310 lines
40 KiB
XML
310 lines
40 KiB
XML
<document id="517C415EB64A26B6CEACC3B34493E3B5" ID-CLB-Dataset="24204" ID-DOI="10.11646/zootaxa.4730.1.1" ID-GBIF-Dataset="bcc0e0fc-b6a5-4d5a-8008-7b714c106a7c" ID-ISSN="1175-5326" ID-Zenodo-Dep="3634270" ID-ZooBank="7F4CD129-9FF9-4593-A8A4-DB999B3E402F" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1580713876401" checkinUser="plazi" docAuthor="Blake, James A." docDate="2020" docId="757F0D3E966E2310D8C0FEDCFBF5FE3D" docLanguage="en" docName="zootaxa.4730.1.1.pdf" docOrigin="Zootaxa 4730 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Leitoscoloplos williamsae Blake 2020, new species" docType="treatment" docUuid="8CBB5BCD-91E0-4077-9803-095212E75A95" docUuidSource="ZooBank" docVersion="8" lastPageNumber="19" masterDocId="89467546967E2302D857FFACFFABFF92" masterDocTitle="New species and records of deep-water Orbiniidae (Annelida, Polychaeta) from the Eastern Pacific continental slope, abyssal Pacific Ocean, and the South China Sea" masterLastPageNumber="61" masterPageNumber="1" pageNumber="17" updateTime="1698780308397" updateUser="ExternalLinkService">
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<mods:title id="3ED5D21A9A5FABF4E6189FC18EF58258">New species and records of deep-water Orbiniidae (Annelida, Polychaeta) from the Eastern Pacific continental slope, abyssal Pacific Ocean, and the South China Sea</mods:title>
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<mods:roleTerm id="6C710A1BE1E51BC557802E8D5700401D">Author</mods:roleTerm>
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<mods:namePart id="DF03F6D967D064E82703735638DF8D6B">Blake, James A.</mods:namePart>
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<mods:typeOfResource id="7882A22798FD2CE9D375B45E685A961A">text</mods:typeOfResource>
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<mods:title id="570D52F0C3C389BE515BF3C87BE54C0B">Zootaxa</mods:title>
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<mods:part id="F852D96AE2E00CAF41A058877E6BD02F">
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<mods:date id="EA6E5100D540C66B4414FFF3E0F74352">2020</mods:date>
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<mods:number id="D2F90D797CA56A154191D3AC3DE2C4BA">2020-02-03</mods:number>
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<mods:number id="1C9E37AD2F0BE99BA3F85DB0319779DC">4730</mods:number>
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<mods:detail id="C7339B7B81AADEBB41EE3396E5641CF3" type="issue">
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<mods:identifier id="913DE1E3A60950970E5617CFA871D8E0" type="DOI">10.11646/zootaxa.4730.1.1</mods:identifier>
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<treatment id="757F0D3E966E2310D8C0FEDCFBF5FE3D" ID-GBIF-Taxon="161938995" LSID="urn:lsid:zoobank.org:act:8CBB5BCD-91E0-4077-9803-095212E75A95" httpUri="http://treatment.plazi.org/id/757F0D3E966E2310D8C0FEDCFBF5FE3D" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17">
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<subSubSection id="B5CCEFA3966E2312D8C0FEDCFDC8FE18" box="[151,611,368,395]" pageId="16" pageNumber="17" type="nomenclature">
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<paragraph id="FD69BC28966E2312D8C0FEDCFDC8FE18" blockId="16.[151,959,368,467]" box="[151,611,368,395]" pageId="16" pageNumber="17">
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<emphasis id="CFA2603A966E2312D8C0FEDCFDC8FE18" bold="true" box="[151,611,368,395]" pageId="16" pageNumber="17">
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<taxonomicName id="3AD6C7AB966E2312D8C0FEDCFE67FE19" ID-CoL="85RHH" authority="Blake, 2020" authorityName="Blake" authorityYear="2020" box="[151,460,368,395]" class="Polychaeta" family="Orbiniidae" genus="Leitoscoloplos" kingdom="Animalia" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="williamsae" status="sp. nov.">
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<emphasis id="CFA2603A966E2312D8C0FEDCFE67FE19" bold="true" box="[151,460,368,395]" italics="true" pageId="16" pageNumber="17">Leitoscoloplos williamsae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D491DD41966E2312D984FEDDFDC8FE18" box="[467,611,368,395]" pageId="16" pageNumber="17" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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<subSubSection id="B5CCEFA3966E2312D8C0FE38FC14FE41" pageId="16" pageNumber="17" type="description">
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<paragraph id="FD69BC28966E2312D8C0FE38FEB6FE3D" blockId="16.[151,959,368,467]" box="[151,285,404,431]" pageId="16" pageNumber="17">
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<figureCitation id="65EDA0AD966E2312D8C0FE38FEB6FE3D" box="[151,285,404,431]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStartId-0="17.[151,250,1945,1970]" captionStartId-1="18.[151,250,1164,1189]" captionTargetBox-0="[194,1388,189,1911]" captionTargetBox-1="[156,1433,458,1134]" captionTargetId-0="figure@17.[184,1403,181,1920]" captionTargetId-1="figure@18.[151,1436,453,1139]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="FIGURE 8. Leitoscoloplos williamsae n. sp. Holotype (LACM-AHF Poly 11658):A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior end, right lateral view; D, thoracic setiger 7, right posterior view." captionText-1="FIGURE 9. Leitoscoloplos williamsae n. sp. Holotype (LACM-AHF Poly 11658): A, abdominal setiger 20, left side, anterior view; B, abdominal setiger 32, right side, anterior view." figureDoi-0="http://doi.org/10.5281/zenodo.3634286" figureDoi-1="http://doi.org/10.5281/zenodo.3634288" httpUri-0="https://zenodo.org/record/3634286/files/figure.png" httpUri-1="https://zenodo.org/record/3634288/files/figure.png" pageId="16" pageNumber="17">Figures 8–9</figureCitation>
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</paragraph>
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<paragraph id="FD69BC28966E2312D8C0FE14FC14FE41" blockId="16.[151,959,368,467]" box="[151,959,440,467]" pageId="16" pageNumber="17">
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<uri id="8947B02A966E2312D8C0FE14FC14FE41" box="[151,959,440,467]" pageId="16" pageNumber="17">
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urn:lsid:zoobank.org:act:
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<uuid id="897086FD966E2312D9FEFE14FC14FE41" box="[425,959,440,467]" pageId="16" pageNumber="17">8CBB5BCD-91E0-4077-9803-095212E75A95</uuid>
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</paragraph>
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<subSubSection id="B5CCEFA3966E2312D8C0FE51FD74FD87" box="[151,735,508,533]" pageId="16" pageNumber="17" type="reference_group">
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<paragraph id="FD69BC28966E2312D8C0FE51FD74FD87" blockId="16.[151,735,508,533]" box="[151,735,508,533]" pageId="16" pageNumber="17">
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<treatmentCitationGroup id="DDC69B06966E2312D8C0FE51FD74FD87" box="[151,735,508,533]" pageId="16" pageNumber="17">
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<taxonomicName id="3AD6C7AB966E2312D8C0FE51FE80FD87" ID-CoL="84PL5" authorityName="Day" authorityYear="1977" box="[151,299,509,533]" class="Polychaeta" family="Orbiniidae" genus="Leitoscoloplos" kingdom="Animalia" pageId="16" pageNumber="17" phylum="Annelida" rank="genus">
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<emphasis id="CFA2603A966E2312D8C0FE51FE80FD87" box="[151,299,509,533]" italics="true" pageId="16" pageNumber="17">Leitoscoloplos</emphasis>
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</taxonomicName>
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sp. 3 and sp. 4:
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<treatmentCitation id="7C779A39966E2312D987FE50FD70FD87" author="Hilbig, B. & Blake, J. A." box="[464,731,508,533]" page="265" pageId="16" pageNumber="17" year="2006">
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<bibRefCitation id="9947C1D9966E2312D987FE50FD70FD87" author="Hilbig, B. & Blake, J. A." box="[464,731,508,533]" pageId="16" pageNumber="17" pagination="243 - 269" refId="ref37496" refString="Hilbig, B. & Blake, J. A. (2006) Deep-sea polychaete communities in the Northeast Pacific Ocean off the Gulf of the Farallones, California. Bulletin of Marine Science, 78, 243 - 269. [https: // www. ingentaconnect. com / content / umrsmas / bullmar / 2006 / 00000078 / 00000002 / art 00002]" type="journal article" year="2006">Hilbig & Blake 2006: 265</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<subSubSection id="B5CCEFA3966E2312D8C0FDEEFCFFFD5B" pageId="16" pageNumber="17" type="materials_examined">
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<paragraph id="FD69BC28966E2312D8C0FDEEFCFBFD5B" blockId="16.[151,1437,578,2046]" pageId="16" pageNumber="17">
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<emphasis id="CFA2603A966E2312D8C0FDEEFED3FDCF" bold="true" box="[151,376,578,605]" pageId="16" pageNumber="17">Material examined</emphasis>
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.
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<emphasis id="CFA2603A966E2312D9D2FDEEFD5FFDCF" bold="true" box="[389,756,578,605]" pageId="16" pageNumber="17">Continental slope, off northern</emphasis>
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<materialsCitation id="4DBEB675966E2312DAACFDEEFD0EFD37" ID-GBIF-Occurrence="2571581955" collectingDate="1991-09-16" collectingDateMax="1991-09-17" collectingDateMin="1991-09-16" collectionCode="R, V, LACM-AHF" collectorName="Half Moon Bay, U. S & Site & Survey & Point Sur & J. A. Blake & Sta." county="Half Moon Bay" elevation="1760" latitude="37.434166" location="North of Pioneer Canyon" longLatPrecision="11" longitude="-123.33317" municipality="Pioneer Canyon" pageId="16" pageNumber="17" specimenCount="1" stateProvince="California" typeStatus="holotype">
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<collectingRegion id="3F1272CA966E2312DAACFDEEFCDDFDCF" box="[763,886,578,605]" country="United States of America" name="California" pageId="16" pageNumber="17">
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<emphasis id="CFA2603A966E2312DAACFDEEFCDDFDCF" bold="true" box="[763,886,578,605]" pageId="16" pageNumber="17">California</emphasis>
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</collectingRegion>
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,
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<emphasis id="CFA2603A966E2312DBD5FDEEFBCFFDCF" bold="true" box="[898,1124,578,605]" pageId="16" pageNumber="17">
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off
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<collectorName id="5023D9FE966E2312DBFEFDEEFBCFFDCF" box="[937,1124,578,605]" pageId="16" pageNumber="17">
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<collectingCounty id="1408C4A4966E2312DBFEFDEEFBF4FDCF" box="[937,1119,578,605]" pageId="16" pageNumber="17">Half Moon Bay</collectingCounty>
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,
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</collectorName>
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</emphasis>
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U.S EPA
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<collectorName id="5023D9FE966E2312DC85FDEFFB55FDCF" box="[1234,1278,579,605]" pageId="16" pageNumber="17">Site</collectorName>
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selection
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<collectorName id="5023D9FE966E2312DD3AFDEFFF6BFD10" pageId="16" pageNumber="17">Survey</collectorName>
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,
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<collectionCode id="9BC724ED966E2312D89BFDCBFF75FD13" box="[204,222,615,641]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="16" pageNumber="17">R</collectionCode>
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/
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<collectionCode id="9BC724ED966E2312D8B1FDCBFF52FD13" box="[230,249,615,641]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="17" type="Museum">V</collectionCode>
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<emphasis id="CFA2603A966E2312D956FDC4FEC0FD10" box="[257,363,616,642]" italics="true" pageId="16" pageNumber="17">
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<collectorName id="5023D9FE966E2312D956FDC4FEC0FD10" box="[257,363,616,642]" pageId="16" pageNumber="17">Point Sur</collectorName>
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</emphasis>
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, coll.
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<collectorName id="5023D9FE966E2312D9E6FDCBFD82FD13" box="[433,553,615,641]" pageId="16" pageNumber="17">J.A. Blake</collectorName>
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,
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<emphasis id="CFA2603A966E2312DA62FDCBFCF1FD13" bold="true" box="[565,858,614,641]" pageId="16" pageNumber="17">
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south of
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<collectingMunicipality id="1D0D2652966E2312DACCFDCAFCF1FD13" box="[667,858,614,641]" pageId="16" pageNumber="17">Pioneer Canyon</collectingMunicipality>
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</emphasis>
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,
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<collectorName id="5023D9FE966E2312DB3EFDCBFC3FFD13" box="[873,916,615,641]" pageId="16" pageNumber="17">Sta.</collectorName>
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4-10,
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<date id="89689AE8966E2312DB8BFDCBFBC1FD13" box="[988,1130,615,641]" pageId="16" pageNumber="17" value="1991-09-17">
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<collectingDate id="992C6300966E2312DB8BFDCBFBC1FD13" box="[988,1130,615,641]" pageId="16" pageNumber="17" value="1991-09-17">17 Sep 1991</collectingDate>
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</date>
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,
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<geoCoordinate id="98E2DAEF966E2312DC21FDCAFB53FD13" box="[1142,1272,614,641]" degrees="37" direction="north" minutes="11.26" orientation="latitude" pageId="16" pageNumber="17" precision="9" value="37.187668">37°11.26′N</geoCoordinate>
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,
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<geoCoordinate id="98E2DAEF966E2312DD54FDCAFA33FD13" box="[1283,1432,614,641]" degrees="123" direction="west" minutes="15.26" orientation="longitude" pageId="16" pageNumber="17" precision="9" value="-123.25433">123°15.26′W</geoCoordinate>
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,
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<quantity id="3A2E11CD966E2312D8C0FD27FF5AFD34" box="[151,241,651,678]" metricMagnitude="3" metricUnit="m" metricValue="1.76" pageId="16" pageNumber="17" unit="m" value="1760.0">
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<elevation id="76FB5B1B966E2312D8C0FD27FF5AFD34" box="[151,241,651,678]" metricMagnitude="3" metricUnit="m" metricValue="1.76" pageId="16" pageNumber="17" unit="m" value="1760.0">1760 m</elevation>
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</quantity>
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,
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<emphasis id="CFA2603A966E2312D957FD27FECCFD37" bold="true" box="[256,359,651,677]" pageId="16" pageNumber="17">
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<typeStatus id="226D028A966E2312D957FD27FECCFD37" box="[256,359,651,677]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
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</emphasis>
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(
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<collectionCode id="9BC724ED966E2312D92FFD27FDA7FD37" box="[376,524,651,677]" pageId="16" pageNumber="17">LACM-AHF</collectionCode>
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Poly 11658)
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="4DBEB675966E2312DAE4FD27FCFBFD5B" ID-GBIF-Occurrence="2571581941" collectingDate="1991-09-16" collectingDateMax="1991-09-17" collectingDateMin="1991-09-16" collectionCode="R, V, LACM-AHF" collectorName="Half Moon Bay, U. S & Site & Survey & Point Sur & J. A. Blake & Sta." county="Half Moon Bay" elevation="1760" latitude="37.434166" location="North of Pioneer Canyon" longLatPrecision="11" longitude="-123.33317" municipality="Pioneer Canyon" pageId="16" pageNumber="17" specimenCount="1" stateProvince="California" typeStatus="holotype">
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<emphasis id="CFA2603A966E2312DAE4FD27FC43FD37" bold="true" box="[691,1000,650,677]" pageId="16" pageNumber="17">
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<location id="F809EAF3966E2312DAE4FD27FC43FD37" LSID="urn:lsid:plazi:treatment:757F0D3E966E2310D8C0FEDCFBF5FE3D:F809EAF3966E2312DAE4FD27FC43FD37" box="[691,1000,650,677]" county="Half Moon Bay" latitude="37.434166" longLatPrecision="11" longitude="-123.33317" municipality="Pioneer Canyon" name="North of Pioneer Canyon" pageId="16" pageNumber="17" stateProvince="California">North of Pioneer Canyon</location>
|
||
</emphasis>
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||
, Sta. 3-19,
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||
<date id="89689AE8966E2312DC25FD27FAADFD37" box="[1138,1286,651,677]" pageId="16" pageNumber="17" value="1991-09-16">
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<collectingDate id="992C6300966E2312DC25FD27FAADFD37" box="[1138,1286,651,677]" pageId="16" pageNumber="17" value="1991-09-16">16 Sep 1991</collectingDate>
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</date>
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,
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<geoCoordinate id="98E2DAEF966E2312DD42FD26FA3CFD37" box="[1301,1431,650,677]" degrees="37" direction="north" minutes="26.05" orientation="latitude" pageId="16" pageNumber="17" precision="9" value="37.434166">37°26.05′N</geoCoordinate>
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,
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<geoCoordinate id="98E2DAEF966E2312D8C0FD02FE87FD5B" box="[151,300,686,713]" degrees="123" direction="west" minutes="19.99" orientation="longitude" pageId="16" pageNumber="17" precision="9" value="-123.33317">123°19.99′W</geoCoordinate>
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,
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<quantity id="3A2E11CD966E2312D960FD03FE25FD58" box="[311,398,687,714]" metricMagnitude="3" metricUnit="m" metricValue="1.76" pageId="16" pageNumber="17" unit="m" value="1760.0">
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<elevation id="76FB5B1B966E2312D960FD03FE25FD58" box="[311,398,687,714]" metricMagnitude="3" metricUnit="m" metricValue="1.76" pageId="16" pageNumber="17" unit="m" value="1760.0">1760 m</elevation>
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</quantity>
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||
,
|
||
<specimenCount id="EBD077A1966E2312D9CDFD03FDB1FD5B" box="[410,538,687,713]" pageId="16" pageNumber="17" type="generic" typeStatus="paratype">
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1
|
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<emphasis id="CFA2603A966E2312D9F8FD03FDB1FD5B" bold="true" box="[431,538,687,713]" pageId="16" pageNumber="17">paratype</emphasis>
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</specimenCount>
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(
|
||
<collectionCode id="9BC724ED966E2312DA7EFD03FD16FD5B" box="[553,701,687,713]" pageId="16" pageNumber="17">LACM-AHF</collectionCode>
|
||
Poly 11659)
|
||
</materialsCitation>
|
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</paragraph>
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<paragraph id="FD69BC28966E2312DB07FD02FCFFFD5B" blockId="16.[151,1437,578,2046]" box="[848,852,686,713]" pageId="16" pageNumber="17">.</paragraph>
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</subSubSection>
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<subSubSection id="B5CCEFA3966E2312D890FD7EFA21F893" pageId="16" pageNumber="17" type="description">
|
||
<paragraph id="FD69BC28966E2312D890FD7EFCC8FB9C" blockId="16.[151,1437,578,2046]" pageId="16" pageNumber="17">
|
||
<emphasis id="CFA2603A966E2312D890FD7EFEFAFD7F" bold="true" box="[199,337,722,749]" pageId="16" pageNumber="17">Description</emphasis>
|
||
. A moderate-sized species,
|
||
<typeStatus id="226D028A966E2312DAD3FD7FFD4DFD7F" box="[644,742,723,749]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
|
||
complete,
|
||
<quantity id="3A2E11CD966E2312DB37FD7EFC68FD7C" box="[864,963,722,750]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="16" pageNumber="17" unit="mm" value="10.5">10.5 mm</quantity>
|
||
long,
|
||
<quantity id="3A2E11CD966E2312DC50FD7FFBF7FD7C" box="[1031,1116,723,750]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="16" pageNumber="17" unit="mm" value="0.6">0.6 mm</quantity>
|
||
across thoracic segments for 65 setigers;
|
||
<typeStatus id="226D028A966E2312D976FD54FE2AFC80" box="[289,385,760,786]" pageId="16" pageNumber="17" type="paratype">paratype</typeStatus>
|
||
mostly complete,
|
||
<quantity id="3A2E11CD966E2312DA03FD5AFD00FC80" box="[596,683,758,786]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="16" pageNumber="17" unit="mm" value="5.7">5.7 mm</quantity>
|
||
long,
|
||
<quantity id="3A2E11CD966E2312DAA4FD5AFCF2FC80" box="[755,857,758,786]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="16" pageNumber="17" unit="mm" value="0.25">0.25 mm</quantity>
|
||
across thorax for 40 setigers. Body elongate, more or less consistently same width throughout, tapering to narrow posterior end. Thoracic segments about 3.5 times as wide as long, each separated from one another by prominent intersegmental grooves, grooves with an elevated transverse dorsal ridge or hump (
|
||
<figureCitation id="65EDA0AD966E2312DA42FCCFFDFCFCEF" box="[533,599,867,893]" captionStart="FIGURE 8" captionStartId="17.[151,250,1945,1970]" captionTargetBox="[194,1388,189,1911]" captionTargetId="figure@17.[184,1403,181,1920]" captionTargetPageId="17" captionText="FIGURE 8. Leitoscoloplos williamsae n. sp. Holotype (LACM-AHF Poly 11658):A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior end, right lateral view; D, thoracic setiger 7, right posterior view." figureDoi="http://doi.org/10.5281/zenodo.3634286" httpUri="https://zenodo.org/record/3634286/files/figure.png" pageId="16" pageNumber="17">Fig. 8</figureCitation>
|
||
A–B) producing a biannulate appearance; lateral and ventral grooves not as conspicuous. Anterior abdominal segments narrow, becoming longer, about twice as wide as long in middle and posterior segments. Anterior and middle abdominal segments with intersegmental groove transformed into a flattened band producing a biannulate appearance, band obscured in posterior abdominal segments. Longitudinal grooves or ridges along body absent. Color in alcohol: light tan.
|
||
</paragraph>
|
||
<paragraph id="FD69BC28966E2312D890FBBBFDF0FB0F" blockId="16.[151,1437,578,2046]" pageId="16" pageNumber="17">
|
||
Pre-setiger region triangular, longer than wide, narrowing anteriorly (
|
||
<figureCitation id="65EDA0AD966E2312DB90FBBBFBA3FBA3" box="[967,1032,1047,1073]" captionStart="FIGURE 8" captionStartId="17.[151,250,1945,1970]" captionTargetBox="[194,1388,189,1911]" captionTargetId="figure@17.[184,1403,181,1920]" captionTargetPageId="17" captionText="FIGURE 8. Leitoscoloplos williamsae n. sp. Holotype (LACM-AHF Poly 11658):A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior end, right lateral view; D, thoracic setiger 7, right posterior view." figureDoi="http://doi.org/10.5281/zenodo.3634286" httpUri="https://zenodo.org/record/3634286/files/figure.png" pageId="16" pageNumber="17">Fig. 8</figureCitation>
|
||
A–B). Prostomium conical, tapering to narrow apex (
|
||
<figureCitation id="65EDA0AD966E2312D902FB97FE1AFBC7" box="[341,433,1083,1109]" captionStart="FIGURE 8" captionStartId="17.[151,250,1945,1970]" captionTargetBox="[194,1388,189,1911]" captionTargetId="figure@17.[184,1403,181,1920]" captionTargetPageId="17" captionText="FIGURE 8. Leitoscoloplos williamsae n. sp. Holotype (LACM-AHF Poly 11658):A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior end, right lateral view; D, thoracic setiger 7, right posterior view." figureDoi="http://doi.org/10.5281/zenodo.3634286" httpUri="https://zenodo.org/record/3634286/files/figure.png" pageId="16" pageNumber="17">Fig. 8A</figureCitation>
|
||
); nuchal organs narrow, crescent-shaped on posterior lateral margin (
|
||
<figureCitation id="65EDA0AD966E2312DC9DFB97FA8FFBC7" box="[1226,1316,1083,1109]" captionStart="FIGURE 8" captionStartId="17.[151,250,1945,1970]" captionTargetBox="[194,1388,189,1911]" captionTargetId="figure@17.[184,1403,181,1920]" captionTargetPageId="17" captionText="FIGURE 8. Leitoscoloplos williamsae n. sp. Holotype (LACM-AHF Poly 11658):A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior end, right lateral view; D, thoracic setiger 7, right posterior view." figureDoi="http://doi.org/10.5281/zenodo.3634286" httpUri="https://zenodo.org/record/3634286/files/figure.png" pageId="16" pageNumber="17">Fig. 8B</figureCitation>
|
||
); eyespots absent. Peristomium composed of a single, smooth achaetous ring, ventrally surrounding mouth, inflated dorsally, with no transverse grooves (
|
||
<figureCitation id="65EDA0AD966E2312D984FB2FFDBEFB0F" box="[467,533,1155,1181]" captionStart="FIGURE 8" captionStartId="17.[151,250,1945,1970]" captionTargetBox="[194,1388,189,1911]" captionTargetId="figure@17.[184,1403,181,1920]" captionTargetPageId="17" captionText="FIGURE 8. Leitoscoloplos williamsae n. sp. Holotype (LACM-AHF Poly 11658):A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior end, right lateral view; D, thoracic setiger 7, right posterior view." figureDoi="http://doi.org/10.5281/zenodo.3634286" httpUri="https://zenodo.org/record/3634286/files/figure.png" pageId="16" pageNumber="17">Fig. 8</figureCitation>
|
||
A–B).
|
||
</paragraph>
|
||
<paragraph id="FD69BC28966E2312D890FB0BFAE1FA0B" blockId="16.[151,1437,578,2046]" pageId="16" pageNumber="17">
|
||
Thorax with nine setigers on
|
||
<typeStatus id="226D028A966E2312DA46FB0BFDDDFB53" box="[529,630,1191,1217]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
|
||
, transitioning to abdominal segments over 1–2 segments;
|
||
<typeStatus id="226D028A966E2312DD54FB04FAC8FB50" box="[1283,1379,1192,1218]" pageId="16" pageNumber="17" type="paratype">paratype</typeStatus>
|
||
with eight thoracic setigers. Boundary between last thoracic and first abdominal setiger demarcated by loss of most neurosetae and increase in size of neuropodial lobe. Thoracic notopodial and neuropodial lobes elongate, digitiform, both similar in form arising from broad rounded base (
|
||
<figureCitation id="65EDA0AD966E2312DB1BFABFFC03FABF" box="[844,936,1299,1325]" captionStart="FIGURE 8" captionStartId="17.[151,250,1945,1970]" captionTargetBox="[194,1388,189,1911]" captionTargetId="figure@17.[184,1403,181,1920]" captionTargetPageId="17" captionText="FIGURE 8. Leitoscoloplos williamsae n. sp. Holotype (LACM-AHF Poly 11658):A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior end, right lateral view; D, thoracic setiger 7, right posterior view." figureDoi="http://doi.org/10.5281/zenodo.3634286" httpUri="https://zenodo.org/record/3634286/files/figure.png" pageId="16" pageNumber="17">Fig. 8D</figureCitation>
|
||
); notopodia of anterior abdominal segments becoming thickened, elongate (
|
||
<figureCitation id="65EDA0AD966E2312D9ADFA9BFDFEFAC3" box="[506,597,1335,1361]" captionStart="FIGURE 9" captionStartId="18.[151,250,1164,1189]" captionTargetBox="[156,1433,458,1134]" captionTargetId="figure@18.[151,1436,453,1139]" captionTargetPageId="18" captionText="FIGURE 9. Leitoscoloplos williamsae n. sp. Holotype (LACM-AHF Poly 11658): A, abdominal setiger 20, left side, anterior view; B, abdominal setiger 32, right side, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3634288" httpUri="https://zenodo.org/record/3634288/files/figure.png" pageId="16" pageNumber="17">Fig. 9A</figureCitation>
|
||
); posterior notopodia narrower, tapering to pointed tip (
|
||
<figureCitation id="65EDA0AD966E2312DC87FA9BFA81FAC3" box="[1232,1322,1335,1361]" captionStart="FIGURE 9" captionStartId="18.[151,250,1164,1189]" captionTargetBox="[156,1433,458,1134]" captionTargetId="figure@18.[151,1436,453,1139]" captionTargetPageId="18" captionText="FIGURE 9. Leitoscoloplos williamsae n. sp. Holotype (LACM-AHF Poly 11658): A, abdominal setiger 20, left side, anterior view; B, abdominal setiger 32, right side, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3634288" httpUri="https://zenodo.org/record/3634288/files/figure.png" pageId="16" pageNumber="17">Fig. 9B</figureCitation>
|
||
). Anterior abdominal neuropodia thickened, with rounded terminal lobe and short ventral cirrus (
|
||
<figureCitation id="65EDA0AD966E2312DC34FAF7FB16FAE7" box="[1123,1213,1371,1397]" captionStart="FIGURE 9" captionStartId="18.[151,250,1164,1189]" captionTargetBox="[156,1433,458,1134]" captionTargetId="figure@18.[151,1436,453,1139]" captionTargetPageId="18" captionText="FIGURE 9. Leitoscoloplos williamsae n. sp. Holotype (LACM-AHF Poly 11658): A, abdominal setiger 20, left side, anterior view; B, abdominal setiger 32, right side, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3634288" httpUri="https://zenodo.org/record/3634288/files/figure.png" pageId="16" pageNumber="17">Fig. 9A</figureCitation>
|
||
); middle and posterior abdominal neuropodia longer, narrow, with short ventral cirrus and bulbous subpodial flange (
|
||
<figureCitation id="65EDA0AD966E2312DCB3FAD3FA96FA0B" box="[1252,1341,1407,1433]" captionStart="FIGURE 9" captionStartId="18.[151,250,1164,1189]" captionTargetBox="[156,1433,458,1134]" captionTargetId="figure@18.[151,1436,453,1139]" captionTargetPageId="18" captionText="FIGURE 9. Leitoscoloplos williamsae n. sp. Holotype (LACM-AHF Poly 11658): A, abdominal setiger 20, left side, anterior view; B, abdominal setiger 32, right side, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3634288" httpUri="https://zenodo.org/record/3634288/files/figure.png" pageId="16" pageNumber="17">Fig. 9B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FD69BC28966E2312D890FA0FFAA1F997" blockId="16.[151,1437,578,2046]" pageId="16" pageNumber="17">
|
||
Branchiae first present from setiger
|
||
<quantity id="3A2E11CD966E2312DA34FA0FFD35FA2F" box="[611,670,1443,1469]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" pageId="16" pageNumber="17" unit="in" value="17.0">17 in</quantity>
|
||
<typeStatus id="226D028A966E2312DAF1FA0FFCA3FA2F" box="[678,776,1443,1469]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
|
||
and setiger
|
||
<quantity id="3A2E11CD966E2312DBC1FA0FFC7BFA2F" box="[918,976,1443,1469]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="16" pageNumber="17" unit="in" value="18.0">18 in</quantity>
|
||
<typeStatus id="226D028A966E2312DB8EFA08FB96FA2C" box="[985,1085,1444,1470]" pageId="16" pageNumber="17" type="paratype">paratype</typeStatus>
|
||
; branchiae moderately sized at first, triangular in shape (
|
||
<figureCitation id="65EDA0AD966E2312D9F9FA6BFDACFA73" box="[430,519,1479,1505]" captionStart="FIGURE 9" captionStartId="18.[151,250,1164,1189]" captionTargetBox="[156,1433,458,1134]" captionTargetId="figure@18.[151,1436,453,1139]" captionTargetPageId="18" captionText="FIGURE 9. Leitoscoloplos williamsae n. sp. Holotype (LACM-AHF Poly 11658): A, abdominal setiger 20, left side, anterior view; B, abdominal setiger 32, right side, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3634288" httpUri="https://zenodo.org/record/3634288/files/figure.png" pageId="16" pageNumber="17">Fig. 9A</figureCitation>
|
||
), becoming longer and thicker in middle abdominal segments (
|
||
<figureCitation id="65EDA0AD966E2312DC97FA6BFAB3FA73" box="[1216,1304,1479,1505]" captionStart="FIGURE 9" captionStartId="18.[151,250,1164,1189]" captionTargetBox="[156,1433,458,1134]" captionTargetId="figure@18.[151,1436,453,1139]" captionTargetPageId="18" captionText="FIGURE 9. Leitoscoloplos williamsae n. sp. Holotype (LACM-AHF Poly 11658): A, abdominal setiger 20, left side, anterior view; B, abdominal setiger 32, right side, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3634288" httpUri="https://zenodo.org/record/3634288/files/figure.png" pageId="16" pageNumber="17">Fig. 9B</figureCitation>
|
||
). Branchiae of some far posterior abdominal segments of
|
||
<typeStatus id="226D028A966E2312DAC4FA47FD5EF997" box="[659,757,1515,1541]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
|
||
asymmetrical, with a lateral flaglike extension.
|
||
</paragraph>
|
||
<paragraph id="FD69BC28966E2312D890F9BCFC0BF92B" blockId="16.[151,1437,578,2046]" pageId="16" pageNumber="17">
|
||
Notosetae include camerated capillaries throughout; furcate and flail setae absent. Thoracic notosetae numbering about 25 per fascicle, arranged in two or three rows. Abdominal notosetae narrow camerated capillaries numbering
|
||
<quantity id="3A2E11CD966E2312D94EF9FBFEC4F9E3" box="[281,367,1623,1649]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" metricValueMax="3.048" metricValueMin="2.032" pageId="16" pageNumber="17" unit="in" value="10.0" valueMax="12.0" valueMin="8.0">8–12 in</quantity>
|
||
anterior abdominal segments, reduced to 5–10 posteriorly. Thoracic neurosetae all capillaries, numbering about 20–25 per fascicle in at least two rows. Abdominal neurosetae include 3–5 thin capillaries and 1–2 short protruding aciculae, these minute, with rounded apex (
|
||
<figureCitation id="65EDA0AD966E2312DB6EF933FC38F92B" box="[825,915,1695,1721]" captionStart="FIGURE 9" captionStartId="18.[151,250,1164,1189]" captionTargetBox="[156,1433,458,1134]" captionTargetId="figure@18.[151,1436,453,1139]" captionTargetPageId="18" captionText="FIGURE 9. Leitoscoloplos williamsae n. sp. Holotype (LACM-AHF Poly 11658): A, abdominal setiger 20, left side, anterior view; B, abdominal setiger 32, right side, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3634288" httpUri="https://zenodo.org/record/3634288/files/figure.png" pageId="16" pageNumber="17">Fig. 9A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FD69BC28966E2312D890F96FFB40F94F" blockId="16.[151,1437,578,2046]" box="[199,1259,1731,1758]" pageId="16" pageNumber="17">
|
||
Pygidium bluntly rounded, bearing two thin elongate cirri dorsal to the anal opening (
|
||
<figureCitation id="65EDA0AD966E2312DCD1F96FFB75F94F" box="[1158,1246,1731,1757]" captionStart="FIGURE 8" captionStartId="17.[151,250,1945,1970]" captionTargetBox="[194,1388,189,1911]" captionTargetId="figure@17.[184,1403,181,1920]" captionTargetPageId="17" captionText="FIGURE 8. Leitoscoloplos williamsae n. sp. Holotype (LACM-AHF Poly 11658):A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior end, right lateral view; D, thoracic setiger 7, right posterior view." figureDoi="http://doi.org/10.5281/zenodo.3634286" httpUri="https://zenodo.org/record/3634286/files/figure.png" pageId="16" pageNumber="17">Fig. 8C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FD69BC28966E2312D890F94BFA21F893" blockId="16.[151,1437,578,2046]" box="[199,1418,1766,1794]" pageId="16" pageNumber="17">
|
||
<emphasis id="CFA2603A966E2312D890F94BFE04F893" bold="true" box="[199,431,1766,1793]" pageId="16" pageNumber="17">Methyl Green stain</emphasis>
|
||
. No pattern; prostomium not staining, rest of body staining uniformly, de-stains rapidly.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B5CCEFA3966E2310D890F8A7FEF2FF69" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17" type="discussion">
|
||
<paragraph id="FD69BC28966E2310D890F8A7FEF2FF69" blockId="16.[151,1437,578,2046]" lastBlockId="18.[151,1437,153,431]" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17">
|
||
<emphasis id="CFA2603A966E2312D890F8A7FE9FF8B7" bold="true" box="[199,308,1803,1829]" pageId="16" pageNumber="17">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="3AD6C7AB966E2312D915F8A7FDCDF8B7" authority="Blake, 2020" authorityName="Blake" authorityYear="2020" box="[322,614,1803,1829]" class="Polychaeta" family="Orbiniidae" genus="Leitoscoloplos" kingdom="Animalia" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="williamsae" status="sp. nov.">
|
||
<emphasis id="CFA2603A966E2312D915F8A7FDCDF8B7" box="[322,614,1803,1829]" italics="true" pageId="16" pageNumber="17">Leitoscoloplos williamsae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CFA2603A966E2312DA3AF8A7FD07F8B7" bold="true" box="[621,684,1803,1829]" pageId="16" pageNumber="17">
|
||
<taxonomicNameLabel id="D491DD41966E2312DA3AF8A7FD07F8B7" box="[621,684,1803,1829]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
superficially resembles
|
||
<taxonomicName id="3AD6C7AB966E2312DBE9F8A0FB9EF8B7" authority="Blake, 2020" authorityName="Blake" authorityYear="2020" box="[958,1077,1803,1829]" class="Polychaeta" family="Orbiniidae" genus="Leitoscoloplos" kingdom="Animalia" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="cliffordi" status="sp. nov.">
|
||
<emphasis id="CFA2603A966E2312DBE9F8A0FB9EF8B7" box="[958,1077,1803,1829]" italics="true" pageId="16" pageNumber="17">L. cliffordi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CFA2603A966E2312DC6BF8A7FBD0F8B7" bold="true" box="[1084,1147,1803,1829]" pageId="16" pageNumber="17">
|
||
<taxonomicNameLabel id="D491DD41966E2312DC6BF8A7FBD0F8B7" box="[1084,1147,1803,1829]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having a narrow, elongate pre-setiger region and maximally nine thoracic setigers. However, there are several important differences between the two species:
|
||
<taxonomicName id="3AD6C7AB966E2312D9EBF8F8FD9EF8FF" authority="Blake, 2020" authorityName="Blake" authorityYear="2020" box="[444,565,1875,1901]" class="Polychaeta" family="Orbiniidae" genus="Leitoscoloplos" kingdom="Animalia" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="cliffordi" status="sp. nov.">
|
||
<emphasis id="CFA2603A966E2312D9EBF8F8FD9EF8FF" box="[444,565,1875,1901]" italics="true" pageId="16" pageNumber="17">L. cliffordi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CFA2603A966E2312DA6AF8FFFDD5F8FF" bold="true" box="[573,638,1875,1901]" pageId="16" pageNumber="17">
|
||
<taxonomicNameLabel id="D491DD41966E2312DA6AF8FFFDD5F8FF" box="[573,638,1875,1901]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has lateral grooves on the peristomium; a longitudinal ventral groove along the body; thoracic parapodial postsetal lobes are short and digitiform; branchiae are first present from setigers 15–16; notopodial furcate setae with blunt, unequal tynes are present; abdominal neuropodia become short and thick in posterior segments and abdominal neuropodia lack a subpodial flange;
|
||
<taxonomicName id="3AD6C7AB966E2312DC4BF86CFB1FF84B" authority="Blake, 2020" authorityName="Blake" authorityYear="2020" box="[1052,1204,1983,2009]" class="Polychaeta" family="Orbiniidae" genus="Leitoscoloplos" kingdom="Animalia" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="williamsae" status="sp. nov.">
|
||
<emphasis id="CFA2603A966E2312DC4BF86CFB1FF84B" box="[1052,1204,1983,2009]" italics="true" pageId="16" pageNumber="17">L. williamsae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CFA2603A966E2312DCEBF813FB57F84B" bold="true" box="[1212,1276,1983,2009]" pageId="16" pageNumber="17">
|
||
<taxonomicNameLabel id="D491DD41966E2312DCEBF813FB57F84B" box="[1212,1276,1983,2009]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has a peristomium with no lateral grooves; longitudinal grooves along the body are absent; thoracic postsetal lobes are long and digitiform; branchiae are first present from setigers 17–18; notopodial furcate setae are absent; abdominal neuropodia are long and digitate throughout; and abdominal neuropodia of middle and posterior segments have a bulbous subpodial flange.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="A9A9ECA0966F2313D8C0F835FBFEF844" ID-DOI="http://doi.org/10.5281/zenodo.3634286" ID-Zenodo-Dep="3634286" httpUri="https://zenodo.org/record/3634286/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,250,1945,1970]" targetBox="[194,1388,189,1911]" targetPageId="17">
|
||
<paragraph id="FD69BC28966F2313D8C0F835FBFEF844" blockId="17.[151,1436,1945,2006]" pageId="17" pageNumber="18">
|
||
<emphasis id="CFA2603A966F2313D8C0F835FEA7F820" bold="true" box="[151,268,1945,1970]" pageId="17" pageNumber="18">FIGURE 8</emphasis>
|
||
.
|
||
<taxonomicName id="3AD6C7AB966F2313D94FF836FDB4F820" authority="Blake, 2020" authorityName="Blake" authorityYear="2020" box="[280,543,1946,1970]" class="Polychaeta" family="Orbiniidae" genus="Leitoscoloplos" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="williamsae" status="sp. nov.">
|
||
<emphasis id="CFA2603A966F2313D94FF836FDB4F820" box="[280,543,1946,1970]" italics="true" pageId="17" pageNumber="18">Leitoscoloplos williamsae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D491DD41966F2313DA73F836FDF7F823" box="[548,604,1946,1970]" pageId="17" pageNumber="18" rank="species">
|
||
<emphasis id="CFA2603A966F2313DA73F836FDFDF820" bold="true" box="[548,598,1946,1970]" pageId="17" pageNumber="18">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
Holotype (LACM-AHF Poly 11658): A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior end, right lateral view; D, thoracic setiger 7, right posterior view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="B5CCEFA3966C2310D890FEA9FC59FE19" pageId="18" pageNumber="19" type="etymology">
|
||
<paragraph id="FD69BC28966C2310D890FEA9FC59FE19" blockId="18.[151,1437,153,431]" pageId="18" pageNumber="19">
|
||
<emphasis id="CFA2603A966C2310D890FEA9FEE3FE8D" bold="true" box="[199,328,261,287]" pageId="18" pageNumber="19">Etymology</emphasis>
|
||
. This species is named for my friend and long-time colleague, Isabelle P. (Izzie) Williams. Ms. Williams was part of the field team for the site-selection survey on which this species was collected and most of the following monitoring surveys at the SF-DODS site. Ms. Williams was also senior taxonomist on these projects, identifying the crustaceans and molluscs, and managed the benthic database.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B5CCEFA3966C2310D890FE38FBF5FE3D" box="[199,1118,404,431]" pageId="18" pageNumber="19" type="distribution">
|
||
<paragraph id="FD69BC28966C2310D890FE38FBF5FE3D" blockId="18.[151,1437,153,431]" box="[199,1118,404,431]" pageId="18" pageNumber="19">
|
||
<emphasis id="CFA2603A966C2310D890FE38FEF2FE3D" bold="true" box="[199,345,404,431]" pageId="18" pageNumber="19">Distribution</emphasis>
|
||
. Off northern
|
||
<collectingRegion id="3F1272CA966C2310D9ACFE39FDC6FE3D" box="[507,621,405,431]" country="United States of America" name="California" pageId="18" pageNumber="19">California</collectingRegion>
|
||
in middle continental slope depths,
|
||
<quantity id="3A2E11CD966C2310DC56FE39FBF2FE3D" box="[1025,1113,405,431]" metricMagnitude="3" metricUnit="m" metricValue="1.76" pageId="18" pageNumber="19" unit="m" value="1760.0">1760 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |