245 lines
28 KiB
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245 lines
28 KiB
XML
<document ID-CLB-Dataset="94519" ID-DOI="http://dx.doi.org/10.3897/zookeys.1112.85752" ID-GBIF-Dataset="5b4d920a-898b-49da-b958-8ee8a0774e8a" ID-PMC="PMC9848642" ID-Pensoft-Pub="1313-2970-1112-11" ID-Pensoft-UUID="59936DE5F02E5E1D8346772863977AEA" ID-PubMed="36760626" ID-ZooBank="F9F1E9A2CB9249CB8D4CFB3FAC592F96" ModsDocID="1313-2970-1112-11" checkinTime="1657650131167" checkinUser="pensoft" docAuthor="Sanchez, Enrico Gerard S., Delocado, Emmanuel D. & Freitag, Hendrik" docDate="2022" docId="779A4B5041735BF39C0F09B3E8386E7C" docLanguage="en" docName="ZooKeys 1112: 11-25" docOrigin="ZooKeys 1112" docPubDate="2022-07-12" docSource="http://dx.doi.org/10.3897/zookeys.1112.85752" docTitle="Anacaena angatbuhay Sanchez & Delocado & Freitag 2022, sp. nov." docType="treatment" docUuid="A237568F-6B56-4F69-A5F0-08D4D2311949" docUuidSource="ZooBank" docVersion="5" id="59936DE5F02E5E1D8346772863977AEA" lastPageNumber="11" masterDocId="59936DE5F02E5E1D8346772863977AEA" masterDocTitle="Two new species of Anacaena Thomson, 1859 (Coleoptera, Hydrophilidae) from Northern Luzon, Philippines" masterLastPageNumber="25" masterPageNumber="11" pageNumber="11" updateTime="1732844981149" updateUser="ExternalLinkService">
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<mods:titleInfo id="55512AF4FFB7FD88C538969ACA5053D9">
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<mods:title id="ACA2F11A31EF5405E7AB1FF265DF2F44">Two new species of Anacaena Thomson, 1859 (Coleoptera, Hydrophilidae) from Northern Luzon, Philippines</mods:title>
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<mods:name id="74A8CF22F5B310FFE5D90DEF082DA44C" type="personal">
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<mods:roleTerm id="1B2218B88F41F9773793E1BD65395D37">Author</mods:roleTerm>
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<mods:namePart id="6646D9965CFE065893DAE921CAFDAAC8">Sanchez, Enrico Gerard S.</mods:namePart>
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<mods:affiliation id="AFC89BD7FEDEAE611B110A58B9DA9B74">Ateneo Biodiversity Research Laboratory, Department of Biology, School of Science and Engineering, Ateneo de Manila University, Quezon City, 1108 Philippines</mods:affiliation>
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<mods:nameIdentifier id="0E1A327E97087271283E7DD997134DF7" type="email">enrico.sanchez@obf.ateneo.edu</mods:nameIdentifier>
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<mods:namePart id="29B27594FE6702377943B22EE88BBFB9">Delocado, Emmanuel D.</mods:namePart>
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<mods:nameIdentifier id="5615F6B1330326FC7A2E0F5252C5D1BD" type="ORCID">https://orcid.org/0000-0001-7731-3476</mods:nameIdentifier>
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<mods:affiliation id="7021FB05A127E238A4412F5C5FCDBF46">Ateneo Biodiversity Research Laboratory, Department of Biology, School of Science and Engineering, Ateneo de Manila University, Quezon City, 1108 Philippines</mods:affiliation>
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<mods:namePart id="28D2F8F507152B4E6D6EB64BA6CDA9D8">Freitag, Hendrik</mods:namePart>
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<mods:nameIdentifier id="92C43C3411E4D37F03FFC2532A869990" type="ORCID">https://orcid.org/0000-0002-1325-0979</mods:nameIdentifier>
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<mods:affiliation id="89ADB9318CC7248728AB61DF71A7228D">Ateneo Biodiversity Research Laboratory, Department of Biology, School of Science and Engineering, Ateneo de Manila University, Quezon City, 1108 Philippines</mods:affiliation>
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<mods:typeOfResource id="6E6A895791502889B5F47FFC83DAE508">text</mods:typeOfResource>
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<mods:title id="4E8757D63C77B2E3930A837AFE8D95B1">ZooKeys</mods:title>
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<mods:part id="B649B3FB7B2270A9DF4349EA8545D01F">
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<mods:date id="DF8D77C39DE0690F644AD01B0871EE64">2022</mods:date>
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<mods:number id="47FE4469C9EAA83AFDAE91BF3B6CCBCD">2022-07-12</mods:number>
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<mods:number id="560CB64C7DB25A86840AC5D33E4195E9">1112</mods:number>
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<mods:url id="692ABF9A80F6CA67BE1B329D6BB510E2">http://dx.doi.org/10.3897/zookeys.1112.85752</mods:url>
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<mods:classification id="94772BE165E08E1B55793EF183106052">journal article</mods:classification>
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<mods:identifier id="19BC5E1ED93EB68FF0FB6B4D65FBAE8C" type="DOI">http://dx.doi.org/10.3897/zookeys.1112.85752</mods:identifier>
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<treatment id="779A4B5041735BF39C0F09B3E8386E7C" ID-GBIF-Taxon="197228274" LSID="urn:lsid:zoobank.org:act:A237568F-6B56-4F69-A5F0-08D4D2311949" httpUri="http://treatment.plazi.org/id/779A4B5041735BF39C0F09B3E8386E7C" lastPageNumber="11" pageId="0" pageNumber="11" scope_family="Hydrophilidae" scope_order="Coleoptera">
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<subSubSection id="CA45A400D4892969028372D475A1B5F2" pageId="0" pageNumber="11" type="nomenclature">
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<paragraph id="1535AF272F7EDCC988E20AE48D335FEF" pageId="0" pageNumber="11">
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<taxonomicName id="68B0CA421958947C75839B94C7143D50" ID-CoL="9GDLC" LSID="http://zoobank.org/A237568F-6B56-4F69-A5F0-08D4D2311949" authority="Sanchez & Delocado & Freitag, 2022" authorityName="Sanchez & Delocado & Freitag" authorityYear="2022" class="Insecta" family="Hydrophilidae" genus="Anacaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anacaena angatbuhay" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="angatbuhay" status="sp. nov.">Anacaena angatbuhay</taxonomicName>
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<taxonomicNameLabel id="B118F2CA177A96F4A5671BFD2811322B" pageId="0" pageNumber="11">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="DD557B7DD1A3235F5DB619C58A16C22D" pageId="0" pageNumber="11" type="description">
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<paragraph id="735255EAC9943CF620498403F1B28C52" pageId="0" pageNumber="11">
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<figureCitation id="7564309AA0095BE767366AFB68B00C12" captionStart="Figure 1–4" captionStartId="F1" captionText="Figure 1 - 4. Habitus of new Anacaena species 1, 2 Anacaena angatbuhay sp. nov. 3, 4 Anacaena auxilium sp. nov. in (1, 3) dorsal and (2, 4) lateral views. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1112.85752.figure1-4" httpUri="https://binary.pensoft.net/fig/713647" pageId="0" pageNumber="11">Figs 1-2</figureCitation>
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<figureCitation id="D0F52BFBF173998DC4ADD36ED4F6294A" captionStart="Figure 5–6" captionStartId="F2" captionText="Figure 5 - 6. Aedeagi of 5 Anacaena angatbuhay sp. nov. holotype (GS 076) 6 Anacaena auxilium sp. nov. holotype (GS 066) in dorsal view. Scale bar: 0.25 mm." figureDoi="10.3897/zookeys.1112.85752.figure5-6" httpUri="https://binary.pensoft.net/fig/713648" pageId="0" pageNumber="11">, 5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="3B68C5B2ABA733487AF84367EE2D33DB" pageId="0" pageNumber="11" type="type locality">
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<paragraph id="E622DB9C57E7710185A908664194ECD2" pageId="0" pageNumber="11">Type locality.</paragraph>
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<paragraph id="1B81B815A46D4A8155F69D769CA73DE0" pageId="0" pageNumber="11">Philippines • Luzon, Ifugao, Banaue, Sumigar Bridge; mountain creek, secondary forest; 16°59'37"N, 121°02'51"E; ca. 1700 m a.s.l.</paragraph>
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</subSubSection>
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<subSubSection id="7987ED273BCDC574B34F0E512FCF4FAD" pageId="0" pageNumber="11" type="type material">
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<paragraph id="FD98EF015F715CB3BB186988CFE9B9E2" pageId="0" pageNumber="11">Type material.</paragraph>
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<paragraph id="B4FB15A966A91A7A2B472F6ADD274510" pageId="0" pageNumber="11">
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<emphasis id="F7F4F3273FF5F63119DF0473D622D1D5" bold="true" italics="true" pageId="0" pageNumber="11">Holotype</emphasis>
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: Philippines • ♂ (PNM: GS076), "PHIL.: Luzon, Ifugao, Banaue, \ Sumigar Bridge; mt. creek, sec. forest; \
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<geoCoordinate id="774B0A2CB8099F7242842A5BDE259F25" degrees="16" direction="north" minutes="59" orientation="latitude" precision="15" seconds="37" value="16.993612">16°59'37"N</geoCoordinate>
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;
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<geoCoordinate id="5A5C81F671847EC62DAE31C9FCBFF22C" degrees="121" direction="east" minutes="02" orientation="longitude" precision="15" seconds="51" value="121.0475">121°02'51"E</geoCoordinate>
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; ca. 1700 m a.s.l.; \ Nov. 1997, leg. Mey (455)L"; GS076, specimen and terminal parts of the abdomen, including genitalia, were glued separately on the entomological card.
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<emphasis id="21701D9FBBCF005421FDEED9606ED74A" bold="true" italics="true" pageId="0" pageNumber="11">Paratypes</emphasis>
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: Philippines • 4♂♂ (ADMU: GS068, GS075; ZMB: GS069): same data as holotype.
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</paragraph>
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</subSubSection>
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<subSubSection id="CB27ACCF5C275F3FBEF98B362528B893" pageId="0" pageNumber="11" type="description">
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<paragraph id="0F373F249545363529B1C5011F00E1BD" pageId="0" pageNumber="11">Description.</paragraph>
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<paragraph id="F3999EA1A34E4057A9C496A026E1C0E2" pageId="0" pageNumber="11">
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(Fig.
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<figureCitation id="DD8C0D42B893F8FB3CDF199F451A64EE" captionStart="Figure 1–4" captionStartId="F1" captionText="Figure 1 - 4. Habitus of new Anacaena species 1, 2 Anacaena angatbuhay sp. nov. 3, 4 Anacaena auxilium sp. nov. in (1, 3) dorsal and (2, 4) lateral views. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1112.85752.figure1-4" httpUri="https://binary.pensoft.net/fig/713647" pageId="0" pageNumber="11">1</figureCitation>
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). TL 2.4 mm (2.1-2.5 mm); TW 1.5 mm (1.3-1.5 mm); EL 1.7 mm (1.5-1.9 mm). Body form oval, moderately convex (Fig.
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<figureCitation id="9B5F8C606B01086299F1464EE9CD6E36" captionStart="Figure 1–4" captionStartId="F1" captionText="Figure 1 - 4. Habitus of new Anacaena species 1, 2 Anacaena angatbuhay sp. nov. 3, 4 Anacaena auxilium sp. nov. in (1, 3) dorsal and (2, 4) lateral views. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1112.85752.figure1-4" httpUri="https://binary.pensoft.net/fig/713647" pageId="0" pageNumber="11">2</figureCitation>
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); elytra about 3.2 times as long as pronotum (dorsal view).
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</paragraph>
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<caption id="47FA8E37C2E1DC4CEF1DB37578A439A3" doi="10.3897/zookeys.1112.85752.figure1-4" httpUri="https://binary.pensoft.net/fig/713647" pageId="0" pageNumber="11" start="Figure 1–4" startId="F1">
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<paragraph id="744771F18C335473EF1B2D75800A390C" pageId="0" pageNumber="11">
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<emphasis id="EF16BE1FC668F67A4B48F2A6C68993AF" bold="true" pageId="0" pageNumber="11">Figure 1-4.</emphasis>
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Habitus of new
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<taxonomicName id="D8BDB40B4C6C682AE978370A3E3838FE" authorityName="C.G. Thomson" authorityYear="1859" class="Insecta" family="Hydrophilidae" genus="Anacaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anacaena" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis id="874853CEC161F8CD95278C196EBB1B6E" italics="true" pageId="0" pageNumber="11">Anacaena</emphasis>
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</taxonomicName>
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species
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<emphasis id="7D068C2E4DC5B6EC5304071310853A22" bold="true" pageId="0" pageNumber="11">1, 2</emphasis>
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<taxonomicName id="DD972D241714D8CFD3C3E1126B436E26" authorityName="Sanchez & Delocado & Freitag" authorityYear="2022" class="Insecta" family="Hydrophilidae" genus="Anacaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anacaena angatbuhay" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="angatbuhay">
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<emphasis id="07001431DAC1727F5CA17B7BAA5109BA" italics="true" pageId="0" pageNumber="11">Anacaena angatbuhay</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis id="7628F2F70DFDD952D453481F5E043F6A" bold="true" pageId="0" pageNumber="11">3, 4</emphasis>
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<taxonomicName id="4F860747FF45E0B3C5F2F0B672712A9D" authorityName="Sanchez & Delocado & Freitag" authorityYear="2022" class="Insecta" family="Hydrophilidae" genus="Anacaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anacaena auxilium" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="auxilium">
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<emphasis id="E0D127FA43635991C1F0F345C57BFEEA" italics="true" pageId="0" pageNumber="11">Anacaena auxilium</emphasis>
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</taxonomicName>
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sp. nov. in (
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<emphasis id="730CB44ED2181CDF0EAB8557D0544DB2" bold="true" pageId="0" pageNumber="11">1, 3</emphasis>
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) dorsal and (
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<emphasis id="825551228CE97471B1C039F550E9BF9B" bold="true" pageId="0" pageNumber="11">2, 4</emphasis>
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) lateral views. Scale bar: 1 mm.
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</paragraph>
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</caption>
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<paragraph id="68446DE01779A239EB251F7716A41512" pageId="0" pageNumber="11">
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<emphasis id="6015085FA27BCDEAE1EAEAC67DFCA0CC" bold="true" italics="true" pageId="0" pageNumber="11">Head</emphasis>
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: Clypeus dark brown to black, moderately large, anterior margin straight, with indistinct antero-lateral angle. Frons black; frontoclypeal suture visible. Labrum black. Puncturation regular, coarse, strongly impressed throughout; interstices as wide as diameter of one puncture; series of densely arranged punctures along inner margin of eyes absent. Ventral punctures irregular, coarse, shallowly impressed, interstices as wide as diameter of one puncture. Maxillary palpomeres dark yellow to brown; palpomeres 1-3 dark yellow; palpomere 2 moderately inflated, lateral margins black. Palpomere 4 widest toward mid-length wider proximally than distally; inner margin straight; outer margin distinctly convex; proximal half yellow; anterior 0.3 black, but apex yellow. Mentum with fine setae on lateral margins; anterior margin with distinct median incision. Labial palpi stout, not longer than lateral edge of mentum; palpomere 3 about twice as long as palpomere 2. Eyes not constricted anteriorly, dorsal and ventral portions of almost equal size. Antennae capitate, 9-segmented; scape triangular, broadest at base; pedicel oval; antennomere 1 (segment 3) elongate oval, pointed bluntly at apex; antennomeres 2-4 smallest in length and width, paler than adjacent segments, indistinct intermediate segments, slightly longer than antennomere 1, decreasing in size distally; antennomere 5-7 darkest of all antennomeres; antennomeres 5 and 6 of equal length, with numerous, thick, erect, dark brown setae. Segment 7 (first club segment) irregularly globular; segment 8 globular. Terminal segment slightly longer than segments 7+8, widest near mid-length, curved on lateral margins, asymmetrical apicad; setae distributed throughout, dark brown, long, thick, erect, but setae on apical one-third longer. Preocular patches absent.
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</paragraph>
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<paragraph id="36316FF370FBE50ACC0BC719801FFE5A" pageId="0" pageNumber="11">
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<emphasis id="13DFC8FFF291273884C7D57662ABB7CD" bold="true" italics="true" pageId="0" pageNumber="11">Thorax</emphasis>
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: PW/PL = 2.93; PL/PW = 0.34. Pronotum dark brown on the disc, light brown anteriad and laterad, with narrow light brown to yellow margins not wider than diameter of eye. Pronotal punctures large, sparse, deeply impressed on the disc, shallowly impressed laterad; interstices as wide as diameter of 2-5 punctures. Anterior margin curving slightly inwards on both sides behind the eye, gradually curving outwards starting from lateral 0.15-0.2 on both ends. Lateral margins almost straight, with setae more numerous in the anterior portion. Postero-lateral angles slightly rounded, ca. 80-85°. Posterior margin thickly bordered, almost straight. Prosternum flat. EL/EW = 1.2; EL/PL = 4.1; EW/PW = 1.1; TL/EW = 1.8. Elytra dark brown, darkest on disc, with very narrow yellow lateral margins; setae thin, very long, some setae one-fifth length of elytra, unevenly and sparsely distributed, but denser on the disc. Elytral punctures coarse, moderately impressed, arrangement denser on the disc, spacing as wide as punctures; rows of coarser punctures on lateral portion present, but not very distinct. Mesoventrite distinctly elevated medially, with protuberance subtly bulging.
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</paragraph>
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<paragraph id="150A886DC4F74172E1AB9880C9EBB2D4" pageId="0" pageNumber="11">
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<emphasis id="2BB24358596012FC44C07755C170DACA" bold="true" italics="true" pageId="0" pageNumber="11">Legs</emphasis>
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: Light brown, but tarsi amber-coloured. Procoxa with spine-like setae. Metafemur with minute pubescence on proximal one-fourth near outer margin, hairline direction horizontal and recumbent. Tibia with long, thick, brown, spine-like setae along lateral margins, pointed distally; distal end of tibia with coupled pairs of setae distinctly longer than proximal setae, with exterior pair longer than interior pair; setae uneven in length, with terminal pair of setae at least 1.5 as long as adjacent setae; tibiae length ratio (protibia: mesotibia: metatibia) 1.0: 1.3: 2.0. Metatarsus size slightly longer than metatibia. Tarsi length ratio (protarsus: mesotarsus: metatarsus) 1.0: 2.0: 2.4.
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</paragraph>
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<paragraph id="09B466A1939582D66A94AF25AD5E1E36" pageId="0" pageNumber="11">
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<emphasis id="03F3C0A0774109C158BD43622475836B" bold="true" italics="true" pageId="0" pageNumber="11">Aedeagus</emphasis>
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: (Fig.
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<figureCitation id="84879A9DC9D4AAFCCF8BA1A3F03AB36E" captionStart="Figure 5–6" captionStartId="F2" captionText="Figure 5 - 6. Aedeagi of 5 Anacaena angatbuhay sp. nov. holotype (GS 076) 6 Anacaena auxilium sp. nov. holotype (GS 066) in dorsal view. Scale bar: 0.25 mm." figureDoi="10.3897/zookeys.1112.85752.figure5-6" httpUri="https://binary.pensoft.net/fig/713648" pageId="0" pageNumber="11">5</figureCitation>
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) Parameres symmetrical, apices rounded; apical region widens laterally; inner margins slightly concaving until apical one-third, then forming a mesal gap, converging to base; outer margins slightly biconvex with pronounced lateral expansion towards posterior 0.4 then slightly narrowing; combined width of parameres approximately the same or slightly broader as phallobase; basal portion 5 times as wide as apical part. Median lobe with apex rounded, broadens starting in anterior 0.6, abruptly bulging prominently at posterior 0.25, then narrowing before converging at the base; base of median lobe about 4 times as wide as width of apex; basal apophyses extending about one-third into the phallobase, distinctly curving outwards, narrowing basad. Phallobase longer than parameres, longer than wide; median reinforcement or pigmented line absent.
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</paragraph>
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<caption id="FA6ED2113610A4C6AA7C23EC8F5FD5D1" doi="10.3897/zookeys.1112.85752.figure5-6" httpUri="https://binary.pensoft.net/fig/713648" pageId="0" pageNumber="11" start="Figure 5–6" startId="F2">
|
||
<paragraph id="F3DA391B1CAB2AECA9FBE09A623D9439" pageId="0" pageNumber="11">
|
||
<emphasis id="03E5126AE882156964086059E0C55777" bold="true" pageId="0" pageNumber="11">Figure 5-6.</emphasis>
|
||
Aedeagi of
|
||
<emphasis id="4B45BD0F1CB4EAA44BAD395BA6AEB723" bold="true" pageId="0" pageNumber="11">5</emphasis>
|
||
<taxonomicName id="75CED24278728D563C9272CF8B6BEFD9" authorityName="Sanchez & Delocado & Freitag" authorityYear="2022" class="Insecta" family="Hydrophilidae" genus="Anacaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anacaena angatbuhay" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="angatbuhay">
|
||
<emphasis id="70385C20E0851CC1226B3B2E78E5E5E6" italics="true" pageId="0" pageNumber="11">Anacaena angatbuhay</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. holotype (GS076)
|
||
<emphasis id="49AC1FC4D717395F299A8DC9816ABCE3" bold="true" pageId="0" pageNumber="11">6</emphasis>
|
||
<taxonomicName id="BC9CABCDEB80BBC169F68F7507DEDFC6" authorityName="Sanchez & Delocado & Freitag" authorityYear="2022" class="Insecta" family="Hydrophilidae" genus="Anacaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anacaena auxilium" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="auxilium">
|
||
<emphasis id="54D2D09388A503C78AF3DF58F6F04151" italics="true" pageId="0" pageNumber="11">Anacaena auxilium</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. holotype (GS066) in dorsal view. Scale bar: 0.25 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="1C4E9B40FE8A9C75D4986A067D681129" pageId="0" pageNumber="11" type="differential diagnosis">
|
||
<paragraph id="2631F84C6B230A858B8C55606A688110" pageId="0" pageNumber="11">Differential diagnosis.</paragraph>
|
||
<paragraph id="00A3E0262AD59A2EA8C23BEFD0A18160" pageId="0" pageNumber="11">
|
||
<taxonomicName id="34C7895A5B1BDB5151469426380507A0" lsidName="A. angatbuhay" pageId="0" pageNumber="11" rank="species" species="angatbuhay">
|
||
<emphasis id="178FD4A79800044D3299B6FDA0FF58ED" italics="true" pageId="0" pageNumber="11">A. angatbuhay</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (Fig.
|
||
<figureCitation id="AD1C5CAB24B143C256DF3BBB5ACCEA94" captionStart="Figure 1–4" captionStartId="F1" captionText="Figure 1 - 4. Habitus of new Anacaena species 1, 2 Anacaena angatbuhay sp. nov. 3, 4 Anacaena auxilium sp. nov. in (1, 3) dorsal and (2, 4) lateral views. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1112.85752.figure1-4" httpUri="https://binary.pensoft.net/fig/713647" pageId="0" pageNumber="11">1</figureCitation>
|
||
) resembles
|
||
<taxonomicName id="FAED166532D48D01EEFFD97DF3FB2B50" lsidName="A. philippina" pageId="0" pageNumber="11" rank="species" species="philippina">
|
||
<emphasis id="A11BC9C144D85C6A704B646EFF41156F" italics="true" pageId="0" pageNumber="11">A. philippina</emphasis>
|
||
</taxonomicName>
|
||
Komarek & Freitag, 2014 and
|
||
<taxonomicName id="1E8E96EFE9256E8C8CFF53BCB91E3D10" lsidName="A. zamboangana" pageId="0" pageNumber="11" rank="species" species="zamboangana">
|
||
<emphasis id="AE1BB71DE53D0971A046C096ED6F37EE" italics="true" pageId="0" pageNumber="11">A. zamboangana</emphasis>
|
||
</taxonomicName>
|
||
Komarek & Freitag, 2014 in terms of colour and size. However, the new species is different from these congeners in that its labial palpi are stout compared to the slender labial palpi of
|
||
<taxonomicName id="27301D95A57A96127E5EA4CDB59EA981" lsidName="A. philippina" pageId="0" pageNumber="11" rank="species" species="philippina">
|
||
<emphasis id="1B36B0D9E731DC4F1FE59BF292519A44" italics="true" pageId="0" pageNumber="11">A. philippina</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="382D6CE4104851056955E9803BE0BF4A" lsidName="A. zamboangana" pageId="0" pageNumber="11" rank="species" species="zamboangana">
|
||
<emphasis id="3C15F91D967AAEA257A67B0B57A5938D" italics="true" pageId="0" pageNumber="11">A. zamboangana</emphasis>
|
||
</taxonomicName>
|
||
. Additionally, the new species has spine-like setae on procoxa unlike
|
||
<taxonomicName id="DC4775A60B99C949E1C882DF55E4BC84" lsidName="A. philippina" pageId="0" pageNumber="11" rank="species" species="philippina">
|
||
<emphasis id="933E17C8A02E964F3C9DCC3D96BC0204" italics="true" pageId="0" pageNumber="11">A. philippina</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="AB96041A90BD55F84B4A1D0F8C1B2225" lsidName="A. zamboangana" pageId="0" pageNumber="11" rank="species" species="zamboangana">
|
||
<emphasis id="65B64269F4254F0C3C33954ADF088331" italics="true" pageId="0" pageNumber="11">A. zamboangana</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="C7A6493C476C30E530E9F5FCEDB0090F" pageId="0" pageNumber="11">
|
||
Moreover, in terms of the male genitalia,
|
||
<taxonomicName id="0D133DB753C98D4BFA21757E5EAC5276" lsidName="A. angatbuhay" pageId="0" pageNumber="11" rank="species" species="angatbuhay">
|
||
<emphasis id="8A5F92354EFDF932E41F2B297D350468" italics="true" pageId="0" pageNumber="11">A. angatbuhay</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName id="3B1E77AF791BC432EDF1EE064C479086" lsidName="A. philippina" pageId="0" pageNumber="11" rank="species" species="philippina">
|
||
<emphasis id="0100BED38B4F8AE2793C580EC6F6D83C" italics="true" pageId="0" pageNumber="11">A. philippina</emphasis>
|
||
</taxonomicName>
|
||
exhibit similarities in the apical portion of their median lobes. While the median lobe of
|
||
<taxonomicName id="5176CBDC1973B012855C12091869E548" lsidName="A. philippina" pageId="0" pageNumber="11" rank="species" species="philippina">
|
||
<emphasis id="2CCAE9F6C102108D434B6C4BC0073DA7" italics="true" pageId="0" pageNumber="11">A. philippina</emphasis>
|
||
</taxonomicName>
|
||
has straight margins, the median lobe of the new species has a distinctly pointed lateral expansion towards the middle, loosely resembling a diamond shape. Additionally, the phallobase of
|
||
<taxonomicName id="8AA22130B6C55C60FD1DD30A5C3ED6A7" lsidName="A. angatbuhay" pageId="0" pageNumber="11" rank="species" species="angatbuhay">
|
||
<emphasis id="C9E676038377C2186CAD28EA82C74736" italics="true" pageId="0" pageNumber="11">A. angatbuhay</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is considerably broader than that of
|
||
<taxonomicName id="6A87398BF39E7F3343D41C26EF18C520" lsidName="A. philippina" pageId="0" pageNumber="11" rank="species" species="philippina">
|
||
<emphasis id="D6392DF320CF3B93C07B9750C920BE64" italics="true" pageId="0" pageNumber="11">A. philippina</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4D435096859B5C0502894E181954E506" pageId="0" pageNumber="11" type="distribution">
|
||
<paragraph id="54852A397B0DA756903F182B1B81B06D" pageId="0" pageNumber="11">Distribution.</paragraph>
|
||
<paragraph id="37AD0403EE51A2C754F582144415DDF8" pageId="0" pageNumber="11">
|
||
The species is only known from the type locality (Fig.
|
||
<figureCitation id="00D67B2CCD4F37C8B3D061FF8C33BA17" captionStart="Figure 7" captionStartId="F3" captionText="Figure 7. Map showing the updated distribution of Anacaena in Northern Luzon, including Komarek & Freitag (2014) data." figureDoi="10.3897/zookeys.1112.85752.figure7" httpUri="https://binary.pensoft.net/fig/713649" pageId="0" pageNumber="11">7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DA9B9968C99F374C40858CB7C54A992A" doi="10.3897/zookeys.1112.85752.figure7" httpUri="https://binary.pensoft.net/fig/713649" pageId="0" pageNumber="11" start="Figure 7" startId="F3">
|
||
<paragraph id="7D02C0C9E195513470662AEBC3D633B1" pageId="0" pageNumber="11">
|
||
<emphasis id="3F8BE6FFBE2AC9C6C617D13CA9CC11B1" bold="true" pageId="0" pageNumber="11">Figure 7.</emphasis>
|
||
Map showing the updated distribution of
|
||
<taxonomicName id="73D5DAAE8D4B506FEB5F015932159529" authorityName="C.G. Thomson" authorityYear="1859" class="Insecta" family="Hydrophilidae" genus="Anacaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anacaena" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="355478DE2DCDDB39138373BD3A9CC0C7" italics="true" pageId="0" pageNumber="11">Anacaena</emphasis>
|
||
</taxonomicName>
|
||
in Northern Luzon, including Komarek & Freitag (2014) data.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="7A7C7C7A1A6468D9A93C4D22B1A030A9" pageId="0" pageNumber="11" type="remarks">
|
||
<paragraph id="A1331BCCA0C9B763E8AC8E233C690A05" pageId="0" pageNumber="11">Remarks.</paragraph>
|
||
<paragraph id="075C6DEBB0D47C574B85BEB706529EC8" pageId="0" pageNumber="11">No external sexual dimorphism is observed.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D5F33452A260B89844B7B1719A29FEE6" pageId="0" pageNumber="11" type="etymology">
|
||
<paragraph id="77272E4008AF1C8860C398FCA27808BC" pageId="0" pageNumber="11">Etymology.</paragraph>
|
||
<paragraph id="56448522FD2F5AB336BC793003C52942" pageId="0" pageNumber="11">
|
||
The species epithet alludes to the Angat Buhay (literal translation: lifting lives) anti-poverty flagship programme of Her Excellency Maria Leonor
|
||
<normalizedToken id="929DE337EBAA20F4CF927B0278D853C2" originalValue="“Leni”">"Leni"</normalizedToken>
|
||
Gerona Robredo, the 14th Vice President of the Republic of the Philippines who concluded her term this year. It is in honour of the exemplary service of her office, with emphasis on engaging with local communities, uplifting the marginalised and heeding the needs of the health care sector, especially in the time of the pandemic. The term is used as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |