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<document id="783E7AB3DDEC57150412C9438AF0995A" ID-DOI="10.11646/zootaxa.5496.3.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="13346901" ID-ZooBank="58D78B70-604A-42F5-86D4-4395CDB17A41" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1724133547272" checkinUser="plazi" docAuthor="Shah, Bismillah, Shao, Yisa, Du, Hong, Wang, Yang &amp; Huang, Junhao" docDate="2024" docId="766187C56737FFE6FF76FE86F19DFDBA" docLanguage="en" docName="zootaxa.5496.3.5.pdf" docOrigin="Zootaxa 5496 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5496.3.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zygoneura (Pharetratula) yangi Shah &amp; Shao &amp; Du &amp; Wang &amp; Huang 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="394" masterDocId="8A58FFBD6727FFF7FFE1FFE9F30AFFDB" masterDocTitle="Taxonomy and DNA barcoding of the dark-winged fungus gnat genus Zygoneura Meigen (Diptera: Sciaridae) from China, with revision of the type materials" masterLastPageNumber="400" masterPageNumber="377" pageNumber="393" updateTime="1724134118297" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B21392B816C2C70C65E5B622D1CB80E6">Taxonomy and DNA barcoding of the dark-winged fungus gnat genus Zygoneura Meigen (Diptera: Sciaridae) from China, with revision of the type materials</mods:title>
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<mods:namePart id="A811C45B4C11D591405FB0DBDBDE2BF4">Shah, Bismillah</mods:namePart>
<mods:nameIdentifier id="1902E04248CE6DD75E00CF1FA2A6697A" type="ORCID">0000-0002-8407-8627</mods:nameIdentifier>
<mods:affiliation id="CC1FCEF4727D425A3882D5C1F393C7AA">Department of Forestry Protection, School of Forestry and Biotechnology, Zhejiang A &amp; F University, 666 Wusu street, Linan, Hangzhou, Zhejiang 311300, P. R. China</mods:affiliation>
<mods:nameIdentifier id="1C68A0EC6C099B7681073ED89D12365E" type="email">bismillahshah1990@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="E55B1CF7626FF6272D55811296334F0A">Shao, Yisa</mods:namePart>
<mods:nameIdentifier id="B7F256985963C003432FC60CAD7ED49A" type="ORCID">0009-0008-5493-6409</mods:nameIdentifier>
<mods:affiliation id="360BABCF37D1D955D3910D52B1653F94">Department of Forestry Protection, School of Forestry and Biotechnology, Zhejiang A &amp; F University, 666 Wusu street, Linan, Hangzhou, Zhejiang 311300, P. R. China</mods:affiliation>
<mods:nameIdentifier id="6070ED1CEB41321B44A8C9D92FA989B2" type="email">2909187740@qq.com</mods:nameIdentifier>
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<mods:namePart id="5B4473906DF8EB88C87EC43EA8531681">Du, Hong</mods:namePart>
<mods:nameIdentifier id="34E9FCD84088F98FE7C5822E0C2288FC" type="ORCID">0009-0009-4843-4899</mods:nameIdentifier>
<mods:affiliation id="1340607FBD65C42BDF0124A3BA6772F3">Department of Forestry Protection, School of Forestry and Biotechnology, Zhejiang A &amp; F University, 666 Wusu street, Linan, Hangzhou, Zhejiang 311300, P. R. China</mods:affiliation>
<mods:nameIdentifier id="3F757FE492A49BD1BAD1C463CD274317" type="email">983468026@qq.com</mods:nameIdentifier>
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<mods:namePart id="DB055C404E931E279FF30BE5DAC38778">Wang, Yang</mods:namePart>
<mods:nameIdentifier id="5B1F811E1C6C753043C6A97C001081C1" type="ORCID">0009-0002-2897-8193</mods:nameIdentifier>
<mods:affiliation id="00DDF88EF33CC4600A9C565C308A1D6B">Comprehensive Service Center of Public Institutions in Qiantan Town, Jiande, Zhejiang 311602, P. R. China</mods:affiliation>
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<mods:namePart id="E320F8DBCEF89A3763BE1945BE68C16A">Huang, Junhao</mods:namePart>
<mods:nameIdentifier id="F625D55871FBF69A92B7F08EEA230291" type="ORCID">0000-0002-5038-3535</mods:nameIdentifier>
<mods:affiliation id="0574B98B66D46302CF88818E1B57FFCF">Department of Forestry Protection, School of Forestry and Biotechnology, Zhejiang A &amp; F University, 666 Wusu street, Linan, Hangzhou, Zhejiang 311300, P. R. China</mods:affiliation>
<mods:nameIdentifier id="5670C22C6C2EA99267A0F8BCED7B9957" type="email">huangjh@zafu.edu.cn</mods:nameIdentifier>
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<mods:date id="198D5BD9BCFC9C22A721A3B5C096EEDF">2024</mods:date>
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<mods:number id="440003F54B91FE104619E3C6519BA961">5496</mods:number>
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<treatment id="766187C56737FFE6FF76FE86F19DFDBA" ID-DOI="http://doi.org/10.5281/zenodo.13346955" ID-Zenodo-Dep="13346955" LSID="urn:lsid:plazi:treatment:766187C56737FFE6FF76FE86F19DFDBA" httpUri="http://treatment.plazi.org/id/766187C56737FFE6FF76FE86F19DFDBA" lastPageId="17" lastPageNumber="394" pageId="16" pageNumber="393">
<subSubSection id="B6D265586737FFE7FF76FE86F049FE52" box="[151,835,366,393]" pageId="16" pageNumber="393" type="nomenclature">
<paragraph id="FE7736D36737FFE7FF76FE86F049FE52" blockId="16.[151,835,366,429]" box="[151,835,366,393]" pageId="16" pageNumber="393">
<heading id="A53F81BF6737FFE7FF76FE86F049FE52" bold="true" box="[151,835,366,393]" fontSize="11" level="1" pageId="16" pageNumber="393" reason="1">
<emphasis id="CCBCEAC16737FFE7FF76FE86F049FE52" bold="true" box="[151,835,366,393]" pageId="16" pageNumber="393">
<taxonomicName id="39C84D506737FFE7FF76FE86F1D4FE52" authority="Shah &amp; Huang" authorityName="Shah &amp; Shao &amp; Du &amp; Wang &amp; Huang" authorityYear="2024" box="[151,734,366,393]" class="Insecta" family="Sciaridae" genus="Zygoneura" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="393" phylum="Arthropoda" rank="species" species="yangi" status="sp. nov." subGenus="Pharetratula">
<emphasis id="CCBCEAC16737FFE7FF76FE86F211FE52" bold="true" box="[151,283,367,393]" italics="true" pageId="16" pageNumber="393">Zygoneura</emphasis>
(
<emphasis id="CCBCEAC16737FFE7FECDFE87F2CDFE52" bold="true" box="[300,455,366,393]" italics="true" pageId="16" pageNumber="393">Pharetratula</emphasis>
)
<emphasis id="CCBCEAC16737FFE7FE39FE86F111FE52" bold="true" box="[472,539,367,393]" italics="true" pageId="16" pageNumber="393">yangi</emphasis>
Shah &amp; Huang
</taxonomicName>
<taxonomicNameLabel id="D78F57BA6737FFE7FD04FE86F049FE52" box="[741,835,367,393]" pageId="16" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="B6D265586737FFE7FF76FE7AF23EFE76" box="[151,308,403,429]" pageId="16" pageNumber="393" type="description">
<paragraph id="FE7736D36737FFE7FF76FE7AF23EFE76" blockId="16.[151,835,366,429]" box="[151,308,403,429]" pageId="16" pageNumber="393">
(
<figureCitation id="66F32A566737FFE7FF7EFE7AF226FE76" box="[159,300,403,429]" captionStart="FIGURE 13" captionStartId="16.[152,255,1698,1722]" captionTargetBox="[199,1387,570,1666]" captionTargetId="figure-116@16.[190,1397,561,1674]" captionTargetPageId="16" captionText="FIGURE 13. Zygoneura (Pharetratula) yangi Shah &amp; Huang sp. nov. A: head, frontal view; B: fourth antennal flagellomere; C: hypopygium, ventral view; D: tegmen and megasetae / spines of gonostylus." figureDoi="http://doi.org/10.5281/zenodo.13346931" httpUri="https://zenodo.org/record/13346931/files/figure.png" pageId="16" pageNumber="393">Figs 13A–D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B6D265586737FFE7FF76FE33F037FDC2" pageId="16" pageNumber="393" type="materials_examined">
<paragraph id="FE7736D36737FFE7FF76FE33F037FDC2" blockId="16.[151,1437,473,537]" pageId="16" pageNumber="393">
<emphasis id="CCBCEAC16737FFE7FF76FE33F289FE2F" bold="true" box="[151,387,474,500]" pageId="16" pageNumber="393">Material examined.</emphasis>
<materialsCitation id="4EA03C8E6737FFE7FE6DFE32F037FDC2" collectingDate="2019-06-01" collectionCode="ZAFUXZ" collectorName="Qicheng Yang" country="China" elevation="1049" latitude="29.331001" location="Renqingbenglukou" longLatPrecision="1" longitude="95.238" pageId="16" pageNumber="393" specimenCode="ZAFUXZ272" specimenCount="1" specimenCount-male="1" stateProvince="Xizang" typeStatus="holotype">
<typeStatus id="217388716737FFE7FE6DFE32F2F3FE2E" box="[396,505,475,501]" pageId="16" pageNumber="393" type="holotype">Holotype</typeStatus>
:
<specimenCount id="E8CEFD5A6737FFE7FDE7FE32F120FE2E" box="[518,554,475,501]" count="1" pageId="16" pageNumber="393" type="male">1♂</specimenCount>
,
<collectingCountry id="86DF76436737FFE7FDD8FE32F175FE2E" box="[569,639,475,501]" name="China" pageId="16" pageNumber="393">China</collectingCountry>
,
<collectingRegion id="3C0CF8316737FFE7FD6DFE32F046FE2E" box="[652,844,475,501]" country="China" name="Xizang" pageId="16" pageNumber="393">Xizang Province</collectingRegion>
,
<location id="FB1760086737FFE7FCBBFE32F738FE2E" LSID="urn:lsid:plazi:treatment:766187C56737FFE6FF76FE86F19DFDBA:FB1760086737FFE7FCBBFE32F738FE2E" box="[858,1074,475,501]" country="China" latitude="29.331001" longLatPrecision="1" longitude="95.238" name="Renqingbenglukou" pageId="16" pageNumber="393" stateProvince="Xizang">Renqingbenglukou</location>
,
<geoCoordinate id="9BFC50146737FFE7FBA1FE32F7EDFE2E" box="[1088,1255,473,501]" degrees="29" direction="north" minutes="19" orientation="latitude" pageId="16" pageNumber="393" precision="1" seconds="51.6" value="29.331001">29°19’51.6’’ N</geoCoordinate>
,
<geoCoordinate id="9BFC50146737FFE7FB15FE33F69DFE2E" box="[1268,1431,473,501]" degrees="95" direction="east" minutes="14" orientation="longitude" pageId="16" pageNumber="393" precision="1" seconds="16.8" value="95.238">95°14’16.8’’ E</geoCoordinate>
,
<elevation id="75E5D1E06737FFE7FF76FE16F223FDC2" box="[151,297,511,537]" metricMagnitude="3" metricUnit="m" metricValue="1.049" pageId="16" pageNumber="393" unit="m" value="1049.0">
<quantity id="39309B366737FFE7FF76FE16F3E6FDC2" box="[151,236,511,537]" metricMagnitude="3" metricUnit="m" metricValue="1.049" pageId="16" pageNumber="393" unit="m" value="1049.0">1049 m</quantity>
a.s.l.
</elevation>
,
<date id="8A7610136737FFE7FED4FE16F2A3FDC2" box="[309,425,511,537]" pageId="16" pageNumber="393" value="2019-06-01">
<collectingDate id="9A32E9FB6737FFE7FED4FE16F2A3FDC2" box="[309,425,511,537]" pageId="16" pageNumber="393" value="2019-06-01">1.VI.2019</collectingDate>
</date>
, leg.
<collectorName id="533D53056737FFE7FE04FE16F18EFDC2" box="[485,644,511,537]" pageId="16" pageNumber="393">Qicheng Yang</collectorName>
[
<specimenCode id="AE6E9EA86737FFE7FD70FE16F039FDC2" box="[657,819,511,537]" collectionCode="ZAFUXZ" pageId="16" pageNumber="393">ZAFUXZ272</specimenCode>
].
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="AAB7665B6737FFE7FF79F94BF0B6F905" ID-DOI="http://doi.org/10.5281/zenodo.13346931" ID-Zenodo-Dep="13346931" httpUri="https://zenodo.org/record/13346931/files/figure.png" pageId="16" pageNumber="393" startId="16.[152,255,1698,1722]" targetBox="[199,1387,570,1666]" targetPageId="16" targetType="figure">
<paragraph id="FE7736D36737FFE7FF79F94BF0B6F905" blockId="16.[151,1437,1698,1758]" pageId="16" pageNumber="393">
<emphasis id="CCBCEAC16737FFE7FF79F94BF22CF961" bold="true" box="[152,294,1698,1722]" pageId="16" pageNumber="393">FIGURE 13.</emphasis>
<taxonomicName id="39C84D506737FFE7FECDF94AF018F961" authority="Shah &amp; Huang" authorityName="Shah &amp; Shao &amp; Du &amp; Wang &amp; Huang" authorityYear="2024" box="[300,786,1698,1722]" class="Insecta" family="Sciaridae" genus="Zygoneura" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="393" phylum="Arthropoda" rank="species" species="yangi" status="sp. nov." subGenus="Pharetratula">
<emphasis id="CCBCEAC16737FFE7FECDF94AF290F961" box="[300,410,1699,1722]" italics="true" pageId="16" pageNumber="393">Zygoneura</emphasis>
(
<emphasis id="CCBCEAC16737FFE7FE48F94BF121F961" box="[425,555,1698,1722]" italics="true" pageId="16" pageNumber="393">Pharetratula</emphasis>
)
<emphasis id="CCBCEAC16737FFE7FDDBF94BF178F962" box="[570,626,1698,1721]" italics="true" pageId="16" pageNumber="393">yangi</emphasis>
Shah &amp; Huang
</taxonomicName>
<emphasis id="CCBCEAC16737FFE7FCF9F94BF063F961" bold="true" box="[792,873,1698,1722]" pageId="16" pageNumber="393">
<taxonomicNameLabel id="D78F57BA6737FFE7FCF9F94BF063F961" box="[792,873,1698,1722]" pageId="16" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A: head, frontal view; B: fourth antennal flagellomere; C: hypopygium, ventral view; D: tegmen and megasetae / spines of gonostylus.
</paragraph>
</caption>
<subSubSection id="B6D265586737FFE6FF26F8C1F2CAFEBE" lastPageId="17" lastPageNumber="394" pageId="16" pageNumber="393" type="description">
<paragraph id="FE7736D36737FFE6FF26F8C1F2CAFEBE" blockId="16.[151,1437,1832,2039]" lastBlockId="17.[151,1437,151,609]" lastPageId="17" lastPageNumber="394" pageId="16" pageNumber="393">
<emphasis id="CCBCEAC16737FFE7FF26F8C1F253F899" bold="true" box="[199,345,1832,1858]" pageId="16" pageNumber="393">Description.</emphasis>
Male. Head (
<figureCitation id="66F32A566737FFE7FE13F8C1F151F899" box="[498,603,1832,1858]" captionStart="FIGURE 13" captionStartId="16.[152,255,1698,1722]" captionTargetBox="[199,1387,570,1666]" captionTargetId="figure-116@16.[190,1397,561,1674]" captionTargetPageId="16" captionText="FIGURE 13. Zygoneura (Pharetratula) yangi Shah &amp; Huang sp. nov. A: head, frontal view; B: fourth antennal flagellomere; C: hypopygium, ventral view; D: tegmen and megasetae / spines of gonostylus." figureDoi="http://doi.org/10.5281/zenodo.13346931" httpUri="https://zenodo.org/record/13346931/files/figure.png" pageId="16" pageNumber="393">Fig. 13A</figureCitation>
). Head and antenna light brown, maxillary palpus light brown. Eye bridge broad, with 4–5 rows of facets. Face with 5–6 setae. Maxillary palpus with three segments, basal segment with four setae, with an indistinct sensory fossa; 2nd segment with five setae; 3rd segment long with six setae (
<figureCitation id="66F32A566737FFE7FA8EF899F3C1F875" captionStart="FIGURE 13" captionStartId="16.[152,255,1698,1722]" captionTargetBox="[199,1387,570,1666]" captionTargetId="figure-116@16.[190,1397,561,1674]" captionTargetPageId="16" captionText="FIGURE 13. Zygoneura (Pharetratula) yangi Shah &amp; Huang sp. nov. A: head, frontal view; B: fourth antennal flagellomere; C: hypopygium, ventral view; D: tegmen and megasetae / spines of gonostylus." figureDoi="http://doi.org/10.5281/zenodo.13346931" httpUri="https://zenodo.org/record/13346931/files/figure.png" pageId="16" pageNumber="393">Fig. 13A</figureCitation>
). Fourth antennal flagellomere much longer than wide, length/width = 2.65, body with mostly short setae and with a row of distinctly elongated setae near the neck (
<figureCitation id="66F32A566737FFE7FCF7F851F075F809" box="[790,895,1976,2002]" captionStart="FIGURE 13" captionStartId="16.[152,255,1698,1722]" captionTargetBox="[199,1387,570,1666]" captionTargetId="figure-116@16.[190,1397,561,1674]" captionTargetPageId="16" captionText="FIGURE 13. Zygoneura (Pharetratula) yangi Shah &amp; Huang sp. nov. A: head, frontal view; B: fourth antennal flagellomere; C: hypopygium, ventral view; D: tegmen and megasetae / spines of gonostylus." figureDoi="http://doi.org/10.5281/zenodo.13346931" httpUri="https://zenodo.org/record/13346931/files/figure.png" pageId="16" pageNumber="393">Fig. 13C</figureCitation>
). Wing with anterior veins slightly darker than posterior veins, membrane without macrotrichia. M1, M2, CuA1, CuA2, r-m with no setae. stM faintly visible in the specimen. Legs stramineous. Apex of fore tibia with a short comb-like row of setae. Tarsal claws three-toothed. Hypopygium (
<figureCitation id="66F32A566736FFE6FEDBFF52F2CFFF0E" box="[314,453,187,213]" captionStart="FIGURE 13" captionStartId="16.[152,255,1698,1722]" captionTargetBox="[199,1387,570,1666]" captionTargetId="figure-116@16.[190,1397,561,1674]" captionTargetPageId="16" captionText="FIGURE 13. Zygoneura (Pharetratula) yangi Shah &amp; Huang sp. nov. A: head, frontal view; B: fourth antennal flagellomere; C: hypopygium, ventral view; D: tegmen and megasetae / spines of gonostylus." figureDoi="http://doi.org/10.5281/zenodo.13346931" httpUri="https://zenodo.org/record/13346931/files/figure.png" pageId="17" pageNumber="394">Figs 13C–D</figureCitation>
) yellowish brown. The gonocoxae almost as long as gonostyli, outer and inner margins sparsely setose, ventral medial margin with two groups of weak setae. The gonostylus (
<figureCitation id="66F32A566736FFE6FB64FF36F618FF22" box="[1157,1298,223,249]" captionStart="FIGURE 13" captionStartId="16.[152,255,1698,1722]" captionTargetBox="[199,1387,570,1666]" captionTargetId="figure-116@16.[190,1397,561,1674]" captionTargetPageId="16" captionText="FIGURE 13. Zygoneura (Pharetratula) yangi Shah &amp; Huang sp. nov. A: head, frontal view; B: fourth antennal flagellomere; C: hypopygium, ventral view; D: tegmen and megasetae / spines of gonostylus." figureDoi="http://doi.org/10.5281/zenodo.13346931" httpUri="https://zenodo.org/record/13346931/files/figure.png" pageId="17" pageNumber="394">Figs 13C–D</figureCitation>
) stout, with two megasetae of almost equal length; one on dorsal side at the apical third and the other on ventral side near the middle, apex with dense short setosity. Tegmen membranous, wider than high, apically rounded. Aedeagus dark brown, slender (
<figureCitation id="66F32A566736FFE6FEACFEA2F2B9FEBE" box="[333,435,331,357]" captionStart="FIGURE 13" captionStartId="16.[152,255,1698,1722]" captionTargetBox="[199,1387,570,1666]" captionTargetId="figure-116@16.[190,1397,561,1674]" captionTargetPageId="16" captionText="FIGURE 13. Zygoneura (Pharetratula) yangi Shah &amp; Huang sp. nov. A: head, frontal view; B: fourth antennal flagellomere; C: hypopygium, ventral view; D: tegmen and megasetae / spines of gonostylus." figureDoi="http://doi.org/10.5281/zenodo.13346931" httpUri="https://zenodo.org/record/13346931/files/figure.png" pageId="17" pageNumber="394">Fig. 13C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B6D265586736FFE6FF26FE86F607FE52" box="[199,1293,367,393]" pageId="17" pageNumber="394" type="etymology">
<paragraph id="FE7736D36736FFE6FF26FE86F607FE52" blockId="17.[151,1437,151,609]" box="[199,1293,367,393]" pageId="17" pageNumber="394">
<emphasis id="CCBCEAC16736FFE6FF26FE86F244FE52" bold="true" box="[199,334,367,393]" pageId="17" pageNumber="394">Etymology.</emphasis>
The species epithet
<taxonomicName id="39C84D506736FFE6FDD3FE86F170FE53" authorityName="Shah &amp; Shao &amp; Du &amp; Wang &amp; Huang" authorityYear="2024" box="[562,634,367,393]" class="Insecta" family="Sciaridae" genus="Zygoneura" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="394" phylum="Arthropoda" rank="species" species="yangi">
‘
<emphasis id="CCBCEAC16736FFE6FDDDFE86F170FE53" box="[572,634,367,392]" italics="true" pageId="17" pageNumber="394">yangi</emphasis>
</taxonomicName>
’ refers to Qicheng Yang, who collected the
<typeStatus id="217388716736FFE6FB84FE86F79FFE52" box="[1125,1173,367,393]" pageId="17" pageNumber="394">type</typeStatus>
specimen.
</paragraph>
</subSubSection>
<subSubSection id="B6D265586736FFE6FF26FE7AF181FDE6" pageId="17" pageNumber="394" type="discussion">
<paragraph id="FE7736D36736FFE6FF26FE7AF181FDE6" blockId="17.[151,1437,151,609]" pageId="17" pageNumber="394">
<emphasis id="CCBCEAC16736FFE6FF26FE7AF231FE76" bold="true" box="[199,315,403,429]" pageId="17" pageNumber="394">Remarks.</emphasis>
The new species can easily be distinguished from other closely related species of the subgenus by the typical gonostylus with two megasetae of almost equal length; one on the dorsal side at the apical third and the other on the ventral side near the middle (
<figureCitation id="66F32A566736FFE6FD87FE32F1FEFE2E" box="[614,756,475,501]" captionStart="FIGURE 13" captionStartId="16.[152,255,1698,1722]" captionTargetBox="[199,1387,570,1666]" captionTargetId="figure-116@16.[190,1397,561,1674]" captionTargetPageId="16" captionText="FIGURE 13. Zygoneura (Pharetratula) yangi Shah &amp; Huang sp. nov. A: head, frontal view; B: fourth antennal flagellomere; C: hypopygium, ventral view; D: tegmen and megasetae / spines of gonostylus." figureDoi="http://doi.org/10.5281/zenodo.13346931" httpUri="https://zenodo.org/record/13346931/files/figure.png" pageId="17" pageNumber="394">Figs 13C–D</figureCitation>
). In addition, the
<typeStatus id="217388716736FFE6FC57FE32F712FE2E" box="[950,1048,475,501]" pageId="17" pageNumber="394" type="holotype">holotype</typeStatus>
of this species is in poor condition (especially wings and legs), and all possible diagnostic characters were carefully examined, and their images were added to the figure plate for visual evidence.
</paragraph>
</subSubSection>
<subSubSection id="B6D265586736FFE6FF26FDAFF19DFDBA" box="[199,663,582,609]" pageId="17" pageNumber="394" type="distribution">
<paragraph id="FE7736D36736FFE6FF26FDAFF19DFDBA" blockId="17.[151,1437,151,609]" box="[199,663,582,609]" pageId="17" pageNumber="394">
<emphasis id="CCBCEAC16736FFE6FF26FDAFF26BFDBB" bold="true" box="[199,353,582,608]" pageId="17" pageNumber="394">Distribution.</emphasis>
<collectingCountry id="86DF76436736FFE6FE86FDAEF2A4FDBA" box="[359,430,583,609]" name="China" pageId="17" pageNumber="394">China</collectingCountry>
:
<collectingRegion id="3C0CF8316736FFE6FE58FDAEF100FDBA" box="[441,522,583,609]" country="China" name="Xizang" pageId="17" pageNumber="394">Xizang</collectingRegion>
(this study).
</paragraph>
</subSubSection>
</treatment>
</document>
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