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<document ID-CLB-Dataset="53584" ID-DOI="http://dx.doi.org/10.3897/phytokeys.227.101907" ID-GBIF-Dataset="34586947-79c0-4255-a6db-f236107ac2fe" ID-PMC="PMC10251418" ID-Pensoft-Pub="1314-2003-227-43" ID-Pensoft-UUID="AD08E6F7EEA15D67BB611F0B8B730623" ID-PubMed="37303593" ModsDocID="1314-2003-227-43" checkinTime="1685632887257" checkinUser="pensoft" docAuthor="Arista, Jessy Patricia, Hagsater, Eric, Santiago, Elizabeth, Edquen, Jose D., Pariente, Eli, Oliva, Manuel &amp; Salazar, Gerardo A." docDate="2023" docId="75274AB695F15C078A3114BA19DB86D5" docLanguage="en" docName="PhytoKeys 227: 43-87" docOrigin="PhytoKeys 227" docPubDate="2023-06-01" docSource="http://dx.doi.org/10.3897/phytokeys.227.101907" docTitle="Epidendrum echinatiantherum Hagsater, E. Santiago, J. P. Arista &amp; Edquen 2023, sp. nov." docType="treatment" docVersion="4" id="AD08E6F7EEA15D67BB611F0B8B730623" lastPageNumber="43" masterDocId="AD08E6F7EEA15D67BB611F0B8B730623" masterDocTitle="New and noteworthy species of the genus Epidendrum (Orchidaceae, Laeliinae) from the Area de Conservacion Privada La Pampa del Burro, Amazonas, Peru" masterLastPageNumber="87" masterPageNumber="43" pageNumber="43" updateTime="1732479553626" updateUser="ExternalLinkService">
<mods:mods id="2EAE5ADAE67C75AD5EC35B24B38A7A64" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B39A7401DAC6913D0951E293BB18EFE4">
<mods:title id="4D5D8DBA76E4CA0FE290ADFB50075DF5">New and noteworthy species of the genus Epidendrum (Orchidaceae, Laeliinae) from the Area de Conservacion Privada La Pampa del Burro, Amazonas, Peru</mods:title>
</mods:titleInfo>
<mods:name id="BD78D22830116369DBF02D9E43BE4F47" type="personal">
<mods:role id="55CB559526452550A727F210460F2808">
<mods:roleTerm id="37E2DFBA83255F0C029C39BDFDFB80BE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="995458713E1A7A4861EF135341165981">Arista, Jessy Patricia</mods:namePart>
<mods:nameIdentifier id="8A63448645FD5493EED6625CCE07BCDC" type="ORCID">https://orcid.org/0000-0001-8334-5538</mods:nameIdentifier>
<mods:affiliation id="8C16EA92FAFAD22C72C7B2803CE23AFB">Instituto de Investigacion, Innovacion y Desarrollo del Sector Agrario y Agroindustrial (IIDAA), Universidad Nacional Toribio Rodriguez de Mendoza de Amazonas, Chachapoyas, Amazonas, Peru</mods:affiliation>
</mods:name>
<mods:name id="DB760B09D580C265960D0D54AE605EF8" type="personal">
<mods:role id="B7A591E011753B0A0556F5A72136E16F">
<mods:roleTerm id="10FF4B7A36F5058A7ACF42AC3B15E49D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FF7F71C6645895DF5C59D472C108F7B9">Hagsater, Eric</mods:namePart>
<mods:nameIdentifier id="1704D9B54F52486B9C780296F13CF856" type="ORCID">https://orcid.org/0000-0002-2371-9427</mods:nameIdentifier>
<mods:affiliation id="304AA47A0D4E5142CFF735156E6F29E9">Escuela de Posgrado, Universidad Nacional Toribio Rodriguez de Mendoza de Amazonas, Chachapoyas, Amazonas, Peru</mods:affiliation>
</mods:name>
<mods:name id="85A59B2BD573DCA3F66C4EA0738B04CF" type="personal">
<mods:role id="12CFC6C818E85D9848176EF7942B4F6E">
<mods:roleTerm id="FF6269BB803BECE677C81B0E65130D47">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7BC13B469712E7EB3BBC081005D1B114">Santiago, Elizabeth</mods:namePart>
<mods:nameIdentifier id="E09BA887318AB3F99CAA9D7BAB1EC855" type="ORCID">https://orcid.org/0000-0002-1368-9025</mods:nameIdentifier>
<mods:affiliation id="113EE9FEE0DBA84C7E68C1F8F7A007D2">Escuela de Posgrado, Universidad Nacional Toribio Rodriguez de Mendoza de Amazonas, Chachapoyas, Amazonas, Peru</mods:affiliation>
</mods:name>
<mods:name id="195EEFB71F29A5C011EAAD9B27383A05" type="personal">
<mods:role id="BF1E818F34AED0C864794BADCEF51170">
<mods:roleTerm id="4A6D74313A8937610C652D3D3A534A46">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="13017EC009ECE9F707982D073C66D800">Edquen, Jose D.</mods:namePart>
<mods:nameIdentifier id="7E93104C587533793CC4AC7CA21314AD" type="ORCID">https://orcid.org/0000-0002-7289-3974</mods:nameIdentifier>
<mods:affiliation id="F91D7F89BCE82A1AB41DC0EB507F092B">Instituto de Investigacion, Innovacion y Desarrollo del Sector Agrario y Agroindustrial (IIDAA), Universidad Nacional Toribio Rodriguez de Mendoza de Amazonas, Chachapoyas, Amazonas, Peru</mods:affiliation>
</mods:name>
<mods:name id="19371269D99CC76888CABF288142EE8B" type="personal">
<mods:role id="92DDA689744813CFC88653257C26FA51">
<mods:roleTerm id="D3373691D508B6A9F325B85BA4606059">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="203313FE60C9695C8FE5B6FEADA03680">Pariente, Eli</mods:namePart>
<mods:nameIdentifier id="DEFF6FF80570614B5AC6C14E535D8F75" type="ORCID">https://orcid.org/0000-0002-9197-0218</mods:nameIdentifier>
<mods:affiliation id="34BBB91FB869C185E059EB89FB3AB339">Instituto de Investigacion, Innovacion y Desarrollo del Sector Agrario y Agroindustrial (IIDAA), Universidad Nacional Toribio Rodriguez de Mendoza de Amazonas, Chachapoyas, Amazonas, Peru</mods:affiliation>
</mods:name>
<mods:name id="8CF0677BFA798A72FAB780554172D539" type="personal">
<mods:role id="B19F0AF80DF74AFF868DA2988C990911">
<mods:roleTerm id="0900D4164ED20BF3D37CC51A6C28805C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3D3859B8E5FE8B77C731BF72B3AB4D1E">Oliva, Manuel</mods:namePart>
<mods:nameIdentifier id="0AA41243E435E40166F2EF05FA75FDB0" type="ORCID">https://orcid.org/0000-0002-9670-0970</mods:nameIdentifier>
<mods:affiliation id="317FD16AAA1F27237F842233663D683D">Instituto de Investigacion, Innovacion y Desarrollo del Sector Agrario y Agroindustrial (IIDAA), Universidad Nacional Toribio Rodriguez de Mendoza de Amazonas, Chachapoyas, Amazonas, Peru</mods:affiliation>
</mods:name>
<mods:name id="8FFF9F59665349EE6FB502DB6D07E532" type="personal">
<mods:role id="3A37692D5BC8FDF2ADD854E98935092F">
<mods:roleTerm id="FD4E5DD5959BB7037D0BA490B0AFA86A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1BE245D9A7512B4D6B1DEF5CBE718E80">Salazar, Gerardo A.</mods:namePart>
<mods:nameIdentifier id="39A0C79D1AEB10689C526A4CE3F4DF25" type="ORCID">https://orcid.org/0000-0002-5203-5374</mods:nameIdentifier>
<mods:affiliation id="5C74551C387EB5679E74C690A6488548">Herbario AMO, Montanas Calizas 490, Lomas de Chapultepec. Miguel Hidalgo, Mexico City, 11000, Mexico</mods:affiliation>
<mods:nameIdentifier id="A1CD7336E5225F2278EC3DA9E0C46F7B" type="email">gasc@ib.unam.mx</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="C6D526087164F928F3C03CFCC5B099C5">text</mods:typeOfResource>
<mods:relatedItem id="EAB290EC5199C3259BBEC01FDB4C0E32" type="host">
<mods:titleInfo id="02164417E4B11FA6DE3C486C820A16BF">
<mods:title id="4BB544F1A07A8E7CB31FE7A3A6F6D472">PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part id="B57B99A1345A2A38B01C02A3E38E50C8">
<mods:date id="8A85036B7604B57E0C6048867289E7C5">2023</mods:date>
<mods:detail id="F23E9ECB35AB62181026D9AA0F5C9A49" type="pubDate">
<mods:number id="C126537E69AB0120A7C6C1750C6A717A">2023-06-01</mods:number>
</mods:detail>
<mods:detail id="DC1C59E07BBE389B565AB4D2AA359D6B" type="volume">
<mods:number id="F73C9E42E421B05690107EBF0007624E">227</mods:number>
</mods:detail>
<mods:extent id="C9A0762CA951E4FB989AA36F94D538CE" unit="page">
<mods:start id="E79AEEB1CD3D75885E8689C51D64BF71">43</mods:start>
<mods:end id="5538F63C2B084CBB7052EA42B2EE3C14">87</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="8B1CC787CAD1FD1A1482599E3BC775DE">
<mods:url id="180C469E3B090398C228849EADCCA145">http://dx.doi.org/10.3897/phytokeys.227.101907</mods:url>
</mods:location>
<mods:classification id="41B8741C11556A10B38319F54891F102">journal article</mods:classification>
<mods:identifier id="0797B60F4F03701110B56E34E39C2001" type="DOI">http://dx.doi.org/10.3897/phytokeys.227.101907</mods:identifier>
<mods:identifier id="B2E27928078575B63CC09D14F330F1BF" type="Pensoft-Pub">1314-2003-227-43</mods:identifier>
<mods:identifier id="AF07981F5196618223344AA4F455378D" type="Pensoft-UUID">AD08E6F7EEA15D67BB611F0B8B730623</mods:identifier>
</mods:mods>
<treatment id="75274AB695F15C078A3114BA19DB86D5" ID-GBIF-Taxon="209665289" LSID="urn:lsid:plazi:treatment:75274AB695F15C078A3114BA19DB86D5" httpUri="http://treatment.plazi.org/id/75274AB695F15C078A3114BA19DB86D5" lastPageNumber="43" pageId="0" pageNumber="43" scope_family="Orchidaceae">
<subSubSection id="047D9B4365707CD042C33A03F88E6585" pageId="0" pageNumber="43" type="nomenclature">
<paragraph id="3592E69B966598174069675223D628FC" pageId="0" pageNumber="43">
<taxonomicName id="891F2BC313AFA3C9B4731E14B6B60A5A" ID-CoL="CDC7H" LSID="75274AB6-95F1-5C07-8A31-14BA19DB86D5" authority="Hágsater, E. Santiago, J. P. Arista &amp; Edquén" authorityName="Hagsater, E. Santiago, J. P. Arista &amp; Edquen" authorityYear="2023" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Epidendrum echinatiantherum" order="Asparagales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="echinatiantherum" status="sp. nov.">
Epidendrum echinatiantherum
<normalizedToken id="DCE48EAEFC114213DDE26A49787F36F4" originalValue="Hágsater">Hagsater</normalizedToken>
, E.Santiago, J.P.Arista &amp;
<normalizedToken id="D4A9E20A749E2CD38EF0E0141FB4022E" originalValue="Edquén">Edquen</normalizedToken>
</taxonomicName>
<taxonomicNameLabel id="C38E592182F1821AD2BCC515178B56CE" pageId="0" pageNumber="43">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="3D4316852311A73EFDFFD6AFD38F8535" pageId="0" pageNumber="43" type="description">
<paragraph id="CF22586995E11DF716A2AFF549A1CEC7" pageId="0" pageNumber="43">
<figureCitation id="D9A43C40956D43D300AF1AEDBF8A9DD2" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Epidendrum echinatiantherum from Arista et al. 151 A habit B flower and floral bract C dissected perianth with dorsal and one lateral sepal shown from above and below, not spread D inflorescence at apex of stem E longitudinal section of flower F column and ovary-pedicel from below (left) and side (right) G anther from above (up) and below (down), and pollinarium H floral bract (Photographs by J. D. Edquen; plate prepared by A. Cisneros)." figureDoi="10.3897/phytokeys.227.101907.figure6" httpUri="https://binary.pensoft.net/fig/857082" pageId="0" pageNumber="43">Fig. 6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="E834CB916B0A37172BBCF92B067B7188" pageId="0" pageNumber="43" type="type material">
<paragraph id="D88054CE4B0C917F8BC477D9228296DB" pageId="0" pageNumber="43">Type material.</paragraph>
<paragraph id="075CEF3C343B6DE900A97FEC48A4748E" pageId="0" pageNumber="43">
<emphasis id="F3F012D273761D1F2FF278E4186966F9" bold="true" pageId="0" pageNumber="43">Peru. Amazonas</emphasis>
: Prov.
<normalizedToken id="20F9C82D0D0046751E0EA588E9969C4F" originalValue="Bongará">Bongara</normalizedToken>
: Distr. Yambrasbamba: Perla del Imaza,
<normalizedToken id="A329AD03D366C5D78F8C97F7A54FA154" originalValue="Área">Area</normalizedToken>
de
<normalizedToken id="2B70799C70B9AF78BDAC947127CD2D7D" originalValue="Conservación">Conservacion</normalizedToken>
Privada La Pampa del Burro, Bosque de Piedra, 1682 m, 28 Aug. 2021,
<emphasis id="608CEE0DE16635D5E2F72E0EC39E59E1" italics="true" pageId="0" pageNumber="43">J. P. Arista</emphasis>
,
<emphasis id="AF6148FF3FE58478B23398F14CF7EDC6" italics="true" pageId="0" pageNumber="43">
J. D.
<normalizedToken id="69C8859C616BF6DFE17834EF7C963149" originalValue="Edquén">Edquen</normalizedToken>
</emphasis>
,
<emphasis id="DFB7263C5DEB2764F756000AEE5D64D3" italics="true" pageId="0" pageNumber="43">
E.
<normalizedToken id="82602832F9CBBA72EC3DA357081EC0F5" originalValue="Yrigoín">Yrigoin</normalizedToken>
&amp; L. Iliquin 151
</emphasis>
(holotype: KUELAP!).
</paragraph>
</subSubSection>
<subSubSection id="EC251966FEEF5D5ADFD2D445321D2109" pageId="0" pageNumber="43" type="diagnosis">
<paragraph id="EEDFBA5B610EAD5A220FDADBCC46723A" pageId="0" pageNumber="43">Diagnosis.</paragraph>
<paragraph id="5D41B35EA378182F47867C82CBECCFEB" pageId="0" pageNumber="43">
Similar to
<taxonomicName id="C9C1979D27C5F5EFA549421776D5DAB7" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Epidendrum madsenii" order="Asparagales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="madsenii">
<emphasis id="DFFFA7B73CD6F552B65A86134E22604A" italics="true" pageId="0" pageNumber="43">Epidendrum madsenii</emphasis>
</taxonomicName>
<normalizedToken id="BFD4DBEF486B47435CF8202414464886" originalValue="Hágsater">Hagsater</normalizedToken>
&amp; Dodson, both vegetatively and florally, but the main distinguishing feature is the dark vinaceous anther with the central green vertical rib with a heavily echinate apical, elongate, pyramidal, truncate process, covered by numerous disorganized white bristles (vs. dark brown-black anther without a green vertical rib, and an unornamented apical process, which is laminar, forming an erect, semi-tubular laminar flap, the margins dentate).
</paragraph>
</subSubSection>
<subSubSection id="B9C311373F00B5917B182A101EA95FE5" pageId="0" pageNumber="43" type="description">
<paragraph id="D0C004AEFD20A38387BCD7BFF20101C8" pageId="0" pageNumber="43">Description.</paragraph>
<paragraph id="F7F2AEDA8C280490E279A6FB08D8F4A3" pageId="0" pageNumber="43">
Epiphytic, sympodial, branching, pendulous,
<emphasis id="F18AC58C014C3FCC2C2047EB32F64E8B" bold="true" italics="true" pageId="0" pageNumber="43">herb</emphasis>
, 8-13 cm long.
<emphasis id="548B446BA69E5B0AD10E6EAB5BB247D1" bold="true" italics="true" pageId="0" pageNumber="43">Roots</emphasis>
ca. 0.7 mm in diameter very thin, basal on primary stems.
<emphasis id="79C1ED84195C84206D7290CAE248669E" bold="true" italics="true" pageId="0" pageNumber="43">Stems</emphasis>
4.5-13
<normalizedToken id="31944805F74A2341954DEE05F1DAF28C" originalValue="×">x</normalizedToken>
2-3 cm, laterally compressed, new stems produced from sub-apical node of previous stem, sometimes from base of primary stem.
<emphasis id="EBEA3150269A4BE14EB284FF7D016BEF" bold="true" italics="true" pageId="0" pageNumber="43">Leaves</emphasis>
10-25 per stem, distributed throughout the length of the stems, articulate, twisted at base so as to be on same plane of stem as in
<emphasis id="A476401280486E44A3E1BF885D5A3491" italics="true" pageId="0" pageNumber="43">Dichaea</emphasis>
; sheaths 0.4-0.6
<normalizedToken id="A13F88ADE2D2D5CAD1D48EBB94395D1C" originalValue="×">x</normalizedToken>
0.35-0.45 cm, tubular, laterally compressed, ancipitose, minutely rugose, green; leaves 1.1-4.3
<normalizedToken id="2DC9BFCB3BE387A7D1D5A0EE4E8BEDC6" originalValue="×">x</normalizedToken>
0.6-1.2 cm, ovate to lanceolate, acute, fleshy, succulent, margins entire, spreading, medium green.
<emphasis id="91B4A5FF3EE80206B4A8B4898E88752E" bold="true" italics="true" pageId="0" pageNumber="43">Spathe</emphasis>
lacking.
<emphasis id="DEFDF13BBABB2A9D86D2B9307A0ABF96" bold="true" italics="true" pageId="0" pageNumber="43">Inflorescence</emphasis>
apical, racemose, sub-corymbose, pendulous, from mature stem, sessile, 3-flowered.
<emphasis id="9664458B42A0A263820F63DE7C3F8B0E" bold="true" italics="true" pageId="0" pageNumber="43">Floral bracts</emphasis>
6-8
<normalizedToken id="07BC4E17B074DCCDEABD39E62419B26F" originalValue="×">x</normalizedToken>
6-7 mm, conduplicate, widely cordiform when spread, dorsally carinate, especially toward apex, embracing.
<emphasis id="A2F94B3A3F7D7999E3AA1BAB3D94CB7D" bold="true" italics="true" pageId="0" pageNumber="43">Flowers</emphasis>
3, more or less simultaneous, pendulous, pale green, sepals tinged pale brown, calli white; fragrance not recorded.
<emphasis id="583B4EA8DF8BAF7F74B7959ADEB1EAA9" bold="true" italics="true" pageId="0" pageNumber="43">Sepals</emphasis>
9.5
<normalizedToken id="3DEA144844899B7EBBE4494DEFBAC24D" originalValue="×">x</normalizedToken>
3 mm, nearly spreading, free, narrowly lanceolate, acute, 3-veined, lateral sepals dorsally carinate, apex aristate, margins entire, somewhat revolute.
<emphasis id="793DFDD3678E18B4F5E744182D0B3E18" bold="true" italics="true" pageId="0" pageNumber="43">Petals</emphasis>
9
<normalizedToken id="465CC3273B77CDB0F777706080707F94" originalValue="×">x</normalizedToken>
1 mm, free, nearly spreading, linear-lanceolate, acute, 1-veined, margins entire, slightly revolute.
<emphasis id="8FCBCF8B7DADE86E5795D7644744F906" bold="true" italics="true" pageId="0" pageNumber="43">Lip</emphasis>
7.5
<normalizedToken id="F3569071AFCF0AB8BD8CB945AB91EF6A" originalValue="×">x</normalizedToken>
6.0 mm, united to basal half of column, deeply 3-lobed, fleshy, thick, calli 1.1 mm long, basal, digitiform, divergent, disc unornamented, lateral lobes 4.0
<normalizedToken id="D0B99DF13E5580B6D6C9C6623F289B3D" originalValue="×">x</normalizedToken>
2.2 mm, transversely ovate, apex narrowly rounded, nearly spreading flat in natural position, at a 45° angle to midline of lip; mid-lobe 4.5
<normalizedToken id="7A26FF4676DF263DB41364ECB16913A0" originalValue="×">x</normalizedToken>
2.6 mm, semi-elliptic, apex rounded.
<emphasis id="9C98E9611EF1398C571DF3F27282C770" bold="true" italics="true" pageId="0" pageNumber="43">Column</emphasis>
5.5-6.2 mm long dorsally including clinandrium hood, straight, body of column (to rostellum) about half as long as hood.
<emphasis id="696C9946306834715A7083C4219C7F30" bold="true" italics="true" pageId="0" pageNumber="43">Clinandrium hood</emphasis>
prominent, semi-tubular, somewhat funnel-shaped, margin erose-dentate, anther deep within.
<emphasis id="0657B7E2DCE7AEBCA2F547AB6F379431" bold="true" italics="true" pageId="0" pageNumber="43">Anther</emphasis>
reniform, dark vinaceous with central green vertical rib, with an elongate pyramidal, truncate flap at apex, covered by numerous disorganized white bristles, heavily echinate.
<emphasis id="3AEB8898307A61D030A98D8F8BBE3CB1" bold="true" italics="true" pageId="0" pageNumber="43">Pollinia</emphasis>
4, lentil-shaped, caudicles soft and granulose.
<emphasis id="F476EC45A26D1533EE813097E03E729F" bold="true" italics="true" pageId="0" pageNumber="43">Rostellum</emphasis>
apical, nearly at an 80° angle with axis of column, slit; viscarium semi-liquid.
<emphasis id="FA40FBB17F0359B901154853E7313F07" bold="true" italics="true" pageId="0" pageNumber="43">Lateral lobes of stigma</emphasis>
large, transverse, triangular with stigmatic cavity vertical, transverse.
<emphasis id="70D35EB161C6B290508C2BD4FC1DFC95" bold="true" italics="true" pageId="0" pageNumber="43">Cuniculus</emphasis>
penetrating nearly to base of pedicellate ovary, somewhat wide, forming an elongate ventral vesicle in pedicel.
<emphasis id="F53CF50031864914A6CBADBBA3237E27" bold="true" italics="true" pageId="0" pageNumber="43">Ovary</emphasis>
7.5-10
<normalizedToken id="569B92309C0B36227931E90A64613A35" originalValue="×">x</normalizedToken>
2-3 mm including pedicel, terete, straight, inflated nearly to base by elongate vesicle, slightly more inflated basally.
<emphasis id="595FA5CD4EF5887CFAEDB637096E467C" bold="true" italics="true" pageId="0" pageNumber="43">Capsule</emphasis>
not seen.
</paragraph>
<caption id="A3EC6D25BAA43C04E5B79B1FA0600056" doi="10.3897/phytokeys.227.101907.figure6" httpUri="https://binary.pensoft.net/fig/857082" pageId="0" pageNumber="43" start="Figure 6" startId="F6">
<paragraph id="4A3E6AF9E6E93E5FC2F89FFFBF853DEE" pageId="0" pageNumber="43">
<emphasis id="6952ED0F99431D0464DAFD36EE849346" bold="true" pageId="0" pageNumber="43">Figure 6.</emphasis>
<taxonomicName id="A1E244E6E579BC8C1FAEA94C8935B4A0" authorityName="Hagsater, E. Santiago, J. P. Arista &amp; Edquen" authorityYear="2023" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Epidendrum echinatiantherum" order="Asparagales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="echinatiantherum">
<emphasis id="67F8E20DEC268355133B107BD2E88DF1" italics="true" pageId="0" pageNumber="43">Epidendrum echinatiantherum</emphasis>
</taxonomicName>
from
<emphasis id="5390961EDB96F4A71FED3974DAF1972A" italics="true" pageId="0" pageNumber="43">Arista et al. 151</emphasis>
<emphasis id="A9B684A761ECC2D4E3B9B82D0F177661" bold="true" pageId="0" pageNumber="43">A</emphasis>
habit
<emphasis id="FBBF7BA81C3754613A35D00A0931686A" bold="true" pageId="0" pageNumber="43">B</emphasis>
flower and floral bract
<emphasis id="7CD4129422CCD52523F0BA7324B184D1" bold="true" pageId="0" pageNumber="43">C</emphasis>
dissected perianth with dorsal and one lateral sepal shown from above and below, not spread
<emphasis id="7D5C9F4D473039CA984634D16D915EEE" bold="true" pageId="0" pageNumber="43">D</emphasis>
inflorescence at apex of stem
<emphasis id="CFEB1D856F60C997177258F6D04025AE" bold="true" pageId="0" pageNumber="43">E</emphasis>
longitudinal section of flower
<emphasis id="43C5A639BD22F8F9D9A536CE0CF55B0E" bold="true" pageId="0" pageNumber="43">F</emphasis>
column and ovary-pedicel from below (left) and side (right)
<emphasis id="A9F0AE5AEC868BF22D3B8CF30BB631EC" bold="true" pageId="0" pageNumber="43">G</emphasis>
anther from above (up) and below (down), and pollinarium
<emphasis id="C5FE5AE4BB3E00BB0659490DB8BCFE3E" bold="true" pageId="0" pageNumber="43">H</emphasis>
floral bract (Photographs by J. D.
<normalizedToken id="4A7ACC5D991AB89280655D4A5739ED55" originalValue="Edquén">Edquen</normalizedToken>
; plate prepared by A. Cisneros).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C006F7F818633432FA8BEC2C230FE8DA" pageId="0" pageNumber="43" type="distribution">
<paragraph id="298207B252D80B00E33B8A8D318C4551" pageId="0" pageNumber="43">Distribution.</paragraph>
<paragraph id="6AF36CDF7A64AB7929FDE167E4A17D92" pageId="0" pageNumber="43">Known only from the type collection, in Amazonas, Peru, on the eastern slope of the Andes at 1682 m altitude.</paragraph>
</subSubSection>
<subSubSection id="7C99ECBDAE217D9D521CE8ABC5778456" pageId="0" pageNumber="43" type="habitat">
<paragraph id="2094A27020EFB7DBAAE959F3D5592483" pageId="0" pageNumber="43">Habitat and ecology.</paragraph>
<paragraph id="713405D932FFB85B36A4E0F00A89510D" pageId="0" pageNumber="43">
Epiphytic in wet montane forest on white sand, covered by accumulated organic material, dominated by palms, with
<taxonomicName id="C14695CB55124D5C98F9A76496B928E2" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Rubiaceae" genus="Cinchona" kingdom="Plantae" lsidName="Cinchona" order="Gentianales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="7A5ED6ED1DA8CBFCBA0D0D1629DACE07" italics="true" pageId="0" pageNumber="43">Cinchona</emphasis>
</taxonomicName>
sp. and
<taxonomicName id="BC2700D518E6953F25903C68C04C58F2" class="Liliopsida" family="Bromeliaceae" genus="Cedrela" kingdom="Plantae" lsidName="Cedrela" order="Poales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="A5A92BB027D360DEFF2CB649D640DF8D" italics="true" pageId="0" pageNumber="43">Cedrela</emphasis>
</taxonomicName>
sp. Presence of many
<taxonomicName id="ED86BED12ABE948861D6AB4C34A2112D" class="Liliopsida" family="Bromeliaceae" kingdom="Plantae" lsidName="" order="Poales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="family">Bromeliaceae</taxonomicName>
, mosses. Growing on dry hanging thin branch covered by moss.
</paragraph>
</subSubSection>
<subSubSection id="4DE35BBD388579E1FE46CEC55606FA5D" pageId="0" pageNumber="43" type="phenology">
<paragraph id="3FA5A50689C287867D60C1085CB6A695" pageId="0" pageNumber="43">Phenology.</paragraph>
<paragraph id="C76638956331F4DE467929EE91AA173B" pageId="0" pageNumber="43">Flowering in August.</paragraph>
</subSubSection>
<subSubSection id="597DBDEDF6031E05EE45916C284B1E85" pageId="0" pageNumber="43" type="taxonomic notes">
<paragraph id="9802F3E4D44227B2D1DCEF88B9E210A7" pageId="0" pageNumber="43">Taxonomic notes.</paragraph>
<paragraph id="124FDF7034FDD502A3259DE3AD0B8355" pageId="0" pageNumber="43">
<taxonomicName id="17BAA5E77C27B7F3F4235824D58A0743" authorityName="Hagsater, E. Santiago, J. P. Arista &amp; Edquen" authorityYear="2023" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Epidendrum echinatiantherum" order="Asparagales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="echinatiantherum">
<emphasis id="2F14AEA98C8FFD98392C8B147D8397D6" italics="true" pageId="0" pageNumber="43">Epidendrum echinatiantherum</emphasis>
</taxonomicName>
belongs to the Nanum Group, which is characterized by the
<emphasis id="B2ADF033831882EE880AF72748318B49" italics="true" pageId="0" pageNumber="43">Dichaea</emphasis>
-like horizontal or pendulous stems and the inflorescence produced by pairs of opposite flowers without spathes, but with prominent floral bracts. The new species is characterized by the few flowers, the cuniculus forming a long ventral swollen vesicle nearly reaching the base of the pedicellate ovary, relatively large, the sepals 9.5 mm long, the deeply 3-lobed lip with prominent ovate lateral lobes on the lip, the semi-elliptic mid-lobe and the prominent clinandrium hood, somewhat funnel-shaped and especially the reniform anther, dark wine-red with central green vertical rib, with an elongate pyramidal, truncate flap at apex, covered by numerous disorganized white bristles, heavily echinate. It is most similar to
<taxonomicName id="DB7585417FBF534130EFC5D5A811EBF4" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Epidendrum madsenii" order="Asparagales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="madsenii">
<emphasis id="2D5A61D1820E64D384DC91FDE91E0FC4" italics="true" pageId="0" pageNumber="43">Epidendrum madsenii</emphasis>
</taxonomicName>
, which also has a prominent, very long swollen vesicle at the base of the ovary, nearly as long as the ovary itself, the pale green flowers, the prominent 3-lobed, erose clinandrium hood, the triangular, acute mid-lobe of the lip, and the ornamentation of the anther consisting of a transverse, dentate process in front. It closely resembles
<taxonomicName id="C494C51FC3352DCEF65083FD26ADFB3E" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Epidendrum lueri" order="Asparagales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="lueri">
<emphasis id="151B01791ED171B559DEDA8FBCE99462" italics="true" pageId="0" pageNumber="43">Epidendrum lueri</emphasis>
</taxonomicName>
Dodson &amp;
<normalizedToken id="5D2A3911FB6A65ACB5555A3628BA8D78" originalValue="Hágsater">Hagsater</normalizedToken>
, which has pinkish yellow flowers with a bright yellow lip, a denticulate clinandrium-hood, but it is neither 3-lobed nor fimbriate.
</paragraph>
</subSubSection>
<subSubSection id="D6F2D92712AD85377D9E8B13C6DDAE3D" pageId="0" pageNumber="43" type="etymology">
<paragraph id="6F0C4726CC395EDA05FCBE98DA3FEEEA" pageId="0" pageNumber="43">Etymology.</paragraph>
<paragraph id="E1E606070CEF371A65E01872C7D99F51" pageId="0" pageNumber="43">
From Latin
<emphasis id="30368475C4724430844A18F5C3B0DBE8" italics="true" pageId="0" pageNumber="43">echinatus</emphasis>
, bristly, furnished with numerous rigid hairs, or straight prickles, and
<emphasis id="120229A84F44EF9EC2AF9915235D2AFD" italics="true" pageId="0" pageNumber="43">anthera</emphasis>
, the cover of the pollinarium at the apex of the column, which has an appendage heavily covered with white bristles, a rare and prominent feature of this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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