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<mods:title id="0755B3ED1F60CEEA202883FC16B2282D">New species and records of deep-water Orbiniidae (Annelida, Polychaeta) from the Eastern Pacific continental slope, abyssal Pacific Ocean, and the South China Sea</mods:title>
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<subSubSection id="B5CCEFA39662231ED8C0FE15FDE4FE41" box="[151,591,441,468]" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="FD69BC289662231ED8C0FE15FDE4FE41" blockId="28.[151,591,441,468]" box="[151,591,441,468]" pageId="28" pageNumber="29">
<heading id="A6210B449662231ED8C0FE15FDE4FE41" bold="true" box="[151,591,441,468]" fontSize="11" level="1" pageId="28" pageNumber="29" reason="1">
<emphasis id="CFA2603A9662231ED8C0FE15FDE4FE41" bold="true" box="[151,591,441,468]" pageId="28" pageNumber="29">
Genus
<taxonomicName id="3AD6C7AB9662231ED8B9FE15FDE4FE41" ID-CoL="9CM26" authority="de Blainville, 1828" authorityName="de Blainville" authorityYear="1828" box="[238,591,441,468]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Annelida" rank="genus">
<emphasis id="CFA2603A9662231ED8B9FE15FEC9FE46" bold="true" box="[238,354,441,468]" italics="true" pageId="28" pageNumber="29">Scoloplos</emphasis>
de
<bibRefCitation id="9947C1D99662231ED9D9FE15FDE4FE41" author="Blainville, H. de" box="[398,591,441,467]" pageId="28" pageNumber="29" pagination="368 - 501" refId="ref35358" refString="Blainville, H. de (1828) Dictionnaire des Sciences naturelles, 57, 368 - 501. Available from: https: // biodiversitylibrary. org / page / 25316890 (accessed 12 July 2019)" type="journal article" year="1828">Blainville, 1828</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B5CCEFA39662231ED8C0FE51FB53FDA6" pageId="28" pageNumber="29" type="type_taxon">
<paragraph id="FD69BC289662231ED8C0FE51FCB8FD84" blockId="28.[151,1272,508,564]" box="[151,787,508,534]" pageId="28" pageNumber="29">
<emphasis id="CFA2603A9662231ED8C0FE51FE8DFD87" bold="true" box="[151,294,508,533]" pageId="28" pageNumber="29">
<typeStatus id="226D028A9662231ED8C0FE51FF67FD87" box="[151,204,509,533]" pageId="28" pageNumber="29">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="3AD6C7AB9662231ED97BFE51FDD7FD87" authority="Muller, 1776" authorityName="Muller" authorityYear="1776" box="[300,636,508,533]" class="Polychaeta" family="Orbiniidae" genus="Lumbricus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Annelida" rank="species" species="armiger">
<emphasis id="CFA2603A9662231ED97BFE51FE5AFD87" box="[300,497,509,533]" italics="true" pageId="28" pageNumber="29">Lumbricus armiger</emphasis>
<bibRefCitation id="9947C1D99662231ED9A0FE50FDD7FD87" author="Muller, O. F." box="[503,636,508,533]" pageId="28" pageNumber="29" refId="ref38554" refString="Muller, O. F. (1776) Zoologiae Danicae Prodromus, seu Animalium Daniae et Norvegiae indigenarum characteres, nomina, et synonyma imprimis popularium. typis Hallageriis, Havniae, xxxii + 282 pp. https: // doi. org / 10.5962 / bhl. title. 13268" type="book" year="1776">Müller, 1776</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
<paragraph id="FD69BC289662231ED8C0FDB7FB53FDA6" blockId="28.[151,1272,508,564]" box="[151,1272,539,564]" pageId="28" pageNumber="29">
<emphasis id="CFA2603A9662231ED8C0FDB7FEAEFDA6" bold="true" box="[151,261,539,564]" pageId="28" pageNumber="29">Synonym:</emphasis>
<taxonomicName id="3AD6C7AB9662231ED95CFDB0FE5AFDA6" ID-CoL="0463366b-c7a4-441e-9d34-549aaa913a08" authority="Eisig, 1914" authorityName="Eisig" authorityYear="1914" box="[267,497,539,564]" class="Polychaeta" family="Orbiniidae" genus="Scolaricia" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Annelida" rank="genus">
<emphasis id="CFA2603A9662231ED95CFDB0FED8FDA6" box="[267,371,540,564]" italics="true" pageId="28" pageNumber="29">Scolaricia</emphasis>
<bibRefCitation id="9947C1D99662231ED92DFDB7FE5AFDA6" author="Eisig, H." box="[378,497,539,564]" pageId="28" pageNumber="29" pagination="153 - 600" refId="ref36564" refString="Eisig, H. (1914) Zur Systematik, Anatomie und Morphologie der Ariciiden nebst Beitragen zur generallen Systematik. Mitteilungen aus der Zoologischen Station Neapel, 21, 153 - 600, pls. 10 - 27, 23 figures. [https: // biodiversitylibrary. org / page / 47117120]" type="book chapter" year="1914">Eisig, 1914</bibRefCitation>
</taxonomicName>
. Type-species:
<taxonomicName id="3AD6C7AB9662231EDADCFDB0FC16FDA6" authority="Eisig, 1914" authorityName="Eisig" authorityYear="1914" box="[651,957,539,564]" class="Polychaeta" family="Orbiniidae" genus="Scolaricia" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Annelida" rank="species" species="typicus">
<emphasis id="CFA2603A9662231EDADCFDB0FCEAFDA6" box="[651,833,540,564]" italics="true" pageId="28" pageNumber="29">Scolaricia typicus</emphasis>
<bibRefCitation id="9947C1D99662231EDB10FDB7FC16FDA6" author="Eisig, H." box="[839,957,539,564]" pageId="28" pageNumber="29" pagination="153 - 600" refId="ref36564" refString="Eisig, H. (1914) Zur Systematik, Anatomie und Morphologie der Ariciiden nebst Beitragen zur generallen Systematik. Mitteilungen aus der Zoologischen Station Neapel, 21, 153 - 600, pls. 10 - 27, 23 figures. [https: // biodiversitylibrary. org / page / 47117120]" type="book chapter" year="1914">Eisig, 1914</bibRefCitation>
</taxonomicName>
, by monotypy.
<emphasis id="CFA2603A9662231EDC0EFDB0FB2CFDA6" box="[1113,1159,540,564]" italics="true" pageId="28" pageNumber="29">Fide</emphasis>
<bibRefCitation id="9947C1D99662231EDCDAFDB0FB5FFDA6" author="Day, J. H." box="[1165,1268,540,564]" pageId="28" pageNumber="29" pagination="1 - 140" refId="ref36420" refString="Day, J. H. (1973) New Polychaeta from Beaufort, with a key to all species recorded from North Carolina. NOAA Technical Report, National Marine Fisheries Circular, 375, 1 - 140. [https: // repository. library. noaa. gov / view / noaa / 3030] https: // doi. org / 10.5962 / bhl. title. 62852" type="journal article" year="1973">Day 1973</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B5CCEFA39662231ED8C0FDCDFE7BFCC6" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="FD69BC289662231ED8C0FDCDFE7BFCC6" blockId="28.[151,1437,609,1609]" pageId="28" pageNumber="29">
<emphasis id="CFA2603A9662231ED8C0FDCDFEA1FDEE" bold="true" box="[151,266,609,636]" pageId="28" pageNumber="29">Diagnosis</emphasis>
. (Emended). Prostomium pointed, usually prolonged; single achaetous peristomial ring. Branchiae first present from middle or posterior thoracic setigers or from abdominal setigers (826). Posterior thoracic setigers with 02 postsetal lobes and 02 subpodial lobes, never more than four lobes of both
<typeStatus id="226D028A9662231EDC00FD07FB39FD57" box="[1111,1170,683,709]" pageId="28" pageNumber="29">types</typeStatus>
combined; not forming ventral fringes. Thoracic neurosetae including blunt, inconspicuous uncini, few or many in distinct rows; accompanied by few to many crenulated capillaries; furcate setae usually present; heavy spear-like spines and bristle-topped setae absent. Abdominal neuropodia with embedded, non-projecting acicula. Abdominal noto- and or neuropodial flail setae present or absent.
</paragraph>
</subSubSection>
<subSubSection id="B5CCEFA39662231ED890FCF2FAA2F9DA" pageId="28" pageNumber="29" type="discussion">
<paragraph id="FD69BC289662231ED890FCF2FD2FFB73" blockId="28.[151,1437,609,1609]" pageId="28" pageNumber="29">
<emphasis id="CFA2603A9662231ED890FCF2FE90FCEA" bold="true" box="[199,315,862,888]" pageId="28" pageNumber="29">Remarks.</emphasis>
The type-species of
<taxonomicName id="3AD6C7AB9662231EDA7FFCF2FD3EFCEA" authorityName="de Blainville" authorityYear="1828" box="[552,661,862,888]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Annelida" rank="genus">
<emphasis id="CFA2603A9662231EDA7FFCF2FD3EFCEA" box="[552,661,862,888]" italics="true" pageId="28" pageNumber="29">Scoloplos</emphasis>
</taxonomicName>
,
<taxonomicName id="3AD6C7AB9662231EDAF3FCF3FCB0FCEA" authorityName="Muller" authorityYear="1776" box="[676,795,863,889]" class="Polychaeta" family="Orbiniidae" genus="Lumbricus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Annelida" rank="species" species="armiger">
<emphasis id="CFA2603A9662231EDAF3FCF3FCB0FCEA" box="[676,795,863,889]" italics="true" pageId="28" pageNumber="29">S. armiger</emphasis>
</taxonomicName>
has been reported globally despite there being no recent description of the species from the type-locality in
<collectingCountry id="85C1FCB89662231EDA8CFC2FFC9CFC0F" box="[731,823,899,925]" name="Norway" pageId="28" pageNumber="29">Norway</collectingCountry>
. Although most descriptions of
<taxonomicName id="3AD6C7AB9662231EDCF7FC2FFABCFC0E" authorityName="Muller" authorityYear="1776" box="[1184,1303,899,925]" class="Polychaeta" family="Orbiniidae" genus="Lumbricus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Annelida" rank="species" species="armiger">
<emphasis id="CFA2603A9662231EDCF7FC2FFABCFC0E" box="[1184,1303,899,925]" italics="true" pageId="28" pageNumber="29">S. armiger</emphasis>
</taxonomicName>
include the presence of subpodial papillae along the body, there have been two different patterns reported for the neuropodial spines or uncini that occur in thoracic neuropodia. One pattern is where there are numerous rows of uncini and fewer capillaries in thoracic neuropodia (
<bibRefCitation id="9947C1D99662231EDA09FC42FCA8FB9A" author="Hartman, O." box="[606,771,1005,1032]" pageId="28" pageNumber="29" pagination="211 - 393" refId="ref36915" refString="Hartman, O. (1957) Orbiniidae, Apistobranchidae, Paraonidae and Longosomidae. Allan Hancock Pacific Expeditions, 15 (3), 211 - 393, pls. 20 - 44, 1 chart. [https: // biodiversitylibrary. org / page / 4160176]" type="journal article" year="1957">Hartman 1957</bibRefCitation>
;
<bibRefCitation id="9947C1D99662231EDB59FC42FB84FB9B" author="Hartmann-Schroder, G." box="[782,1071,1006,1033]" pageId="28" pageNumber="29" pagination="1 - 648" refId="ref37384" refString="Hartmann-Schroder, G. (1996) Annelida, Borstenwurmer, Polychaeta. Die Tierwelt Deutschlands, 58, 1 - 648. [Second Edition]" type="journal article" year="1996">Hartmann-Schröder 1996</bibRefCitation>
). A second pattern is where there are only a few uncini that typically occur in the ventral-most part of the fascicle and in the first or second setal row (
<bibRefCitation id="9947C1D99662231ED8C8FB9AFEE4FBC2" author="McIntosh, W. C." box="[159,335,1078,1104]" pageId="28" pageNumber="29" pagination="235 - 524" refId="ref38113" refString="McIntosh, W. C. (1910) A Monograph of the British Marine Annelids. Vol. II. Part II. Polychaeta: Syllidae to Arciidae. Ray Society, London, 290 pp., pls. XLIII-LXXXVII. [pp. 235 - 524] [https: // biodiversitylibrary. org / page / 38533042]" type="book chapter" year="1910">McIntosh 1910</bibRefCitation>
;
<bibRefCitation id="9947C1D99662231ED90BFB9AFDA2FBC3" author="Pettibone, M. H." box="[348,521,1078,1105]" pageId="28" pageNumber="29" pagination="1 - 346" refId="ref38950" refString="Pettibone, M. H. (1963) Marine polychaete worms of the New England region. Bulletin of the United States National Museum, 227, 1 - 346. https: // doi. org / 10.5479 / si. 03629236.227.1" type="journal article" year="1963">Pettibone 1963</bibRefCitation>
); capillaries are numerous in this second pattern and the uncini may not occur in all thoracic neuropodia. Both of these patterns have been reported widely and it is not apparent which of these actually refers to
<taxonomicName id="3AD6C7AB9662231ED97DFBD3FE34FB0A" authorityName="Muller" authorityYear="1776" box="[298,415,1151,1177]" class="Polychaeta" family="Orbiniidae" genus="Lumbricus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Annelida" rank="species" species="armiger">
<emphasis id="CFA2603A9662231ED97DFBD3FE34FB0A" box="[298,415,1151,1177]" italics="true" pageId="28" pageNumber="29">S. armiger</emphasis>
</taxonomicName>
from the type-locality in
<collectingCountry id="85C1FCB89662231EDAECFBD3FCBCFB0B" box="[699,791,1151,1177]" name="Norway" pageId="28" pageNumber="29">Norway</collectingCountry>
. Recent ecological investigations and results of molecular sequence data in northern Europe suggest that at least three distinct populations or species of
<taxonomicName id="3AD6C7AB9662231EDCF9FB0EFAB0FB2E" authorityName="de Blainville" authorityYear="1828" box="[1198,1307,1186,1212]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Annelida" rank="genus">
<emphasis id="CFA2603A9662231EDCF9FB0EFAB0FB2E" box="[1198,1307,1186,1212]" italics="true" pageId="28" pageNumber="29">Scoloplos</emphasis>
</taxonomicName>
are present (
<bibRefCitation id="9947C1D99662231ED8C8FB6AFE2CFB72" author="Kruse, I. &amp; Reise, K." box="[159,391,1222,1248]" pageId="28" pageNumber="29" pagination="511 - 517" refId="ref37795" refString="Kruse, I. &amp; Reise, K. (2003) Reproductive isolation between intertidal and subtidal Scoloplos armiger (Polychaeta, Orbiniidae) indicates sibling species in the North Sea. Marine Biology, 143, 511 - 517. https: // doi. org / 10.1007 / s 00227 - 003 - 1112 - x" type="journal article" year="2003">Kruse &amp; Reise 2003</bibRefCitation>
;
<bibRefCitation id="9947C1D99662231ED9C4FB6AFDD3FB73" author="Bleidorn, C. &amp; Kruse, I. &amp; Albrecht, S. &amp; Bartolomaeus, T." box="[403,632,1222,1249]" pageId="28" pageNumber="29" pagination="1 - 13" refId="ref36000" refString="Bleidorn, C., Kruse, I., Albrecht, S. &amp; Bartolomaeus, T. (2006) Mitochondrial sequence data expose the putative cosmopolitan polychaete Scoloplos armiger (Annelida, Orbiniidae) as a species complex. BMC Evolutionary Biology, 6 (47), 1 - 13. https: // doi. org / 10.1186 / 1471 - 2148 - 6 - 47" type="journal article" year="2006">
Bleidorn
<emphasis id="CFA2603A9662231ED9ABFB6BFD86FB72" box="[508,557,1222,1248]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
. 2006
</bibRefCitation>
).
</paragraph>
<paragraph id="FD69BC289662231ED890FB46FAFCFAE3" blockId="28.[151,1437,609,1609]" pageId="28" pageNumber="29">
<bibRefCitation id="9947C1D99662231ED890FB46FE11FA96" author="Bleidorn, C. &amp; Kruse, I. &amp; Albrecht, S. &amp; Bartolomaeus, T." box="[199,442,1258,1285]" pageId="28" pageNumber="29" pagination="1 - 13" refId="ref36000" refString="Bleidorn, C., Kruse, I., Albrecht, S. &amp; Bartolomaeus, T. (2006) Mitochondrial sequence data expose the putative cosmopolitan polychaete Scoloplos armiger (Annelida, Orbiniidae) as a species complex. BMC Evolutionary Biology, 6 (47), 1 - 13. https: // doi. org / 10.1186 / 1471 - 2148 - 6 - 47" type="journal article" year="2006">
Bleidorn
<emphasis id="CFA2603A9662231ED965FB47FECDFA96" box="[306,358,1258,1284]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
. (2006
</bibRefCitation>
,
<bibRefCitation id="9947C1D99662231ED99FFB46FDA9FA96" author="Bleidorn, C. &amp; Hill, N. &amp; Erseus, C. &amp; Tiedemann, R." box="[456,514,1258,1284]" pageId="28" pageNumber="29" pagination="57 - 69" refId="ref36071" refString="Bleidorn, C., Hill, N., Erseus, C. &amp; Tiedemann, R. (2009) On the role of character loss in orbiniid phylogeny (Annelida): Molecules vs. Morphology. Molecular Phylogenetics and Evolution, 52, 57 - 69. https: // doi. org / 10.1016 / j. ympev. 2009.03.022" type="journal article" year="2009">2009</bibRefCitation>
) also provided molecular results on some orbiniids from the eastern Pacific that suggested that specimens identified as
<taxonomicName id="3AD6C7AB9662231EDA07FAA3FD6CFABA" authorityName="Muller" authorityYear="1776" box="[592,711,1295,1321]" class="Polychaeta" family="Orbiniidae" genus="Lumbricus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Annelida" rank="species" species="armiger">
<emphasis id="CFA2603A9662231EDA07FAA3FD6CFABA" box="[592,711,1295,1321]" italics="true" pageId="28" pageNumber="29">S. armiger</emphasis>
</taxonomicName>
from
<collectingRegion id="3F1272CA9662231EDB59FAA2FC2BFABA" box="[782,896,1294,1320]" country="United States of America" name="California" pageId="28" pageNumber="29">California</collectingRegion>
are distinct from the European populations and represent additional undescribed species. To date, none of the various species suggested by these studies in Europe and
<collectingRegion id="3F1272CA9662231ED890FAFAFE92FAE2" box="[199,313,1366,1392]" country="United States of America" name="California" pageId="28" pageNumber="29">California</collectingRegion>
have been described or redescribed, including specimens from the
<typeStatus id="226D028A9662231EDC7DFAFBFBF1FAE3" box="[1066,1114,1367,1393]" pageId="28" pageNumber="29">type</typeStatus>
locality of
<taxonomicName id="3AD6C7AB9662231EDC8DFAFBFAFBFAE2" authorityName="Muller" authorityYear="1776" box="[1242,1360,1367,1393]" class="Polychaeta" family="Orbiniidae" genus="Lumbricus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Annelida" rank="species" species="armiger">
<emphasis id="CFA2603A9662231EDC8DFAFBFAFBFAE2" box="[1242,1360,1367,1393]" italics="true" pageId="28" pageNumber="29">S. armiger</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="FD69BC289662231ED890FAD6FAA2F9DA" blockId="28.[151,1437,609,1609]" pageId="28" pageNumber="29">
Among approximately 28 known species of
<taxonomicName id="3AD6C7AB9662231EDAEAFAD6FC81FA06" authorityName="de Blainville" authorityYear="1828" box="[701,810,1402,1428]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Annelida" rank="genus">
<emphasis id="CFA2603A9662231EDAEAFAD6FC81FA06" box="[701,810,1402,1428]" italics="true" pageId="28" pageNumber="29">Scoloplos</emphasis>
</taxonomicName>
, only three have been recorded from deep-sea habitats:
<taxonomicName id="3AD6C7AB9662231ED8C0FA33FE25FA2A" authority="Blake, 1985" authorityName="Blake" authorityYear="1985" box="[151,398,1437,1465]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Annelida" rank="species" species="ehlersi">
<emphasis id="CFA2603A9662231ED8C0FA33FF54FA2A" box="[151,255,1438,1465]" italics="true" pageId="28" pageNumber="29">S. ehlersi</emphasis>
<bibRefCitation id="9947C1D99662231ED950FA32FE25FA2A" author="Blake, J. A." box="[263,398,1437,1464]" pageId="28" pageNumber="29" pagination="67 - 101" refId="ref35396" refString="Blake, J. A. (1985) Polychaeta from the vicinity of deep-sea geothermal vents in the eastern Pacific. I: Euphrosinidae, Phyllodocidae, Hesionidae, Nereididae, Glyceridae, Dorvilleidae, Orbiniidae and Maldanidae. In: Jones, M. L. (Ed.), The Hydrothermal Vents of the Eastern Pacific: An Overview. Bulletin of the Biological Society of Washington, No. 6, pp. 67 - 101. [https: // www. researchgate. net / publication / 279962731 _ Polychaeta _ from _ the _ vicinity _ of _ deep-sea _ geothermal _ vents _ in _ the _ eastern _ Pacific _ I _ Euphrosinidae _ Phyllodocidae _ Hesionidae _ Nereididae _ Glyceridae _ Dorvilleidae _ Orbiniidae _ and _ Maldanidae]" type="book chapter" year="1985">Blake, 1985</bibRefCitation>
</taxonomicName>
from the Galápagos Rift hydrothermal mounds,
<taxonomicName id="3AD6C7AB9662231EDBE5FA33FB79FA2A" authority="Blake, 2017" authorityName="Blake" authorityYear="2017" box="[946,1234,1438,1465]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Annelida" rank="species" species="bathytatus">
<emphasis id="CFA2603A9662231EDBE5FA33FBE8FA2A" box="[946,1091,1438,1465]" italics="true" pageId="28" pageNumber="29">S. bathytatus</emphasis>
<bibRefCitation id="9947C1D99662231EDC1DFA32FB79FA2A" author="Blake, J. A." box="[1098,1234,1438,1464]" pageId="28" pageNumber="29" pagination="1 - 145" refId="ref35753" refString="Blake, J. A. (2017) Polychaeta Orbiniidae from Antarctica, the Southern Ocean, the abyssal Pacific Ocean, and off South Amer- ica. Zootaxa, 4218 (1), 1 - 145. https: // doi. org / 10.11646 / zootaxa. 4218.1.1" type="journal article" year="2017">Blake, 2017</bibRefCitation>
</taxonomicName>
from the Weddell Sea,
<collectingCountry id="85C1FCB89662231ED89CFA6EFEE8FA4E" box="[203,323,1474,1500]" name="Antarctica" pageId="28" pageNumber="29">Antarctica</collectingCountry>
, and
<taxonomicName id="3AD6C7AB9662231ED92CFA6FFD71FA4E" authority="(Hartman, 1965)" baseAuthorityName="Hartman" baseAuthorityYear="1965" box="[379,730,1473,1501]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Annelida" rank="species" species="intermedius">
<emphasis id="CFA2603A9662231ED92CFA6FFDB1FA4E" box="[379,538,1474,1501]" italics="true" pageId="28" pageNumber="29">S. intermedius</emphasis>
(
<bibRefCitation id="9947C1D99662231EDA7FFA6EFD7AFA4E" author="Hartman, O." box="[552,721,1473,1500]" pageId="28" pageNumber="29" pagination="1 - 378" refId="ref37054" refString="Hartman, O. (1965) Deep-water benthic polychaetous annelids off New England to Bermuda and other North Atlantic areas. Allan Hancock Foundation Occasional Paper, 28, 1 - 378. [http: // digitallibrary. usc. edu / cdm / ref / collection / p 15799 coll 82 / id / 20299]" type="journal article" year="1965">Hartman, 1965</bibRefCitation>
)
</taxonomicName>
from the western North Atlantic. Two new species,
<taxonomicName id="3AD6C7AB9662231EDD40FA6FFF64F992" authority="Blake, 2020" authorityName="Blake" authorityYear="2020" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Annelida" rank="species" species="californiensis" status="sp. nov.">
<emphasis id="CFA2603A9662231EDD40FA6FFA8EFA4F" box="[1303,1317,1475,1501]" italics="true" pageId="28" pageNumber="29">S</emphasis>
.
<emphasis id="CFA2603A9662231EDD65FA6EFF64F992" italics="true" pageId="28" pageNumber="29">californiensis</emphasis>
</taxonomicName>
<emphasis id="CFA2603A9662231ED880FA4AFEBCF992" bold="true" box="[215,279,1510,1536]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="D491DD419662231ED880FA4AFEBCF992" box="[215,279,1510,1536]" pageId="28" pageNumber="29" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
from off northern California and
<taxonomicName id="3AD6C7AB9662231EDACFFA4BFCE4F992" authority="Blake, 2020" authorityName="Blake" authorityYear="2020" box="[664,847,1510,1537]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Annelida" rank="species" species="sparsaciculus" status="sp. nov.">
<emphasis id="CFA2603A9662231EDACFFA4BFCE4F992" box="[664,847,1510,1537]" italics="true" pageId="28" pageNumber="29">S. sparsaciculus</emphasis>
</taxonomicName>
<emphasis id="CFA2603A9662231EDB00FA4AFC3CF992" bold="true" box="[855,919,1510,1536]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="D491DD419662231EDB00FA4AFC3CF992" box="[855,919,1510,1536]" pageId="28" pageNumber="29" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
from off
<collectingCountry id="85C1FCB89662231EDC51FA4AFBF9F992" box="[1030,1106,1510,1536]" name="Brunei" pageId="28" pageNumber="29">Brunei</collectingCountry>
in the South
<collectingCountry id="85C1FCB89662231EDCBBFA4AFA84F992" box="[1260,1327,1510,1536]" name="China" pageId="28" pageNumber="29">China</collectingCountry>
Sea have been discovered in the deep-water samples treated in the present study and are described. Both of these new deepwater species belong to the group having a reduced number of thoracic uncini in thoracic neuropodia.
</paragraph>
</subSubSection>
</treatment>
</document>