259 lines
37 KiB
XML
259 lines
37 KiB
XML
<document id="8CA7120193D7B1C8D2D1CDF72469FE4B" ID-CLB-Dataset="24204" ID-DOI="10.11646/zootaxa.4730.1.1" ID-GBIF-Dataset="bcc0e0fc-b6a5-4d5a-8008-7b714c106a7c" ID-ISSN="1175-5326" ID-Zenodo-Dep="3634270" ID-ZooBank="7F4CD129-9FF9-4593-A8A4-DB999B3E402F" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1580713876401" checkinUser="plazi" docAuthor="Blake, James A." docDate="2020" docId="757F0D3E9677230BD8C0FF35FE68F8BE" docLanguage="en" docName="zootaxa.4730.1.1.pdf" docOrigin="Zootaxa 4730 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Leitoscoloplos gordaensis Blake 2020, new species" docType="treatment" docUuid="03E8D6B6-DE48-42A5-9087-703A3E571C24" docUuidSource="ZooBank" docVersion="8" lastPageNumber="10" masterDocId="89467546967E2302D857FFACFFABFF92" masterDocTitle="New species and records of deep-water Orbiniidae (Annelida, Polychaeta) from the Eastern Pacific continental slope, abyssal Pacific Ocean, and the South China Sea" masterLastPageNumber="61" masterPageNumber="1" pageNumber="10" updateTime="1698780308397" updateUser="ExternalLinkService">
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<mods:title id="30D02229A138FD18320D184326C78CBA">New species and records of deep-water Orbiniidae (Annelida, Polychaeta) from the Eastern Pacific continental slope, abyssal Pacific Ocean, and the South China Sea</mods:title>
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<mods:roleTerm id="9171BBBC1EDDF2E1D32ADDF8D8210E2A">Author</mods:roleTerm>
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<mods:namePart id="FFACA6124775BD9D619650DE78F7C7E3">Blake, James A.</mods:namePart>
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<mods:typeOfResource id="AA1CF0D73F050648852CE54335FDA179">text</mods:typeOfResource>
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<mods:title id="C7F75D921E680E82C7DE020C32EC0BEC">Zootaxa</mods:title>
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<mods:part id="CE2C84E0D941798AD4545AF4B3F432A9">
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<mods:date id="60C69D9EDC1749F6BC02E343890E813D">2020</mods:date>
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<mods:number id="DBF2B930136C449C4D2397C12DD8BCF3">2020-02-03</mods:number>
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<mods:number id="89D862CEA0C53071A72A408ACF895D04">4730</mods:number>
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<mods:identifier id="ED2FE6DCCAA31D4B1E1DC3AAEB2B4FF9" type="DOI">10.11646/zootaxa.4730.1.1</mods:identifier>
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<mods:identifier id="9D0139FFE25AC8D51F6781CD19873A74" type="ISSN">1175-5326</mods:identifier>
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<treatment id="757F0D3E9677230BD8C0FF35FE68F8BE" ID-GBIF-Taxon="161938992" LSID="urn:lsid:zoobank.org:act:03E8D6B6-DE48-42A5-9087-703A3E571C24" httpUri="http://treatment.plazi.org/id/757F0D3E9677230BD8C0FF35FE68F8BE" lastPageNumber="10" pageId="9" pageNumber="10">
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<subSubSection id="B5CCEFA39677230BD8C0FF35FDCEFF21" box="[151,613,153,180]" pageId="9" pageNumber="10" type="nomenclature">
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<paragraph id="FD69BC289677230BD8C0FF35FDCEFF21" blockId="9.[151,960,153,252]" box="[151,613,153,180]" pageId="9" pageNumber="10">
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<heading id="A6210B449677230BD8C0FF35FDCEFF21" bold="true" box="[151,613,153,180]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
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<emphasis id="CFA2603A9677230BD8C0FF35FDCEFF21" bold="true" box="[151,613,153,180]" pageId="9" pageNumber="10">
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<taxonomicName id="3AD6C7AB9677230BD8C0FF35FE66FF26" ID-CoL="85RHC" authority="Blake, 2020" authorityName="Blake" authorityYear="2020" box="[151,461,153,180]" class="Polychaeta" family="Orbiniidae" genus="Leitoscoloplos" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="gordaensis" status="sp. nov.">
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<emphasis id="CFA2603A9677230BD8C0FF35FE66FF26" bold="true" box="[151,461,153,180]" italics="true" pageId="9" pageNumber="10">Leitoscoloplos gordaensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D491DD419677230BD982FF36FDCEFF21" box="[469,613,153,180]" pageId="9" pageNumber="10" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="B5CCEFA39677230BD8C0FF11FC6BFF6E" pageId="9" pageNumber="10" type="description">
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<paragraph id="FD69BC289677230BD8C0FF11FEB7FF45" blockId="9.[151,960,153,252]" box="[151,284,189,215]" pageId="9" pageNumber="10">
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<figureCitation id="65EDA0AD9677230BD8C0FF11FEB7FF45" box="[151,284,189,215]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="10.[151,250,1945,1970]" captionStartId-1="11.[151,249,1420,1445]" captionTargetBox-0="[202,1375,189,1913]" captionTargetBox-1="[152,1435,182,1394]" captionTargetId-0="figure@10.[196,1391,181,1920]" captionTargetId-1="figure@11.[151,1436,181,1394]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURE 3. Leitoscoloplos gordaensis n. sp. Holotype (MCZ 153564): A, anterior end, dorsal view. Paratype (MCZ 153565): B, posterior end, dorsal view; C, thoracic setiger, anterior view; D, anterior abdominal setiger, posterior view; E, posterior abdominal setiger, posterior view." captionText-1="FIGURE 4. Leitoscoloplos gordaensis n. sp. Holotype (MCZ 153564):A, anterior end, left lateral view; B, posterior thoracic and anterior abdominal setigers, left lateral view, setigers numbered; C, middle abdominal setigers, left lateral view, arrows denote tri-annular rings on venter. Paratype (MCZ 153565): D, posterior abdominal setiger, anterior view. Stained with Shirlastain A." figureDoi-0="http://doi.org/10.5281/zenodo.3634276" figureDoi-1="http://doi.org/10.5281/zenodo.3634278" httpUri-0="https://zenodo.org/record/3634276/files/figure.png" httpUri-1="https://zenodo.org/record/3634278/files/figure.png" pageId="9" pageNumber="10">Figures 3–4</figureCitation>
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</paragraph>
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<paragraph id="FD69BC289677230BD8C0FF4DFC6BFF6E" blockId="9.[151,960,153,252]" box="[151,960,225,252]" pageId="9" pageNumber="10">
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<uri id="8947B02A9677230BD8C0FF4DFC6BFF6E" box="[151,960,225,252]" pageId="9" pageNumber="10">
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urn:lsid:zoobank.org:act:
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<uuid id="897086FD9677230BD9FAFF4DFC6BFF6E" box="[429,960,225,252]" pageId="9" pageNumber="10">03E8D6B6-DE48-42A5-9087-703A3E571C24</uuid>
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</uri>
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</subSubSection>
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<subSubSection id="B5CCEFA39677230BD8C0FE85FC38FE1E" pageId="9" pageNumber="10" type="materials_examined">
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<paragraph id="FD69BC289677230BD8C0FE85FC38FE1E" blockId="9.[151,1437,297,1836]" pageId="9" pageNumber="10">
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<emphasis id="CFA2603A9677230BD8C0FE85FED6FED6" bold="true" box="[151,381,297,324]" pageId="9" pageNumber="10">Material examined</emphasis>
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.
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<emphasis id="CFA2603A9677230BD9D8FE85FD7EFED6" bold="true" box="[399,725,297,324]" pageId="9" pageNumber="10">Gorda Ridge, off northern</emphasis>
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<materialsCitation id="4DBEB6759677230BDAB7FE85FBDEFEFA" ID-GBIF-Occurrence="2562076929" collectingDate="1988-05-12" collectionCode="MCZ" collectorName="J. F. Grassle" elevation="3271" latitude="41.006668" location="California" longLatPrecision="115" longitude="-127.488335" pageId="9" pageNumber="10" specimenCode="MCZ 153564" specimenCount="1" stateProvince="California" typeStatus="holotype">
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<emphasis id="CFA2603A9677230BDAB7FE85FBE9FED6" bold="true" box="[736,1090,297,324]" pageId="9" pageNumber="10">
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<collectingRegion id="3F1272CA9677230BDAB7FE85FCF6FED6" box="[736,861,297,324]" country="United States of America" name="California" pageId="9" pageNumber="10">California</collectingRegion>
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, Escanaba Trough
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</emphasis>
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, HOV
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<emphasis id="CFA2603A9677230BDCCCFE85FB7DFED1" box="[1179,1238,297,323]" italics="true" pageId="9" pageNumber="10">Alvin</emphasis>
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||
Dive 2042, coll.
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<collectorName id="5023D9FE9677230BD8C0FEE2FEB5FEF5" box="[151,286,333,360]" pageId="9" pageNumber="10">J.F. Grassle</collectorName>
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,
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<date id="89689AE89677230BD97BFEE1FE62FEF5" box="[300,457,333,360]" pageId="9" pageNumber="10" value="1988-05-12">
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<collectingDate id="992C63009677230BD97BFEE1FE62FEF5" box="[300,457,333,360]" pageId="9" pageNumber="10" value="1988-05-12">12 May 1988</collectingDate>
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</date>
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,
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<geoCoordinate id="98E2DAEF9677230BD980FEE1FDE6FEFA" box="[471,589,333,360]" degrees="41" direction="north" minutes="00.4" orientation="latitude" pageId="9" pageNumber="10" precision="92" value="41.006668">41°00.4′N</geoCoordinate>
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,
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||
<geoCoordinate id="98E2DAEF9677230BDA0DFEE1FD4AFEFA" box="[602,737,333,360]" degrees="127" direction="west" minutes="29.3" orientation="longitude" pageId="9" pageNumber="10" precision="92" value="-127.488335">127°29.3′W</geoCoordinate>
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,
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||
<quantity id="3A2E11CD9677230BDAB8FEE1FCE2FEFA" box="[751,841,333,360]" metricMagnitude="3" metricUnit="m" metricValue="3.271" pageId="9" pageNumber="10" unit="m" value="3271.0">
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||
<elevation id="76FB5B1B9677230BDAB8FEE1FCE2FEFA" box="[751,841,333,360]" metricMagnitude="3" metricUnit="m" metricValue="3.271" pageId="9" pageNumber="10" unit="m" value="3271.0">3271 m</elevation>
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</quantity>
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,
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<emphasis id="CFA2603A9677230BDB00FEE2FC15FEFA" bold="true" box="[855,958,334,360]" pageId="9" pageNumber="10">
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||
<typeStatus id="226D028A9677230BDB00FEE2FC15FEFA" box="[855,958,334,360]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
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||
</emphasis>
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||
(
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||
<specimenCode id="AD7014539677230BDB98FEE1FBC6FEFA" box="[975,1133,333,360]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="9" pageNumber="10" type="Museum">MCZ 153564</specimenCode>
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||
)
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||
</materialsCitation>
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||
<materialsCitation id="4DBEB6759677230BDC29FEE1FF53FE1E" ID-GBIF-Occurrence="2562076943" collectionCode="MCZ" pageId="9" pageNumber="10" specimenCode="MCZ 153565" specimenCount="30" stateProvince="California" typeStatus="paratype">
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||
and
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||
<specimenCount id="EBD077A19677230BDCE6FEE1FAE7FEFA" box="[1201,1356,333,360]" pageId="9" pageNumber="10" type="generic" typeStatus="paratypes">
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||
30
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<emphasis id="CFA2603A9677230BDC81FEE2FAE7FEFA" bold="true" box="[1238,1356,334,360]" pageId="9" pageNumber="10">paratypes</emphasis>
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||
</specimenCount>
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||
(
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<specimenCode id="AD7014539677230BDD09FEE1FF5BFE1E" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="9" pageNumber="10" type="Museum">MCZ 153565</specimenCode>
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||
)
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</materialsCitation>
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||
;
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<materialsCitation id="4DBEB6759677230BD954FEDEFC24FE1E" ID-GBIF-Occurrence="2562076922" box="[259,911,369,396]" collectionCode="MCZ" pageId="9" pageNumber="10" specimenCode="MCZ 153566" specimenCount="2" stateProvince="California" typeStatus="paratype">
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same station, clam washings,
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<specimenCount id="EBD077A19677230BDA07FEDDFD71FE19" box="[592,730,369,395]" pageId="9" pageNumber="10" type="generic">2 specimens</specimenCount>
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||
(
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<specimenCode id="AD7014539677230BDABFFEDDFC2CFE1E" box="[744,903,369,396]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="9" pageNumber="10" type="Museum">MCZ 153566</specimenCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="B5CCEFA39677230BD890FE39FC96FA51" pageId="9" pageNumber="10" type="description">
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<paragraph id="FD69BC289677230BD890FE39FAADFD61" blockId="9.[151,1437,297,1836]" pageId="9" pageNumber="10">
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<emphasis id="CFA2603A9677230BD890FE39FEF2FE22" bold="true" box="[199,345,405,432]" pageId="9" pageNumber="10">Description.</emphasis>
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A moderate-sized species,
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<typeStatus id="226D028A9677230BDAD2FE39FD4CFE3D" box="[645,743,405,431]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
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incomplete,
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<quantity id="3A2E11CD9677230BDB20FE39FC6EFE22" box="[887,965,405,432]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="10" unit="mm" value="23.0">23 mm</quantity>
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||
long and
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||
<quantity id="3A2E11CD9677230BDC65FE39FBD9FE22" box="[1074,1138,405,432]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="10" unit="mm" value="1.0">1 mm</quantity>
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||
wide for 63 setigers; some
|
||
<typeStatus id="226D028A9677230BD8C0FE16FEA9FE46" box="[151,258,442,468]" pageId="9" pageNumber="10" type="paratype">paratypes</typeStatus>
|
||
complete, up to
|
||
<quantity id="3A2E11CD9677230BD9EBFE15FDA1FE46" box="[444,522,441,468]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="10" unit="mm" value="20.0">20 mm</quantity>
|
||
long and
|
||
<quantity id="3A2E11CD9677230BDA2FFE15FD66FE46" box="[632,717,441,468]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="10" unit="mm" value="1.7">1.7 mm</quantity>
|
||
wide for approximately 90 setigers. Body elongate, with narrow crowded segments throughout (
|
||
<figureCitation id="65EDA0AD9677230BD9ABFE71FDE2FE65" box="[508,585,477,503]" captionStart="FIGURE 3" captionStartId="10.[151,250,1945,1970]" captionTargetBox="[202,1375,189,1913]" captionTargetId="figure@10.[196,1391,181,1920]" captionTargetPageId="10" captionText="FIGURE 3. Leitoscoloplos gordaensis n. sp. Holotype (MCZ 153564): A, anterior end, dorsal view. Paratype (MCZ 153565): B, posterior end, dorsal view; C, thoracic setiger, anterior view; D, anterior abdominal setiger, posterior view; E, posterior abdominal setiger, posterior view." figureDoi="http://doi.org/10.5281/zenodo.3634276" httpUri="https://zenodo.org/record/3634276/files/figure.png" pageId="9" pageNumber="10">Figs. 3</figureCitation>
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||
A–B, 4A–C); thoracic and 1–2 transitional segments about six times wider than long; middle and posterior segments narrower, more crowded. Transition from thoracic to abdominal segments marked by development of three annular rings ventral to neuropodia, consisting initially of a broad medial ring joined on subsequent segments by two thin lateral intersegmental rings (
|
||
<figureCitation id="65EDA0AD9677230BDB97FDE5FBA8FDF1" box="[960,1027,585,611]" captionStart="FIGURE 4" captionStartId="11.[151,249,1420,1445]" captionTargetBox="[152,1435,182,1394]" captionTargetId="figure@11.[151,1436,181,1394]" captionTargetPageId="11" captionText="FIGURE 4. Leitoscoloplos gordaensis n. sp. Holotype (MCZ 153564):A, anterior end, left lateral view; B, posterior thoracic and anterior abdominal setigers, left lateral view, setigers numbered; C, middle abdominal setigers, left lateral view, arrows denote tri-annular rings on venter. Paratype (MCZ 153565): D, posterior abdominal setiger, anterior view. Stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.3634278" httpUri="https://zenodo.org/record/3634278/files/figure.png" pageId="9" pageNumber="10">Fig. 4</figureCitation>
|
||
B–C), thus providing ventral surface of each abdominal segment with three rings and an intersegmental groove between each segment. These rings most prominent in middle body segments (
|
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<figureCitation id="65EDA0AD9677230BDA13FD3DFD35FD39" box="[580,670,657,683]" captionStart="FIGURE 4" captionStartId="11.[151,249,1420,1445]" captionTargetBox="[152,1435,182,1394]" captionTargetId="figure@11.[151,1436,181,1394]" captionTargetPageId="11" captionText="FIGURE 4. Leitoscoloplos gordaensis n. sp. Holotype (MCZ 153564):A, anterior end, left lateral view; B, posterior thoracic and anterior abdominal setigers, left lateral view, setigers numbered; C, middle abdominal setigers, left lateral view, arrows denote tri-annular rings on venter. Paratype (MCZ 153565): D, posterior abdominal setiger, anterior view. Stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.3634278" httpUri="https://zenodo.org/record/3634278/files/figure.png" pageId="9" pageNumber="10">Fig. 4C</figureCitation>
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), becoming shallower and less evident in posterior segments. Both dorsal and ventral surfaces smooth, lacking longitudinal grooves or ridges. Some
|
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<typeStatus id="226D028A9677230BDC68FD1AFB01FD42" box="[1087,1194,694,720]" pageId="9" pageNumber="10" type="paratype">paratypes</typeStatus>
|
||
mature females with large irregularly shaped eggs, ranging from
|
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<quantity id="3A2E11CD9677230BDAD4FD75FCB9FD66" box="[643,786,729,756]" metricMagnitude="-1" metricUnit="m" metricValue="3.2" metricValueMax="3.5" metricValueMin="2.9" pageId="9" pageNumber="10" unit="mm" value="320.0" valueMax="350.0" valueMin="290.0">290–350 µm</quantity>
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in diameter. Color in alcohol: opaque white.
|
||
</paragraph>
|
||
<paragraph id="FD69BC289677230BD890FD51FBC4FC35" blockId="9.[151,1437,297,1836]" pageId="9" pageNumber="10">
|
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Prostomium triangular, short (
|
||
<figureCitation id="65EDA0AD9677230BDA4EFD51FDDDFC85" box="[537,630,765,791]" captionStart="FIGURE 3" captionStartId="10.[151,250,1945,1970]" captionTargetBox="[202,1375,189,1913]" captionTargetId="figure@10.[196,1391,181,1920]" captionTargetPageId="10" captionText="FIGURE 3. Leitoscoloplos gordaensis n. sp. Holotype (MCZ 153564): A, anterior end, dorsal view. Paratype (MCZ 153565): B, posterior end, dorsal view; C, thoracic setiger, anterior view; D, anterior abdominal setiger, posterior view; E, posterior abdominal setiger, posterior view." figureDoi="http://doi.org/10.5281/zenodo.3634276" httpUri="https://zenodo.org/record/3634276/files/figure.png" pageId="9" pageNumber="10">Figs 3A</figureCitation>
|
||
,
|
||
<figureCitation id="65EDA0AD9677230BDAD4FD51FD02FC85" box="[643,681,765,791]" captionStart="FIGURE 4" captionStartId="11.[151,249,1420,1445]" captionTargetBox="[152,1435,182,1394]" captionTargetId="figure@11.[151,1436,181,1394]" captionTargetPageId="11" captionText="FIGURE 4. Leitoscoloplos gordaensis n. sp. Holotype (MCZ 153564):A, anterior end, left lateral view; B, posterior thoracic and anterior abdominal setigers, left lateral view, setigers numbered; C, middle abdominal setigers, left lateral view, arrows denote tri-annular rings on venter. Paratype (MCZ 153565): D, posterior abdominal setiger, anterior view. Stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.3634278" httpUri="https://zenodo.org/record/3634278/files/figure.png" pageId="9" pageNumber="10">4A</figureCitation>
|
||
), with nuchal organs elongate slits on posterior lateral margin (
|
||
<figureCitation id="65EDA0AD9677230BDD38FD51FF16FCA9" captionStart="FIGURE 4" captionStartId="11.[151,249,1420,1445]" captionTargetBox="[152,1435,182,1394]" captionTargetId="figure@11.[151,1436,181,1394]" captionTargetPageId="11" captionText="FIGURE 4. Leitoscoloplos gordaensis n. sp. Holotype (MCZ 153564):A, anterior end, left lateral view; B, posterior thoracic and anterior abdominal setigers, left lateral view, setigers numbered; C, middle abdominal setigers, left lateral view, arrows denote tri-annular rings on venter. Paratype (MCZ 153565): D, posterior abdominal setiger, anterior view. Stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.3634278" httpUri="https://zenodo.org/record/3634278/files/figure.png" pageId="9" pageNumber="10">Fig. 4A</figureCitation>
|
||
), eyespots absent. Peristomium with one or two annular rings, first longer and narrower than second when two present (
|
||
<figureCitation id="65EDA0AD9677230BD8A0FCE9FEF0FCCD" box="[247,347,837,863]" captionStart="FIGURE 3" captionStartId="10.[151,250,1945,1970]" captionTargetBox="[202,1375,189,1913]" captionTargetId="figure@10.[196,1391,181,1920]" captionTargetPageId="10" captionText="FIGURE 3. Leitoscoloplos gordaensis n. sp. Holotype (MCZ 153564): A, anterior end, dorsal view. Paratype (MCZ 153565): B, posterior end, dorsal view; C, thoracic setiger, anterior view; D, anterior abdominal setiger, posterior view; E, posterior abdominal setiger, posterior view." figureDoi="http://doi.org/10.5281/zenodo.3634276" httpUri="https://zenodo.org/record/3634276/files/figure.png" pageId="9" pageNumber="10">Figs. 3A</figureCitation>
|
||
,
|
||
<figureCitation id="65EDA0AD9677230BD930FCE9FE26FCCD" box="[359,397,837,863]" captionStart="FIGURE 4" captionStartId="11.[151,249,1420,1445]" captionTargetBox="[152,1435,182,1394]" captionTargetId="figure@11.[151,1436,181,1394]" captionTargetPageId="11" captionText="FIGURE 4. Leitoscoloplos gordaensis n. sp. Holotype (MCZ 153564):A, anterior end, left lateral view; B, posterior thoracic and anterior abdominal setigers, left lateral view, setigers numbered; C, middle abdominal setigers, left lateral view, arrows denote tri-annular rings on venter. Paratype (MCZ 153565): D, posterior abdominal setiger, anterior view. Stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.3634278" httpUri="https://zenodo.org/record/3634278/files/figure.png" pageId="9" pageNumber="10">4A</figureCitation>
|
||
); presence of one or two peristomial rings not size related. Thorax with ten setigers followed by 1–2 transitional setigers (
|
||
<figureCitation id="65EDA0AD9677230BD987FCC5FD8CFC11" box="[464,551,873,899]" captionStart="FIGURE 4" captionStartId="11.[151,249,1420,1445]" captionTargetBox="[152,1435,182,1394]" captionTargetId="figure@11.[151,1436,181,1394]" captionTargetPageId="11" captionText="FIGURE 4. Leitoscoloplos gordaensis n. sp. Holotype (MCZ 153564):A, anterior end, left lateral view; B, posterior thoracic and anterior abdominal setigers, left lateral view, setigers numbered; C, middle abdominal setigers, left lateral view, arrows denote tri-annular rings on venter. Paratype (MCZ 153565): D, posterior abdominal setiger, anterior view. Stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.3634278" httpUri="https://zenodo.org/record/3634278/files/figure.png" pageId="9" pageNumber="10">Fig. 4B</figureCitation>
|
||
); boundary between last thoracic setiger and first abdominal demarcated by loss of most neurosetae and increase in size of neuropodial lobe on setigers 11–12 (
|
||
<figureCitation id="65EDA0AD9677230BDC5EFC21FBC9FC35" box="[1033,1122,909,935]" captionStart="FIGURE 4" captionStartId="11.[151,249,1420,1445]" captionTargetBox="[152,1435,182,1394]" captionTargetId="figure@11.[151,1436,181,1394]" captionTargetPageId="11" captionText="FIGURE 4. Leitoscoloplos gordaensis n. sp. Holotype (MCZ 153564):A, anterior end, left lateral view; B, posterior thoracic and anterior abdominal setigers, left lateral view, setigers numbered; C, middle abdominal setigers, left lateral view, arrows denote tri-annular rings on venter. Paratype (MCZ 153565): D, posterior abdominal setiger, anterior view. Stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.3634278" httpUri="https://zenodo.org/record/3634278/files/figure.png" pageId="9" pageNumber="10">Fig. 4B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FD69BC289677230BD890FC1DFBCBFBC9" blockId="9.[151,1437,297,1836]" pageId="9" pageNumber="10">
|
||
Thoracic notopodial and neuropodial lobes elongate with triangular base, both similar in form (
|
||
<figureCitation id="65EDA0AD9677230BDD5FFC1DFAC9FC59" box="[1288,1378,945,971]" captionStart="FIGURE 3" captionStartId="10.[151,250,1945,1970]" captionTargetBox="[202,1375,189,1913]" captionTargetId="figure@10.[196,1391,181,1920]" captionTargetPageId="10" captionText="FIGURE 3. Leitoscoloplos gordaensis n. sp. Holotype (MCZ 153564): A, anterior end, dorsal view. Paratype (MCZ 153565): B, posterior end, dorsal view; C, thoracic setiger, anterior view; D, anterior abdominal setiger, posterior view; E, posterior abdominal setiger, posterior view." figureDoi="http://doi.org/10.5281/zenodo.3634276" httpUri="https://zenodo.org/record/3634276/files/figure.png" pageId="9" pageNumber="10">Fig. 3C</figureCitation>
|
||
); abdominal notopodia becoming narrower, more elongate, shifting dorsally in position (
|
||
<figureCitation id="65EDA0AD9677230BDC0EFC79FB0EFC7D" box="[1113,1189,981,1007]" captionStart="FIGURE 3" captionStartId="10.[151,250,1945,1970]" captionTargetBox="[202,1375,189,1913]" captionTargetId="figure@10.[196,1391,181,1920]" captionTargetPageId="10" captionText="FIGURE 3. Leitoscoloplos gordaensis n. sp. Holotype (MCZ 153564): A, anterior end, dorsal view. Paratype (MCZ 153565): B, posterior end, dorsal view; C, thoracic setiger, anterior view; D, anterior abdominal setiger, posterior view; E, posterior abdominal setiger, posterior view." figureDoi="http://doi.org/10.5281/zenodo.3634276" httpUri="https://zenodo.org/record/3634276/files/figure.png" pageId="9" pageNumber="10">Figs. 3</figureCitation>
|
||
D–E, 4D); abdominal neuropodia thickened, with two terminal lobes (
|
||
<figureCitation id="65EDA0AD9677230BDAE7FC55FD57FB81" box="[688,764,1017,1043]" captionStart="FIGURE 3" captionStartId="10.[151,250,1945,1970]" captionTargetBox="[202,1375,189,1913]" captionTargetId="figure@10.[196,1391,181,1920]" captionTargetPageId="10" captionText="FIGURE 3. Leitoscoloplos gordaensis n. sp. Holotype (MCZ 153564): A, anterior end, dorsal view. Paratype (MCZ 153565): B, posterior end, dorsal view; C, thoracic setiger, anterior view; D, anterior abdominal setiger, posterior view; E, posterior abdominal setiger, posterior view." figureDoi="http://doi.org/10.5281/zenodo.3634276" httpUri="https://zenodo.org/record/3634276/files/figure.png" pageId="9" pageNumber="10">Figs. 3</figureCitation>
|
||
D–E, 4D); subpodial flanges and papillae absent. Branchiae first present from setiger 14–16 as rudiments; gradually increasing in size over middle abdominal segments, becoming broad and flattened, about one-third longer than notopodial lobe (
|
||
<figureCitation id="65EDA0AD9677230BDBC8FBEDFC41FBC9" box="[927,1002,1089,1115]" captionStart="FIGURE 3" captionStartId="10.[151,250,1945,1970]" captionTargetBox="[202,1375,189,1913]" captionTargetId="figure@10.[196,1391,181,1920]" captionTargetPageId="10" captionText="FIGURE 3. Leitoscoloplos gordaensis n. sp. Holotype (MCZ 153564): A, anterior end, dorsal view. Paratype (MCZ 153565): B, posterior end, dorsal view; C, thoracic setiger, anterior view; D, anterior abdominal setiger, posterior view; E, posterior abdominal setiger, posterior view." figureDoi="http://doi.org/10.5281/zenodo.3634276" httpUri="https://zenodo.org/record/3634276/files/figure.png" pageId="9" pageNumber="10">Figs. 3</figureCitation>
|
||
D–E, 4D).
|
||
</paragraph>
|
||
<paragraph id="FD69BC289677230BD890FBC9FE75FAA1" blockId="9.[151,1437,297,1836]" pageId="9" pageNumber="10">
|
||
All notosetae camerated capillaries; thoracic notosetae arranged in a tight fascicle with capillaries appearing to arise from a spiral, numbering about 30–40 per fascicle; abdominal capillaries numbering 15–20 per notopodium in anterior and middle setigers, reduced to
|
||
<quantity id="3A2E11CD9677230BDA21FB01FD72FB55" box="[630,729,1197,1223]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" metricValueMax="4.572" metricValueMin="2.54" pageId="9" pageNumber="10" unit="in" value="14.0" valueMax="18.0" valueMin="10.0">10–18 in</quantity>
|
||
posterior setigers. Thoracic neurosetae all capillaries, with one prominent anterior row and 2–3 poorly organized secondary rows, numbering about 25–30 fascicle; abdominal neurosetae include 2–5 capillaries and 1–3 narrow aciculae with tips emergent, each short, smooth with rounded tip; furcate and flail setae absent.
|
||
</paragraph>
|
||
<paragraph id="FD69BC289677230BD890FA91FC68FAC5" blockId="9.[151,1437,297,1836]" box="[199,963,1341,1368]" pageId="9" pageNumber="10">
|
||
Pygidium bluntly rounded, bearing two thin elongate cirri (
|
||
<figureCitation id="65EDA0AD9677230BDB09FA91FC1EFAC5" box="[862,949,1341,1367]" captionStart="FIGURE 3" captionStartId="10.[151,250,1945,1970]" captionTargetBox="[202,1375,189,1913]" captionTargetId="figure@10.[196,1391,181,1920]" captionTargetPageId="10" captionText="FIGURE 3. Leitoscoloplos gordaensis n. sp. Holotype (MCZ 153564): A, anterior end, dorsal view. Paratype (MCZ 153565): B, posterior end, dorsal view; C, thoracic setiger, anterior view; D, anterior abdominal setiger, posterior view; E, posterior abdominal setiger, posterior view." figureDoi="http://doi.org/10.5281/zenodo.3634276" httpUri="https://zenodo.org/record/3634276/files/figure.png" pageId="9" pageNumber="10">Fig. 3B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FD69BC289677230BD890FACDFB34FA0D" blockId="9.[151,1437,297,1836]" pageId="9" pageNumber="10">
|
||
<emphasis id="CFA2603A9677230BD890FACDFEECFAEE" bold="true" box="[199,327,1377,1404]" pageId="9" pageNumber="10">Variability</emphasis>
|
||
. The separation of the two peristomial rings is sometimes apparent only laterally, with the groove not crossing the dorsum. The anal cirri are usually lost, best observed on smaller specimens.
|
||
</paragraph>
|
||
<paragraph id="FD69BC289677230BD890FA06FC96FA51" blockId="9.[151,1437,297,1836]" box="[199,829,1449,1476]" pageId="9" pageNumber="10">
|
||
<emphasis id="CFA2603A9677230BD890FA06FE04FA56" bold="true" box="[199,431,1449,1476]" pageId="9" pageNumber="10">Methyl Green stain</emphasis>
|
||
. No pattern, body stains uniformly.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B5CCEFA39677230BD890FA62FDBFF952" pageId="9" pageNumber="10" type="discussion">
|
||
<paragraph id="FD69BC289677230BD890FA62FDBFF952" blockId="9.[151,1437,297,1836]" pageId="9" pageNumber="10">
|
||
<emphasis id="CFA2603A9677230BD890FA62FE90FA7A" bold="true" box="[199,315,1486,1512]" pageId="9" pageNumber="10">Remarks.</emphasis>
|
||
<taxonomicName id="3AD6C7AB9677230BD915FA61FDCDFA75" authority="Blake, 2020" authorityName="Blake" authorityYear="2020" box="[322,614,1485,1511]" class="Polychaeta" family="Orbiniidae" genus="Leitoscoloplos" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="gordaensis" status="sp. nov.">
|
||
<emphasis id="CFA2603A9677230BD915FA61FDCDFA75" box="[322,614,1485,1511]" italics="true" pageId="9" pageNumber="10">Leitoscoloplos gordaensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CFA2603A9677230BDA3AFA62FD00FA7A" bold="true" box="[621,683,1486,1512]" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="D491DD419677230BDA3AFA62FD00FA7A" box="[621,683,1486,1512]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
belongs to the
|
||
<taxonomicName id="3AD6C7AB9677230BDB01FA62FC55FA75" baseAuthorityName="McIntosh" baseAuthorityYear="1885" box="[854,1022,1485,1511]" class="Polychaeta" family="Orbiniidae" genus="Leitoscoloplos" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="kerguelensis">
|
||
<emphasis id="CFA2603A9677230BDB01FA62FC55FA75" box="[854,1022,1485,1511]" italics="true" pageId="9" pageNumber="10">L. kerguelensis</emphasis>
|
||
</taxonomicName>
|
||
group in having branchiae first present from anterior abdominal setigers (
|
||
<bibRefCitation id="9947C1D99677230BDA6CFA5DFD69F999" author="Blake, J. A." box="[571,706,1521,1547]" pageId="9" pageNumber="10" pagination="1 - 145" refId="ref35753" refString="Blake, J. A. (2017) Polychaeta Orbiniidae from Antarctica, the Southern Ocean, the abyssal Pacific Ocean, and off South Amer- ica. Zootaxa, 4218 (1), 1 - 145. https: // doi. org / 10.11646 / zootaxa. 4218.1.1" type="journal article" year="2017">Blake 2017</bibRefCitation>
|
||
). The five species in this group from the eastern North Pacific are separated from the others in the key above and by additional characteristics in
|
||
<tableCitation id="B05489939677230BDC55F9B9FBFDF9BD" box="[1026,1110,1557,1583]" captionStart="TABLE 1" captionStartId="51.[146,234,475,501]" captionTargetPageId="51" captionText="TABLE 1. Comparison of 13 Species of Leitoscoloplos from deep-sea habitats greater than 750 m." httpUri="http://table.plazi.org/id/A9A9ECA0964D2331D8C5FE77FB19FE67" pageId="9" pageNumber="10" tableUuid="A9A9ECA0964D2331D8C5FE77FB19FE67">Table 1</tableCitation>
|
||
. Of the four species that lack furcate setae,
|
||
<taxonomicName id="3AD6C7AB9677230BD964F996FE61F9C1" authority="Blake, 2020" authorityName="Blake" authorityYear="2020" box="[307,458,1593,1619]" class="Polychaeta" family="Orbiniidae" genus="Leitoscoloplos" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="gordaensis" status="sp. nov.">
|
||
<emphasis id="CFA2603A9677230BD964F996FE61F9C1" box="[307,458,1593,1619]" italics="true" pageId="9" pageNumber="10">L. gordaensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CFA2603A9677230BD985F996FDBAF9C6" bold="true" box="[466,529,1594,1620]" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="D491DD419677230BD985F996FDBAF9C6" box="[466,529,1594,1620]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="3AD6C7AB9677230BDA1FF996FD13F9C1" authority="Blake, 2020" authorityName="Blake" authorityYear="2020" box="[584,696,1593,1619]" class="Polychaeta" family="Orbiniidae" genus="Leitoscoloplos" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="lunulus" status="sp. nov.">
|
||
<emphasis id="CFA2603A9677230BDA1FF996FD13F9C1" box="[584,696,1593,1619]" italics="true" pageId="9" pageNumber="10">L. lunulus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CFA2603A9677230BDA97F996FD54F9C6" bold="true" box="[704,767,1594,1620]" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="D491DD419677230BDA97F996FD54F9C6" box="[704,767,1594,1620]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
each have a relatively short pre-setal region where the prostomium and peristomium are separated from one another by a distinct groove. The two species differ significantly from one another in that
|
||
<taxonomicName id="3AD6C7AB9677230BD9FFF92EFDBCF909" authority="Blake, 2020" authorityName="Blake" authorityYear="2020" box="[424,535,1665,1691]" class="Polychaeta" family="Orbiniidae" genus="Leitoscoloplos" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="lunulus" status="sp. nov.">
|
||
<emphasis id="CFA2603A9677230BD9FFF92EFDBCF909" box="[424,535,1665,1691]" italics="true" pageId="9" pageNumber="10">L. lunulus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CFA2603A9677230BDA49F92EFDF0F90E" bold="true" box="[542,603,1666,1692]" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="D491DD419677230BDA49F92EFDF0F90E" box="[542,603,1666,1692]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has a ventral cirrus and subpodial flange on the abdominal neuropodia that are lacking in
|
||
<taxonomicName id="3AD6C7AB9677230BD960F90AFE65F92D" authority="Blake, 2020" authorityName="Blake" authorityYear="2020" box="[311,462,1701,1727]" class="Polychaeta" family="Orbiniidae" genus="Leitoscoloplos" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="gordaensis" status="sp. nov.">
|
||
<emphasis id="CFA2603A9677230BD960F90AFE65F92D" box="[311,462,1701,1727]" italics="true" pageId="9" pageNumber="10">L. gordaensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CFA2603A9677230BD982F90AFDBFF952" bold="true" box="[469,532,1702,1728]" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="D491DD419677230BD982F90AFDBFF952" box="[469,532,1702,1728]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B5CCEFA39677230BD890F966FC51F971" box="[199,1018,1737,1764]" pageId="9" pageNumber="10" type="etymology">
|
||
<paragraph id="FD69BC289677230BD890F966FC51F971" blockId="9.[151,1437,297,1836]" box="[199,1018,1737,1764]" pageId="9" pageNumber="10">
|
||
<emphasis id="CFA2603A9677230BD890F966FEE3F976" bold="true" box="[199,328,1738,1764]" pageId="9" pageNumber="10">Etymology</emphasis>
|
||
. The species is named for its occurrence on the Gorda Ridge.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B5CCEFA39677230BD890F941FE68F8BE" pageId="9" pageNumber="10" type="distribution">
|
||
<paragraph id="FD69BC289677230BD890F941FE68F8BE" blockId="9.[151,1437,297,1836]" pageId="9" pageNumber="10">
|
||
<emphasis id="CFA2603A9677230BD890F941FEF2F89A" bold="true" box="[199,345,1773,1800]" pageId="9" pageNumber="10">Distribution</emphasis>
|
||
. Eastern Pacific, off northern
|
||
<collectingRegion id="3F1272CA9677230BDAF0F941FCB2F895" box="[679,793,1773,1799]" country="United States of America" name="California" pageId="9" pageNumber="10">California</collectingRegion>
|
||
in the vicinity of hydrothermal vents on the Gorda Ridge, Escanaba Trough,
|
||
<quantity id="3A2E11CD9677230BD930F8BDFE14F8BE" box="[359,447,1809,1836]" metricMagnitude="3" metricUnit="m" metricValue="3.271" pageId="9" pageNumber="10" unit="m" value="3271.0">3271 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |