329 lines
44 KiB
XML
329 lines
44 KiB
XML
<document id="84D69C055BEC3F560930CEA72A31735F" ID-CLB-Dataset="24204" ID-DOI="10.11646/zootaxa.4730.1.1" ID-GBIF-Dataset="bcc0e0fc-b6a5-4d5a-8008-7b714c106a7c" ID-ISSN="1175-5326" ID-Zenodo-Dep="3634270" ID-ZooBank="7F4CD129-9FF9-4593-A8A4-DB999B3E402F" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1580713876401" checkinUser="plazi" docAuthor="Blake, James A." docDate="2020" docId="757F0D3E965C2326D8C0FAB5FBB5FAC5" docLanguage="en" docName="zootaxa.4730.1.1.pdf" docOrigin="Zootaxa 4730 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Leodamas bathyalis Blake 2020, new species" docType="treatment" docUuid="981586BA-A3D9-4383-8856-1AB2BBAD8913" docUuidSource="ZooBank" docVersion="8" lastPageNumber="37" masterDocId="89467546967E2302D857FFACFFABFF92" masterDocTitle="New species and records of deep-water Orbiniidae (Annelida, Polychaeta) from the Eastern Pacific continental slope, abyssal Pacific Ocean, and the South China Sea" masterLastPageNumber="61" masterPageNumber="1" pageNumber="35" updateTime="1698780308397" updateUser="ExternalLinkService">
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<mods:title id="D8C65716118BA6EA4BA932569D97078D">New species and records of deep-water Orbiniidae (Annelida, Polychaeta) from the Eastern Pacific continental slope, abyssal Pacific Ocean, and the South China Sea</mods:title>
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<mods:namePart id="2883025B345D1E08561FA5F4FCDA0FF6">Blake, James A.</mods:namePart>
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<mods:title id="5AD4CFA3410E812CB5F1E52C6AA35739">Zootaxa</mods:title>
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<mods:part id="5DBB59162C88B720B40081EFF210493F">
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<mods:date id="F60EB21098DD3E9AD8DBF980E8251D09">2020</mods:date>
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<taxonomicName id="3AD6C7AB965C2320D8C0FAB5FE2CFAA6" ID-CoL="85RHL" authority="Blake, 2020" authorityName="Blake" authorityYear="2020" box="[151,391,1305,1332]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="34" pageNumber="35" phylum="Annelida" rank="species" species="bathyalis" status="sp. nov.">
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<emphasis id="CFA2603A965C2320D8C0FAB5FE2CFAA6" bold="true" box="[151,391,1305,1332]" italics="true" pageId="34" pageNumber="35">Leodamas bathyalis</emphasis>
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, box corer,
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(
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;
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Sta. 28,
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|
||
<specimenCount id="EBD077A1965C2320D8ECF9A2FE81F9BA" box="[187,298,1550,1576]" pageId="34" pageNumber="35" type="juvenile">1 juvenile</specimenCount>
|
||
specimen (
|
||
<specimenCode id="AD701453965C2320D9FEF9A2FDE3F9BA" box="[425,584,1549,1576]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="34" pageNumber="35" type="Museum">MCZ 153571</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="FD69BC28965C2320DA07F9A1FDFFF9BA" blockId="34.[151,1437,1441,2045]" box="[592,596,1549,1576]" pageId="34" pageNumber="35">.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B5CCEFA3965C2326D890F99DFB5AFBA5" lastPageId="36" lastPageNumber="37" pageId="34" pageNumber="35" type="description">
|
||
<paragraph id="FD69BC28965C2320D890F99DFCB3F8B7" blockId="34.[151,1437,1441,2045]" pageId="34" pageNumber="35">
|
||
<emphasis id="CFA2603A965C2320D890F99DFEFAF9DE" bold="true" box="[199,337,1585,1612]" pageId="34" pageNumber="35">Description</emphasis>
|
||
. A moderately sized species, all specimens incomplete.
|
||
<typeStatus id="226D028A965C2320DBE9F99EFB8DF9DE" box="[958,1062,1586,1612]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
|
||
largest, 32 setigers,
|
||
<quantity id="3A2E11CD965C2320DD5EF99EFAF5F9DF" box="[1289,1374,1586,1613]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="34" pageNumber="35" unit="mm" value="6.7">6.7 mm</quantity>
|
||
long,
|
||
<quantity id="3A2E11CD965C2320D8C0F9FAFF51F9E3" box="[151,250,1622,1649]" metricMagnitude="-4" metricUnit="m" metricValue="8.7" pageId="34" pageNumber="35" unit="mm" value="0.87">0.87 mm</quantity>
|
||
wide across thoracic segments;
|
||
<typeStatus id="226D028A965C2320DA09F9FBFD15F9E3" box="[606,702,1623,1649]" pageId="34" pageNumber="35" type="paratype">paratype</typeStatus>
|
||
23 setigers,
|
||
<quantity id="3A2E11CD965C2320DB1FF9FAFC36F9E3" box="[840,925,1622,1649]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="34" pageNumber="35" unit="mm" value="2.3">2.3 mm</quantity>
|
||
long,
|
||
<quantity id="3A2E11CD965C2320DBB6F9FAFBEFF9E3" box="[993,1092,1622,1649]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="34" pageNumber="35" unit="mm" value="0.44">0.44 mm</quantity>
|
||
wide; juvenile, 13 setigers,
|
||
<quantity id="3A2E11CD965C2320DD2EF9FAFF68F907" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="34" pageNumber="35" unit="mm" value="1.4">1.4 mm</quantity>
|
||
long,
|
||
<quantity id="3A2E11CD965C2320D95FF9D6FEF6F907" box="[264,349,1658,1685]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="34" pageNumber="35" unit="mm" value="0.3">0.3 mm</quantity>
|
||
wide. Body with narrow thoracic segments (
|
||
<figureCitation id="65EDA0AD965C2320DB1AF9D6FC1FF906" box="[845,948,1658,1684]" captionStart="FIGURES 17" captionStartId="35.[151,264,1945,1970]" captionTargetBox="[206,1376,189,1905]" captionTargetId="figure@35.[199,1389,181,1920]" captionTargetPageId="35" captionText="FIGURES 17. Leodamas bathyalis n. sp. A, anterior end dorsal view; B, thoracic setiger 9, right, posterior view; C, middle abdominal setiger, anterior view; D, neuropodial uncini (holotype); E, neuropodial uncini (paratype); F, furcate setae. A–D, F, holotype (MCZ 153569); E, paratype (MCZ 153700)." figureDoi="http://doi.org/10.5281/zenodo.3634304" httpUri="https://zenodo.org/record/3634304/files/figure.png" pageId="34" pageNumber="35">Fig. 17A</figureCitation>
|
||
), flattened dorsally, rounded on venter, with lateral parapodia; thoracic segments about seven times wider than long, with prominent intersegmental furrow (
|
||
<figureCitation id="65EDA0AD965C2320DD38F932FF67F94E" captionStart="FIGURES 17" captionStartId="35.[151,264,1945,1970]" captionTargetBox="[206,1376,189,1905]" captionTargetId="figure@35.[199,1389,181,1920]" captionTargetPageId="35" captionText="FIGURES 17. Leodamas bathyalis n. sp. A, anterior end dorsal view; B, thoracic setiger 9, right, posterior view; C, middle abdominal setiger, anterior view; D, neuropodial uncini (holotype); E, neuropodial uncini (paratype); F, furcate setae. A–D, F, holotype (MCZ 153569); E, paratype (MCZ 153700)." figureDoi="http://doi.org/10.5281/zenodo.3634304" httpUri="https://zenodo.org/record/3634304/files/figure.png" pageId="34" pageNumber="35">Fig. 17A</figureCitation>
|
||
). abdominal segments narrower, with parapodia dorsally elevated. Number of thoracic segments size related:
|
||
<typeStatus id="226D028A965C2320D8C0F94AFF52F892" box="[151,249,1766,1792]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
|
||
with 13 thoracic setigers, smaller
|
||
<typeStatus id="226D028A965C2320DA20F94BFD7CF893" box="[631,727,1767,1793]" pageId="34" pageNumber="35" type="paratype">paratype</typeStatus>
|
||
with 10 thoracic setigers; juvenile with six thoracic setigers, all with branchiae from setiger 6. Color in alcohol: light tan.
|
||
</paragraph>
|
||
<paragraph id="FD69BC28965C2320D890F882FE75F826" blockId="34.[151,1437,1441,2045]" pageId="34" pageNumber="35">
|
||
Pre-setiger region triangular. Prostomium narrowing to rounded tip (
|
||
<figureCitation id="65EDA0AD965C2320DB83F882FB95F8DA" box="[980,1086,1838,1864]" captionStart="FIGURES 17" captionStartId="35.[151,264,1945,1970]" captionTargetBox="[206,1376,189,1905]" captionTargetId="figure@35.[199,1389,181,1920]" captionTargetPageId="35" captionText="FIGURES 17. Leodamas bathyalis n. sp. A, anterior end dorsal view; B, thoracic setiger 9, right, posterior view; C, middle abdominal setiger, anterior view; D, neuropodial uncini (holotype); E, neuropodial uncini (paratype); F, furcate setae. A–D, F, holotype (MCZ 153569); E, paratype (MCZ 153700)." figureDoi="http://doi.org/10.5281/zenodo.3634304" httpUri="https://zenodo.org/record/3634304/files/figure.png" pageId="34" pageNumber="35">Fig. 17A</figureCitation>
|
||
); nuchal organs distinct lateral grooves (
|
||
<figureCitation id="65EDA0AD965C2320D8A8F8FEFECCF8FE" box="[255,359,1874,1900]" captionStart="FIGURES 17" captionStartId="35.[151,264,1945,1970]" captionTargetBox="[206,1376,189,1905]" captionTargetId="figure@35.[199,1389,181,1920]" captionTargetPageId="35" captionText="FIGURES 17. Leodamas bathyalis n. sp. A, anterior end dorsal view; B, thoracic setiger 9, right, posterior view; C, middle abdominal setiger, anterior view; D, neuropodial uncini (holotype); E, neuropodial uncini (paratype); F, furcate setae. A–D, F, holotype (MCZ 153569); E, paratype (MCZ 153700)." figureDoi="http://doi.org/10.5281/zenodo.3634304" httpUri="https://zenodo.org/record/3634304/files/figure.png" pageId="34" pageNumber="35">Fig. 17A</figureCitation>
|
||
); eyespots absent. Peristomium longer, but narrower than first setiger, with a single annular ring (
|
||
<figureCitation id="65EDA0AD965C2320D8F7F8DAFEADF802" box="[160,262,1910,1936]" captionStart="FIGURES 17" captionStartId="35.[151,264,1945,1970]" captionTargetBox="[206,1376,189,1905]" captionTargetId="figure@35.[199,1389,181,1920]" captionTargetPageId="35" captionText="FIGURES 17. Leodamas bathyalis n. sp. A, anterior end dorsal view; B, thoracic setiger 9, right, posterior view; C, middle abdominal setiger, anterior view; D, neuropodial uncini (holotype); E, neuropodial uncini (paratype); F, furcate setae. A–D, F, holotype (MCZ 153569); E, paratype (MCZ 153700)." figureDoi="http://doi.org/10.5281/zenodo.3634304" httpUri="https://zenodo.org/record/3634304/files/figure.png" pageId="34" pageNumber="35">Fig. 17A</figureCitation>
|
||
) bearing dorsolateral grooves suggesting a simple dorsal crest. Proboscis everted on
|
||
<typeStatus id="226D028A965C2320DC9FF8DAFA86F802" box="[1224,1325,1910,1936]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
|
||
, dendritic with numerous narrow lobes.
|
||
</paragraph>
|
||
<paragraph id="FD69BC28965C2326D890F812FEB8FED1" blockId="34.[151,1437,1441,2045]" lastBlockId="36.[151,1437,153,1367]" lastPageId="36" lastPageNumber="37" pageId="34" pageNumber="35">
|
||
Thoracic notopodia low mounds, with long, narrow, digitiform postsetal lobe (
|
||
<figureCitation id="65EDA0AD965C2320DC10F812FB33F84A" box="[1095,1176,1982,2008]" captionStart="FIGURES 17" captionStartId="35.[151,264,1945,1970]" captionTargetBox="[206,1376,189,1905]" captionTargetId="figure@35.[199,1389,181,1920]" captionTargetPageId="35" captionText="FIGURES 17. Leodamas bathyalis n. sp. A, anterior end dorsal view; B, thoracic setiger 9, right, posterior view; C, middle abdominal setiger, anterior view; D, neuropodial uncini (holotype); E, neuropodial uncini (paratype); F, furcate setae. A–D, F, holotype (MCZ 153569); E, paratype (MCZ 153700)." figureDoi="http://doi.org/10.5281/zenodo.3634304" httpUri="https://zenodo.org/record/3634304/files/figure.png" pageId="34" pageNumber="35">Fig. 17</figureCitation>
|
||
A–B). Thoracic neuropodia broadly swollen, bearing up to three short postsetal lobes (
|
||
<figureCitation id="65EDA0AD965C2320DB3FF84EFC65F86E" box="[872,974,2018,2044]" captionStart="FIGURES 17" captionStartId="35.[151,264,1945,1970]" captionTargetBox="[206,1376,189,1905]" captionTargetId="figure@35.[199,1389,181,1920]" captionTargetPageId="35" captionText="FIGURES 17. Leodamas bathyalis n. sp. A, anterior end dorsal view; B, thoracic setiger 9, right, posterior view; C, middle abdominal setiger, anterior view; D, neuropodial uncini (holotype); E, neuropodial uncini (paratype); F, furcate setae. A–D, F, holotype (MCZ 153569); E, paratype (MCZ 153700)." figureDoi="http://doi.org/10.5281/zenodo.3634304" httpUri="https://zenodo.org/record/3634304/files/figure.png" pageId="34" pageNumber="35">Fig. 17B</figureCitation>
|
||
); initially a single lobe increasing to two, and then three by setiger
|
||
<quantity id="3A2E11CD965A2326D9E8FF35FE46FF21" box="[447,493,153,179]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="36" pageNumber="37" unit="in" value="7.0">7 in</quantity>
|
||
<typeStatus id="226D028A965A2326D9A1FF35FDF7FF21" box="[502,604,153,179]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
|
||
;
|
||
<typeStatus id="226D028A965A2326DA3EFF35FD62FF21" box="[617,713,153,179]" pageId="36" pageNumber="37" type="paratype">paratype</typeStatus>
|
||
with up to two thoracic postsetal lobes; juvenile with a single thoracic postsetal lobe. Transition to abdominal segments abrupt, with fewer noto- and neurosetae. Notopodium of abdominal segments similar to those of thoracic setigers with long, digitiform postsetal lobe (
|
||
<figureCitation id="65EDA0AD965A2326DCEDFF4DFA8AFF69" box="[1210,1313,225,251]" captionStart="FIGURES 17" captionStartId="35.[151,264,1945,1970]" captionTargetBox="[206,1376,189,1905]" captionTargetId="figure@35.[199,1389,181,1920]" captionTargetPageId="35" captionText="FIGURES 17. Leodamas bathyalis n. sp. A, anterior end dorsal view; B, thoracic setiger 9, right, posterior view; C, middle abdominal setiger, anterior view; D, neuropodial uncini (holotype); E, neuropodial uncini (paratype); F, furcate setae. A–D, F, holotype (MCZ 153569); E, paratype (MCZ 153700)." figureDoi="http://doi.org/10.5281/zenodo.3634304" httpUri="https://zenodo.org/record/3634304/files/figure.png" pageId="36" pageNumber="37">Fig. 17C</figureCitation>
|
||
); neuropodium becoming narrow, elongate, shifted dorsally; with narrow rounded setal lobe and a short lateral ventral cirrus (
|
||
<figureCitation id="65EDA0AD965A2326D8F7FE85FEADFED1" box="[160,262,297,323]" captionStart="FIGURES 17" captionStartId="35.[151,264,1945,1970]" captionTargetBox="[206,1376,189,1905]" captionTargetId="figure@35.[199,1389,181,1920]" captionTargetPageId="35" captionText="FIGURES 17. Leodamas bathyalis n. sp. A, anterior end dorsal view; B, thoracic setiger 9, right, posterior view; C, middle abdominal setiger, anterior view; D, neuropodial uncini (holotype); E, neuropodial uncini (paratype); F, furcate setae. A–D, F, holotype (MCZ 153569); E, paratype (MCZ 153700)." figureDoi="http://doi.org/10.5281/zenodo.3634304" httpUri="https://zenodo.org/record/3634304/files/figure.png" pageId="36" pageNumber="37">Fig. 17C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="A9A9ECA0965D2321D8C0F835FD12F868" ID-DOI="http://doi.org/10.5281/zenodo.3634304" ID-Zenodo-Dep="3634304" httpUri="https://zenodo.org/record/3634304/files/figure.png" pageId="35" pageNumber="36" startId="35.[151,264,1945,1970]" targetBox="[206,1376,189,1905]" targetPageId="35">
|
||
<paragraph id="FD69BC28965D2321D8C0F835FD12F868" blockId="35.[151,1436,1945,2042]" pageId="35" pageNumber="36">
|
||
<emphasis id="CFA2603A965D2321D8C0F835FE84F820" bold="true" box="[151,303,1945,1970]" pageId="35" pageNumber="36">FIGURES 17.</emphasis>
|
||
<taxonomicName id="3AD6C7AB965D2321D960F836FDAFF820" authority="Blake, 2020" authorityName="Blake" authorityYear="2020" box="[311,516,1946,1970]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="35" pageNumber="36" phylum="Annelida" rank="species" species="bathyalis" status="sp. nov.">
|
||
<emphasis id="CFA2603A965D2321D960F836FDAFF820" box="[311,516,1946,1970]" italics="true" pageId="35" pageNumber="36">Leodamas bathyalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CFA2603A965D2321DA5BF836FDECF820" bold="true" box="[524,583,1946,1970]" pageId="35" pageNumber="36">
|
||
<taxonomicNameLabel id="D491DD41965D2321DA5BF836FDECF820" box="[524,583,1946,1970]" pageId="35" pageNumber="36" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A, anterior end dorsal view; B, thoracic setiger 9, right, posterior view; C, middle abdominal setiger, anterior view; D, neuropodial uncini (holotype); E, neuropodial uncini (paratype); F, furcate setae. A–D, F, holotype (MCZ 153569); E, paratype (MCZ 153700).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FD69BC28965A2326D890FEE1FDA6FE3D" blockId="36.[151,1437,153,1367]" pageId="36" pageNumber="37">
|
||
Branchiae from sixth thoracic setiger (
|
||
<figureCitation id="65EDA0AD965A2326DA2DFEE1FD49FEF5" box="[634,738,333,359]" captionStart="FIGURES 17" captionStartId="35.[151,264,1945,1970]" captionTargetBox="[206,1376,189,1905]" captionTargetId="figure@35.[199,1389,181,1920]" captionTargetPageId="35" captionText="FIGURES 17. Leodamas bathyalis n. sp. A, anterior end dorsal view; B, thoracic setiger 9, right, posterior view; C, middle abdominal setiger, anterior view; D, neuropodial uncini (holotype); E, neuropodial uncini (paratype); F, furcate setae. A–D, F, holotype (MCZ 153569); E, paratype (MCZ 153700)." figureDoi="http://doi.org/10.5281/zenodo.3634304" httpUri="https://zenodo.org/record/3634304/files/figure.png" pageId="36" pageNumber="37">Fig. 17A</figureCitation>
|
||
), continuing along body through posterior setigers. Branchiae of thoracic setigers narrow, triangular in shape (
|
||
<figureCitation id="65EDA0AD965A2326DAFEFEDDFCA4FE19" box="[681,783,369,395]" captionStart="FIGURES 17" captionStartId="35.[151,264,1945,1970]" captionTargetBox="[206,1376,189,1905]" captionTargetId="figure@35.[199,1389,181,1920]" captionTargetPageId="35" captionText="FIGURES 17. Leodamas bathyalis n. sp. A, anterior end dorsal view; B, thoracic setiger 9, right, posterior view; C, middle abdominal setiger, anterior view; D, neuropodial uncini (holotype); E, neuropodial uncini (paratype); F, furcate setae. A–D, F, holotype (MCZ 153569); E, paratype (MCZ 153700)." figureDoi="http://doi.org/10.5281/zenodo.3634304" httpUri="https://zenodo.org/record/3634304/files/figure.png" pageId="36" pageNumber="37">Fig. 17B</figureCitation>
|
||
); abdominal branchiae becoming larger, thickened basally, tapering to narrow tip (
|
||
<figureCitation id="65EDA0AD965A2326D9CDFE39FDABFE3D" box="[410,512,405,431]" captionStart="FIGURES 17" captionStartId="35.[151,264,1945,1970]" captionTargetBox="[206,1376,189,1905]" captionTargetId="figure@35.[199,1389,181,1920]" captionTargetPageId="35" captionText="FIGURES 17. Leodamas bathyalis n. sp. A, anterior end dorsal view; B, thoracic setiger 9, right, posterior view; C, middle abdominal setiger, anterior view; D, neuropodial uncini (holotype); E, neuropodial uncini (paratype); F, furcate setae. A–D, F, holotype (MCZ 153569); E, paratype (MCZ 153700)." figureDoi="http://doi.org/10.5281/zenodo.3634304" httpUri="https://zenodo.org/record/3634304/files/figure.png" pageId="36" pageNumber="37">Fig. 17C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FD69BC28965A2326D890FE15FCA4FC85" blockId="36.[151,1437,153,1367]" pageId="36" pageNumber="37">
|
||
Thoracic notosetae numerous camerated capillaries arranged in 3–4 rows. Thoracic neurosetae include heavy spines or uncini among numerous camerated capillaries (
|
||
<figureCitation id="65EDA0AD965A2326DB45FE71FCD3FE65" box="[786,888,477,503]" captionStart="FIGURES 17" captionStartId="35.[151,264,1945,1970]" captionTargetBox="[206,1376,189,1905]" captionTargetId="figure@35.[199,1389,181,1920]" captionTargetPageId="35" captionText="FIGURES 17. Leodamas bathyalis n. sp. A, anterior end dorsal view; B, thoracic setiger 9, right, posterior view; C, middle abdominal setiger, anterior view; D, neuropodial uncini (holotype); E, neuropodial uncini (paratype); F, furcate setae. A–D, F, holotype (MCZ 153569); E, paratype (MCZ 153700)." figureDoi="http://doi.org/10.5281/zenodo.3634304" httpUri="https://zenodo.org/record/3634304/files/figure.png" pageId="36" pageNumber="37">Fig. 17B</figureCitation>
|
||
). Uncini in two rows, with first row curving ventrally and joining second row in a U-shaped pattern. Capillaries of first four segments reduced to a fascicle dorsal to uncini; subsequent setigers with multiple rows of numerous capillaries accompanying uncini through thoracic setigers. Uncini of larger
|
||
<typeStatus id="226D028A965A2326D9CBFDE5FE55FDF1" box="[412,510,585,611]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
|
||
thick, with blunt tip covered with a thin cloak of fibrils, shaft with transverse rows of ribs worn down to low ridge (
|
||
<figureCitation id="65EDA0AD965A2326D9A9FDC1FDCEFD15" box="[510,613,621,647]" captionStart="FIGURES 17" captionStartId="35.[151,264,1945,1970]" captionTargetBox="[206,1376,189,1905]" captionTargetId="figure@35.[199,1389,181,1920]" captionTargetPageId="35" captionText="FIGURES 17. Leodamas bathyalis n. sp. A, anterior end dorsal view; B, thoracic setiger 9, right, posterior view; C, middle abdominal setiger, anterior view; D, neuropodial uncini (holotype); E, neuropodial uncini (paratype); F, furcate setae. A–D, F, holotype (MCZ 153569); E, paratype (MCZ 153700)." figureDoi="http://doi.org/10.5281/zenodo.3634304" httpUri="https://zenodo.org/record/3634304/files/figure.png" pageId="36" pageNumber="37">Fig. 17D</figureCitation>
|
||
); smaller
|
||
<typeStatus id="226D028A965A2326DA85FDC1FC99FD15" box="[722,818,621,647]" pageId="36" pageNumber="37" type="paratype">paratype</typeStatus>
|
||
with uncini narrower with transverse rows of ribs more prominent, resembling low barbs (
|
||
<figureCitation id="65EDA0AD965A2326DA43FD3DFDD3FD39" box="[532,632,657,683]" captionStart="FIGURES 17" captionStartId="35.[151,264,1945,1970]" captionTargetBox="[206,1376,189,1905]" captionTargetId="figure@35.[199,1389,181,1920]" captionTargetPageId="35" captionText="FIGURES 17. Leodamas bathyalis n. sp. A, anterior end dorsal view; B, thoracic setiger 9, right, posterior view; C, middle abdominal setiger, anterior view; D, neuropodial uncini (holotype); E, neuropodial uncini (paratype); F, furcate setae. A–D, F, holotype (MCZ 153569); E, paratype (MCZ 153700)." figureDoi="http://doi.org/10.5281/zenodo.3634304" httpUri="https://zenodo.org/record/3634304/files/figure.png" pageId="36" pageNumber="37">Fig. 17E</figureCitation>
|
||
). Abdominal notosetae 2–3 long, relatively smooth capillaries with short barbs along with one edge and 1–2 furcate setae; flail setae absent. Furcate setae with unequal, blunt-tipped tynes, connected by thin web of stiff fibrils (
|
||
<figureCitation id="65EDA0AD965A2326DA14FD75FD0CFD61" box="[579,679,729,755]" captionStart="FIGURES 17" captionStartId="35.[151,264,1945,1970]" captionTargetBox="[206,1376,189,1905]" captionTargetId="figure@35.[199,1389,181,1920]" captionTargetPageId="35" captionText="FIGURES 17. Leodamas bathyalis n. sp. A, anterior end dorsal view; B, thoracic setiger 9, right, posterior view; C, middle abdominal setiger, anterior view; D, neuropodial uncini (holotype); E, neuropodial uncini (paratype); F, furcate setae. A–D, F, holotype (MCZ 153569); E, paratype (MCZ 153700)." figureDoi="http://doi.org/10.5281/zenodo.3634304" httpUri="https://zenodo.org/record/3634304/files/figure.png" pageId="36" pageNumber="37">Fig. 17F</figureCitation>
|
||
). Abdominal neurosetae with 2–3 long, smooth capillaries; internal acicula present but not emergent in available specimens.
|
||
</paragraph>
|
||
<paragraph id="FD69BC28965A2326D890FC8DFE78FCA9" blockId="36.[151,1437,153,1367]" box="[199,467,801,827]" pageId="36" pageNumber="37">Pygidium not observed.</paragraph>
|
||
<paragraph id="FD69BC28965A2326D890FCE8FC16FC59" blockId="36.[151,1437,153,1367]" pageId="36" pageNumber="37">
|
||
<emphasis id="CFA2603A965A2326D890FCE8FEECFCCD" bold="true" box="[199,327,836,863]" pageId="36" pageNumber="37">Variability</emphasis>
|
||
. The three specimens available represent a size range with the most obvious difference being the number of thoracic setigers:
|
||
<quantity id="3A2E11CD965A2326D9B3FCC5FDB9FC11" box="[484,530,873,899]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="36" pageNumber="37" unit="in" value="6.0">6 in</quantity>
|
||
the small juvenile,
|
||
<quantity id="3A2E11CD965A2326DAAEFCC5FC9EFC11" box="[761,821,873,899]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="36" pageNumber="37" unit="in" value="10.0">10 in</quantity>
|
||
the
|
||
<typeStatus id="226D028A965A2326DB3CFCC5FC60FC11" box="[875,971,873,899]" pageId="36" pageNumber="37" type="paratype">paratype</typeStatus>
|
||
and
|
||
<quantity id="3A2E11CD965A2326DC5FFCC5FBEFFC11" box="[1032,1092,873,899]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="36" pageNumber="37" unit="in" value="13.0">13 in</quantity>
|
||
the larger
|
||
<typeStatus id="226D028A965A2326DC92FCC5FA80FC11" box="[1221,1323,873,899]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
|
||
. All three specimens, however, have branchiae first present from setiger 6. These results suggest that the number of thoracic setigers is size-related, but the first occurrence of branchiae is invariant.
|
||
</paragraph>
|
||
<paragraph id="FD69BC28965A2326D890FC79FDF6FB81" blockId="36.[151,1437,153,1367]" pageId="36" pageNumber="37">
|
||
The thoracic uncini of the
|
||
<typeStatus id="226D028A965A2326D9BAFC79FDE6FC7D" box="[493,589,981,1007]" pageId="36" pageNumber="37" type="paratype">paratype</typeStatus>
|
||
and juvenile exhibit distinct low barbs along the shaft; these are worn down and barely visible in the larger
|
||
<typeStatus id="226D028A965A2326D9A4FC55FDF2FB81" box="[499,601,1017,1043]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="FD69BC28965A2326D890FBB1FB5AFBA5" blockId="36.[151,1437,153,1367]" box="[199,1265,1052,1079]" pageId="36" pageNumber="37">
|
||
<emphasis id="CFA2603A965A2326D890FBB1FE04FBA5" bold="true" box="[199,431,1052,1079]" pageId="36" pageNumber="37">Methyl Green stain</emphasis>
|
||
. Prostomium not staining; rest of body stains uniformly; de-stains rapidly.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B5CCEFA3965A2326D890FBEDFA21FA9D" pageId="36" pageNumber="37" type="discussion">
|
||
<paragraph id="FD69BC28965A2326D890FBEDFA21FA9D" blockId="36.[151,1437,153,1367]" pageId="36" pageNumber="37">
|
||
<emphasis id="CFA2603A965A2326D890FBEDFE9FFBC9" bold="true" box="[199,308,1089,1115]" pageId="36" pageNumber="37">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="3AD6C7AB965A2326D913FBEDFD8CFBC9" authority="Blake, 2020" authorityName="Blake" authorityYear="2020" box="[324,551,1089,1115]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Annelida" rank="species" species="bathyalis" status="sp. nov.">
|
||
<emphasis id="CFA2603A965A2326D913FBEDFD8CFBC9" box="[324,551,1089,1115]" italics="true" pageId="36" pageNumber="37">Leodamas bathyalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CFA2603A965A2326DA78FBEDFDDBFBC9" bold="true" box="[559,624,1089,1115]" pageId="36" pageNumber="37">
|
||
<taxonomicNameLabel id="D491DD41965A2326DA78FBEDFDDBFBC9" box="[559,624,1089,1115]" pageId="36" pageNumber="37" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is a distinctive species in the nature of the thoracic neuropodial uncini and the presence of up to three short postsetal lobes in thoracic neuropodia. The reduced number of capillaries in the first 3–4 thoracic neuropodia accentuates the visibility of the uncini, which are more visible than in the following setigers where they are obscured by the numerous capillaries. Superficially the more obvious neuropodial uncini in a few anterior setigers are reminiscent of species of
|
||
<taxonomicName id="3AD6C7AB965A2326DA91FB7DFCBFFB79" authorityName="Hartman" authorityYear="1957" box="[710,788,1233,1259]" class="Polychaeta" family="Orbiniidae" genus="Califia" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Annelida" rank="genus">
|
||
<emphasis id="CFA2603A965A2326DA91FB7DFCBFFB79" box="[710,788,1233,1259]" italics="true" pageId="36" pageNumber="37">Califia</emphasis>
|
||
</taxonomicName>
|
||
, in which heavy uncini of the first three setigers provide a prominent armature. Unlike species of
|
||
<taxonomicName id="3AD6C7AB965A2326DA1BFB59FD31FA9D" authorityName="Hartman" authorityYear="1957" box="[588,666,1269,1295]" class="Polychaeta" family="Orbiniidae" genus="Califia" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Annelida" rank="genus">
|
||
<emphasis id="CFA2603A965A2326DA1BFB59FD31FA9D" box="[588,666,1269,1295]" italics="true" pageId="36" pageNumber="37">Califia</emphasis>
|
||
</taxonomicName>
|
||
, however, these same uncini continue through all thoracic setigers.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B5CCEFA3965A2326D890FAB5FCC9FAA1" box="[199,866,1305,1331]" pageId="36" pageNumber="37" type="etymology">
|
||
<paragraph id="FD69BC28965A2326D890FAB5FCC9FAA1" blockId="36.[151,1437,153,1367]" box="[199,866,1305,1331]" pageId="36" pageNumber="37">
|
||
<emphasis id="CFA2603A965A2326D890FAB5FEE3FAA1" bold="true" box="[199,328,1305,1331]" pageId="36" pageNumber="37">Etymology</emphasis>
|
||
. The epithet is from the Greek,
|
||
<emphasis id="CFA2603A965A2326DAF9FAB5FD5CFAA1" box="[686,759,1305,1331]" italics="true" pageId="36" pageNumber="37">bathys</emphasis>
|
||
for deep.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B5CCEFA3965A2326D890FA90FBB5FAC5" box="[199,1054,1340,1367]" pageId="36" pageNumber="37" type="distribution">
|
||
<paragraph id="FD69BC28965A2326D890FA90FBB5FAC5" blockId="36.[151,1437,153,1367]" box="[199,1054,1340,1367]" pageId="36" pageNumber="37">
|
||
<emphasis id="CFA2603A965A2326D890FA90FEF2FAC5" bold="true" box="[199,345,1340,1367]" pageId="36" pageNumber="37">Distribution</emphasis>
|
||
. South
|
||
<collectingCountry id="85C1FCB8965A2326D9E7FA91FE58FAC5" box="[432,499,1341,1367]" name="China" pageId="36" pageNumber="37">China</collectingCountry>
|
||
Sea, off
|
||
<collectingCountry id="85C1FCB8965A2326DA00FA91FD0DFAC5" box="[599,678,1341,1367]" name="Brunei" pageId="36" pageNumber="37">Brunei</collectingCountry>
|
||
, Island of Borneo,
|
||
<quantity id="3A2E11CD965A2326DB2BFA90FBB1FAC5" box="[892,1050,1340,1367]" metricMagnitude="3" metricUnit="m" metricValue="1.4084999999999999" metricValueMax="1.557" metricValueMin="1.26" pageId="36" pageNumber="37" unit="m" value="1408.5" valueMax="1557.0" valueMin="1260.0">1260–1557 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |