id="374B9B46C44B9C38FF573C3DFEFA9DCD" box="[181,259,633,654]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="7" pageNumber="168" unit="mm" value="6.9">6.9 mm</quantity>
, shell width
<quantity id="374B9B46C44B9C38FE613C3DFE289DCD" box="[387,465,633,654]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="7" pageNumber="168" unit="mm" value="9.6">9.6 mm</quantity>
, 7 whorls
</materialsCitation>
.
<materialsCitation id="40DB3CFEC44B9C38FDA23C3DFE469D84" collectionCode="CUMZ" pageId="7" pageNumber="168" specimenCode="CUMZ 5020" specimenCount="2" typeStatus="paratype">
<typeStatus id="2F088801C44B9C38FDA23C3DFD589DCD" box="[576,673,633,654]" pageId="7" pageNumber="168" type="paratype">Paratypes</typeStatus>
:
<specimenCode id="A0159ED8C44B9C38FD4E3C3DFF589DE9" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="7" pageNumber="168">CUMZ 5020</specimenCode>
(3 shells), 6200 (60 shells;
<figureCitation id="68882A26C44B9C38FE5B3CD1FDDC9DE9" box="[441,549,661,682]" captionStart="Fig" captionStartId="7.[113,148,2005,2026]" captionTargetBox="[113,1453,825,1982]" captionTargetId="figure-13@7.[113,1453,825,1982]" captionTargetPageId="7" captionText="Fig. 6. A, Shell of Stemmatopsis poirieri lectotype MNHN 26980; B–E, Shells of Carinartemis vesperus, new species, (B) holotype CUMZ 5019 with apertural dentition, (C, D) paratype CUMZ 6200 from Khao Kling, Khao Yoi District, Phetchaburi, and (E) specimen NMW 1955.158.25233 from Khao Yoi, Petchaburi; F, G, Carinartemis striatus, new species, (F) holotype CUMZ 6203 with apertural dentition, and (G) paratype CUMZ 6117 from Tam Kra Sae, Sai Yok, Kanchanaburi." figureDoi="http://doi.org/10.5281/zenodo.4504087" httpUri="https://zenodo.org/record/4504087/files/figure.png" pageId="7" pageNumber="168">Fig. 6C, D</figureCitation>
), 6201 (
<specimenCount id="E6B5FD2AC44B9C38FD9A3CD1FD089DE9" box="[632,753,661,682]" pageId="7" pageNumber="168" type="generic">2 specimens</specimenCount>
in ethanol;
<figureCitation id="68882A26C44B9C38FF3D3CF6FED49D84" box="[223,301,690,711]" captionStart="Fig" captionStartId="2.[836,872,1975,1996]" captionTargetBox="[836,1473,561,1952]" captionTargetId="figure-361@2.[836,1473,561,1952]" captionTargetPageId="2" captionText="Fig. 2. Living snails of: A, Indoartemon medius, new species, paratype CUMZ 5017 (shell width about 8 mm); B, Carinartemis vesperus, new species, paratype CUMZ 6201 (shell width about 10 mm); C, Carinartemis striatus, new species, paratype CUMZ 6205 (shell width about 12 mm)." figureDoi="http://doi.org/10.5281/zenodo.4504079" httpUri="https://zenodo.org/record/4504079/files/figure.png" pageId="7" pageNumber="168">Figs 2B</figureCitation>
,
<figureCitation id="68882A26C44B9C38FED53CF6FE8A9D84" box="[311,371,690,711]" captionStart="Fig" captionStartId="8.[136,171,2035,2056]" captionTargetBox="[146,1462,502,2007]" captionTargetId="figure-66@8.[131,1478,489,2021]" captionTargetPageId="8" captionText="Fig. 7. Genitalia of Carinartemis spp: A, B, Carinartemis vesperus, new species, paratype CUMZ 6201, (A) reproductive system, and (B) insertion of vas deferens into penial sheath; C, D, Carinartemis striatus, new species, paratype CUMZ 6205, (C) reproductive system, and (D) attachment of vas deferens into distal end of the penial sheath." figureDoi="http://doi.org/10.5281/zenodo.4504089" httpUri="https://zenodo.org/record/4504089/files/figure.png" pageId="7" pageNumber="168">7A, B</figureCitation>
,
<figureCitation id="68882A26C44B9C38FE9C3CF6FE709D84" box="[382,393,690,711]" captionStart="Fig" captionStartId="9.[114,149,1977,1998]" captionTargetBox="[114,1451,379,1954]" captionTargetId="figure-54@9.[108,1458,374,1960]" captionTargetPageId="9" captionText="Fig. 8. Internal sculpture of genitalia of Carinartemis spp: A–F, Carinartemis vesperus, new species, paratype CUMZ 6201, (A) details of atrial pore on the atrium surface, (B) high magnification of penial hooks, (C) top view of penial hook, (D) arrangement of vaginal folds with hooks, (E) high magnification of vaginal hook, and (F) radula morphology. G–K, Carinartemis striatus, new species, paratype CUMZ 6205, (G) details of atrium surface, (H) low magnification shows arrangement of penial hooks, (I) low magnification shows arrangement of vaginal folds, (J) high magnification of penial hooks, and (K) radula morphology." figureDoi="http://doi.org/10.5281/zenodo.4504091" httpUri="https://zenodo.org/record/4504091/files/figure.png" pageId="7" pageNumber="168">8</figureCitation>
A–F)
</materialsCitation>
,
<materialsCitation id="40DB3CFEC44B9C38FE2B3CF6FD149D84" box="[457,749,690,711]" collectionCode="NHMUK" pageId="7" pageNumber="168" specimenCode="NHMUK 20130077" specimenCount="1" typeStatus="paratype">
<specimenCode id="A0159ED8C44B9C38FE2B3CF6FD729D84" box="[457,651,690,711]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="7" pageNumber="168" type="Museum">NHMUK 20130077</specimenCode>
(2 shells)
</materialsCitation>
,
<collectionCode id="96A2AE66C44B9C38FF933C8BFF589DA7" box="[113,161,719,740]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="7" pageNumber="168">SMF</collectionCode>
(2 shells), and
<collectionCode id="96A2AE66C44B9C38FEDB3C8BFE919DA7" box="[313,360,719,740]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="168">ZRC</collectionCode>
(2 shells).
</paragraph>
<paragraph id="F00C36A3C44B9C38FCCE3EF9FB139E4F" blockId="7.[812,1454,189,268]" pageId="7" pageNumber="168">
<materialsCitation id="40DB3CFEC44B9C38FCCE3EF9FB1F9E4F" country="Thailand" county="Rock" elevation="100" latitude="13.2601385" location="Khao Kling" longLatPrecision="1" longitude="99.804115" municipality="Khao Yoi District" pageId="7" pageNumber="168" specimenCount="1" stateProvince="Phetchaburi" typeStatus="holotype">
<emphasis id="C2C7EAB1C44B9C38FCCE3EF9FC569F91" bold="true" box="[812,943,189,210]" pageId="7" pageNumber="168">
<typeStatus id="2F088801C44B9C38FCCE3EF9FCA79F91" box="[812,862,189,210]" pageId="7" pageNumber="168">Type</typeStatus>
locality.
</emphasis>
<location id="F56C6078C44B9C38FC563EF9FBDD9F91" LSID="urn:lsid:plazi:treatment:781A87B5C44B9C35FF9D3FB9FC1C9DB6:F56C6078C44B9C38FC563EF9FBDD9F91" box="[948,1060,189,210]" country="Thailand" county="Rock" latitude="13.2601385" longLatPrecision="1" longitude="99.804115" municipality="Khao Yoi District" name="Khao Kling" pageId="7" pageNumber="168" stateProvince="Phetchaburi">Khao Kling</location>
,
<collectingMunicipality id="1068ACD9C44B9C38FBCE3EF9FB289F91" box="[1068,1233,189,210]" pageId="7" pageNumber="168">Khao Yoi District</collectingMunicipality>
,
<collectingRegion id="3277F841C44B9C38FB3B3EF9FAB59F91" box="[1241,1356,189,210]" country="Thailand" name="Phetchaburi" pageId="7" pageNumber="168">Phetchaburi</collectingRegion>
,
<collectingCountry id="88A47633C44B9C38FAB13EF9FA539F91" box="[1363,1450,189,210]" name="Thailand" pageId="7" pageNumber="168">Thailand</collectingCountry>
.
<collectingCounty id="196D4E2FC44B9C38FCCE3E9EFCA79FAC" box="[812,862,218,239]" pageId="7" pageNumber="168">Rock</collectingCounty>
crevices on isolated limestone hill reaching about
<quantity id="374B9B46C44B9C38FAD73E9EFA899FAC" box="[1333,1392,218,239]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="168" unit="m" value="100.0">
<elevation id="7B9ED190C44B9C38FAD73E9EFA899FAC" box="[1333,1392,218,239]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="168" unit="m" value="100.0">100 m</elevation>
</quantity>
above mean sea level (
<geoCoordinate id="95875064C44B9C38FC283EB3FBA99E4F" box="[970,1104,247,268]" degrees="13" direction="north" minutes="15" orientation="latitude" pageId="7" pageNumber="168" precision="1" seconds="36.5" value="13.2601385">13°15'36.5&quot;N</geoCoordinate>
,
<geoCoordinate id="95875064C44B9C38FBB93EB3FB269E4F" box="[1115,1247,247,268]" degrees="99" direction="east" minutes="48" orientation="longitude" pageId="7" pageNumber="168" precision="1" seconds="14.8" value="99.804115">99°48'14.8&quot;E</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
<paragraph id="F00C36A3C44B9C38FCCE3F74FC7B9E3C" blockId="7.[812,1453,304,383]" pageId="7" pageNumber="168">
<emphasis id="C2C7EAB1C44B9C38FCCE3F74FBD29E06" bold="true" box="[812,1067,304,325]" pageId="7" pageNumber="168">Other material examined.</emphasis>
Khao Yoi, Khao Yoi District,
<collectingRegion id="3277F841C44B9C38FADF3F74FA549E06" box="[1341,1453,304,325]" country="Thailand" name="Phetchaburi" pageId="7" pageNumber="168">Phetchaburi</collectingRegion>
(
<collectionCode id="96A2AE66C44B9C38FCD03F09FC839E21" box="[818,890,333,354]" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="7" pageNumber="168">CUMZ</collectionCode>
6202). Khao Yoi,
<collectingRegion id="3277F841C44B9C38FBDF3F09FB4A9E21" box="[1085,1203,333,354]" country="Thailand" name="Phetchaburi" pageId="7" pageNumber="168">Phetchaburi</collectingRegion>
(
<collectionCode id="96A2AE66C44B9C38FB203F09FAFA9E21" box="[1218,1283,333,354]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="7" pageNumber="168">NMW</collectionCode>
.1955.158.25233,
<figureCitation id="68882A26C44B9C38FCCE3F2EFC8E9E3C" box="[812,887,362,383]" captionStart="Fig" captionStartId="7.[113,148,2005,2026]" captionTargetBox="[113,1453,825,1982]" captionTargetId="figure-13@7.[113,1453,825,1982]" captionTargetPageId="7" captionText="Fig. 6. A, Shell of Stemmatopsis poirieri lectotype MNHN 26980; B–E, Shells of Carinartemis vesperus, new species, (B) holotype CUMZ 5019 with apertural dentition, (C, D) paratype CUMZ 6200 from Khao Kling, Khao Yoi District, Phetchaburi, and (E) specimen NMW 1955.158.25233 from Khao Yoi, Petchaburi; F, G, Carinartemis striatus, new species, (F) holotype CUMZ 6203 with apertural dentition, and (G) paratype CUMZ 6117 from Tam Kra Sae, Sai Yok, Kanchanaburi." figureDoi="http://doi.org/10.5281/zenodo.4504087" httpUri="https://zenodo.org/record/4504087/files/figure.png" pageId="7" pageNumber="168">Fig. 6E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B8A96528C44B9C38FCCE3FECFBE89EBC" pageId="7" pageNumber="168" type="etymology">
<paragraph id="F00C36A3C44B9C38FCCE3FECFBE89EBC" blockId="7.[812,1453,424,511]" pageId="7" pageNumber="168">
<emphasis id="C2C7EAB1C44B9C38FCCE3FECFC579EFC" bold="true" box="[812,942,424,447]" pageId="7" pageNumber="168">Etymology.</emphasis>
The specific epithet “
<taxonomicName id="37B34D20C44B9C38FB4B3FECFAF19EFC" authorityName="Siriboon &amp; Sutcharit &amp; Naggs &amp; Rowson &amp; Panha" authorityYear="2014" box="[1193,1288,424,447]" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="168" phylum="Mollusca" rank="species" species="vesperus">
<emphasis id="C2C7EAB1C44B9C38FB4B3FECFAF19EFC" box="[1193,1288,424,447]" italics="true" pageId="7" pageNumber="168">vesperus</emphasis>
</taxonomicName>
” from a Latin term for “west” refers to the locality of the new species in the west of
<collectingCountry id="88A47633C44B9C38FC493FACFBF49EBC" box="[939,1037,488,511]" name="Thailand" pageId="7" pageNumber="168">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B8A96528C44B9C35FCCE3C6CFF059E76" lastPageId="10" lastPageNumber="171" pageId="7" pageNumber="168" type="diagnosis">
<paragraph id="F00C36A3C44B9C37FCCE3C6CFECF9FB7" blockId="7.[812,1453,552,735]" lastBlockId="8.[136,775,189,244]" lastPageId="8" lastPageNumber="169" pageId="7" pageNumber="168">
<emphasis id="C2C7EAB1C44B9C38FCCE3C6CFC5F9D7C" bold="true" box="[812,934,552,575]" pageId="7" pageNumber="168">Diagnosis.</emphasis>
<taxonomicName id="37B34D20C44B9C38FC533C6CFB4A9D7C" authority="Siriboon &amp; Sutcharit &amp; Naggs &amp; Rowson &amp; Panha, 2014" authorityName="Siriboon &amp; Sutcharit &amp; Naggs &amp; Rowson &amp; Panha" authorityYear="2014" box="[945,1203,552,575]" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="168" phylum="Mollusca" rank="species" species="vesperus" status="sp. nov.">
<emphasis id="C2C7EAB1C44B9C38FC533C6CFB4A9D7C" box="[945,1203,552,575]" italics="true" pageId="7" pageNumber="168">Carinartemis vesperus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="D9F457CAC44B9C38FB273C6CFAAB9D7C" box="[1221,1362,552,575]" pageId="7" pageNumber="168" rank="species">new species</taxonomicNameLabel>
, can be distinguished from
<taxonomicName id="37B34D20C44B9C38FBED3C0CFA759D1C" authority="(Mollendorff, 1894)" baseAuthorityName="Mollendorff" baseAuthorityYear="1894" box="[1039,1420,584,607]" class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="168" phylum="Mollusca" rank="species" species="mirificus">
<emphasis id="C2C7EAB1C44B9C38FBED3C0CFB639D1C" box="[1039,1178,584,607]" italics="true" pageId="7" pageNumber="168">H. mirificus</emphasis>
(
<bibRefCitation id="94224B52C44B9C38FB4F3C0CFA7D9D1C" author="von Mollendorff O" box="[1197,1412,584,607]" pageId="7" pageNumber="168" pagination="146 - 156" refId="ref8963" refString="von Mollendorff O (1894) On a Collection of Land-Shells from the Samui Islands, Gulf of Siam. Proceedings of the Zoological Society of London, 1894: 146 - 156, pl. 16." type="journal article" year="1894">Möllendorff, 1894</bibRefCitation>
)
</taxonomicName>
in having a higher spire, peripheral keel around nearly the entire penultimate whorl, a subcircular aperture, and in lacking apertural lamellae. While
<taxonomicName id="37B34D20C44B9C38FBB13CECFB2E9DFC" baseAuthorityName="Mollendorff" baseAuthorityYear="1894" box="[1107,1239,680,703]" class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="168" phylum="Mollusca" rank="species" species="mirificus">
<emphasis id="C2C7EAB1C44B9C38FBB13CECFB2E9DFC" box="[1107,1239,680,703]" italics="true" pageId="7" pageNumber="168">H. mirificus</emphasis>
</taxonomicName>
has a lower spire, a peripheral keel around less than half of the penultimate whorl, a triangular aperture, and a strong and triangular parietal lamella.
</paragraph>
<caption id="A4CC662BC44B9C38FF933991FCA99702" ID-DOI="http://doi.org/10.5281/zenodo.4504087" ID-Zenodo-Dep="4504087" httpUri="https://zenodo.org/record/4504087/files/figure.png" pageId="7" pageNumber="168" startId="7.[113,148,2005,2026]" targetBox="[113,1453,825,1982]" targetPageId="7">
<paragraph id="F00C36A3C44B9C38FF933991FCA99702" blockId="7.[113,1452,2005,2113]" pageId="7" pageNumber="168">
Fig. 6. A, Shell of
<taxonomicName id="37B34D20C44B9C38FEC03991FE0C98A9" authorityName="Mabille" authorityYear="1887" box="[290,501,2005,2026]" class="Gastropoda" family="Streptaxidae" genus="Stemmatopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="168" phylum="Mollusca" rank="species" species="poirieri">
<emphasis id="C2C7EAB1C44B9C38FEC03991FE0C98A9" box="[290,501,2005,2026]" italics="true" pageId="7" pageNumber="168">Stemmatopsis poirieri</emphasis>
</taxonomicName>
lectotype MNHN 26980; B–E, Shells of
<taxonomicName id="37B34D20C44B9C38FC633991FBA298A9" authority="Siriboon &amp; Sutcharit &amp; Naggs &amp; Rowson &amp; Panha, 2014" authorityName="Siriboon &amp; Sutcharit &amp; Naggs &amp; Rowson &amp; Panha" authorityYear="2014" box="[897,1115,2005,2026]" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="168" phylum="Mollusca" rank="species" species="vesperus" status="sp. nov.">
<emphasis id="C2C7EAB1C44B9C38FC633991FBA298A9" box="[897,1115,2005,2026]" italics="true" pageId="7" pageNumber="168">Carinartemis vesperus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="D9F457CAC44B9C38FB853991FB2498A9" box="[1127,1245,2005,2026]" pageId="7" pageNumber="168" rank="species">new species</taxonomicNameLabel>
, (B) holotype CUMZ 5019 with apertural dentition, (C, D) paratype CUMZ 6200 from Khao Kling, Khao Yoi District, Phetchaburi, and (E) specimen NMW 1955.158.25233 from Khao Yoi, Petchaburi; F, G,
<taxonomicName id="37B34D20C44B9C38FDBD364BFCD69767" authority="Siriboon &amp; Sutcharit &amp; Naggs &amp; Rowson &amp; Panha, 2014" authorityName="Siriboon &amp; Sutcharit &amp; Naggs &amp; Rowson &amp; Panha" authorityYear="2014" box="[607,815,2063,2084]" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="168" phylum="Mollusca" rank="species" species="striatus" status="sp. nov.">
<emphasis id="C2C7EAB1C44B9C38FDBD364BFCD69767" box="[607,815,2063,2084]" italics="true" pageId="7" pageNumber="168">Carinartemis striatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="D9F457CAC44B9C38FCDE364BFC4A9767" box="[828,947,2063,2084]" pageId="7" pageNumber="168" rank="species">new species</taxonomicNameLabel>
, (F) holotype CUMZ 6203 with apertural dentition, and (G) paratype CUMZ 6117 from Tam Kra Sae, Sai Yok, Kanchanaburi.
</paragraph>
</caption>
<paragraph id="F00C36A3C4449C37FF6A3F59FADD9E17" blockId="8.[136,776,285,436]" lastBlockId="8.[836,1474,189,340]" pageId="8" pageNumber="169">
Compared with
<taxonomicName id="37B34D20C4449C37FEDA3F59FDA59E77" authority="(Gould, 1856)" baseAuthorityName="Gould" baseAuthorityYear="1856" box="[312,604,285,308]" class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="169" phylum="Mollusca" rank="species" species="exacutus">
<emphasis id="C2C7EAB1C4449C37FEDA3F59FE419E77" box="[312,440,285,308]" italics="true" pageId="8" pageNumber="169">H. exacutus</emphasis>
(
<bibRefCitation id="94224B52C4449C37FE2B3F59FDAD9E77" author="Gould AA" box="[457,596,285,308]" pageId="8" pageNumber="169" pagination="11 - 16" refId="ref8658" refString="Gould AA (1856) Descriptions of new species of shells. Proceedings of the Boston Society of Natural History, 6: 11 - 16." type="journal article" year="1856">Gould, 1856</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="37B34D20C4449C37FD763F59FEC99E17" authority="(Benson, 1859)" baseAuthorityName="Benson" baseAuthorityYear="1859" class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="169" phylum="Mollusca" rank="species" species="sankeyi">
<emphasis id="C2C7EAB1C4449C37FD763F59FCFE9E77" box="[660,775,285,308]" italics="true" pageId="8" pageNumber="169">H. sankeyi</emphasis>
(
<bibRefCitation id="94224B52C4449C37FF6D3F79FED19E17" author="Benson WH" box="[143,296,317,340]" pageId="8" pageNumber="169" pagination="471 - 474" refId="ref8197" refString="Benson WH (1859) Characters of a new Burmese Streptaxis and of two forms belonging to a peculiar section of Helix collected by Captain Richard H. Sankey, Madras Engineers. The Annals and Magazine of Natural History, Series 3, 3: 471 - 474." type="journal article" year="1859">Benson, 1859</bibRefCitation>
)
</taxonomicName>
, this new species differs by having a strong peripheral keel around nearly the entire penultimate whorl, having the last whorl less inflated and much deviated from the vertical axis, and in lacking strong apertural lamellae. Whereas, these two species exhibit a strong peripheral keel over less than half of the penultimate whorl, and have the last whorl inflated and less deviated from the vertical axis. The apertural dentition of
<taxonomicName id="37B34D20C4449C37FBCD3F59FB549E77" baseAuthorityName="Gould" baseAuthorityYear="1856" box="[1071,1197,285,308]" class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="169" phylum="Mollusca" rank="species" species="exacutus">
<emphasis id="C2C7EAB1C4449C37FBCD3F59FB549E77" box="[1071,1197,285,308]" italics="true" pageId="8" pageNumber="169">H. exacutus</emphasis>
</taxonomicName>
has two parietal lamellae and
<taxonomicName id="37B34D20C4449C37FC903F79FC1D9E17" baseAuthorityName="Benson" baseAuthorityYear="1859" box="[882,996,317,340]" class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="169" phylum="Mollusca" rank="species" species="sankeyi">
<emphasis id="C2C7EAB1C4449C37FC903F79FC1D9E17" box="[882,996,317,340]" italics="true" pageId="8" pageNumber="169">H. sankeyi</emphasis>
</taxonomicName>
has a strong parietal lamella.
</paragraph>
<paragraph id="F00C36A3C4449C35FCA63F39FF059E76" blockId="8.[836,1474,381,436]" lastBlockId="10.[136,775,190,309]" lastPageId="10" lastPageNumber="171" pageId="8" pageNumber="169">
<taxonomicName id="37B34D20C4449C37FCA63F39FBCD9ED7" authority="Siriboon &amp; Sutcharit &amp; Naggs &amp; Rowson &amp; Panha, 2014" authorityName="Siriboon &amp; Sutcharit &amp; Naggs &amp; Rowson &amp; Panha" authorityYear="2014" box="[836,1076,381,404]" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="169" phylum="Mollusca" rank="species" species="vesperus" status="sp. nov.">
<emphasis id="C2C7EAB1C4449C37FCA63F39FBCD9ED7" box="[836,1076,381,404]" italics="true" pageId="8" pageNumber="169">Carinartemis vesperus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="D9F457CAC4449C37FBA23F39FB389ED7" box="[1088,1217,381,404]" pageId="8" pageNumber="169" rank="species">new species</taxonomicNameLabel>
, differs from
<taxonomicName id="37B34D20C4449C37FAAC3F39FA429ED7" authority="Siriboon &amp; Sutcharit &amp; Naggs &amp; Rowson &amp; Panha, 2014" authorityName="Siriboon &amp; Sutcharit &amp; Naggs &amp; Rowson &amp; Panha" authorityYear="2014" box="[1358,1467,381,404]" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="169" phylum="Mollusca" rank="species" species="striatus" status="sp. nov.">
<emphasis id="C2C7EAB1C4449C37FAAC3F39FA429ED7" box="[1358,1467,381,404]" italics="true" pageId="8" pageNumber="169">C. striatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="D9F457CAC4449C37FCA63FD9FC3D9EF7" box="[836,964,413,436]" pageId="8" pageNumber="169" rank="species">new species</taxonomicNameLabel>
, by having a smaller shell with weaker transverse ridges, the area below the periphery of the penultimate whorl smooth, a short longitudinal furrow in the umbilical area, and in lacking apertural lamellae. The genitalia have a short penis and vagina, and a thick penial sheath with the vas deferens passing it. Penial papillae are absent, and vaginal hooks are present. Whereas
<taxonomicName id="37B34D20C4459C36FC0B3E9AFBA19FB6" authority="Siriboon &amp; Sutcharit &amp; Naggs &amp; Rowson &amp; Panha, 2014" authorityName="Siriboon &amp; Sutcharit &amp; Naggs &amp; Rowson &amp; Panha" authorityYear="2014" box="[1001,1112,222,245]" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="170" phylum="Mollusca" rank="species" species="striatus" status="sp. nov.">
<emphasis id="C2C7EAB1C4459C36FC0B3E9AFBA19FB6" box="[1001,1112,222,245]" italics="true" pageId="9" pageNumber="170">C. striatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="D9F457CAC4459C36FB873E9AFB119FB6" box="[1125,1256,222,245]" pageId="9" pageNumber="170" rank="species">new species</taxonomicNameLabel>
, has a larger shell, strong transverse ridges over the entire shell, no longitudinal furrow in the umbilical area, and has a prominent parietal lamella. The genitalia have a long penis and vagina and a very thin penial sheath with the vas deferens attached to its distal end. Penial papillae are present, and vaginal hooks are absent.
</paragraph>
</subSubSection>
<caption id="A4CC662BC4449C37FF6A39B7FCBC9701" ID-DOI="http://doi.org/10.5281/zenodo.4504089" ID-Zenodo-Dep="4504089" httpUri="https://zenodo.org/record/4504089/files/figure.png" pageId="8" pageNumber="169" startId="8.[136,171,2035,2056]" targetBox="[146,1462,502,2007]" targetPageId="8">
<paragraph id="F00C36A3C4449C37FF6A39B7FCBC9701" blockId="8.[136,1474,2035,2114]" pageId="8" pageNumber="169">
Fig. 7. Genitalia of
<taxonomicName id="37B34D20C4449C37FEA539B7FE3E974B" authorityName="Siriboon &amp; Sutcharit &amp; Naggs &amp; Rowson &amp; Panha" authorityYear="2014" box="[327,455,2035,2056]" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="169" phylum="Mollusca" rank="genus">
<emphasis id="C2C7EAB1C4449C37FEA539B7FE3E974B" box="[327,455,2035,2056]" italics="true" pageId="8" pageNumber="169">Carinartemis</emphasis>
</taxonomicName>
spp: A, B,
<taxonomicName id="37B34D20C4449C37FDD439B7FCE8974B" authority="Siriboon &amp; Sutcharit &amp; Naggs &amp; Rowson &amp; Panha, 2014" authorityName="Siriboon &amp; Sutcharit &amp; Naggs &amp; Rowson &amp; Panha" authorityYear="2014" box="[566,785,2035,2056]" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="169" phylum="Mollusca" rank="species" species="vesperus" status="sp. nov.">
<emphasis id="C2C7EAB1C4449C37FDD439B7FCE8974B" box="[566,785,2035,2056]" italics="true" pageId="8" pageNumber="169">Carinartemis vesperus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="D9F457CAC4449C37FCFC39B7FC6C974B" box="[798,917,2035,2056]" pageId="8" pageNumber="169" rank="species">new species</taxonomicNameLabel>
, paratype CUMZ 6201, (A) reproductive system, and (B) insertion of vas deferens into penial sheath; C, D,
<taxonomicName id="37B34D20C4449C37FD603654FCAB9766" authority="Siriboon &amp; Sutcharit &amp; Naggs &amp; Rowson &amp; Panha, 2014" authorityName="Siriboon &amp; Sutcharit &amp; Naggs &amp; Rowson &amp; Panha" authorityYear="2014" box="[642,850,2064,2085]" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="169" phylum="Mollusca" rank="species" species="striatus" status="sp. nov.">
<emphasis id="C2C7EAB1C4449C37FD603654FCAB9766" box="[642,850,2064,2085]" italics="true" pageId="8" pageNumber="169">Carinartemis striatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="D9F457CAC4449C37FC833654FC239766" box="[865,986,2064,2085]" pageId="8" pageNumber="169" rank="species">new species</taxonomicNameLabel>
, paratype CUMZ 6205, (C) reproductive system, and (D) attachment of vas deferens into distal end of the penial sheath.
</paragraph>
</caption>
<caption id="A4CC662BC4459C36FF9039FDFC499702" ID-DOI="http://doi.org/10.5281/zenodo.4504091" ID-Zenodo-Dep="4504091" httpUri="https://zenodo.org/record/4504091/files/figure.png" pageId="9" pageNumber="170" startId="9.[114,149,1977,1998]" targetBox="[114,1451,379,1954]" targetPageId="9">
<paragraph id="F00C36A3C4459C36FF9039FDFC499702" blockId="9.[113,1451,1977,2113]" pageId="9" pageNumber="170">
Fig. 8. Internal sculpture of genitalia of
<taxonomicName id="37B34D20C4459C36FE1439FDFD8F988D" authorityName="Siriboon &amp; Sutcharit &amp; Naggs &amp; Rowson &amp; Panha" authorityYear="2014" box="[502,630,1977,1998]" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="170" phylum="Mollusca" rank="genus">
<emphasis id="C2C7EAB1C4459C36FE1439FDFD8F988D" box="[502,630,1977,1998]" italics="true" pageId="9" pageNumber="170">Carinartemis</emphasis>
</taxonomicName>
spp: A–F,
<taxonomicName id="37B34D20C4459C36FD3D39FDFC43988D" authority="Siriboon &amp; Sutcharit &amp; Naggs &amp; Rowson &amp; Panha, 2014" authorityName="Siriboon &amp; Sutcharit &amp; Naggs &amp; Rowson &amp; Panha" authorityYear="2014" box="[735,954,1977,1998]" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="170" phylum="Mollusca" rank="species" species="vesperus" status="sp. nov.">
<emphasis id="C2C7EAB1C4459C36FD3D39FDFC43988D" box="[735,954,1977,1998]" italics="true" pageId="9" pageNumber="170">Carinartemis vesperus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="D9F457CAC4459C36FC2539FDFBC7988D" box="[967,1086,1977,1998]" pageId="9" pageNumber="170" rank="species">new species</taxonomicNameLabel>
, paratype CUMZ 6201, (A) details of atrial pore on the atrium surface, (B) high magnification of penial hooks, (C) top view of penial hook, (D) arrangement of vaginal folds with hooks, (E) high magnification of vaginal hook, and (F) radula morphology. G–K,
<taxonomicName id="37B34D20C4459C36FC5139B7FB78974B" authority="Siriboon &amp; Sutcharit &amp; Naggs &amp; Rowson &amp; Panha, 2014" authorityName="Siriboon &amp; Sutcharit &amp; Naggs &amp; Rowson &amp; Panha" authorityYear="2014" box="[947,1153,2035,2056]" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="170" phylum="Mollusca" rank="species" species="striatus" status="sp. nov.">
<emphasis id="C2C7EAB1C4459C36FC5139B7FB78974B" box="[947,1153,2035,2056]" italics="true" pageId="9" pageNumber="170">Carinartemis striatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="D9F457CAC4459C36FB6C39B7FAFD974B" box="[1166,1284,2035,2056]" pageId="9" pageNumber="170" rank="species">new species</taxonomicNameLabel>
, paratype CUMZ 6205, (G) details of atrium surface, (H) low magnification shows arrangement of penial hooks, (I) low magnification shows arrangement of vaginal folds, (J) high magnification of penial hooks, and (K) radula morphology.
</paragraph>
</caption>
<subSubSection id="B8A96528C4469C35FF6A3F1AFEEB9756" pageId="10" pageNumber="171" type="description">
<paragraph id="F00C36A3C4469C35FF6A3F1AFDF59DB6" blockId="10.[136,776,350,757]" pageId="10" pageNumber="171">
<emphasis id="C2C7EAB1C4469C35FF6A3F1AFED99E36" bold="true" box="[136,288,350,373]" pageId="10" pageNumber="171">Description.</emphasis>
Shell suboblique-heliciform, white and translucent; whorls 7, spire conical with distinct suture. Shell surface glossy, with thin and fine transverse ridges; ridges disappear below periphery of penultimate whorl and around umbilicus. Embryonic shell with about 2½ whorls and smooth surface; following whorls regularly expanding. Shell periphery wide and sharply keeled around nearly entire penultimate whorl; last whorl axially deflected. Umbilicus widely open and deep with short longitudinal furrow. Aperture subcircular, peristome discontinuous, thickened, and expanded. Apertural dentition usually without lamellae, but sometimes with small parietal lamella, and rarely with very small other lamellae (
<figureCitation id="68882A26C4469C35FE483C9AFE069DB6" box="[426,511,734,757]" captionStart="Fig" captionStartId="7.[113,148,2005,2026]" captionTargetBox="[113,1453,825,1982]" captionTargetId="figure-13@7.[113,1453,825,1982]" captionTargetPageId="7" captionText="Fig. 6. A, Shell of Stemmatopsis poirieri lectotype MNHN 26980; B–E, Shells of Carinartemis vesperus, new species, (B) holotype CUMZ 5019 with apertural dentition, (C, D) paratype CUMZ 6200 from Khao Kling, Khao Yoi District, Phetchaburi, and (E) specimen NMW 1955.158.25233 from Khao Yoi, Petchaburi; F, G, Carinartemis striatus, new species, (F) holotype CUMZ 6203 with apertural dentition, and (G) paratype CUMZ 6117 from Tam Kra Sae, Sai Yok, Kanchanaburi." figureDoi="http://doi.org/10.5281/zenodo.4504087" httpUri="https://zenodo.org/record/4504087/files/figure.png" pageId="10" pageNumber="171">Fig. 6B</figureCitation>
).
</paragraph>
<paragraph id="F00C36A3C4469C35FF6A3D5AFF169CB6" blockId="10.[136,775,798,1013]" pageId="10" pageNumber="171">
<emphasis id="C2C7EAB1C4469C35FF6A3D5AFF1B9C76" bold="true" box="[136,226,798,821]" pageId="10" pageNumber="171">Radula.</emphasis>
Teeth arranged in anteriorly V-shaped rows, each row containing 47–49 teeth with the formula (23-24)-1-(23- 24); central tooth small, short, triangular with pointed cusp. Lateral and marginal teeth undifferentiated, unicuspid, and lanceolate; lateral teeth gradually reduced in length and size with outer teeth much smaller and shorter than inner teeth (
<figureCitation id="68882A26C4469C35FF723D9AFF1B9CB6" box="[144,226,990,1013]" captionStart="Fig" captionStartId="9.[114,149,1977,1998]" captionTargetBox="[114,1451,379,1954]" captionTargetId="figure-54@9.[108,1458,374,1960]" captionTargetPageId="9" captionText="Fig. 8. Internal sculpture of genitalia of Carinartemis spp: A–F, Carinartemis vesperus, new species, paratype CUMZ 6201, (A) details of atrial pore on the atrium surface, (B) high magnification of penial hooks, (C) top view of penial hook, (D) arrangement of vaginal folds with hooks, (E) high magnification of vaginal hook, and (F) radula morphology. G–K, Carinartemis striatus, new species, paratype CUMZ 6205, (G) details of atrium surface, (H) low magnification shows arrangement of penial hooks, (I) low magnification shows arrangement of vaginal folds, (J) high magnification of penial hooks, and (K) radula morphology." figureDoi="http://doi.org/10.5281/zenodo.4504091" httpUri="https://zenodo.org/record/4504091/files/figure.png" pageId="10" pageNumber="171">Fig. 8F</figureCitation>
).
</paragraph>
<paragraph id="F00C36A3C4469C35FF6A3A5AFD189A16" blockId="10.[136,775,1054,1365]" pageId="10" pageNumber="171">
<emphasis id="C2C7EAB1C4469C35FF6A3A5AFEC19B76" bold="true" box="[136,312,1054,1077]" pageId="10" pageNumber="171">Genital organs.</emphasis>
Atrium (at) short. Proximal penis (p) long, slender; becoming slightly broader with short pouch before tapering distally. Penial sheath (ps) thick, extending about fourth-fifths of total penis length; penial sheath retractor muscle (psr) very thin, originating at atrium and body wall (
<figureCitation id="68882A26C4469C35FF723AFAFF1D9B96" box="[144,228,1214,1237]" captionStart="Fig" captionStartId="8.[136,171,2035,2056]" captionTargetBox="[146,1462,502,2007]" captionTargetId="figure-66@8.[131,1478,489,2021]" captionTargetPageId="8" captionText="Fig. 7. Genitalia of Carinartemis spp: A, B, Carinartemis vesperus, new species, paratype CUMZ 6201, (A) reproductive system, and (B) insertion of vas deferens into penial sheath; C, D, Carinartemis striatus, new species, paratype CUMZ 6205, (C) reproductive system, and (D) attachment of vas deferens into distal end of the penial sheath." figureDoi="http://doi.org/10.5281/zenodo.4504089" httpUri="https://zenodo.org/record/4504089/files/figure.png" pageId="10" pageNumber="171">Fig. 7A</figureCitation>
). Vas deferens (vd) long, slender; slightly broadened before tapering distally; passes through about one-tenth of penial sheath length before entering into penis distally (
<figureCitation id="68882A26C4469C35FD3E3ABAFF539A76" captionStart="Fig" captionStartId="8.[136,171,2035,2056]" captionTargetBox="[146,1462,502,2007]" captionTargetId="figure-66@8.[131,1478,489,2021]" captionTargetPageId="8" captionText="Fig. 7. Genitalia of Carinartemis spp: A, B, Carinartemis vesperus, new species, paratype CUMZ 6201, (A) reproductive system, and (B) insertion of vas deferens into penial sheath; C, D, Carinartemis striatus, new species, paratype CUMZ 6205, (C) reproductive system, and (D) attachment of vas deferens into distal end of the penial sheath." figureDoi="http://doi.org/10.5281/zenodo.4504089" httpUri="https://zenodo.org/record/4504089/files/figure.png" pageId="10" pageNumber="171">Fig. 7B</figureCitation>
). Penial retractor muscle (pr) thin and very long, inserting distally on penis at the penis and vas deferens junction.
</paragraph>
<paragraph id="F00C36A3C4469C35FF6A3B3AFDF29956" blockId="10.[136,775,1406,1557]" pageId="10" pageNumber="171">
Internal wall of atrium smooth with numerous atrial pores (
<figureCitation id="68882A26C4469C35FF723BDAFF119AF6" box="[144,232,1438,1461]" captionStart="Fig" captionStartId="9.[114,149,1977,1998]" captionTargetBox="[114,1451,379,1954]" captionTargetId="figure-54@9.[108,1458,374,1960]" captionTargetPageId="9" captionText="Fig. 8. Internal sculpture of genitalia of Carinartemis spp: A–F, Carinartemis vesperus, new species, paratype CUMZ 6201, (A) details of atrial pore on the atrium surface, (B) high magnification of penial hooks, (C) top view of penial hook, (D) arrangement of vaginal folds with hooks, (E) high magnification of vaginal hook, and (F) radula morphology. G–K, Carinartemis striatus, new species, paratype CUMZ 6205, (G) details of atrium surface, (H) low magnification shows arrangement of penial hooks, (I) low magnification shows arrangement of vaginal folds, (J) high magnification of penial hooks, and (K) radula morphology." figureDoi="http://doi.org/10.5281/zenodo.4504091" httpUri="https://zenodo.org/record/4504091/files/figure.png" pageId="10" pageNumber="171">Fig. 8A</figureCitation>
). Penial papillae absent; pale brown penial hooks present, about 40 hooks/200 μm
<superScript id="07C69BEBC4469C35FE3E3BF8FE1D9A89" attach="left" box="[476,484,1468,1482]" fontSize="6" pageId="10" pageNumber="171">2</superScript>
. Penial hooks short (&lt;
<quantity id="374B9B46C4469C35FD3A3BFAFF4B9AB6" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="171" unit="mm" value="0.03">0.03 mm</quantity>
in length), expanding at base, tips obtuse and curved towards genital orifice (
<figureCitation id="68882A26C4469C35FE6B3BBAFE069956" box="[393,511,1534,1557]" captionStart="Fig" captionStartId="9.[114,149,1977,1998]" captionTargetBox="[114,1451,379,1954]" captionTargetId="figure-54@9.[108,1458,374,1960]" captionTargetPageId="9" captionText="Fig. 8. Internal sculpture of genitalia of Carinartemis spp: A–F, Carinartemis vesperus, new species, paratype CUMZ 6201, (A) details of atrial pore on the atrium surface, (B) high magnification of penial hooks, (C) top view of penial hook, (D) arrangement of vaginal folds with hooks, (E) high magnification of vaginal hook, and (F) radula morphology. G–K, Carinartemis striatus, new species, paratype CUMZ 6205, (G) details of atrium surface, (H) low magnification shows arrangement of penial hooks, (I) low magnification shows arrangement of vaginal folds, (J) high magnification of penial hooks, and (K) radula morphology." figureDoi="http://doi.org/10.5281/zenodo.4504091" httpUri="https://zenodo.org/record/4504091/files/figure.png" pageId="10" pageNumber="171">Fig. 8B, C</figureCitation>
).
</paragraph>
<paragraph id="F00C36A3C4469C35FF6A387AFD749816" blockId="10.[136,775,1598,1877]" pageId="10" pageNumber="171">
Vagina (v) very short and stout. Gametolytic duct (gd) long, extending as far as albumin gland; gametolytic sac ovate (gs). Free oviduct (fo) long, proximally large with equivalent diameter with vagina, and tapering to smaller tube distally. Oviduct (ov) enlarged and folded; prostate gland inconspicuous and bound to oviduct. Talon (ta) small, short and club shaped. Hermaphroditic duct (hd) bearing seminal vesicle (sv) about as long as the length from talon to branching point of seminal vesicle (
<figureCitation id="68882A26C4469C35FDCB397AFD799816" box="[553,640,1854,1877]" captionStart="Fig" captionStartId="8.[136,171,2035,2056]" captionTargetBox="[146,1462,502,2007]" captionTargetId="figure-66@8.[131,1478,489,2021]" captionTargetPageId="8" captionText="Fig. 7. Genitalia of Carinartemis spp: A, B, Carinartemis vesperus, new species, paratype CUMZ 6201, (A) reproductive system, and (B) insertion of vas deferens into penial sheath; C, D, Carinartemis striatus, new species, paratype CUMZ 6205, (C) reproductive system, and (D) attachment of vas deferens into distal end of the penial sheath." figureDoi="http://doi.org/10.5281/zenodo.4504089" httpUri="https://zenodo.org/record/4504089/files/figure.png" pageId="10" pageNumber="171">Fig. 7A</figureCitation>
).
</paragraph>
<paragraph id="F00C36A3C4469C35FF6A393AFEEB9756" blockId="10.[136,775,1918,2069]" pageId="10" pageNumber="171">
Vaginal wall with transparent vaginal hooks about 8 hooks/200 μm
<superScript id="07C69BEBC4469C35FEC739D8FED498E9" attach="left" box="[293,301,1948,1962]" fontSize="6" pageId="10" pageNumber="171">2</superScript>
; hooks located on oblique parallel vaginal folds. Vaginal hooks short (&lt;
<quantity id="374B9B46C4469C35FE2239FAFDE79896" box="[448,542,1982,2005]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="171" unit="mm" value="0.04">0.04 mm</quantity>
in length), expanding at base, tips obtuse slightly curving away from genital orifice (
<figureCitation id="68882A26C4469C35FF7239BAFEFF9756" box="[144,262,2046,2069]" captionStart="Fig" captionStartId="9.[114,149,1977,1998]" captionTargetBox="[114,1451,379,1954]" captionTargetId="figure-54@9.[108,1458,374,1960]" captionTargetPageId="9" captionText="Fig. 8. Internal sculpture of genitalia of Carinartemis spp: A–F, Carinartemis vesperus, new species, paratype CUMZ 6201, (A) details of atrial pore on the atrium surface, (B) high magnification of penial hooks, (C) top view of penial hook, (D) arrangement of vaginal folds with hooks, (E) high magnification of vaginal hook, and (F) radula morphology. G–K, Carinartemis striatus, new species, paratype CUMZ 6205, (G) details of atrium surface, (H) low magnification shows arrangement of penial hooks, (I) low magnification shows arrangement of vaginal folds, (J) high magnification of penial hooks, and (K) radula morphology." figureDoi="http://doi.org/10.5281/zenodo.4504091" httpUri="https://zenodo.org/record/4504091/files/figure.png" pageId="10" pageNumber="171">Fig. 8D, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B8A96528C4469C35FCA13EFAFBEF9E76" pageId="10" pageNumber="171" type="distribution">
<paragraph id="F00C36A3C4469C35FCA13EFAFBEF9E76" blockId="10.[835,1474,190,309]" pageId="10" pageNumber="171">
<emphasis id="C2C7EAB1C4469C35FCA13EFAFC2F9F96" bold="true" box="[835,982,190,213]" pageId="10" pageNumber="171">Distribution.</emphasis>
This new species seems to be the restricted to limestone in western
<collectingCountry id="88A47633C4469C35FBA53E9AFB5F9FB6" box="[1095,1190,222,245]" name="Thailand" pageId="10" pageNumber="171">Thailand</collectingCountry>
at the
<typeStatus id="2F088801C4469C35FB103E9AFAD99FB6" box="[1266,1312,222,245]" pageId="10" pageNumber="171">type</typeStatus>
locality and at Khao Yoi,
<collectingRegion id="3277F841C4469C35FC563EBAFBCF9E56" box="[948,1078,254,277]" country="Thailand" name="Phetchaburi" pageId="10" pageNumber="171">Phetchaburi</collectingRegion>
, the limestone karst about
<quantity id="374B9B46C4469C35FAB23EBAFA6B9E56" box="[1360,1426,254,277]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="171" unit="km" value="10.0">10 km</quantity>
east of the
<typeStatus id="2F088801C4469C35FC683F5AFC419E76" box="[906,952,286,309]" pageId="10" pageNumber="171">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="B8A96528C4469C35FCA13F1AFC1C9DB6" pageId="10" pageNumber="171" type="discussion">
<paragraph id="F00C36A3C4469C35FCA13F1AFC1C9DB6" blockId="10.[835,1475,350,757]" pageId="10" pageNumber="171">
<emphasis id="C2C7EAB1C4469C35FCA13F1AFC4B9E36" bold="true" box="[835,946,350,373]" pageId="10" pageNumber="171">Remarks.</emphasis>
Shell variation can be observed in five
<typeStatus id="2F088801C4469C35FABE3F1AFA3B9E36" box="[1372,1474,350,373]" pageId="10" pageNumber="171" type="paratype">paratypes</typeStatus>
by having very small parietal lamella and the detectable upper palatal, palatal, basal and columellar lamellae (
<figureCitation id="68882A26C4469C35FA753FDAFC9E9E96" captionStart="Fig" captionStartId="7.[113,148,2005,2026]" captionTargetBox="[113,1453,825,1982]" captionTargetId="figure-13@7.[113,1453,825,1982]" captionTargetPageId="7" captionText="Fig. 6. A, Shell of Stemmatopsis poirieri lectotype MNHN 26980; B–E, Shells of Carinartemis vesperus, new species, (B) holotype CUMZ 5019 with apertural dentition, (C, D) paratype CUMZ 6200 from Khao Kling, Khao Yoi District, Phetchaburi, and (E) specimen NMW 1955.158.25233 from Khao Yoi, Petchaburi; F, G, Carinartemis striatus, new species, (F) holotype CUMZ 6203 with apertural dentition, and (G) paratype CUMZ 6117 from Tam Kra Sae, Sai Yok, Kanchanaburi." figureDoi="http://doi.org/10.5281/zenodo.4504087" httpUri="https://zenodo.org/record/4504087/files/figure.png" pageId="10" pageNumber="171">Fig. 6D</figureCitation>
). In addition, a single shell from Khao Yoi,
<collectingRegion id="3277F841C4469C35FAA13FFAFA3B9E96" box="[1347,1474,446,469]" country="Thailand" name="Phetchaburi" pageId="10" pageNumber="171">Phetchaburi</collectingRegion>
(
<figureCitation id="68882A26C4469C35FCA93F9AFC639EB6" box="[843,922,478,501]" captionStart="Fig" captionStartId="7.[113,148,2005,2026]" captionTargetBox="[113,1453,825,1982]" captionTargetId="figure-13@7.[113,1453,825,1982]" captionTargetPageId="7" captionText="Fig. 6. A, Shell of Stemmatopsis poirieri lectotype MNHN 26980; B–E, Shells of Carinartemis vesperus, new species, (B) holotype CUMZ 5019 with apertural dentition, (C, D) paratype CUMZ 6200 from Khao Kling, Khao Yoi District, Phetchaburi, and (E) specimen NMW 1955.158.25233 from Khao Yoi, Petchaburi; F, G, Carinartemis striatus, new species, (F) holotype CUMZ 6203 with apertural dentition, and (G) paratype CUMZ 6117 from Tam Kra Sae, Sai Yok, Kanchanaburi." figureDoi="http://doi.org/10.5281/zenodo.4504087" httpUri="https://zenodo.org/record/4504087/files/figure.png" pageId="10" pageNumber="171">Fig. 6E</figureCitation>
; NMW.1955.158.25233) has a strong parietal lamella and four other very small peripheral lamellae. However, we considered this conspecific based on the characters of thin and fine transverse ridges and broadly expanded apertural lip, and because it was collected within the range of this species. However, further anatomical and molecular data of the newly collected specimen with precise locality will elucidate whether this represents intraspecific variation or a separate taxon.
</paragraph>
</subSubSection>
</treatment>
</document>
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