244 lines
27 KiB
XML
244 lines
27 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.898.39265" ID-GBIF-Dataset="9dbc167e-c273-42f5-82a0-dedb1dfce5bc" ID-PMC="PMC6917625" ID-Pensoft-Pub="1313-2970-898-121" ID-Pensoft-UUID="2A2FB0D263AA5E498C5C85EA4031053A" ID-PubMed="31866734" ID-ZooBank="9B537DC38DB9459E97717687AFA19244" ModsDocID="1313-2970-898-121" checkinTime="1575997087753" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I., Semenyuk, Irina, Efeykin, Boris D. & Panha, Somsak" docDate="2019" docId="77D6A3E35F285A719541BBBA5DFCF21B" docLanguage="en" docName="ZooKeys 898: 121-158" docOrigin="ZooKeys 898" docSource="http://dx.doi.org/10.3897/zookeys.898.39265" docTitle="Orthomorpha caramel Likhitrakarn, Golovatch, Semenyuk, Efeykin & Panha, 2019, sp. nov." docType="treatment" docUuid="0668527A-B7A6-4EE1-B5CD-7BA8B49BC540" docUuidSource="ZooBank" docVersion="5" id="2A2FB0D263AA5E498C5C85EA4031053A" lastPageNumber="121" masterDocId="2A2FB0D263AA5E498C5C85EA4031053A" masterDocTitle="Review of the millipede genus Orthomorpha Bollman, 1893 (Diplopoda, Polydesmida, Paradoxosomatidae) in Vietnam, with several new records and descriptions of two new species" masterLastPageNumber="158" masterPageNumber="121" pageNumber="121" updateTime="1668168219523" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Review of the millipede genus Orthomorpha Bollman, 1893 (Diplopoda, Polydesmida, Paradoxosomatidae) in Vietnam, with several new records and descriptions of two new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Likhitrakarn, Natdanai</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Golovatch, Sergei I.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Semenyuk, Irina</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Efeykin, Boris D.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>898</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>121</mods:start>
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<mods:end>158</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.898.39265</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.898.39265</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-898-121</mods:identifier>
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<mods:identifier type="ZooBank">9B537DC38DB9459E97717687AFA19244</mods:identifier>
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<mods:identifier type="Pensoft-UUID">2A2FB0D263AA5E498C5C85EA4031053A</mods:identifier>
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<treatment ID-GBIF-Taxon="160689311" LSID="urn:lsid:zoobank.org:act:0668527A-B7A6-4EE1-B5CD-7BA8B49BC540" httpUri="http://treatment.plazi.org/id/77D6A3E35F285A719541BBBA5DFCF21B" lastPageNumber="121" pageId="0" pageNumber="121">
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<subSubSection pageId="0" pageNumber="121" type="nomenclature">
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<paragraph pageId="0" pageNumber="121">
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<taxonomicName LSID="77D6A3E3-5F28-5A71-9541-BBBA5DFCF21B" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha caramel" order="Polydesmida" pageId="0" pageNumber="121" phylum="Arthropoda" rank="species" species="caramel">Orthomorpha caramel</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="121">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Orthomorpha caramel sp. nov., ♂ holotype A habitus, live colouration B, C anterior part of body, dorsal and lateral views, respectively D, E segments 10 and 11, dorsal and lateral views, respectively F-H posterior part of body, dorsal, ventral and lateral views, respectively I, J sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.898.39265.figure12" httpUri="https://binary.pensoft.net/fig/363867" pageId="0" pageNumber="121" tableDoi="10.3897/zookeys.898.39265.figure12">Figs 12</figureCitation>
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,
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Orthomorpha caramel sp. nov., ♂ holotype, right gonopod A, B lateral and mesal views, respectively C distal part, suboral view. Scale bars: 0.5 mm" figureDoi="10.3897/zookeys.898.39265.figure13" httpUri="https://binary.pensoft.net/fig/363868" pageId="0" pageNumber="121" tableDoi="10.3897/zookeys.898.39265.figure13">13</figureCitation>
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,
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Orthomorpha caramel sp. nov., ♂ holotype, right gonopod A, B mesal and lateral views, respectively C-G distal part, caudal, mesal, oral, submesal and subcaudal views, respectively." figureDoi="10.3897/zookeys.898.39265.figure14" httpUri="https://binary.pensoft.net/fig/363869" pageId="0" pageNumber="121" tableDoi="10.3897/zookeys.898.39265.figure14">14</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="121" type="type material">
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<paragraph pageId="0" pageNumber="121">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="121">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="121">Holotype</emphasis>
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♂ (ZMUM Rd 4197), Vietnam, Gia Lai Province, Kon Ka Kinh National Park, forest near the village Krong,
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<geoCoordinate degrees="14" direction="north" minutes="18" orientation="latitude" precision="15" seconds="03" value="14.300834">14°18'03"N</geoCoordinate>
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,
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<geoCoordinate degrees="108" direction="east" minutes="26" orientation="longitude" precision="15" seconds="42" value="108.445">108°26'42"E</geoCoordinate>
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, 600 m a.s.l., mixed tropical forest on slopes to a small stream, on tree trunk, daytime, 13.V.2017, I. Semenyuk leg.
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</paragraph>
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<paragraph pageId="0" pageNumber="121">
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<emphasis bold="true" pageId="0" pageNumber="121">
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<emphasis italics="true" pageId="0" pageNumber="121">Paratypes</emphasis>
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.
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</emphasis>
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1 ♂ (ZMUM Rd 4198), same locality, together with holotype; 1 ♂ (ZMUM), same locality,
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<geoCoordinate degrees="14" direction="north" minutes="17" orientation="latitude" precision="15" seconds="46" value="14.296111">14°17'46"N</geoCoordinate>
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,
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<geoCoordinate degrees="108" direction="east" minutes="26" orientation="longitude" precision="15" seconds="41" value="108.444725">108°26'41"E</geoCoordinate>
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, 750 m a.s.l., disturbed forest with dominating bamboo on hills, on tree trunk, daytime 18.V.2017, all leg. I. Semenyuk.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="121" type="name">
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<paragraph pageId="0" pageNumber="121">Name.</paragraph>
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<paragraph pageId="0" pageNumber="121">
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A noun in apposition, to emphasize the general
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<taxonomicName lsidName="caramel" pageId="0" pageNumber="121" rank="species" species="caramel">caramel</taxonomicName>
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colouration of the animals.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="121" type="diagnosis">
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<paragraph pageId="0" pageNumber="121">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="121">
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Using the latest key (
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.131.1921" author="Likhitrakarn, N" journalOrPublisher="ZooKeys" pageId="0" pageNumber="121" pagination="1 - 161" refId="B23" refString="Likhitrakarn, N, Golovatch, SI, Panha, S, 2011. Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 131: 1 - 161" title="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)." url="https://doi.org/10.3897/zookeys.131.1921" volume="131" year="2011">Likhitrakarn et al. 2011</bibRefCitation>
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), distinguished from all known congeners by the tip of the solenophore being very faintly bifid, with a nearly smooth terminal lobe bearing a minute lobule at the base; in the gonopod structure it is similar to
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<taxonomicName lsidName="O. tuberculifera" pageId="0" pageNumber="121" rank="species" species="tuberculifera">
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<emphasis italics="true" pageId="0" pageNumber="121">O. tuberculifera</emphasis>
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</taxonomicName>
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Likhitrakarn, Golovatch & Panha, 2011, but differs in the rather smooth and shining dorsal tegument devoid of tubercles, coupled with the particular colouration.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="121" type="description">
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<paragraph pageId="0" pageNumber="121">Description.</paragraph>
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<paragraph pageId="0" pageNumber="121">Length of holotype 36.5 mm (♂), width of midbody pro- and metazonae 2.8 and 4.2 mm, respectively. Paratype 31.5-34.5 mm long, 2.6-2.8 and 4.0-4.1 mm wide on midbody pro- and metazonae, respectively.</paragraph>
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<paragraph pageId="0" pageNumber="121">
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Colouration of live animals dark chocolate brown (
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Orthomorpha caramel sp. nov., ♂ holotype A habitus, live colouration B, C anterior part of body, dorsal and lateral views, respectively D, E segments 10 and 11, dorsal and lateral views, respectively F-H posterior part of body, dorsal, ventral and lateral views, respectively I, J sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.898.39265.figure12" httpUri="https://binary.pensoft.net/fig/363867" pageId="0" pageNumber="121" tableDoi="10.3897/zookeys.898.39265.figure12">Fig. 12A</figureCitation>
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); metaterga, paraterga and epiproct
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<taxonomicName lsidName="caramel" pageId="0" pageNumber="121" rank="species" species="caramel">caramel</taxonomicName>
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in colour; head, antennae and legs dark brownish; colouration in alcohol after one year of preservation chocolate brown or faded to light brownish (
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Orthomorpha caramel sp. nov., ♂ holotype A habitus, live colouration B, C anterior part of body, dorsal and lateral views, respectively D, E segments 10 and 11, dorsal and lateral views, respectively F-H posterior part of body, dorsal, ventral and lateral views, respectively I, J sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.898.39265.figure12" httpUri="https://binary.pensoft.net/fig/363867" pageId="0" pageNumber="121" tableDoi="10.3897/zookeys.898.39265.figure12">
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Fig. 12
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<normalizedToken originalValue="B–H">B-H</normalizedToken>
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</figureCitation>
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); metaterga, paraterga and epiproct
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<taxonomicName lsidName="caramel" pageId="0" pageNumber="121" rank="species" species="caramel">caramel</taxonomicName>
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to light brownish in colour; legs and sterna light brown to pale yellow; head and antennae dark brownish to brown.
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</paragraph>
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<paragraph pageId="0" pageNumber="121">
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Clypeolabral region densely setose, vertigial region sparsely so; epicranial suture distinct. Antennae rather long (
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Orthomorpha caramel sp. nov., ♂ holotype A habitus, live colouration B, C anterior part of body, dorsal and lateral views, respectively D, E segments 10 and 11, dorsal and lateral views, respectively F-H posterior part of body, dorsal, ventral and lateral views, respectively I, J sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.898.39265.figure12" httpUri="https://binary.pensoft.net/fig/363867" pageId="0" pageNumber="121" tableDoi="10.3897/zookeys.898.39265.figure12">Fig. 12A</figureCitation>
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), reaching the end of body segment 3 when stretched dorsally. In width, head <segment 3 = 4 <collum <segment 5 <2 <6-17, body gently and gradually tapering thereafter. Collum with three transverse rows of setae: 4+4 in anterior, 2+2 in intermediate, and 3+3 in posterior row; a slight furrow laterally in posterior 1/3; caudal corner of paraterga very narrowly rounded, slightly upturned, but not drawn behind rear margin (
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Orthomorpha caramel sp. nov., ♂ holotype A habitus, live colouration B, C anterior part of body, dorsal and lateral views, respectively D, E segments 10 and 11, dorsal and lateral views, respectively F-H posterior part of body, dorsal, ventral and lateral views, respectively I, J sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.898.39265.figure12" httpUri="https://binary.pensoft.net/fig/363867" pageId="0" pageNumber="121" tableDoi="10.3897/zookeys.898.39265.figure12">Fig. 12B, C</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.898.39265.figure12" httpUri="https://binary.pensoft.net/fig/363867" pageId="0" pageNumber="121" start="Figure 12" startId="F12">
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<paragraph pageId="0" pageNumber="121">
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<emphasis bold="true" pageId="0" pageNumber="121">Figure 12.</emphasis>
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha caramel" order="Polydesmida" pageId="0" pageNumber="121" phylum="Arthropoda" rank="species" species="caramel">
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<emphasis italics="true" pageId="0" pageNumber="121">Orthomorpha caramel</emphasis>
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</taxonomicName>
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sp. nov., ♂ holotype
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<emphasis bold="true" pageId="0" pageNumber="121">A</emphasis>
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habitus, live colouration
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<emphasis bold="true" pageId="0" pageNumber="121">B, C</emphasis>
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anterior part of body, dorsal and lateral views, respectively
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<emphasis bold="true" pageId="0" pageNumber="121">D, E</emphasis>
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segments 10 and 11, dorsal and lateral views, respectively
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<emphasis bold="true" pageId="0" pageNumber="121">
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<normalizedToken originalValue="F–H">F-H</normalizedToken>
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</emphasis>
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posterior part of body, dorsal, ventral and lateral views, respectively
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<emphasis bold="true" pageId="0" pageNumber="121">I, J</emphasis>
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sternal cones between coxae 4, subcaudal and sublateral views, respectively.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="121">Tegument shining, prozonae finely shagreened, metaterga rugose to rugulose, surface below paraterga finely microgranulate and rugulose. Postcollum metaterga each with two transverse rows of short, mostly abraded setae traceable only as insertion points, short wrinkles or minute tubercles: anterior (pre-sulcus) row with 2+2 mostly abraded setae traceable only as insertion points; posterior (postsulcus) row with 3+3 setae borne on minute tubercles or short wrinkles. Tergal setae long, slender, ca. 1/3 of metatergal length. Axial line rather clear, especially so on metazonae.</paragraph>
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<paragraph pageId="0" pageNumber="121">
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Paraterga very strongly developed (
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Orthomorpha caramel sp. nov., ♂ holotype A habitus, live colouration B, C anterior part of body, dorsal and lateral views, respectively D, E segments 10 and 11, dorsal and lateral views, respectively F-H posterior part of body, dorsal, ventral and lateral views, respectively I, J sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.898.39265.figure12" httpUri="https://binary.pensoft.net/fig/363867" pageId="0" pageNumber="121" tableDoi="10.3897/zookeys.898.39265.figure12">
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Fig. 12
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<normalizedToken originalValue="B–H">B-H</normalizedToken>
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</figureCitation>
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), mostly upturned, all lying below dorsum, set at ca. upper 1/4 of midbody height, nearly level with dorsum on segments 15-18, caudal corner always spiniform and narrowly rounded, extending beyond rear tergal margin; in lateral view, paraterga thinner in poreless segments and modestly enlarged in pore-bearing ones.
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</paragraph>
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<paragraph pageId="0" pageNumber="121">
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Paraterga 2 broad, anterior edge evidently convex, lateral edge with one larger incision in anterior 1/3, one smaller, but evident incision in the middle and a faint furrow at posterior 1/3; posterior edge oblique (
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Orthomorpha caramel sp. nov., ♂ holotype A habitus, live colouration B, C anterior part of body, dorsal and lateral views, respectively D, E segments 10 and 11, dorsal and lateral views, respectively F-H posterior part of body, dorsal, ventral and lateral views, respectively I, J sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.898.39265.figure12" httpUri="https://binary.pensoft.net/fig/363867" pageId="0" pageNumber="121" tableDoi="10.3897/zookeys.898.39265.figure12">Fig. 12B, D, F</figureCitation>
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). Anterior edges of following paraterga strongly and regularly rounded, lateral edge with only one small, but evident incision in anterior 1/3, posterior edge clearly concave, especially strongly emarginate in segments 17-19 (
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Orthomorpha caramel sp. nov., ♂ holotype A habitus, live colouration B, C anterior part of body, dorsal and lateral views, respectively D, E segments 10 and 11, dorsal and lateral views, respectively F-H posterior part of body, dorsal, ventral and lateral views, respectively I, J sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.898.39265.figure12" httpUri="https://binary.pensoft.net/fig/363867" pageId="0" pageNumber="121" tableDoi="10.3897/zookeys.898.39265.figure12">
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Fig. 12
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<normalizedToken originalValue="F–H">F-H</normalizedToken>
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</figureCitation>
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). Calluses on paraterga 2-4 strongly delimited by a sulcus only dorsally, on following paraterga both dorsally and ventrally.
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</paragraph>
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<paragraph pageId="0" pageNumber="121">
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Ozopores evident, lateral, lying inside an ovoid groove at ca. 1/4 in front of caudal corner. Transverse sulcus usually distinct (
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Orthomorpha caramel sp. nov., ♂ holotype A habitus, live colouration B, C anterior part of body, dorsal and lateral views, respectively D, E segments 10 and 11, dorsal and lateral views, respectively F-H posterior part of body, dorsal, ventral and lateral views, respectively I, J sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.898.39265.figure12" httpUri="https://binary.pensoft.net/fig/363867" pageId="0" pageNumber="121" tableDoi="10.3897/zookeys.898.39265.figure12">Fig. 12C, E, H</figureCitation>
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), complete on metaterga 5-18, incomplete and nearly wanting on segments 4 and 19, wave-shaped, rather shallow, nearly reaching the bases of paraterga, ribbed at bottom. Stricture between pro- and metazona rather wide and deep, beaded at bottom down to base of paraterga (
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Orthomorpha caramel sp. nov., ♂ holotype A habitus, live colouration B, C anterior part of body, dorsal and lateral views, respectively D, E segments 10 and 11, dorsal and lateral views, respectively F-H posterior part of body, dorsal, ventral and lateral views, respectively I, J sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.898.39265.figure12" httpUri="https://binary.pensoft.net/fig/363867" pageId="0" pageNumber="121" tableDoi="10.3897/zookeys.898.39265.figure12">Fig. 12B, D, F</figureCitation>
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). Pleurosternal carinae complete crests with a sharp caudal tooth on segments 2-7(8), thereafter bulged anteriorly and with a small, sharp, caudal tooth on segments 8-10, the tooth gradually reduced into small, caudally roughly granulate crests until segment 12 (
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Orthomorpha caramel sp. nov., ♂ holotype A habitus, live colouration B, C anterior part of body, dorsal and lateral views, respectively D, E segments 10 and 11, dorsal and lateral views, respectively F-H posterior part of body, dorsal, ventral and lateral views, respectively I, J sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.898.39265.figure12" httpUri="https://binary.pensoft.net/fig/363867" pageId="0" pageNumber="121" tableDoi="10.3897/zookeys.898.39265.figure12">Fig. 12C, E, H</figureCitation>
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). Epiproct (
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Orthomorpha caramel sp. nov., ♂ holotype A habitus, live colouration B, C anterior part of body, dorsal and lateral views, respectively D, E segments 10 and 11, dorsal and lateral views, respectively F-H posterior part of body, dorsal, ventral and lateral views, respectively I, J sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.898.39265.figure12" httpUri="https://binary.pensoft.net/fig/363867" pageId="0" pageNumber="121" tableDoi="10.3897/zookeys.898.39265.figure12">Fig. 12F, G</figureCitation>
|
||
) conical, flattened dorsoventrally, with two evident apical papillae; tip subtruncate; pre-apical papillae small, but evident, lying rather close to tip. Hypoproct pentagonal, setiferous knobs at caudal edge evident and well-separated.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.898.39265.figure13" httpUri="https://binary.pensoft.net/fig/363868" pageId="0" pageNumber="121" start="Figure 13" startId="F13">
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<emphasis bold="true" pageId="0" pageNumber="121">Figure 13.</emphasis>
|
||
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha caramel" order="Polydesmida" pageId="0" pageNumber="121" phylum="Arthropoda" rank="species" species="caramel">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Orthomorpha caramel</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., ♂ holotype, right gonopod
|
||
<emphasis bold="true" pageId="0" pageNumber="121">A, B</emphasis>
|
||
lateral and mesal views, respectively
|
||
<emphasis bold="true" pageId="0" pageNumber="121">C</emphasis>
|
||
distal part, suboral view. Scale bars: 0.5 mm
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Sterna sparsely setose, without modifications except for two rather large and long, fully separated, sternal cones between ♂ coxae 4 (
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Orthomorpha caramel sp. nov., ♂ holotype A habitus, live colouration B, C anterior part of body, dorsal and lateral views, respectively D, E segments 10 and 11, dorsal and lateral views, respectively F-H posterior part of body, dorsal, ventral and lateral views, respectively I, J sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.898.39265.figure12" httpUri="https://binary.pensoft.net/fig/363867" pageId="0" pageNumber="121" tableDoi="10.3897/zookeys.898.39265.figure12">Fig. 12I, J</figureCitation>
|
||
). A paramedian pair of evident tubercles in front of gonopod aperture. Legs long and slender, midbody ones ca. 1.2-1.4 times as long as body height, prefemora without modifications, ♂ tarsal brushes present until legs 15.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Gonopods (
|
||
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Orthomorpha caramel sp. nov., ♂ holotype, right gonopod A, B lateral and mesal views, respectively C distal part, suboral view. Scale bars: 0.5 mm" figureDoi="10.3897/zookeys.898.39265.figure13" httpUri="https://binary.pensoft.net/fig/363868" pageId="0" pageNumber="121" tableDoi="10.3897/zookeys.898.39265.figure13">Figs 13</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Orthomorpha caramel sp. nov., ♂ holotype, right gonopod A, B mesal and lateral views, respectively C-G distal part, caudal, mesal, oral, submesal and subcaudal views, respectively." figureDoi="10.3897/zookeys.898.39265.figure14" httpUri="https://binary.pensoft.net/fig/363869" pageId="0" pageNumber="121" tableDoi="10.3897/zookeys.898.39265.figure14">14</figureCitation>
|
||
) simple. Coxa slender and long, with several setae distoventrally. Femorite ca. 2 times as long as prefemoral (= strongly setose) part. Femorite slender, suberect to slightly curved,
|
||
<normalizedToken originalValue="“postfemoral”">"postfemoral"</normalizedToken>
|
||
portion demarcated by an oblique lateral sulcus; solenophore moderately twisted and curved, tip of solenophore very faintly bifid, with a nearly smooth terminal lobe bearing a minute lobule at base; solenomere long and flagelliform, as usual.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="121" type="remarks">
|
||
<paragraph pageId="0" pageNumber="121">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
The biology and behaviour of this species are very similar to those of
|
||
<taxonomicName lsidName="O. vietnamica" pageId="0" pageNumber="121" rank="species" species="vietnamica">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">O. vietnamica</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. During field observations in May 2017, millipedes occurred mainly on tree trunks. Mating was also recorded, but no females were collected.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.898.39265.figure14" httpUri="https://binary.pensoft.net/fig/363869" pageId="0" pageNumber="121" start="Figure 14" startId="F14">
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<emphasis bold="true" pageId="0" pageNumber="121">Figure 14.</emphasis>
|
||
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha caramel" order="Polydesmida" pageId="0" pageNumber="121" phylum="Arthropoda" rank="species" species="caramel">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Orthomorpha caramel</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., ♂ holotype, right gonopod
|
||
<emphasis bold="true" pageId="0" pageNumber="121">A, B</emphasis>
|
||
mesal and lateral views, respectively
|
||
<emphasis bold="true" pageId="0" pageNumber="121">
|
||
<normalizedToken originalValue="C–G">C-G</normalizedToken>
|
||
</emphasis>
|
||
distal part, caudal, mesal, oral, submesal and subcaudal views, respectively.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |