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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1149.99651" ID-Pensoft-Pub="1313-2970-1149-181" ID-Pensoft-UUID="49B1C50F79A8585499B05B33E7367DF2" ID-ZooBank="52FBD43553204F088EBBF18230DC9B62" ModsDocID="1313-2970-1149-181" checkinTime="1677176331870" checkinUser="pensoft" docAuthor="Nguyen, Anh D., Stoev, Pavel, Nguyen, Lien T. P. &amp; Vu, Tam T." docDate="2023" docId="779EF539627B50EC9FB930E0AFFD9B47" docLanguage="en" docName="ZooKeys 1149: 181-195" docOrigin="ZooKeys 1149" docPubDate="2023-02-23" docSource="http://dx.doi.org/10.3897/zookeys.1149.99651" docTitle="Paracortina kyrang Nguyen &amp; Stoev &amp; Nguyen &amp; Vu 2023, sp. nov." docType="treatment" docUuid="22146F0D-BFA6-4F91-84B5-CEB60238183D" docUuidSource="ZooBank" docVersion="1" id="49B1C50F79A8585499B05B33E7367DF2" lastPageNumber="181" masterDocId="49B1C50F79A8585499B05B33E7367DF2" masterDocTitle="A new species of Paracortina from a Vietnamese cave, with remarkable secondary sexual characters in males (Callipodida, Paracortinidae)" masterLastPageNumber="195" masterPageNumber="181" pageNumber="181" updateTime="1677176331870" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of Paracortina from a Vietnamese cave, with remarkable secondary sexual characters in males (Callipodida, Paracortinidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nguyen, Anh D.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9273-0040</mods:nameIdentifier>
<mods:affiliation>Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam</mods:affiliation>
<mods:nameIdentifier type="email">ducanh410@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Stoev, Pavel</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5702-5677</mods:nameIdentifier>
<mods:affiliation>National Museum of Natural History, Bulgarian Academy of Sciences, Tsar Osvoboditel Blvd. 1, 1000 Sofia, Bulgaria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nguyen, Lien T. P.</mods:namePart>
<mods:affiliation>Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vu, Tam T.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1145-975X</mods:nameIdentifier>
<mods:affiliation>Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-23</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1149</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>181</mods:start>
<mods:end>195</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1149.99651</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1149.99651</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1149-181</mods:identifier>
<mods:identifier type="ZooBank">52FBD43553204F088EBBF18230DC9B62</mods:identifier>
<mods:identifier type="Pensoft-UUID">49B1C50F79A8585499B05B33E7367DF2</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:22146F0D-BFA6-4F91-84B5-CEB60238183D" httpUri="http://treatment.plazi.org/id/779EF539627B50EC9FB930E0AFFD9B47" lastPageNumber="181" pageId="0" pageNumber="181">
<subSubSection pageId="0" pageNumber="181" type="nomenclature">
<paragraph pageId="0" pageNumber="181">
<taxonomicName LSID="https://zoobank.org/22146F0D-BFA6-4F91-84B5-CEB60238183D" authority="Nguyen &amp; Stoev &amp; Nguyen &amp; Vu, 2023" authorityName="Nguyen &amp; Stoev &amp; Nguyen &amp; Vu" authorityYear="2023" class="Diplopoda" family="Paracortinidae" genus="Paracortina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracortina kyrang" order="Callipodida" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="kyrang" status="sp. nov.">Paracortina kyrang</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="181">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="181" type="description">
<paragraph pageId="0" pageNumber="181">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paracortina kyrang sp. nov. A entrance of Ky Rang Cave B habitat of the species B, C habitus, in situ. Images not to scale." figureDoi="10.3897/zookeys.1149.99651.figure1" httpUri="https://binary.pensoft.net/fig/814458" pageId="0" pageNumber="181">Figs 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Paracortina kyrang sp. nov., holotype A anterior body in ventral view B anterior body in lateral view C head in dorsal view D segments 6 - 12 in dorsal view. Abbreviation: lp = a large projection on head." figureDoi="10.3897/zookeys.1149.99651.figure2" httpUri="https://binary.pensoft.net/fig/814459" pageId="0" pageNumber="181">, 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paracortina kyrang sp. nov., holotype A anterior part of body in lateral view B posterior part of body in dorsal view C telson in dorsal view D hypoproct, paraprocts, and epiproct in ventral view. Abbreviations: ep = epiproct; pp = paraproct; hp = hypoproct." figureDoi="10.3897/zookeys.1149.99651.figure3" httpUri="https://binary.pensoft.net/fig/814461" pageId="0" pageNumber="181">, 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Paracortina kyrang sp. nov., holotype A right antenna B legs 4 and 5 in posterior view C gonopores in ventral view D female paratype (IEBR-Myr 932), cyphopods in subposterior view." figureDoi="10.3897/zookeys.1149.99651.figure4" httpUri="https://binary.pensoft.net/fig/814462" pageId="0" pageNumber="181">, 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Paracortina kyrang sp. nov., female paratype (IEBR-Myr 935) A head in ventral view B anterior body in lateral view C anterior body in subdorsal view D cyphopods in ventral view." figureDoi="10.3897/zookeys.1149.99651.figure5" httpUri="https://binary.pensoft.net/fig/814463" pageId="0" pageNumber="181">, 5</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paracortina kyrang sp. nov., holotype A leg 7 in posterior view B leg 7 in anterior view C gonopods in posterior view D gonopods in anterior view. Abbreviations: st = short tubercle; rap = rounded anterior process; a = coxal process a; b = coxal process b; te = telopodite; cp = clavate prefemoroidal processes; sp = distomesal spine on telopodite; sl = solenomere." figureDoi="10.3897/zookeys.1149.99651.figure6" httpUri="https://binary.pensoft.net/fig/814464" pageId="0" pageNumber="181">, 6</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Paracortina kyrang sp. nov., holotype A right gonopod in lateral view B right gonopod in ventral view C right gonopod in mesal view D solenomere in ventral view. Abbreviations: a = coxal process a; te = telopodite; cp = clavate prefemoroidal processes; sp = distomesal spine on telopodite; sg = seminal groove; sl = solenomere; ps = parasolenomere." figureDoi="10.3897/zookeys.1149.99651.figure7" httpUri="https://binary.pensoft.net/fig/814465" pageId="0" pageNumber="181">, 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="181" type="materials_examined">
<paragraph pageId="0" pageNumber="181">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="181">
<materialsCitation collectingDate="2021-11-02" collectorName="Anh D. Nguyen." county="Ky Rang Cave" location="Cao Bang Province" municipality="Quoc Toan" specimenCount="1" specimenCount-male="1" typeStatus="Holotype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="181">
<typeStatus>Holotype</typeStatus>
.
</emphasis>
<specimenCount type="male">1 male</specimenCount>
(
<emphasis bold="true" pageId="0" pageNumber="181">IEBR-Myr 921</emphasis>
)
<location LSID="urn:lsid:plazi:treatment:779EF539627B50EC9FB930E0AFFD9B47:AF40BE9784D5BAE86E838E383DCEA346" county="Ky Rang Cave" municipality="Quoc Toan" name="Cao Bang Province">Cao Bang Province</location>
,
<location LSID="urn:lsid:plazi:treatment:779EF539627B50EC9FB930E0AFFD9B47:0FC8905A1551F6BE56CDF4618448F5B6" county="Ky Rang Cave" municipality="Quoc Toan" name="Quang Hoa District">Quang Hoa District</location>
,
<collectingMunicipality>Quoc Toan</collectingMunicipality>
commune,
<collectingCounty>Ky Rang Cave</collectingCounty>
,
<collectingDate value="2021-11-02">2.xi.2021</collectingDate>
, leg.
<collectorName>Anh D. Nguyen.</collectorName>
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="181">
<emphasis bold="true" italics="true" pageId="0" pageNumber="181">
<typeStatus>Paratypes</typeStatus>
.
</emphasis>
<specimenCount type="female">1 female</specimenCount>
,
<specimenCount type="juvenile">1 juvenile</specimenCount>
(
<emphasis bold="true" pageId="0" pageNumber="181">IEBR-Myr 932</emphasis>
),
<materialsCitation collectingDate="2022-03-17" collectorName="Anh D. Nguyen, D. D. Nguyen." specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount type="female">1 female</specimenCount>
(
<emphasis bold="true" pageId="0" pageNumber="181">IEBR-Myr 935</emphasis>
) same locality, but
<collectingDate value="2022-03-17">17.iii.2022</collectingDate>
, leg.
<collectorName>Anh D. Nguyen</collectorName>
&amp;
<collectorName>D.D. Nguyen.</collectorName>
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="181">
<emphasis bold="true" italics="true" pageId="0" pageNumber="181">Non-types.</emphasis>
<specimenCount type="male">1 male</specimenCount>
,
<specimenCount type="male">1 male</specimenCount>
juvenile,
<materialsCitation collectingDate="2018-10-16" collectorName="Alexandre Faille." specimenCount="1" specimenCount-female="1">
<specimenCount type="female">1 female</specimenCount>
juvenile (
<emphasis bold="true" pageId="0" pageNumber="181">IEBR-Myr 954</emphasis>
), same locality, but
<collectingDate value="2018-10-16">16.x.2018</collectingDate>
, leg.
<collectorName>Alexandre Faille.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="181" type="diagnosis">
<paragraph pageId="0" pageNumber="181">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="181">
The new species is well distinguished from all congeners by the strongly modified head in males bearing a unique apically bent projection. Body composed of 68-74 pleurotergites +telson, eyes reduced, composed of 19 or 20 ommatidia in two or three rows. Gonocoxa with an anterior long spiniform process (
<emphasis bold="true" pageId="0" pageNumber="181">a</emphasis>
), as long as ca 80% of telopodite stem, and a rather slender, much shorter, cephalad process (
<emphasis bold="true" pageId="0" pageNumber="181">b</emphasis>
). Process
<emphasis bold="true" pageId="0" pageNumber="181">a</emphasis>
with a cephalad lobe distally, process
<emphasis bold="true" pageId="0" pageNumber="181">b</emphasis>
about 1/3 the length of telopodite. Telopodite with two long, clavate prefemoroidal processes (
<emphasis bold="true" pageId="0" pageNumber="181">cp</emphasis>
), densely covered with long macrosetae apically. Telopodite long, slender, apically twisted laterad, with a distal, reverse, short spine. Distal part of telopodite rather sinuous, narrowed at the base, then smoothly widened at its top, to narrow sharply finally at the solenomere (
<emphasis bold="true" pageId="0" pageNumber="181">sl</emphasis>
) and parasolenomere (
<emphasis bold="true" pageId="0" pageNumber="181">ps</emphasis>
).
</paragraph>
<paragraph pageId="0" pageNumber="181">
The new species can be keyed out into the first branch in Liu et al.'s (2015) key for identification of the species of
<taxonomicName authorityName="Wang &amp; Zhang" authorityYear="1993" class="Diplopoda" family="Paracortinidae" genus="Paracortina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracortina" order="Callipodida" pageId="0" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="181">Paracortina</emphasis>
</taxonomicName>
, with the clustering species having a pair of prefemoroidal clavate processes (
<emphasis bold="true" pageId="0" pageNumber="181">cp</emphasis>
) on the gonopods:
<taxonomicName class="Diplopoda" kingdom="Animalia" lsidName="P. thallina" order="Polydesmida" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="thallina">
<emphasis italics="true" pageId="0" pageNumber="181">P. thallina</emphasis>
</taxonomicName>
,
<taxonomicName class="Diplopoda" kingdom="Animalia" lsidName="P. stimula" order="Polydesmida" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="stimula">
<emphasis italics="true" pageId="0" pageNumber="181">P. stimula</emphasis>
</taxonomicName>
,
<taxonomicName class="Diplopoda" kingdom="Animalia" lsidName="P. leptoclada" order="Polydesmida" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="leptoclada">
<emphasis italics="true" pageId="0" pageNumber="181">P. leptoclada</emphasis>
</taxonomicName>
,
<taxonomicName class="Diplopoda" kingdom="Animalia" lsidName="P. voluta" order="Polydesmida" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="voluta">
<emphasis italics="true" pageId="0" pageNumber="181">P. voluta</emphasis>
</taxonomicName>
,
<taxonomicName class="Diplopoda" kingdom="Animalia" lsidName="P. serrata" order="Polydesmida" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="serrata">
<emphasis italics="true" pageId="0" pageNumber="181">P. serrata</emphasis>
</taxonomicName>
,
<taxonomicName class="Diplopoda" kingdom="Animalia" lsidName="P. viriosa" order="Polydesmida" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="viriosa">
<emphasis italics="true" pageId="0" pageNumber="181">P. viriosa</emphasis>
</taxonomicName>
, and
<taxonomicName class="Diplopoda" kingdom="Animalia" lsidName="P. carinata" order="Polydesmida" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="carinata">
<emphasis italics="true" pageId="0" pageNumber="181">P. carinata</emphasis>
</taxonomicName>
(all from southern China).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="181" type="etymology">
<paragraph pageId="0" pageNumber="181">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="181">
The species epithet &quot;
<emphasis italics="true" pageId="0" pageNumber="181">kyrang</emphasis>
&quot; is a noun in apposition for the type locality, Ky Rang Cave.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="181" type="description">
<paragraph pageId="0" pageNumber="181">Description.</paragraph>
<paragraph pageId="0" pageNumber="181">
<emphasis bold="true" italics="true" pageId="0" pageNumber="181">Male holotype</emphasis>
: Length about 42 mm, width and height of midbody PT 2.3 mm and 2.2 mm, respectively; 68 PT+ telson.
</paragraph>
<paragraph pageId="0" pageNumber="181">
<emphasis bold="true" italics="true" pageId="0" pageNumber="181">Colour</emphasis>
: living specimens greenish white (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paracortina kyrang sp. nov. A entrance of Ky Rang Cave B habitat of the species B, C habitus, in situ. Images not to scale." figureDoi="10.3897/zookeys.1149.99651.figure1" httpUri="https://binary.pensoft.net/fig/814458" pageId="0" pageNumber="181">1C, D</figureCitation>
). Ethanol preserved specimens: generally white-yellowish; posterior part of metazonites with a brown posterior margin; head, pleurotergites, antennae and telson white-yellowish; legs yellow-brownish.
</paragraph>
<caption doi="10.3897/zookeys.1149.99651.figure1" httpUri="https://binary.pensoft.net/fig/814458" pageId="0" pageNumber="181" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="181">
<emphasis bold="true" pageId="0" pageNumber="181">Figure 1.</emphasis>
<taxonomicName authorityName="Nguyen &amp; Stoev &amp; Nguyen &amp; Vu" authorityYear="2023" class="Diplopoda" family="Paracortinidae" genus="Paracortina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracortina kyrang" order="Callipodida" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="kyrang">
<emphasis italics="true" pageId="0" pageNumber="181">Paracortina kyrang</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="181">A</emphasis>
entrance of Ky Rang Cave
<emphasis bold="true" pageId="0" pageNumber="181">B</emphasis>
habitat of the species
<emphasis bold="true" pageId="0" pageNumber="181">B, C</emphasis>
habitus,
<emphasis italics="true" pageId="0" pageNumber="181">in situ</emphasis>
. Images not to scale.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="181">
<emphasis bold="true" italics="true" pageId="0" pageNumber="181">Head</emphasis>
(Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Paracortina kyrang sp. nov., holotype A anterior body in ventral view B anterior body in lateral view C head in dorsal view D segments 6 - 12 in dorsal view. Abbreviation: lp = a large projection on head." figureDoi="10.3897/zookeys.1149.99651.figure2" httpUri="https://binary.pensoft.net/fig/814459" pageId="0" pageNumber="181">2A-C</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paracortina kyrang sp. nov., holotype A anterior part of body in lateral view B posterior part of body in dorsal view C telson in dorsal view D hypoproct, paraprocts, and epiproct in ventral view. Abbreviations: ep = epiproct; pp = paraproct; hp = hypoproct." figureDoi="10.3897/zookeys.1149.99651.figure3" httpUri="https://binary.pensoft.net/fig/814461" pageId="0" pageNumber="181">3A</figureCitation>
) highly modified; frons considerably elongated in large projection (
<emphasis bold="true" pageId="0" pageNumber="181">lp</emphasis>
), which is curved at its end; forehead and vertex concave.
</paragraph>
<caption doi="10.3897/zookeys.1149.99651.figure2" httpUri="https://binary.pensoft.net/fig/814459" pageId="0" pageNumber="181" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="181">
<emphasis bold="true" pageId="0" pageNumber="181">Figure 2.</emphasis>
<taxonomicName authorityName="Nguyen &amp; Stoev &amp; Nguyen &amp; Vu" authorityYear="2023" class="Diplopoda" family="Paracortinidae" genus="Paracortina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracortina kyrang" order="Callipodida" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="kyrang">
<emphasis italics="true" pageId="0" pageNumber="181">Paracortina kyrang</emphasis>
</taxonomicName>
sp. nov., holotype
<emphasis bold="true" pageId="0" pageNumber="181">A</emphasis>
anterior body in ventral view
<emphasis bold="true" pageId="0" pageNumber="181">B</emphasis>
anterior body in lateral view
<emphasis bold="true" pageId="0" pageNumber="181">C</emphasis>
head in dorsal view
<emphasis bold="true" pageId="0" pageNumber="181">D</emphasis>
segments 6-12 in dorsal view. Abbreviation: lp = a large projection on head.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1149.99651.figure3" httpUri="https://binary.pensoft.net/fig/814461" pageId="0" pageNumber="181" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="181">
<emphasis bold="true" pageId="0" pageNumber="181">Figure 3.</emphasis>
<taxonomicName authorityName="Nguyen &amp; Stoev &amp; Nguyen &amp; Vu" authorityYear="2023" class="Diplopoda" family="Paracortinidae" genus="Paracortina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracortina kyrang" order="Callipodida" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="kyrang">
<emphasis italics="true" pageId="0" pageNumber="181">Paracortina kyrang</emphasis>
</taxonomicName>
sp. nov., holotype
<emphasis bold="true" pageId="0" pageNumber="181">A</emphasis>
anterior part of body in lateral view
<emphasis bold="true" pageId="0" pageNumber="181">B</emphasis>
posterior part of body in dorsal view
<emphasis bold="true" pageId="0" pageNumber="181">C</emphasis>
telson in dorsal view
<emphasis bold="true" pageId="0" pageNumber="181">D</emphasis>
hypoproct, paraprocts, and epiproct in ventral view. Abbreviations: ep = epiproct; pp = paraproct; hp = hypoproct.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="181">
<emphasis bold="true" italics="true" pageId="0" pageNumber="181">Antennae</emphasis>
(Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Paracortina kyrang sp. nov., holotype A anterior body in ventral view B anterior body in lateral view C head in dorsal view D segments 6 - 12 in dorsal view. Abbreviation: lp = a large projection on head." figureDoi="10.3897/zookeys.1149.99651.figure2" httpUri="https://binary.pensoft.net/fig/814459" pageId="0" pageNumber="181">2A</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paracortina kyrang sp. nov., holotype A anterior part of body in lateral view B posterior part of body in dorsal view C telson in dorsal view D hypoproct, paraprocts, and epiproct in ventral view. Abbreviations: ep = epiproct; pp = paraproct; hp = hypoproct." figureDoi="10.3897/zookeys.1149.99651.figure3" httpUri="https://binary.pensoft.net/fig/814461" pageId="0" pageNumber="181">3A</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Paracortina kyrang sp. nov., holotype A right antenna B legs 4 and 5 in posterior view C gonopores in ventral view D female paratype (IEBR-Myr 932), cyphopods in subposterior view." figureDoi="10.3897/zookeys.1149.99651.figure4" httpUri="https://binary.pensoft.net/fig/814462" pageId="0" pageNumber="181">4B</figureCitation>
) extremely long, extending beyond the posterior edge of PT 9 when folded backwards; ratio antenna/body length about 1/7; all antennomeres white; length of antennomeres: 1: 0.21 mm, 2: 1.27 mm, 3: 1.26 mm, 4: 1.26 mm, 5: 1.21 mm, 6: 0.67 mm, 7: 0.23 mm; tip of antennomere 7 with four cones protruding well beyond the edge. Eyes (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Paracortina kyrang sp. nov., holotype A anterior body in ventral view B anterior body in lateral view C head in dorsal view D segments 6 - 12 in dorsal view. Abbreviation: lp = a large projection on head." figureDoi="10.3897/zookeys.1149.99651.figure2" httpUri="https://binary.pensoft.net/fig/814459" pageId="0" pageNumber="181">2B, C</figureCitation>
) black, well delineated, composed of 19 ommatidia in three horizontal rows (9+3+7).
<normalizedToken originalValue="Tömösváry’s">Toemoesvary's</normalizedToken>
organ about three times larger than the adjacent ommatidium, placed between the eye and the base of antenna (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Paracortina kyrang sp. nov., holotype A anterior body in ventral view B anterior body in lateral view C head in dorsal view D segments 6 - 12 in dorsal view. Abbreviation: lp = a large projection on head." figureDoi="10.3897/zookeys.1149.99651.figure2" httpUri="https://binary.pensoft.net/fig/814459" pageId="0" pageNumber="181">2B</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="181">Width of PT: 6&gt;7&gt;&gt;8-14&gt;4&gt;3&gt;2&gt;1. PT slightly broader than high; height of 10thPT: 2.19 mm, width 2.32 mm.</paragraph>
<paragraph pageId="0" pageNumber="181">
<emphasis bold="true" italics="true" pageId="0" pageNumber="181">Collum</emphasis>
(Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Paracortina kyrang sp. nov., holotype A anterior body in ventral view B anterior body in lateral view C head in dorsal view D segments 6 - 12 in dorsal view. Abbreviation: lp = a large projection on head." figureDoi="10.3897/zookeys.1149.99651.figure2" httpUri="https://binary.pensoft.net/fig/814459" pageId="0" pageNumber="181">2B, C</figureCitation>
) much narrower than head; pleurotergites 6 and 7 in males strongly enlarged (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Paracortina kyrang sp. nov., holotype A anterior body in ventral view B anterior body in lateral view C head in dorsal view D segments 6 - 12 in dorsal view. Abbreviation: lp = a large projection on head." figureDoi="10.3897/zookeys.1149.99651.figure2" httpUri="https://binary.pensoft.net/fig/814459" pageId="0" pageNumber="181">2D</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paracortina kyrang sp. nov., holotype A anterior part of body in lateral view B posterior part of body in dorsal view C telson in dorsal view D hypoproct, paraprocts, and epiproct in ventral view. Abbreviations: ep = epiproct; pp = paraproct; hp = hypoproct." figureDoi="10.3897/zookeys.1149.99651.figure3" httpUri="https://binary.pensoft.net/fig/814461" pageId="0" pageNumber="181">3A</figureCitation>
). Crests on collum (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Paracortina kyrang sp. nov., holotype A anterior body in ventral view B anterior body in lateral view C head in dorsal view D segments 6 - 12 in dorsal view. Abbreviation: lp = a large projection on head." figureDoi="10.3897/zookeys.1149.99651.figure2" httpUri="https://binary.pensoft.net/fig/814459" pageId="0" pageNumber="181">2C</figureCitation>
) moderately expressed mostly in the posterior part of the segment. Complete crests series appearing from PT2 onwards. Above ozopores, midbody PT with 3+3 primary crests and with 3+3 secondary short crests between primary crests (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Paracortina kyrang sp. nov., holotype A anterior body in ventral view B anterior body in lateral view C head in dorsal view D segments 6 - 12 in dorsal view. Abbreviation: lp = a large projection on head." figureDoi="10.3897/zookeys.1149.99651.figure2" httpUri="https://binary.pensoft.net/fig/814459" pageId="0" pageNumber="181">2D</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paracortina kyrang sp. nov., holotype A anterior part of body in lateral view B posterior part of body in dorsal view C telson in dorsal view D hypoproct, paraprocts, and epiproct in ventral view. Abbreviations: ep = epiproct; pp = paraproct; hp = hypoproct." figureDoi="10.3897/zookeys.1149.99651.figure3" httpUri="https://binary.pensoft.net/fig/814461" pageId="0" pageNumber="181">3B</figureCitation>
); 3rd primary crest strongly enlarged, other primary crests flattened, almost equally broad along the metazonal length; only secondary crests shorter and slightly narrowed posteriorly. Ozopores lying on primary crest 3, visible from sixth to last but two PT (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Paracortina kyrang sp. nov., holotype A anterior body in ventral view B anterior body in lateral view C head in dorsal view D segments 6 - 12 in dorsal view. Abbreviation: lp = a large projection on head." figureDoi="10.3897/zookeys.1149.99651.figure2" httpUri="https://binary.pensoft.net/fig/814459" pageId="0" pageNumber="181">2D</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paracortina kyrang sp. nov., holotype A anterior part of body in lateral view B posterior part of body in dorsal view C telson in dorsal view D hypoproct, paraprocts, and epiproct in ventral view. Abbreviations: ep = epiproct; pp = paraproct; hp = hypoproct." figureDoi="10.3897/zookeys.1149.99651.figure3" httpUri="https://binary.pensoft.net/fig/814461" pageId="0" pageNumber="181">3B</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="181">
Below ozopores, midbody PT with 2+2 primary crests and 2+2 shorter and thinner secondary crests between primary crests, and 8-10 lower crests down to ventral pleurotergal edge (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paracortina kyrang sp. nov., holotype A anterior part of body in lateral view B posterior part of body in dorsal view C telson in dorsal view D hypoproct, paraprocts, and epiproct in ventral view. Abbreviations: ep = epiproct; pp = paraproct; hp = hypoproct." figureDoi="10.3897/zookeys.1149.99651.figure3" httpUri="https://binary.pensoft.net/fig/814461" pageId="0" pageNumber="181">3A</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="181">
Midbody pleurotergal setae 5+5, located at caudal edges of primary crests (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Paracortina kyrang sp. nov., holotype A anterior body in ventral view B anterior body in lateral view C head in dorsal view D segments 6 - 12 in dorsal view. Abbreviation: lp = a large projection on head." figureDoi="10.3897/zookeys.1149.99651.figure2" httpUri="https://binary.pensoft.net/fig/814459" pageId="0" pageNumber="181">2D</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paracortina kyrang sp. nov., holotype A anterior part of body in lateral view B posterior part of body in dorsal view C telson in dorsal view D hypoproct, paraprocts, and epiproct in ventral view. Abbreviations: ep = epiproct; pp = paraproct; hp = hypoproct." figureDoi="10.3897/zookeys.1149.99651.figure3" httpUri="https://binary.pensoft.net/fig/814461" pageId="0" pageNumber="181">3A</figureCitation>
); setal pattern as in below (Chaetotaxy). Axial line rather distinct.
</paragraph>
<paragraph pageId="0" pageNumber="181">
<emphasis bold="true" italics="true" pageId="0" pageNumber="181">Epiproct</emphasis>
(
<emphasis bold="true" pageId="0" pageNumber="181">ep</emphasis>
) (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paracortina kyrang sp. nov., holotype A anterior part of body in lateral view B posterior part of body in dorsal view C telson in dorsal view D hypoproct, paraprocts, and epiproct in ventral view. Abbreviations: ep = epiproct; pp = paraproct; hp = hypoproct." figureDoi="10.3897/zookeys.1149.99651.figure3" httpUri="https://binary.pensoft.net/fig/814461" pageId="0" pageNumber="181">3B-D</figureCitation>
) simple, with 7+7 anterior and 4+4 posterior setiferous knobs in transverse rows. Hypoproct (
<emphasis bold="true" pageId="0" pageNumber="181">hp</emphasis>
) (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Paracortina kyrang sp. nov., holotype A right antenna B legs 4 and 5 in posterior view C gonopores in ventral view D female paratype (IEBR-Myr 932), cyphopods in subposterior view." figureDoi="10.3897/zookeys.1149.99651.figure4" httpUri="https://binary.pensoft.net/fig/814462" pageId="0" pageNumber="181">4D</figureCitation>
) tripartite, medial sclerite largest, subtrapeziform, bearing two paramedian macrosetae; each lateral sclerite with a single macroseta born on a large tubercle. Paraprocts (
<emphasis bold="true" pageId="0" pageNumber="181">pp</emphasis>
) (= anal valves) (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Paracortina kyrang sp. nov., holotype A right antenna B legs 4 and 5 in posterior view C gonopores in ventral view D female paratype (IEBR-Myr 932), cyphopods in subposterior view." figureDoi="10.3897/zookeys.1149.99651.figure4" httpUri="https://binary.pensoft.net/fig/814462" pageId="0" pageNumber="181">4D</figureCitation>
) smooth, each divided into a small upper and large lower sclerites, both with a pair of macrosetae. Spinnerets long and slender, ending with a long seta each. All setae on telson brown, contrasting with the yellowish background.
</paragraph>
<caption doi="10.3897/zookeys.1149.99651.figure4" httpUri="https://binary.pensoft.net/fig/814462" pageId="0" pageNumber="181" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="181">
<emphasis bold="true" pageId="0" pageNumber="181">Figure 4.</emphasis>
<taxonomicName authorityName="Nguyen &amp; Stoev &amp; Nguyen &amp; Vu" authorityYear="2023" class="Diplopoda" family="Paracortinidae" genus="Paracortina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracortina kyrang" order="Callipodida" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="kyrang">
<emphasis italics="true" pageId="0" pageNumber="181">Paracortina kyrang</emphasis>
</taxonomicName>
sp. nov., holotype
<emphasis bold="true" pageId="0" pageNumber="181">A</emphasis>
right antenna
<emphasis bold="true" pageId="0" pageNumber="181">B</emphasis>
legs 4 and 5 in posterior view
<emphasis bold="true" pageId="0" pageNumber="181">C</emphasis>
gonopores in ventral view
<emphasis bold="true" pageId="0" pageNumber="181">D</emphasis>
female paratype (IEBR-Myr 932), cyphopods in subposterior view.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="181">
Male leg-pairs 1 and 2 much shorter, with strong setae on ventral side of femorite and tibia, leg-pair 3 slightly shorter than following legs. Tarsi 1-3 1-segmented, and from tarsus 4 to ultimate pair 2-segmented; tarsal pads large until leg 26, then gradually thinner and eventually absent on subsequent legs. All legs ending with a rather slender, long, curved claw. Coxal sacs present from legs 3-26 (PT 16). Only coxae and tibia finely micropapillate ventrally (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Paracortina kyrang sp. nov., holotype A right antenna B legs 4 and 5 in posterior view C gonopores in ventral view D female paratype (IEBR-Myr 932), cyphopods in subposterior view." figureDoi="10.3897/zookeys.1149.99651.figure4" httpUri="https://binary.pensoft.net/fig/814462" pageId="0" pageNumber="181">4B</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="181">
Coxa 2 with a small anterior process and a posterior gonopore, the latter placed on a small cone (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Paracortina kyrang sp. nov., holotype A right antenna B legs 4 and 5 in posterior view C gonopores in ventral view D female paratype (IEBR-Myr 932), cyphopods in subposterior view." figureDoi="10.3897/zookeys.1149.99651.figure4" httpUri="https://binary.pensoft.net/fig/814462" pageId="0" pageNumber="181">4C</figureCitation>
). Coxa 6 normal, without processes or modifications. Coxa 7 (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paracortina kyrang sp. nov., holotype A leg 7 in posterior view B leg 7 in anterior view C gonopods in posterior view D gonopods in anterior view. Abbreviations: st = short tubercle; rap = rounded anterior process; a = coxal process a; b = coxal process b; te = telopodite; cp = clavate prefemoroidal processes; sp = distomesal spine on telopodite; sl = solenomere." figureDoi="10.3897/zookeys.1149.99651.figure6" httpUri="https://binary.pensoft.net/fig/814464" pageId="0" pageNumber="181">6A, B</figureCitation>
) with a short tubercle (st), and a very strong, rounded anterior process (
<emphasis bold="true" pageId="0" pageNumber="181">rap</emphasis>
). Coxae of the remaining legs normal.
</paragraph>
<paragraph pageId="0" pageNumber="181">
<emphasis bold="true" pageId="0" pageNumber="181">Chaetotaxy</emphasis>
:
</paragraph>
<paragraph pageId="0" pageNumber="181">
<table inLine="true" pageId="0" pageNumber="181">
<tr pageId="0" pageNumber="181">
<td colspan="1" isEmpty="true" pageId="0" pageNumber="181" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="181" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="181">Anterior setae</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="181" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="181">Posterior setae</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="181">
<td colspan="1" pageId="0" pageNumber="181" rowspan="1">Collum</td>
<td colspan="1" pageId="0" pageNumber="181" rowspan="1">4+4</td>
<td colspan="1" pageId="0" pageNumber="181" rowspan="1">2+2</td>
</tr>
<tr pageId="0" pageNumber="181">
<td colspan="1" pageId="0" pageNumber="181" rowspan="1">PT2</td>
<td colspan="1" pageId="0" pageNumber="181" rowspan="1">5+5</td>
<td colspan="1" pageId="0" pageNumber="181" rowspan="1">broken</td>
</tr>
<tr pageId="0" pageNumber="181">
<td colspan="1" pageId="0" pageNumber="181" rowspan="1">PT3</td>
<td colspan="1" pageId="0" pageNumber="181" rowspan="1">5+5</td>
<td colspan="1" pageId="0" pageNumber="181" rowspan="1">broken</td>
</tr>
<tr pageId="0" pageNumber="181">
<td colspan="1" pageId="0" pageNumber="181" rowspan="1">PT4</td>
<td colspan="1" pageId="0" pageNumber="181" rowspan="1">broken</td>
<td colspan="1" pageId="0" pageNumber="181" rowspan="1">5+5</td>
</tr>
<tr pageId="0" pageNumber="181">
<td colspan="1" pageId="0" pageNumber="181" rowspan="1">PT5</td>
<td colspan="1" pageId="0" pageNumber="181" rowspan="1">broken</td>
<td colspan="1" pageId="0" pageNumber="181" rowspan="1">5+5</td>
</tr>
<tr pageId="0" pageNumber="181">
<td colspan="1" pageId="0" pageNumber="181" rowspan="1">PT6-penultimate PT</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="181" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="181" rowspan="1">5+5 (rarely 6+5)</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="181">
Gonopods (Figs
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paracortina kyrang sp. nov., holotype A leg 7 in posterior view B leg 7 in anterior view C gonopods in posterior view D gonopods in anterior view. Abbreviations: st = short tubercle; rap = rounded anterior process; a = coxal process a; b = coxal process b; te = telopodite; cp = clavate prefemoroidal processes; sp = distomesal spine on telopodite; sl = solenomere." figureDoi="10.3897/zookeys.1149.99651.figure6" httpUri="https://binary.pensoft.net/fig/814464" pageId="0" pageNumber="181">6C, D</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Paracortina kyrang sp. nov., holotype A right gonopod in lateral view B right gonopod in ventral view C right gonopod in mesal view D solenomere in ventral view. Abbreviations: a = coxal process a; te = telopodite; cp = clavate prefemoroidal processes; sp = distomesal spine on telopodite; sg = seminal groove; sl = solenomere; ps = parasolenomere." figureDoi="10.3897/zookeys.1149.99651.figure7" httpUri="https://binary.pensoft.net/fig/814465" pageId="0" pageNumber="181">7</figureCitation>
) yellow-brownish, some parts dark brown to black (seminal groove, solenomere, basal part of coxal process
<emphasis bold="true" pageId="0" pageNumber="181">a</emphasis>
). Gonopods protruding well beyond the gonocoel, stems of telopodites long, thin, subparallel, and diverging, pointing posteriad (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paracortina kyrang sp. nov., holotype A leg 7 in posterior view B leg 7 in anterior view C gonopods in posterior view D gonopods in anterior view. Abbreviations: st = short tubercle; rap = rounded anterior process; a = coxal process a; b = coxal process b; te = telopodite; cp = clavate prefemoroidal processes; sp = distomesal spine on telopodite; sl = solenomere." figureDoi="10.3897/zookeys.1149.99651.figure6" httpUri="https://binary.pensoft.net/fig/814464" pageId="0" pageNumber="181">6C, D</figureCitation>
). Coxa with an anterior long spiniform process (
<emphasis bold="true" pageId="0" pageNumber="181">a</emphasis>
), and a rather slender, much shorter, posterior process (
<emphasis bold="true" pageId="0" pageNumber="181">b</emphasis>
) (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paracortina kyrang sp. nov., holotype A leg 7 in posterior view B leg 7 in anterior view C gonopods in posterior view D gonopods in anterior view. Abbreviations: st = short tubercle; rap = rounded anterior process; a = coxal process a; b = coxal process b; te = telopodite; cp = clavate prefemoroidal processes; sp = distomesal spine on telopodite; sl = solenomere." figureDoi="10.3897/zookeys.1149.99651.figure6" httpUri="https://binary.pensoft.net/fig/814464" pageId="0" pageNumber="181">6C</figureCitation>
). Process
<emphasis bold="true" pageId="0" pageNumber="181">a</emphasis>
as long as ca 80% of telopodite stem, with a lobe distally while process
<emphasis bold="true" pageId="0" pageNumber="181">b</emphasis>
about 1/3 the length of telopodite. Telopodite (te) with two long, clavate prefemoroidal processes (
<emphasis bold="true" pageId="0" pageNumber="181">cp</emphasis>
), densely covered with long macrosetae apically. Telopodite long, slender, apically twisted laterad, with a distomesal, reverse, short spine (
<emphasis bold="true" pageId="0" pageNumber="181">sp</emphasis>
) (Figs
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paracortina kyrang sp. nov., holotype A leg 7 in posterior view B leg 7 in anterior view C gonopods in posterior view D gonopods in anterior view. Abbreviations: st = short tubercle; rap = rounded anterior process; a = coxal process a; b = coxal process b; te = telopodite; cp = clavate prefemoroidal processes; sp = distomesal spine on telopodite; sl = solenomere." figureDoi="10.3897/zookeys.1149.99651.figure6" httpUri="https://binary.pensoft.net/fig/814464" pageId="0" pageNumber="181">6C</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Paracortina kyrang sp. nov., holotype A right gonopod in lateral view B right gonopod in ventral view C right gonopod in mesal view D solenomere in ventral view. Abbreviations: a = coxal process a; te = telopodite; cp = clavate prefemoroidal processes; sp = distomesal spine on telopodite; sg = seminal groove; sl = solenomere; ps = parasolenomere." figureDoi="10.3897/zookeys.1149.99651.figure7" httpUri="https://binary.pensoft.net/fig/814465" pageId="0" pageNumber="181">7A, B</figureCitation>
). Distal part of telopodite rather sinuous, narrowed at the base, then smoothly widened at its top, to narrow sharply finally at the solenomere (
<emphasis bold="true" pageId="0" pageNumber="181">sl</emphasis>
) and parasolenomere (
<emphasis bold="true" pageId="0" pageNumber="181">ps</emphasis>
) (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Paracortina kyrang sp. nov., holotype A right gonopod in lateral view B right gonopod in ventral view C right gonopod in mesal view D solenomere in ventral view. Abbreviations: a = coxal process a; te = telopodite; cp = clavate prefemoroidal processes; sp = distomesal spine on telopodite; sg = seminal groove; sl = solenomere; ps = parasolenomere." figureDoi="10.3897/zookeys.1149.99651.figure7" httpUri="https://binary.pensoft.net/fig/814465" pageId="0" pageNumber="181">7D</figureCitation>
). The seminal groove (sg) terminating in the solenomere (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Paracortina kyrang sp. nov., holotype A right gonopod in lateral view B right gonopod in ventral view C right gonopod in mesal view D solenomere in ventral view. Abbreviations: a = coxal process a; te = telopodite; cp = clavate prefemoroidal processes; sp = distomesal spine on telopodite; sg = seminal groove; sl = solenomere; ps = parasolenomere." figureDoi="10.3897/zookeys.1149.99651.figure7" httpUri="https://binary.pensoft.net/fig/814465" pageId="0" pageNumber="181">7D</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1149.99651.figure5" httpUri="https://binary.pensoft.net/fig/814463" pageId="0" pageNumber="181" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="181">
<emphasis bold="true" pageId="0" pageNumber="181">Figure 5.</emphasis>
<taxonomicName authorityName="Nguyen &amp; Stoev &amp; Nguyen &amp; Vu" authorityYear="2023" class="Diplopoda" family="Paracortinidae" genus="Paracortina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracortina kyrang" order="Callipodida" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="kyrang">
<emphasis italics="true" pageId="0" pageNumber="181">Paracortina kyrang</emphasis>
</taxonomicName>
sp. nov., female paratype (IEBR-Myr 935)
<emphasis bold="true" pageId="0" pageNumber="181">A</emphasis>
head in ventral view
<emphasis bold="true" pageId="0" pageNumber="181">B</emphasis>
anterior body in lateral view
<emphasis bold="true" pageId="0" pageNumber="181">C</emphasis>
anterior body in subdorsal view
<emphasis bold="true" pageId="0" pageNumber="181">D</emphasis>
cyphopods in ventral view.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1149.99651.figure6" httpUri="https://binary.pensoft.net/fig/814464" pageId="0" pageNumber="181" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="181">
<emphasis bold="true" pageId="0" pageNumber="181">Figure 6.</emphasis>
<taxonomicName authorityName="Nguyen &amp; Stoev &amp; Nguyen &amp; Vu" authorityYear="2023" class="Diplopoda" family="Paracortinidae" genus="Paracortina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracortina kyrang" order="Callipodida" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="kyrang">
<emphasis italics="true" pageId="0" pageNumber="181">Paracortina kyrang</emphasis>
</taxonomicName>
sp. nov., holotype
<emphasis bold="true" pageId="0" pageNumber="181">A</emphasis>
leg 7 in posterior view
<emphasis bold="true" pageId="0" pageNumber="181">B</emphasis>
leg 7 in anterior view
<emphasis bold="true" pageId="0" pageNumber="181">C</emphasis>
gonopods in posterior view
<emphasis bold="true" pageId="0" pageNumber="181">D</emphasis>
gonopods in anterior view. Abbreviations: st = short tubercle; rap = rounded anterior process; a = coxal process a; b = coxal process b; te = telopodite; cp = clavate prefemoroidal processes; sp = distomesal spine on telopodite; sl = solenomere.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1149.99651.figure7" httpUri="https://binary.pensoft.net/fig/814465" pageId="0" pageNumber="181" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="181">
<emphasis bold="true" pageId="0" pageNumber="181">Figure 7.</emphasis>
<taxonomicName authorityName="Nguyen &amp; Stoev &amp; Nguyen &amp; Vu" authorityYear="2023" class="Diplopoda" family="Paracortinidae" genus="Paracortina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracortina kyrang" order="Callipodida" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="kyrang">
<emphasis italics="true" pageId="0" pageNumber="181">Paracortina kyrang</emphasis>
</taxonomicName>
sp. nov., holotype
<emphasis bold="true" pageId="0" pageNumber="181">A</emphasis>
right gonopod in lateral view
<emphasis bold="true" pageId="0" pageNumber="181">B</emphasis>
right gonopod in ventral view
<emphasis bold="true" pageId="0" pageNumber="181">C</emphasis>
right gonopod in mesal view
<emphasis bold="true" pageId="0" pageNumber="181">D</emphasis>
solenomere in ventral view. Abbreviations: a = coxal process a; te = telopodite; cp = clavate prefemoroidal processes; sp = distomesal spine on telopodite; sg = seminal groove; sl = solenomere; ps = parasolenomere.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="181">
<emphasis bold="true" pageId="0" pageNumber="181">Females.</emphasis>
Head unmodified (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Paracortina kyrang sp. nov., female paratype (IEBR-Myr 935) A head in ventral view B anterior body in lateral view C anterior body in subdorsal view D cyphopods in ventral view." figureDoi="10.3897/zookeys.1149.99651.figure5" httpUri="https://binary.pensoft.net/fig/814463" pageId="0" pageNumber="181">5A-C</figureCitation>
). Length about 54.7 mm. The 10thPT ca 2.62 mm wide and 2.36 mm high; 68-74 PT + telson. Second leg-pair unmodified. Leg-pairs 1-3 with tarsal brushes. Cyphopods small, densely setose, bilobed (Figs
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Paracortina kyrang sp. nov., holotype A right antenna B legs 4 and 5 in posterior view C gonopores in ventral view D female paratype (IEBR-Myr 932), cyphopods in subposterior view." figureDoi="10.3897/zookeys.1149.99651.figure4" httpUri="https://binary.pensoft.net/fig/814462" pageId="0" pageNumber="181">4D</figureCitation>
,
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Paracortina kyrang sp. nov., female paratype (IEBR-Myr 935) A head in ventral view B anterior body in lateral view C anterior body in subdorsal view D cyphopods in ventral view." figureDoi="10.3897/zookeys.1149.99651.figure5" httpUri="https://binary.pensoft.net/fig/814463" pageId="0" pageNumber="181">5A, D</figureCitation>
). Coxae 7 normal, without processes.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="181" type="dna barcode">
<paragraph pageId="0" pageNumber="181">DNA barcode.</paragraph>
<paragraph pageId="0" pageNumber="181">
The new species has a close genetic identity with
<taxonomicName authorityName="Hoffman" authorityYear="1957" class="Diplopoda" family="Abacionidae" genus="Tetracion" kingdom="Animalia" lsidName="Tetracion jonesi" order="Callipodida" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="jonesi">
<emphasis italics="true" pageId="0" pageNumber="181">Tetracion jonesi</emphasis>
</taxonomicName>
Hoffman, 1956 (
<taxonomicName authorityName="Shelley" authorityYear="1979" class="Diplopoda" family="Abacionidae" kingdom="Animalia" lsidName="" order="Callipodida" pageId="0" pageNumber="181" phylum="Arthropoda" rank="family">Abacionidae</taxonomicName>
) from 77.74% to 78.25%.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="181" type="cave habitat">
<paragraph pageId="0" pageNumber="181">Cave habitat.</paragraph>
<paragraph pageId="0" pageNumber="181">
Ky Rang Cave is located in close proximity to Thang Hen Lake in Cao Bang Province, northern Vietnam, at the altitude of 1,000 m a.s.l. The cave entrance is wide, but the only passage is blocked by an artificial door made by the local residents. Because of it, the semi-light part of the cave is missing, and, on entry, the cave is immediately dark (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paracortina kyrang sp. nov. A entrance of Ky Rang Cave B habitat of the species B, C habitus, in situ. Images not to scale." figureDoi="10.3897/zookeys.1149.99651.figure1" httpUri="https://binary.pensoft.net/fig/814458" pageId="0" pageNumber="181">1A</figureCitation>
). The cave is high (15-20 m), wide (15-20 m), and long (700-1,000 m). The floor is mainly wet with clay and there are some small pools (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paracortina kyrang sp. nov. A entrance of Ky Rang Cave B habitat of the species B, C habitus, in situ. Images not to scale." figureDoi="10.3897/zookeys.1149.99651.figure1" httpUri="https://binary.pensoft.net/fig/814458" pageId="0" pageNumber="181">1B</figureCitation>
). Several other millipede species were found in the cave, for example,
<taxonomicName authorityName="Golovatch" authorityYear="2019" class="Diplopoda" family="Paradoxosomatidae" genus="Hylomus" kingdom="Animalia" lsidName="Hylomus srisonchaii" order="Polydesmida" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="srisonchaii">
<emphasis italics="true" pageId="0" pageNumber="181">Hylomus srisonchaii</emphasis>
</taxonomicName>
Golovatch, 2019 and
<taxonomicName class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyleoglomeris alba" order="Glomerida" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="alba">
<emphasis italics="true" pageId="0" pageNumber="181">Hyleoglomeris alba</emphasis>
</taxonomicName>
Kuroda, Nguyen &amp; Eguchi, 2022 (
<bibRefCitation DOI="https://doi.org/10.15298/arthsel.28.4.01" author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="181" pagination="459 - 478" refId="B5" refString="Golovatch, SI, 2019. On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXVII. Arthropoda Selecta 28 (4): 459 - 478, DOI: https://doi.org/10.15298/arthsel.28.4.01" title="On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXVII." url="https://doi.org/10.15298/arthsel.28.4.01" volume="28" year="2019">Golovatch 2019</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.1108.85423" author="Kuroda, M" journalOrPublisher="ZooKeys" pageId="0" pageNumber="181" pagination="161 - 174" refId="B7" refString="Kuroda, M, Eguchi, K, Oguri, E, Nguyen, AD, 2022. Two new cave Hyleoglomeris species (Glomerida, Glomeridae) from northern Vietnam. ZooKeys 1108: 161 - 174, DOI: https://doi.org/10.3897/zookeys.1108.85423" title="Two new cave Hyleoglomeris species (Glomerida, Glomeridae) from northern Vietnam." url="https://doi.org/10.3897/zookeys.1108.85423" volume="1108" year="2022">Kuroda et al. 2022</bibRefCitation>
). The new species was found at a distance of 500 m from the entrance.
</paragraph>
</subSubSection>
</treatment>
</document>
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