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<document ID-CLB-Dataset="35472" ID-DOI="http://dx.doi.org/10.3897/zookeys.245.3416" ID-PMC="PMC3577090" ID-Pensoft-Pub="1313-2970-245-1" ID-Pensoft-UUID="FFFF52503A0AFF882450FFB66D45FF8E" ID-PubMed="23431087" ID-Zenodo-Dep="578462" ModsDocID="1313-2970-245-1" checkinTime="1425642919252" checkinUser="pensoft" docAuthor="Bousquet, Yves" docDate="2012" docId="76AE445BCD89511E8EBD4A17798FC45B" docLanguage="en" docName="ZooKeys 245: 1-1722" docOrigin="ZooKeys 245" docPubDate="2012-11-28" docSource="http://dx.doi.org/10.3897/zookeys.245.3416" docTitle="Bembidium (Cillenus) Samouelle 1819" docType="treatment" docVersion="9" id="FFFF52503A0AFF882450FFB66D45FF8E" lastPageNumber="364" masterDocId="FFFF52503A0AFF882450FFB66D45FF8E" masterDocTitle="Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico" masterLastPageNumber="1722" masterPageNumber="1" pageNumber="364" updateTime="1736396794539" updateUser="felipe">
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<mods:title id="20CCE30D55724067DF36DDA75D429586">Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico</mods:title>
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<mods:namePart id="F6AD6D4B590D3E0A277A53C1AA0C0277">Bousquet, Yves</mods:namePart>
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<treatment id="76AE445BCD89511E8EBD4A17798FC45B" ID-GBIF-Taxon="182238511" LSID="urn:lsid:plazi:treatment:76AE445BCD89511E8EBD4A17798FC45B" httpUri="http://treatment.plazi.org/id/76AE445BCD89511E8EBD4A17798FC45B" lastPageNumber="364" pageId="363" pageNumber="364">
<subSubSection id="46414E6C8CFC998F6DEE9D3EE04FD614" pageId="363" pageNumber="364" type="nomenclature">
<paragraph id="B2D34CF28C926D70B03EB0EA5F527352" pageId="363" pageNumber="364">
Subgenus
<taxonomicName id="FB294C0D6DFEBF0F3D5EBC618E381E26" LSID="76AE445B-CD89-511E-8EBD-4A17798FC45B" authority="Samouelle, 1819" authorityName="Samouelle" authorityYear="1819" class="Insecta" family="Carabidae" genus="Bembidium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="(Cillenus)" order="Coleoptera" pageId="363" pageNumber="364" phylum="Arthropoda" rank="subGenus" subGenus="Cillenus">Cillenus Samouelle, 1819</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="A887E352F71E8AB814B5C61DE4ED04CB" pageId="363" pageNumber="364" type="reference_group">
<paragraph id="E931598B8559845509AFFC65577B8848" pageId="363" pageNumber="364">
<taxonomicName id="78FB4F5600C9C3474E15F74B3734BE4B" class="Insecta" family="Carabidae" genus="Cillenus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="363" pageNumber="364" phylum="Arthropoda" rank="genus">Cillenus</taxonomicName>
Samouelle, 1819: 148. Type species:
<taxonomicName id="314662FABAEADA26CC2A419D99ADFD5D" authorityName="Leach in Samouelle" authorityYear="1819" class="Insecta" family="Carabidae" genus="Cillenus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="363" pageNumber="364" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="03F67F55E8A20C28185A67694A27F62C" italics="true" pageId="363" pageNumber="364">Cillenus lateralis</emphasis>
</taxonomicName>
Samouelle, 1819 by monotypy. Etymology. Unknown [masculine].
</paragraph>
<paragraph id="C926DC2C4701B70D7900C537AB5F358B" pageId="363" pageNumber="364">
<taxonomicName id="7128F38135E5044897076BDE88AAF3B8" class="Insecta" family="Carabidae" genus="Cillenum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="363" pageNumber="364" phylum="Arthropoda" rank="genus">Cillenum</taxonomicName>
Curtis, 1828: plate 200. Unjustified emendation of
<taxonomicName id="75CC0E61DC1A2460D72E62DB1AD8AD3D" class="Insecta" family="Carabidae" genus="Cillenus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="363" pageNumber="364" phylum="Arthropoda" rank="genus">
<emphasis id="1EF04C30F0B14037B3A3EF2283C64F47" italics="true" pageId="363" pageNumber="364">Cillenus</emphasis>
</taxonomicName>
Samouelle, 1819.
</paragraph>
</subSubSection>
<subSubSection id="9FF2CF9F35E532744DC25EC5D474E11F" pageId="363" pageNumber="364" type="diversity">
<paragraph id="D835706BBFA56728251D550372CFB242" pageId="363" pageNumber="364">Diversity.</paragraph>
<paragraph id="C589FD43C835D34421F17E756557F165" pageId="363" pageNumber="364">
Two species, one along the Pacific Coast of North America, the other (
<taxonomicName id="80A31B04983641400A324F372DFD5743" authorityName="Dejean" authorityYear="1831" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="363" pageNumber="364" phylum="Arthropoda" rank="species" species="laterale">
<emphasis id="FDAA60AE13BDCC89273038FDF2820FF2" italics="true" pageId="363" pageNumber="364">Bembidion laterale</emphasis>
</taxonomicName>
Samouelle) along the coasts of Europe and northern Africa. The three other species listed in this subgenus by Lorenz (2005: 216) belong to the subgenus
<taxonomicName id="EA12D1B9803142DC1615A923B9B30498" authorityName="Netolitzky" authorityYear="1942" class="Insecta" family="Carabidae" genus="Desarmatocillenus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="363" pageNumber="364" phylum="Arthropoda" rank="genus">
<emphasis id="C32F440C02BE7A8B96B46431719C73BA" italics="true" pageId="363" pageNumber="364">Desarmatocillenus</emphasis>
</taxonomicName>
Netolitzky (see Lindroth 1980).
</paragraph>
</subSubSection>
<subSubSection id="2CF67E8151D0F72005501645F7B246D3" pageId="363" pageNumber="364" type="identification">
<paragraph id="F366B12D4CFC827BC5999B36281FBE69" pageId="363" pageNumber="364">Identification.</paragraph>
<paragraph id="22E3771E8337A5BEFBA11E90BB6AF6DB" pageId="363" pageNumber="364">
Kavanaugh and Erwin (1992: 317) emended
<normalizedToken id="5C29C8DB9C11B35663B7E8FE649EAEB6" originalValue="Lindroths">Lindroth's</normalizedToken>
(1963b) key to
<taxonomicName id="8301D3BD684225D34A2B007C677DD87F" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="363" pageNumber="364" phylum="Arthropoda" rank="genus">
<emphasis id="414CF953F60034D101D4AFA4AB2410E2" italics="true" pageId="363" pageNumber="364">Bembidion</emphasis>
</taxonomicName>
to accommodate the new species.
</paragraph>
</subSubSection>
<subSubSection id="98715DAE155766A3CC5D1992A3418BF2" pageId="363" pageNumber="364" type="taxonomic note">
<paragraph id="1310416253E56F0AA5813BE3E6995144" pageId="363" pageNumber="364">Taxonomic Note.</paragraph>
<paragraph id="85E2B7F5F1186ED1DE7A114DBB479572" pageId="363" pageNumber="364">
This taxon is ranked as a distinct genus by many authors, with
<taxonomicName id="2BFCDEE0A6C241BFB114E3F0A4725DE8" authorityName="Dupuis" authorityYear="1912" class="Insecta" family="Carabidae" genus="Armatocillenus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="363" pageNumber="364" phylum="Arthropoda" rank="genus">
<emphasis id="1A055E545EF2C860E6B59E5B37323556" italics="true" pageId="363" pageNumber="364">Armatocillenus</emphasis>
</taxonomicName>
Dupuis (three species in Taiwan and New Guinea),
<taxonomicName id="5CD4163249CBA303A66B7B7D53F0C4DA" authorityName="Netolitzky" authorityYear="1942" class="Insecta" family="Carabidae" genus="Desarmatocillenus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="363" pageNumber="364" phylum="Arthropoda" rank="genus">
<emphasis id="AA9A9AEA6EF4F715890FC276082C8675" italics="true" pageId="363" pageNumber="364">Desarmatocillenus</emphasis>
</taxonomicName>
Netolitzky (14 species in southeastern Asia and the Australian Region),
<taxonomicName id="6B2FCCEF0655727B4E956471E2C439B6" authorityName="Netolitzky" authorityYear="1942" class="Insecta" family="Carabidae" genus="Chinocillenus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="363" pageNumber="364" phylum="Arthropoda" rank="genus">
<emphasis id="E7DA260CF0B542693E0521F71F7300F2" italics="true" pageId="363" pageNumber="364">Chinocillenus</emphasis>
</taxonomicName>
Netolitzky (one Chinese species),
<taxonomicName id="5AEF9ED0990791A0FFB795C3710DE64E" authorityName="Ueno" authorityYear="1955" class="Insecta" family="Carabidae" genus="Corallicillenus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="363" pageNumber="364" phylum="Arthropoda" rank="genus">
<emphasis id="CCB83A6777632936B0756000CB6D23C4" italics="true" pageId="363" pageNumber="364">Corallicillenus</emphasis>
</taxonomicName>
<normalizedToken id="FDEA9BF00D396DE494A72FC094B6C401" originalValue="Uéno">Ueno</normalizedToken>
(two Japanese species), and
<taxonomicName id="5A16CDE2AD071E433B0C805EADB5AFE6" authorityName="Ueno &amp; Habu" authorityYear="1955" class="Insecta" family="Carabidae" genus="Novicillenus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="363" pageNumber="364" phylum="Arthropoda" rank="genus">
<emphasis id="3FA66151B157E652D029B028E05BA0A7" italics="true" pageId="363" pageNumber="364">Novicillenus</emphasis>
</taxonomicName>
<normalizedToken id="72864348DF2004F8A01930C946315F5A" originalValue="Uéno">Ueno</normalizedToken>
and Habu (two Japanese species) listed as subgenera (see Lorenz 2005: 216). Recently Sasakawa (2007) synonymized
<taxonomicName id="AB3BA95BF11CFF8C97A92B0AE383DBDA" authorityName="Ueno &amp; Habu" authorityYear="1955" class="Insecta" family="Carabidae" genus="Novicillenus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="363" pageNumber="364" phylum="Arthropoda" rank="genus">
<emphasis id="7B2C806E638820D33E9A7F950292615C" italics="true" pageId="363" pageNumber="364">Novicillenus</emphasis>
</taxonomicName>
and
<taxonomicName id="6A9296A7F4F1025C2C085FD0AC8E64D5" authorityName="Ueno" authorityYear="1955" class="Insecta" family="Carabidae" genus="Corallicillenus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="363" pageNumber="364" phylum="Arthropoda" rank="genus">
<emphasis id="148FC6F85C9B6519B5F9C4F73EA28EF5" italics="true" pageId="363" pageNumber="364">Corallicillenus</emphasis>
</taxonomicName>
with
<taxonomicName id="D66EB1A61F7B01F4AF5A2FC6AD164D9D" authorityName="Netolitzky" authorityYear="1942" class="Insecta" family="Carabidae" genus="Desarmatocillenus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="363" pageNumber="364" phylum="Arthropoda" rank="genus">
<emphasis id="296FCFC7909543B983799C8B305B4835" italics="true" pageId="363" pageNumber="364">Desarmatocillenus</emphasis>
</taxonomicName>
. Recent molecular data analyses conducted by Maddison and Ober (2011: 251) and Maddison (2012: 568) strongly support
<taxonomicName id="044B391D4F8CD9FECB92525DF6C47E72" class="Insecta" family="Carabidae" genus="Cillenus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="363" pageNumber="364" phylum="Arthropoda" rank="genus">
<emphasis id="7D708D0BCE3E33306DE6150339BBF408" italics="true" pageId="363" pageNumber="364">Cillenus</emphasis>
</taxonomicName>
as a member of
<taxonomicName id="2B5E3C5BF7791EE159452519F546FA03" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="363" pageNumber="364" phylum="Arthropoda" rank="genus">
<emphasis id="90E7D2B3BF159176A17152688AB75D0B" italics="true" pageId="363" pageNumber="364">Bembidion</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>