777 lines
88 KiB
XML
777 lines
88 KiB
XML
<document id="7569F6096AE5F39F1D3B9F131C6F7AFE" ID-DOI="10.5281/zenodo.202728" ID-GBIF-Dataset="240e955b-1e76-4082-9e42-d46906041f84" ID-ISSN="1175-5326" ID-Zenodo-Dep="202728" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460241938585" checkinUser="plazi" docAuthor="Bora, Parfait, Ramilijaona, Olga, Raminosoa, Noromalala & Vences, Miguel" docDate="2011" docId="750E906CFF936842FF53FB37F0D7FB22" docLanguage="en" docName="zt02772p060.pdf" docOrigin="Zootaxa 2772" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Mantidactylus paidroa Bora, Ramilijaona, Raminosoa & Vences, 2011, sp. nov." docType="treatment" docVersion="8" lastPageNumber="58" masterDocId="8937E814FF926844FFC4FFF9F432FFBA" masterDocTitle="A new species of Mantidactylus (subgenus Chonomantis) from Ranomafana National Park, eastern Madagascar (Amphibia, Anura, Mantellidae)" masterLastPageNumber="60" masterPageNumber="52" pageNumber="53" updateTime="1698240988888" updateUser="plazi">
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<mods:mods id="0739B39CCFD20E82F8ADDAC3429E8169" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="6A091C2C8070D0C67504FF548B9B62B1">
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<mods:title id="4F0330391ABB8A3DCABD5CEF52FAD572">A new species of Mantidactylus (subgenus Chonomantis) from Ranomafana National Park, eastern Madagascar (Amphibia, Anura, Mantellidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="F38207ED3DD3891BCC1758A926309DD6" type="personal">
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<mods:role id="3B66170EF6FB3C9F38374B5B83E12D17">
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<mods:roleTerm id="E36ECACD677E9790DB8EE6B7DF607EA1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="25779F3C01F821AB5B1509838383320D">Bora, Parfait</mods:namePart>
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</mods:name>
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<mods:name id="C9B7F27A60A02973FD0258338703C785" type="personal">
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<mods:roleTerm id="17D7864428D5CA7B4F0D2154AB55EB68">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="029ECA7444D99817A9AEC61C5C0EACB8">Ramilijaona, Olga</mods:namePart>
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</mods:name>
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<mods:name id="D5ED93DB71D67BC234DB8DCA9DDF02F8" type="personal">
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<mods:role id="617703BA283FA95E6CF465E63D81B367">
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<mods:roleTerm id="06A4A44FB211B6A11DCC7B35D09E0829">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="74975B15BFE06313EF4017501647348F">Raminosoa, Noromalala</mods:namePart>
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</mods:name>
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<mods:name id="16F4B257593B1C6CC233E524005928D6" type="personal">
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<mods:role id="A5E05261EF492CFF97C767D07149BB1C">
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<mods:roleTerm id="F6AF7AD78BD9D4184223AB97E1284185">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C63A0D177241E2600B3ABCCDEBEB77E3">Vences, Miguel</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="B6E640AB0AB7D6C0D34B3D034235D14D">text</mods:typeOfResource>
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<mods:titleInfo id="CB255CBE2B52AC11EAF50CDEDDAEF7E3">
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<mods:title id="5533B44CD289284AE179B58ECC0F90C3">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="4A7D3356063F579A1107FFAA49EA4ECB">
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<mods:date id="8DF98DDD88C54EB82FA90A57B044CBF5">2011</mods:date>
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<mods:detail id="CD6567B5407F77FDC9293589C0C5AEAD" type="volume">
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<mods:number id="FD1B326EC04E9B0C1D07450C7D711A7C">2772</mods:number>
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<mods:start id="6EE025EC540875DF42B8817262C02953">52</mods:start>
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<mods:end id="6BD2A30A81B1F9ACA224F22256E70493">60</mods:end>
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</mods:part>
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<mods:classification id="1792DE3392D3EF32D7D1796DADB83018">journal article</mods:classification>
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<mods:identifier id="A3C7176EC21146E56023218EB87FD57F" type="DOI">10.5281/zenodo.202728</mods:identifier>
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<mods:identifier id="76AE1973EE923023EE18113D5CA7F2BF" type="GBIF-Dataset">240e955b-1e76-4082-9e42-d46906041f84</mods:identifier>
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<mods:identifier id="6DB7EF7F73F20C74A7432030E37AF4D2" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="9159DB5DC4B3FAFDA762B2BE6DC9D3E5" type="Zenodo-Dep">202728</mods:identifier>
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<treatment id="750E906CFF936842FF53FB37F0D7FB22" ID-DOI="http://doi.org/10.5281/zenodo.5620603" ID-GBIF-Taxon="119389869" ID-Zenodo-Dep="5620603" LSID="urn:lsid:plazi:treatment:750E906CFF936842FF53FB37F0D7FB22" httpUri="http://treatment.plazi.org/id/750E906CFF936842FF53FB37F0D7FB22" lastPageId="6" lastPageNumber="58" pageId="1" pageNumber="53">
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<subSubSection id="B5BD72F1FF936845FF53FB37F520FAB3" pageId="1" pageNumber="53" type="nomenclature">
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<paragraph id="FD18217AFF936845FF53FB37F63EFB52" blockId="1.[151,524,1230,1289]" box="[151,524,1230,1256]" pageId="1" pageNumber="53">
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<heading id="A6509616FF936845FF53FB37F63EFB52" bold="true" box="[151,524,1230,1256]" fontSize="11" level="1" pageId="1" pageNumber="53" reason="1">
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<emphasis id="CFD3FD68FF936845FF53FB37F63EFB52" bold="true" box="[151,524,1230,1256]" pageId="1" pageNumber="53">
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||
<taxonomicName id="3AA75AF9FF936845FF53FB37F595FB52" ID-CoL="6R6MG" box="[151,423,1230,1256]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="53" phylum="Chordata" rank="species" species="paidroa" status="sp. nov.">
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<emphasis id="CFD3FD68FF936845FF53FB37F595FB52" bold="true" box="[151,423,1230,1256]" italics="true" pageId="1" pageNumber="53">Mantidactylus paidroa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D4E04013FF936845FE6BFB37F63EFB52" box="[431,524,1230,1256]" pageId="1" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="FD18217AFF936845FF53FB08F520FAB3" blockId="1.[151,524,1230,1289]" box="[151,274,1265,1289]" pageId="1" pageNumber="53">
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||
(
|
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<figureCitation id="659C3DFFFF936845FF5BFB08F4ECFAB3" box="[159,222,1265,1289]" captionStart="FIGURE 1" captionStartId="2.[151,250,1959,1982]" captionTargetBox="[151,1436,337,1938]" captionTargetId="figure@2.[151,1436,337,1938]" captionTargetPageId="2" captionText="FIGURE 1. (a), (b) and (c), Photographs of male holotype of Mantidactylus paidroa (ZSM 1777 / 2008) in life (SVL 22.3 mm), in dorsolateral and ventral views." httpUri="https://zenodo.org/record/202729/files/figure.png" pageId="1" pageNumber="53">Fig. 1</figureCitation>
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a–c)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="B5BD72F1FF936845FF53FAC3F4D3FACF" pageId="1" pageNumber="53" type="discussion">
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<paragraph id="FD18217AFF936845FF53FAC3F4D3FACF" blockId="1.[151,1437,1337,2009]" pageId="1" pageNumber="53">
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<emphasis id="CFD3FD68FF936845FF53FAC3F539FAE8" bold="true" box="[151,267,1338,1362]" pageId="1" pageNumber="53">Remarks.</emphasis>
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This species has been referred to as
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<taxonomicName id="3AA75AF9FF936845FD63FAC0F775FAEB" box="[679,839,1337,1361]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="53" phylum="Chordata" rank="genus">
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<emphasis id="CFD3FD68FF936845FD63FAC0F775FAEB" box="[679,839,1337,1361]" italics="true" pageId="1" pageNumber="53">Mantidactylus</emphasis>
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</taxonomicName>
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sp. 59 by
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<bibRefCitation id="99365C8BFF936845FC78FAC0F0A8FAEB" box="[956,1178,1337,1361]" pageId="1" pageNumber="60" refString="Vieites, D. R., Wollenberg, K. C., Andreone, F., Kohler, J., Glaw, F. & Vences, M. (2009) Vast underestimation of Madagascar's biodiversity evidenced by an integrative amphibian inventory. Proceedings of the National Academy of Sciences of the U. S. A., 106, 8267 - 8272." type="journal article">
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Vieites
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<emphasis id="CFD3FD68FF936845FBD4FAC3F07BFAEB" box="[1040,1097,1337,1361]" italics="true" pageId="1" pageNumber="53">et al.</emphasis>
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(2009)
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</bibRefCitation>
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||
and Grosjean
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<emphasis id="CFD3FD68FF936845FAF9FAC3F144FAEB" box="[1341,1398,1337,1361]" italics="true" pageId="1" pageNumber="53">et al.</emphasis>
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(in press).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="B5BD72F1FF936845FF03FA7BF638F993" pageId="1" pageNumber="53" type="materials_examined">
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||
<paragraph id="FD18217AFF936845FF03FA7BF0F5FA5B" blockId="1.[151,1437,1337,2009]" pageId="1" pageNumber="53">
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||
<emphasis id="CFD3FD68FF936845FF03FA7BF509FA20" bold="true" box="[199,315,1410,1434]" pageId="1" pageNumber="53">
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||
<typeStatus id="221C9FD8FF936845FF03FA7BF504FA20" box="[199,310,1410,1434]" pageId="1" pageNumber="53" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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||
<collectionCode id="9BB6B9BFFF936845FE86FA78F549FA23" box="[322,379,1409,1433]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="1" pageNumber="53">ZSM</collectionCode>
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1777/2008 (field number
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<collectionCode id="9BB6B9BFFF936845FD60FA78F6C7FA23" box="[676,757,1409,1433]" pageId="1" pageNumber="53">ZCMV</collectionCode>
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7108), adult male (
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||
<figureCitation id="659C3DFFFF936845FC17FA78F027FA23" box="[979,1045,1409,1433]" captionStart="FIGURE 1" captionStartId="2.[151,250,1959,1982]" captionTargetBox="[151,1436,337,1938]" captionTargetId="figure@2.[151,1436,337,1938]" captionTargetPageId="2" captionText="FIGURE 1. (a), (b) and (c), Photographs of male holotype of Mantidactylus paidroa (ZSM 1777 / 2008) in life (SVL 22.3 mm), in dorsolateral and ventral views." httpUri="https://zenodo.org/record/202729/files/figure.png" pageId="1" pageNumber="53">Fig. 1</figureCitation>
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), collected by P. Bora, J. Solo and D. Razafindraibe on
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<date id="891907BAFF936845FE42FA5CF671FA07" box="[390,579,1445,1470]" pageId="1" pageNumber="53" value="2008-01-24">24 January 2008</date>
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at a site locally named Ankerana, in Ranomafana National Park, Southern Central East of
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<collectingCountry id="85B061EAFF936845FE82FA30F5FFFA5B" box="[326,461,1481,1505]" name="Madagascar" pageId="1" pageNumber="53">Madagascar</collectingCountry>
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||
(
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<geoCoordinate id="989347BDFF936845FE18FA30F65FFA5B" box="[476,621,1481,1505]" direction="south" orientation="latitude" pageId="1" pageNumber="53" precision="1" value="-21.2597">21°15.582’S</geoCoordinate>
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||
,
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||
<geoCoordinate id="989347BDFF936845FDBCFA30F738FA5B" box="[632,778,1481,1505]" direction="east" orientation="longitude" pageId="1" pageNumber="53" precision="1" value="47.422">47°25.320’E</geoCoordinate>
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||
), approximately
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||
<quantity id="3A5F8C9FFF936845FC03FA30F03CFA5B" box="[967,1038,1481,1505]" metricMagnitude="2" metricUnit="m" metricValue="9.629999999999999" pageId="1" pageNumber="53" unit="m" value="963.0">963 m</quantity>
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above sea level.
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||
</paragraph>
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||
<paragraph id="FD18217AFF936845FF03FA14F638F993" blockId="1.[151,1437,1337,2009]" pageId="1" pageNumber="53">
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||
<emphasis id="CFD3FD68FF936845FF03FA14F574F9BF" bold="true" box="[199,326,1517,1541]" pageId="1" pageNumber="53">
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||
<typeStatus id="221C9FD8FF936845FF03FA14F573F9BF" box="[199,321,1517,1541]" pageId="1" pageNumber="53" type="paratype">Paratypes</typeStatus>
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||
.
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||
</emphasis>
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||
<collectionCode id="9BB6B9BFFF936845FE8AFA14F5B5F9BF" box="[334,391,1517,1541]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="1" pageNumber="53">ZSM</collectionCode>
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||
1778/2008 (field number
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<collectionCode id="9BB6B9BFFF936845FD71FA14F734F9BF" box="[693,774,1517,1541]" pageId="1" pageNumber="53">ZCMV</collectionCode>
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||
7109, male) and
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||
<collectionCode id="9BB6B9BFFF936845FC15FA14F039F9BF" box="[977,1035,1517,1541]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="1" pageNumber="53">ZSM</collectionCode>
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1776/2008 (
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||
<collectionCode id="9BB6B9BFFF936845FB58FA14F0DDF9BF" box="[1180,1263,1517,1541]" pageId="1" pageNumber="53">ZCMV</collectionCode>
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||
7107, female), same collecting data as
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<typeStatus id="221C9FD8FF936845FE65F9E8F635F993" box="[417,519,1553,1577]" pageId="1" pageNumber="53" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="B5BD72F1FF936840FF03F9CCF120FCA0" lastPageId="4" lastPageNumber="56" pageId="1" pageNumber="53" type="diagnosis">
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||
<paragraph id="FD18217AFF936845FF03F9CCF060F8BB" blockId="1.[151,1437,1337,2009]" pageId="1" pageNumber="53">
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||
<emphasis id="CFD3FD68FF936845FF03F9CCF573F9F4" bold="true" box="[199,321,1589,1614]" pageId="1" pageNumber="53">Diagnosis.</emphasis>
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Assigned to the genus
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<taxonomicName id="3AA75AF9FF936845FD8DF9CCF6DBF9F7" box="[585,745,1589,1613]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="53" phylum="Chordata" rank="genus">
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<emphasis id="CFD3FD68FF936845FD8DF9CCF6DBF9F7" box="[585,745,1589,1613]" italics="true" pageId="1" pageNumber="53">Mantidactylus</emphasis>
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</taxonomicName>
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based on the presence of an intercalary element between terminal and subterminal phalanges of fingers and toes (verified by external observation), and of a central depression in femoral glands and presence of a rudimentary femoral gland in the female. Within
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<taxonomicName id="3AA75AF9FF936845FB8DF984F0DBF92F" box="[1097,1257,1661,1685]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="53" phylum="Chordata" rank="genus">
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||
<emphasis id="CFD3FD68FF936845FB8DF984F0DBF92F" box="[1097,1257,1661,1685]" italics="true" pageId="1" pageNumber="53">Mantidactylus</emphasis>
|
||
</taxonomicName>
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||
, assigned to the subgenus
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<taxonomicName id="3AA75AF9FF936845FECDF958F5ADF903" box="[265,415,1697,1721]" class="Amphibia" family="Ranidae" genus="Chonomantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="1" pageNumber="53" phylum="Chordata" rank="genus">
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<emphasis id="CFD3FD68FF936845FECDF958F5ADF903" box="[265,415,1697,1721]" italics="true" pageId="1" pageNumber="53">Chonomantis</emphasis>
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</taxonomicName>
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||
based on: (a) a specialized funnel-mouth tadpole, (b) a sharp dorsolateral colour border between dorsal and lateral coloration, (c) presence of a frenal stripe, (d) diurnal calling activity close to streams, (e) intersexual difference in tympanum size, (f) relatively small size (male SVL
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||
<quantity id="3A5F8C9FFF936845FC36F910F077F8BB" box="[1010,1093,1769,1793]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="53" unit="mm" value="22.0">22 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FD18217AFF936847FF03F8F4F5B9FE3D" blockId="1.[151,1437,1337,2009]" lastBlockId="3.[151,1436,151,392]" lastPageId="3" lastPageNumber="55" pageId="1" pageNumber="53">
|
||
<taxonomicName id="3AA75AF9FF936845FF03F8F4F5FAF89F" box="[199,456,1805,1829]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="53" phylum="Chordata" rank="species" species="paidroa" status="sp. nov.">
|
||
<emphasis id="CFD3FD68FF936845FF03F8F4F5FAF89F" box="[199,456,1805,1829]" italics="true" pageId="1" pageNumber="53">Mantidactylus paidroa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D4E04013FF936845FE14F8F7F611F89F" box="[464,547,1806,1829]" pageId="1" pageNumber="53" rank="species">sp. nov</taxonomicNameLabel>
|
||
is distinguished from all other species in the subgenus
|
||
<taxonomicName id="3AA75AF9FF936845FB65F8F4F104F89F" box="[1185,1334,1805,1829]" class="Amphibia" family="Ranidae" genus="Chonomantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="1" pageNumber="53" phylum="Chordata" rank="genus">
|
||
<emphasis id="CFD3FD68FF936845FB65F8F4F104F89F" box="[1185,1334,1805,1829]" italics="true" pageId="1" pageNumber="53">Chonomantis</emphasis>
|
||
</taxonomicName>
|
||
by presence of serial double notes in advertisement calls (vs. single notes) and a substantial genetic differentiation (see below). In addition, it is distinguished from
|
||
<emphasis id="CFD3FD68FF936845FD49F8ACF177F8D7" box="[653,1349,1877,1901]" italics="true" pageId="1" pageNumber="53">
|
||
<taxonomicName id="3AA75AF9FF936845FD49F8ACF784F8D7" box="[653,950,1877,1901]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="53" phylum="Chordata" rank="species" species="aerumnalis">Mantidactylus aerumnalis</taxonomicName>
|
||
,
|
||
<taxonomicName id="3AA75AF9FF936845FC07F8AFF0B9F8D7" box="[963,1163,1877,1901]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="53" phylum="Chordata" rank="species" species="brevipalmatus">M. brevipalmatus</taxonomicName>
|
||
,
|
||
<taxonomicName id="3AA75AF9FF936845FB5CF8AFF173F8D7" box="[1176,1345,1877,1901]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="53" phylum="Chordata" rank="species" species="charlotteae">M. charlotteae</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="3AA75AF9FF936845FAB9F8AFF4C9F82B" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="53" phylum="Chordata" rank="species" species="delormei">
|
||
<emphasis id="CFD3FD68FF936845FAB9F8AFF4C9F82B" italics="true" pageId="1" pageNumber="53">M. delormei</emphasis>
|
||
</taxonomicName>
|
||
by a bigger ratio of tympanum/eye diameter in males (1.4–1.5 vs. 0.7–1.3); from
|
||
<emphasis id="CFD3FD68FF936845FB4EF883F675F80F" italics="true" pageId="1" pageNumber="53">
|
||
<taxonomicName id="3AA75AF9FF936845FB4EF883F102F82B" box="[1162,1328,1913,1937]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="53" phylum="Chordata" rank="species" species="aerumnalis">M. aerumnalis</taxonomicName>
|
||
,
|
||
<taxonomicName id="3AA75AF9FF936845FAFFF883F4D4F80F" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="53" phylum="Chordata" rank="species" species="charlotteae">M. charlotteae</taxonomicName>
|
||
,
|
||
<taxonomicName id="3AA75AF9FF936845FF36F867F54BF80F" box="[242,377,1950,1973]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="53" phylum="Chordata" rank="species" species="opiparis">M. opiparis</taxonomicName>
|
||
,
|
||
<taxonomicName id="3AA75AF9FF936845FE42F867F675F80F" box="[390,583,1949,1973]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="53" phylum="Chordata" rank="species" species="melanopleura">M. melanopleura</taxonomicName>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="3AA75AF9FF936845FD4CF867F6CFF80F" box="[648,765,1950,1973]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="53" phylum="Chordata" rank="species" species="zipperi">
|
||
<emphasis id="CFD3FD68FF936845FD4CF867F6CFF80F" box="[648,765,1950,1973]" italics="true" pageId="1" pageNumber="53">M. zipperi</emphasis>
|
||
</taxonomicName>
|
||
by a lower amount of webbing on fifth toe (2 vs. 0.75–1.5 phalanges free of web); from
|
||
<taxonomicName id="3AA75AF9FF936845FE2BF83BF684F863" box="[495,694,1985,2009]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="53" phylum="Chordata" rank="species" species="brevipalmatus">
|
||
<emphasis id="CFD3FD68FF936845FE2BF83BF684F863" box="[495,694,1985,2009]" italics="true" pageId="1" pageNumber="53">M. brevipalmatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3AA75AF9FF936845FD36F83BF74CF863" box="[754,894,1985,2009]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="53" phylum="Chordata" rank="species" species="delormei">
|
||
<emphasis id="CFD3FD68FF936845FD36F83BF74CF863" box="[754,894,1985,2009]" italics="true" pageId="1" pageNumber="53">M. delormei</emphasis>
|
||
</taxonomicName>
|
||
by smaller body size (male SVL 22 vs.
|
||
<quantity id="3A5F8C9FFF936846FA92F838F4F6FF15" lastPageId="2" lastPageNumber="54" metricMagnitude="-2" metricUnit="m" metricValue="3.15" metricValueMax="3.5" metricValueMin="2.8" pageId="1" pageNumber="53" unit="mm" value="31.5" valueMax="35.0" valueMin="28.0">28–35 mm</quantity>
|
||
) and by shorter hindlimbs (tibiotarsal articulation reaching anterior eye corner vs. reaching beyond snout tip); from
|
||
<taxonomicName id="3AA75AF9FF906846FF11FF44F56BFF6E" box="[213,345,189,212]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="54" phylum="Chordata" rank="species" species="opiparis">
|
||
<emphasis id="CFD3FD68FF906846FF11FF44F56BFF6E" box="[213,345,189,212]" italics="true" pageId="2" pageNumber="54">M. opiparis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="3AA75AF9FF906846FEADFF44F619FF6E" box="[361,555,188,212]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="54" phylum="Chordata" rank="species" species="melanopleura">
|
||
<emphasis id="CFD3FD68FF906846FEADFF44F619FF6E" box="[361,555,188,212]" italics="true" pageId="2" pageNumber="54">M. melanopleura</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3AA75AF9FF906846FDA0FF44F739FF6E" box="[612,779,188,212]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="54" phylum="Chordata" rank="species" species="charlotteae">
|
||
<emphasis id="CFD3FD68FF906846FDA0FF44F739FF6E" box="[612,779,188,212]" italics="true" pageId="2" pageNumber="54">M. charlotteae</emphasis>
|
||
</taxonomicName>
|
||
by presence of distinct and rather large femoral glands in males (vs. small glands), from
|
||
<taxonomicName id="3AA75AF9FF906846FE32FF19F72FFF42" box="[502,797,224,248]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="54" phylum="Chordata" rank="species" species="aerumnalis">
|
||
<emphasis id="CFD3FD68FF906846FE32FF19F72FFF42" box="[502,797,224,248]" italics="true" pageId="2" pageNumber="54">Mantidactylus aerumnalis</emphasis>
|
||
</taxonomicName>
|
||
by presence of a frenal stripe between forelimb and eye (vs. absence); from
|
||
<emphasis id="CFD3FD68FF906846FEB1FEFCF680FEA6" box="[373,690,260,284]" italics="true" pageId="2" pageNumber="54">
|
||
<taxonomicName id="3AA75AF9FF906846FEB1FEFCF62EFEA6" box="[373,540,260,284]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="54" phylum="Chordata" rank="species" species="aerumnalis">M. aerumnalis</taxonomicName>
|
||
,
|
||
<taxonomicName id="3AA75AF9FF906846FDE3FEFCF69FFEA6" box="[551,685,261,284]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="54" phylum="Chordata" rank="species" species="opiparis">M. opiparis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="3AA75AF9FF906846FD2CFEFCF769FEA6" box="[744,859,261,284]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="54" phylum="Chordata" rank="species" species="zipperi">
|
||
<emphasis id="CFD3FD68FF906846FD2CFEFCF769FEA6" box="[744,859,261,284]" italics="true" pageId="2" pageNumber="54">M. zipperi</emphasis>
|
||
</taxonomicName>
|
||
by a well-delimited narrow extension of the frenal stripe (bending upwards) between eye and nostril (vs. absence), and from
|
||
<emphasis id="CFD3FD68FF916847FC10FF60F175FF0A" box="[980,1351,152,176]" italics="true" pageId="3" pageNumber="55">
|
||
<taxonomicName id="3AA75AF9FF916847FC10FF60F049FF0A" box="[980,1147,152,176]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="55" phylum="Chordata" rank="species" species="aerumnalis">M. aerumnalis</taxonomicName>
|
||
,
|
||
<taxonomicName id="3AA75AF9FF916847FB42FF60F175FF0A" box="[1158,1351,152,176]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="55" phylum="Chordata" rank="species" species="melanopleura">M. melanopleura</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="3AA75AF9FF916847FAB9FF60F4C7FF6E" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="55" phylum="Chordata" rank="species" species="opiparis">
|
||
<emphasis id="CFD3FD68FF916847FAB9FF60F4C7FF6E" italics="true" pageId="3" pageNumber="55">M. opiparis</emphasis>
|
||
</taxonomicName>
|
||
by having the third toe longer than the fifth (vs. fifth toe longer than third).
|
||
<taxonomicName id="3AA75AF9FF916847FB83FF45F174FF6E" box="[1095,1350,188,212]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="55" phylum="Chordata" rank="species" species="paidroa">
|
||
<emphasis id="CFD3FD68FF916847FB83FF45F174FF6E" box="[1095,1350,188,212]" italics="true" pageId="3" pageNumber="55">Mantidactylus paidroa</emphasis>
|
||
</taxonomicName>
|
||
is morphologically most similar to
|
||
<taxonomicName id="3AA75AF9FF916847FE18FF18F6BDFF42" box="[476,655,224,248]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="55" phylum="Chordata" rank="species" species="albofrenatus">
|
||
<emphasis id="CFD3FD68FF916847FE18FF18F6BDFF42" box="[476,655,224,248]" italics="true" pageId="3" pageNumber="55">M. albofrenatus</emphasis>
|
||
</taxonomicName>
|
||
from which it can be mainly distinguished by the throat colouration, which has a continuous median white line (vs. a row of spots in
|
||
<taxonomicName id="3AA75AF9FF916847FCB8FEFCF002FEA6" box="[892,1072,260,284]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="55" phylum="Chordata" rank="species" species="albofrenatus">
|
||
<emphasis id="CFD3FD68FF916847FCB8FEFCF002FEA6" box="[892,1072,260,284]" italics="true" pageId="3" pageNumber="55">M. albofrenatus</emphasis>
|
||
</taxonomicName>
|
||
). Tadpoles of
|
||
<taxonomicName id="3AA75AF9FF916847FB13FEFCF164FEA6" box="[1239,1366,260,284]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="55" phylum="Chordata" rank="species" species="paidroa">
|
||
<emphasis id="CFD3FD68FF916847FB13FEFCF164FEA6" box="[1239,1366,260,284]" italics="true" pageId="3" pageNumber="55">M. paidroa</emphasis>
|
||
</taxonomicName>
|
||
differ from those of all other
|
||
<taxonomicName id="3AA75AF9FF916847FE58FED1F600FEFA" box="[412,562,296,320]" class="Amphibia" family="Ranidae" genus="Chonomantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="CFD3FD68FF916847FE58FED1F600FEFA" box="[412,562,296,320]" italics="true" pageId="3" pageNumber="55">Chonomantis</emphasis>
|
||
</taxonomicName>
|
||
by the presence of very few keratodonts on the upper labium of the oral disk (vs. complete absence; see Grosjean
|
||
<emphasis id="CFD3FD68FF916847FD80FEB4F6B2FEDE" box="[580,640,332,356]" italics="true" pageId="3" pageNumber="55">et al.</emphasis>
|
||
in press).
|
||
<tableCitation id="B02514C1FF916847FD3BFEB5F767FEDE" box="[767,853,332,356]" captionStart="TABLE 1" captionStartId="3.[151,235,440,461]" captionTargetBox="[161,1435,653,1720]" captionTargetPageId="3" captionText="TABLE 1. Morphometric ratios and distinctive morphological and bioacoustic characters in Mantidactylus paidroa sp. nov. compared with other species of Chonomantis, based on data published by Vences & Glaw (2004). Abbreviations used: SVL, snout-vent length; TD, tympanum diameter; ED, eye diameter; FOL, foot length; HAL, hand length; RHL, relative hindlimb length; TT, tibiotarsal articulation. Measurement values are mean ± standard deviation, followed by minimum – maximum values in parentheses. Data for M. delormei are based on the female holotype and photos of additional specimens in Glaw & Vences (2007) for colour in life." httpUri="http://table.plazi.org/id/A9D871F2FF916847FF53FE41F5EEFDD0" pageId="3" pageNumber="55" tableUuid="A9D871F2FF916847FF53FE41F5EEFDD0">Table 1</tableCitation>
|
||
includes a comparison of diagnostic characters in
|
||
<taxonomicName id="3AA75AF9FF916847FF53FE89F51FFE32" box="[151,301,368,392]" class="Amphibia" family="Ranidae" genus="Chonomantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="CFD3FD68FF916847FF53FE89F51FFE32" box="[151,301,368,392]" italics="true" pageId="3" pageNumber="55">Chonomantis</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
</paragraph>
|
||
<caption id="A9D871F2FF906846FF53F85EF5DBF85B" httpUri="https://zenodo.org/record/202729/files/figure.png" pageId="2" pageNumber="54" targetBox="[151,1436,337,1938]" targetPageId="2">
|
||
<paragraph id="FD18217AFF906846FF53F85EF5DBF85B" blockId="2.[151,1436,1959,2017]" pageId="2" pageNumber="54">
|
||
<emphasis id="CFD3FD68FF906846FF53F85EF521F807" bold="true" box="[151,275,1959,1982]" pageId="2" pageNumber="54">FIGURE 1.</emphasis>
|
||
(a), (b) and (c), Photographs of male holotype of
|
||
<taxonomicName id="3AA75AF9FF906846FCCDF851F7DDF804" box="[777,1007,1960,1982]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="54" phylum="Chordata" rank="species" species="paidroa">
|
||
<emphasis id="CFD3FD68FF906846FCCDF851F7DDF804" box="[777,1007,1960,1982]" italics="true" pageId="2" pageNumber="54">Mantidactylus paidroa</emphasis>
|
||
</taxonomicName>
|
||
(ZSM 1777/2008) in life (SVL 22.3 mm), in dorsolateral and ventral views.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="A9D871F2FF916847FF53FE41F5EEFDD0" ID-Table-UUID="A9D871F2FF916847FF53FE41F5EEFDD0" httpUri="http://table.plazi.org/id/A9D871F2FF916847FF53FE41F5EEFDD0" pageId="3" pageNumber="55" targetBox="[161,1435,653,1720]" targetIsTable="true" targetPageId="3">
|
||
<paragraph id="FD18217AFF916847FF53FE41F5EEFDD0" blockId="3.[151,1437,440,619]" pageId="3" pageNumber="55">
|
||
<emphasis id="CFD3FD68FF916847FF53FE41F534FE77" bold="true" box="[151,262,440,461]" pageId="3" pageNumber="55">TABLE 1.</emphasis>
|
||
Morphometric ratios and distinctive morphological and bioacoustic characters in
|
||
<taxonomicName id="3AA75AF9FF916847FB9FFE41F171FE74" box="[1115,1347,440,462]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="55" phylum="Chordata" rank="species" species="paidroa" status="sp. nov.">
|
||
<emphasis id="CFD3FD68FF916847FB9FFE41F171FE74" box="[1115,1347,440,462]" italics="true" pageId="3" pageNumber="55">Mantidactylus paidroa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D4E04013FF916847FA88FE40F1AFFE74" box="[1356,1437,441,462]" pageId="3" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
|
||
compared with other species of
|
||
<emphasis id="CFD3FD68FF916847FE25FE2EF65DFE57" box="[481,623,471,493]" italics="true" pageId="3" pageNumber="55">
|
||
<taxonomicName id="3AA75AF9FF916847FE25FE2EF659FE57" box="[481,619,471,493]" class="Amphibia" family="Ranidae" genus="Chonomantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="55" phylum="Chordata" rank="genus">Chonomantis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
based on data published by Vences & Glaw (2004). Abbreviations used: SVL, snout-vent length; TD, tympanum diameter; ED, eye diameter; FOL, foot length; HAL, hand length; RHL, relative hindlimb length; TT, tibiotarsal articulation. Measurement values are mean ± standard deviation, followed by minimum–maximum values in parentheses. Data for
|
||
<taxonomicName id="3AA75AF9FF916847FE00FDCAF671FDF2" box="[452,579,562,584]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="55" phylum="Chordata" rank="species" species="delormei">
|
||
<emphasis id="CFD3FD68FF916847FE00FDCAF671FDF2" box="[452,579,562,584]" italics="true" pageId="3" pageNumber="55">M. delormei</emphasis>
|
||
</taxonomicName>
|
||
are based on the female holotype and photos of additional specimens in Glaw & Vences (2007) for colour in life.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FD18217AFF916847FEDCFD74F17BF9D2" pageId="3" pageNumber="55">
|
||
<table id="8FA7D3DAFF9197BBFF65FD74F1A9F902" box="[161,1435,653,1720]" gridcols="10" gridrows="12" pageId="3" pageNumber="55">
|
||
<tr id="43972338FF9197BBFF65FD74F1A9FD01" box="[161,1435,653,699]" gridrow="0" pageId="3" pageNumber="55" rowspan-0="1">
|
||
<th id="00464A44FF9197BBFEDCFD74F549FD01" box="[280,379,653,699]" gridcol="1" gridrow="0" pageId="3" pageNumber="55">
|
||
<taxonomicName id="3AA75AF9FF916847FEDCFD74F545FD25" box="[280,375,653,671]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="55" phylum="Chordata" rank="species" species="paidroa" status="sp. nov.">
|
||
<emphasis id="CFD3FD68FF916847FEDCFD74F545FD25" box="[280,375,653,671]" italics="true" pageId="3" pageNumber="55">M. paidroa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D4E04013FF916847FEDCFD51F569FD01" box="[280,347,680,699]" pageId="3" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</th>
|
||
<th id="00464A44FF9197BBFE50FD74F5CAFD01" box="[404,504,653,699]" gridcol="2" gridrow="0" pageId="3" pageNumber="55">
|
||
<taxonomicName id="3AA75AF9FF916847FE50FD74F5C6FD00" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="55" phylum="Chordata" rank="species" species="aerumnalis">
|
||
<emphasis id="CFD3FD68FF916847FE50FD74F5C6FD00" italics="true" pageId="3" pageNumber="55">M. aerumnalis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="00464A44FF9197BBFDD7FD74F64DFD01" box="[531,639,653,699]" gridcol="3" gridrow="0" pageId="3" pageNumber="55">
|
||
<taxonomicName id="3AA75AF9FF916847FDD0FD74F64DFD00" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="55" phylum="Chordata" rank="species" species="albofrenatus">
|
||
<emphasis id="CFD3FD68FF916847FDD0FD74F64DFD00" italics="true" pageId="3" pageNumber="55">M. albofrenatus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="00464A44FF9197BBFD52FD74F73DFD01" box="[662,783,653,699]" gridcol="4" gridrow="0" pageId="3" pageNumber="55">
|
||
<taxonomicName id="3AA75AF9FF916847FD52FD74F73DFD00" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="55" phylum="Chordata" rank="species" species="brevipalmatus">
|
||
<emphasis id="CFD3FD68FF916847FD52FD74F73DFD00" italics="true" pageId="3" pageNumber="55">M. brevipalmatus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="00464A44FF9197BBFCE1FD74F7A1FD01" box="[805,915,653,699]" gridcol="5" gridrow="0" pageId="3" pageNumber="55">
|
||
<taxonomicName id="3AA75AF9FF916847FCE1FD74F7B4FD00" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="55" phylum="Chordata" rank="species" species="charlotteae">
|
||
<emphasis id="CFD3FD68FF916847FCE1FD74F7B4FD00" italics="true" pageId="3" pageNumber="55">M. charlotteae</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="00464A44FF9197BBFC63FD74F7C1FD01" box="[935,1011,653,699]" gridcol="6" gridrow="0" pageId="3" pageNumber="55">
|
||
<taxonomicName id="3AA75AF9FF916847FC63FD74F7C1FD00" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="55" phylum="Chordata" rank="species" species="delormei">
|
||
<emphasis id="CFD3FD68FF916847FC63FD74F7C1FD00" italics="true" pageId="3" pageNumber="55">M. delormei</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="00464A44FF9197BBFBC8FD74F0B5FD01" box="[1036,1159,653,699]" gridcol="7" gridrow="0" pageId="3" pageNumber="55">
|
||
<taxonomicName id="3AA75AF9FF916847FBC8FD74F0B0FD00" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="55" phylum="Chordata" rank="species" species="melanopleura">
|
||
<emphasis id="CFD3FD68FF916847FBC8FD74F0B0FD00" italics="true" pageId="3" pageNumber="55">M. melanopleura</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="00464A44FF9197BBFB64FD74F136FD01" box="[1184,1284,653,699]" gridcol="8" gridrow="0" pageId="3" pageNumber="55">
|
||
<taxonomicName id="3AA75AF9FF916847FB64FD74F0D5FD00" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="55" phylum="Chordata" rank="species" species="opiparis">
|
||
<emphasis id="CFD3FD68FF916847FB64FD74F0D5FD00" italics="true" pageId="3" pageNumber="55">M. opiparis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="00464A44FF9197BBFAD2FD74F1A9FD01" box="[1302,1435,653,699]" gridcol="9" gridrow="0" pageId="3" pageNumber="55">
|
||
<taxonomicName id="3AA75AF9FF916847FAD2FD74F163FD00" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="55" phylum="Chordata" rank="species" species="zipperi">
|
||
<emphasis id="CFD3FD68FF916847FAD2FD74F163FD00" italics="true" pageId="3" pageNumber="55">M. zipperi</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="43972338FF9197BBFF65FD36F1A9FD47" box="[161,1435,719,765]" gridrow="1" pageId="3" pageNumber="55">
|
||
<th id="00464A44FF9197BBFF65FD36F536FD47" box="[161,260,719,765]" gridcol="0" gridrow="1" pageId="3" pageNumber="55">SVL males (mm)</th>
|
||
<td id="00464A44FF9197BBFEDCFD36F549FD47" box="[280,379,719,765]" gridcol="1" gridrow="1" pageId="3" pageNumber="55">22.0–22.3</td>
|
||
<td id="00464A44FF9197BBFE50FD36F5CAFD47" box="[404,504,719,765]" gridcol="2" gridrow="1" pageId="3" pageNumber="55">22.8–26.6</td>
|
||
<td id="00464A44FF9197BBFDD7FD36F64DFD47" box="[531,639,719,765]" gridcol="3" gridrow="1" pageId="3" pageNumber="55">19.3–23.0</td>
|
||
<td id="00464A44FF9197BBFD52FD36F73DFD47" box="[662,783,719,765]" gridcol="4" gridrow="1" pageId="3" pageNumber="55">27.6–35.3</td>
|
||
<td id="00464A44FF9197BBFCE1FD36F7A1FD47" box="[805,915,719,765]" gridcol="5" gridrow="1" pageId="3" pageNumber="55">22.4–26.2</td>
|
||
<td id="00464A44FF9197BBFC63FD36F7C1FD47" box="[935,1011,719,765]" gridcol="6" gridrow="1" pageId="3" pageNumber="55">--</td>
|
||
<td id="00464A44FF9197BBFBC8FD36F0B5FD47" box="[1036,1159,719,765]" gridcol="7" gridrow="1" pageId="3" pageNumber="55">29.9–39.5</td>
|
||
<td id="00464A44FF9197BBFB64FD36F136FD47" box="[1184,1284,719,765]" gridcol="8" gridrow="1" pageId="3" pageNumber="55">23.8–26.1</td>
|
||
<td id="00464A44FF9197BBFAD2FD36F1A9FD47" box="[1302,1435,719,765]" gridcol="9" gridrow="1" pageId="3" pageNumber="55">21.5–23.4</td>
|
||
</tr>
|
||
<tr id="43972338FF9197BBFF65FCEBF1A9FCE0" box="[161,1435,786,858]" gridrow="2" pageId="3" pageNumber="55">
|
||
<th id="00464A44FF9197BBFF65FCEBF536FCE0" box="[161,260,786,858]" gridcol="0" gridrow="2" pageId="3" pageNumber="55">SVL females (mm)</th>
|
||
<td id="00464A44FF9197BBFEDCFCEBF549FCE0" box="[280,379,786,858]" gridcol="1" gridrow="2" pageId="3" pageNumber="55">27.0</td>
|
||
<td id="00464A44FF9197BBFE50FCEBF5CAFCE0" box="[404,504,786,858]" gridcol="2" gridrow="2" pageId="3" pageNumber="55">28.3–31.0</td>
|
||
<td id="00464A44FF9197BBFDD7FCEBF64DFCE0" box="[531,639,786,858]" gridcol="3" gridrow="2" pageId="3" pageNumber="55">25.3–27.1</td>
|
||
<td id="00464A44FF9197BBFD52FCEBF73DFCE0" box="[662,783,786,858]" gridcol="4" gridrow="2" pageId="3" pageNumber="55">34.8–44.9</td>
|
||
<td id="00464A44FF9197BBFCE1FCEBF7A1FCE0" box="[805,915,786,858]" gridcol="5" gridrow="2" pageId="3" pageNumber="55">26.3–32.2</td>
|
||
<td id="00464A44FF9197BBFC63FCEBF7C1FCE0" box="[935,1011,786,858]" gridcol="6" gridrow="2" pageId="3" pageNumber="55">38.9</td>
|
||
<td id="00464A44FF9197BBFBC8FCEBF0B5FCE0" box="[1036,1159,786,858]" gridcol="7" gridrow="2" pageId="3" pageNumber="55">32.3–40.5</td>
|
||
<td id="00464A44FF9197BBFB64FCEBF136FCE0" box="[1184,1284,786,858]" gridcol="8" gridrow="2" pageId="3" pageNumber="55">27.0–33.2</td>
|
||
<td id="00464A44FF9197BBFAD2FCEBF1A9FCE0" box="[1302,1435,786,858]" gridcol="9" gridrow="2" pageId="3" pageNumber="55">29.5</td>
|
||
</tr>
|
||
<tr id="43972338FF9197BBFF65FC97F1A9FC26" box="[161,1435,878,924]" gridrow="3" pageId="3" pageNumber="55">
|
||
<th id="00464A44FF9197BBFF65FC97F536FC26" box="[161,260,878,924]" gridcol="0" gridrow="3" pageId="3" pageNumber="55">TD/ED ratio males</th>
|
||
<td id="00464A44FF9197BBFEDCFC97F549FC26" box="[280,379,878,924]" gridcol="1" gridrow="3" pageId="3" pageNumber="55">1.45 (1.4–1.5)</td>
|
||
<td id="00464A44FF9197BBFE50FC97F5CAFC26" box="[404,504,878,924]" gridcol="2" gridrow="3" pageId="3" pageNumber="55">0.77 ± 0.08 (0.67–0.87)</td>
|
||
<td id="00464A44FF9197BBFDD7FC97F64DFC26" box="[531,639,878,924]" gridcol="3" gridrow="3" pageId="3" pageNumber="55">1.27 ± 0.10 (1.10–1.38)</td>
|
||
<td id="00464A44FF9197BBFD52FC97F73DFC26" box="[662,783,878,924]" gridcol="4" gridrow="3" pageId="3" pageNumber="55">1.17 ± 0.08 (1.05–1.31)</td>
|
||
<td id="00464A44FF9197BBFCE1FC97F7A1FC26" box="[805,915,878,924]" gridcol="5" gridrow="3" pageId="3" pageNumber="55">0.92 ± 0.13 (0.70–1.13)</td>
|
||
<td id="00464A44FF9197BBFC63FC97F7C1FC26" box="[935,1011,878,924]" gridcol="6" gridrow="3" pageId="3" pageNumber="55">--</td>
|
||
<td id="00464A44FF9197BBFBC8FC97F0B5FC26" box="[1036,1159,878,924]" gridcol="7" gridrow="3" pageId="3" pageNumber="55">1.24 ± 0.16 (0.93–1.47)</td>
|
||
<td id="00464A44FF9197BBFB64FC97F136FC26" box="[1184,1284,878,924]" gridcol="8" gridrow="3" pageId="3" pageNumber="55">1.34 ± 0.14 (1.20–1.50)</td>
|
||
<td id="00464A44FF9197BBFAD2FC97F1A9FC26" box="[1302,1435,878,924]" gridcol="9" gridrow="3" pageId="3" pageNumber="55">1.24 ± 0.16 (1.03–1.44)</td>
|
||
</tr>
|
||
<tr id="43972338FF9197BBFF65FC48F1A9FC43" box="[161,1435,945,1017]" gridrow="4" pageId="3" pageNumber="55">
|
||
<th id="00464A44FF9197BBFF65FC48F536FC43" box="[161,260,945,1017]" gridcol="0" gridrow="4" pageId="3" pageNumber="55">TD/ED ratio females</th>
|
||
<td id="00464A44FF9197BBFEDCFC48F549FC43" box="[280,379,945,1017]" gridcol="1" gridrow="4" pageId="3" pageNumber="55">1.04</td>
|
||
<td id="00464A44FF9197BBFE50FC48F5CAFC43" box="[404,504,945,1017]" gridcol="2" gridrow="4" pageId="3" pageNumber="55">0.68 ± 0.05 (0.61–0.74)</td>
|
||
<td id="00464A44FF9197BBFDD7FC48F64DFC43" box="[531,639,945,1017]" gridcol="3" gridrow="4" pageId="3" pageNumber="55">0.73–0.74</td>
|
||
<td id="00464A44FF9197BBFD52FC48F73DFC43" box="[662,783,945,1017]" gridcol="4" gridrow="4" pageId="3" pageNumber="55">0.82 ± 0.07 (0.70–091)</td>
|
||
<td id="00464A44FF9197BBFCE1FC48F7A1FC43" box="[805,915,945,1017]" gridcol="5" gridrow="4" pageId="3" pageNumber="55">0.75 ± 0.08 (0.62–0.84)</td>
|
||
<td id="00464A44FF9197BBFC63FC48F7C1FC43" box="[935,1011,945,1017]" gridcol="6" gridrow="4" pageId="3" pageNumber="55">0.66</td>
|
||
<td id="00464A44FF9197BBFBC8FC48F0B5FC43" box="[1036,1159,945,1017]" gridcol="7" gridrow="4" pageId="3" pageNumber="55">0.80 ± 0.03 (0.76–0.85)</td>
|
||
<td id="00464A44FF9197BBFB64FC48F136FC43" box="[1184,1284,945,1017]" gridcol="8" gridrow="4" pageId="3" pageNumber="55">0.81 ± 0.09 (0.62–0.91)</td>
|
||
<td id="00464A44FF9197BBFAD2FC48F1A9FC43" box="[1302,1435,945,1017]" gridcol="9" gridrow="4" pageId="3" pageNumber="55">0.79</td>
|
||
</tr>
|
||
<tr id="43972338FF9197BBFF65FBF7F1A9FB81" box="[161,1435,1038,1083]" gridrow="5" pageId="3" pageNumber="55">
|
||
<th id="00464A44FF9197BBFF65FBF7F536FB81" box="[161,260,1038,1083]" gridcol="0" gridrow="5" pageId="3" pageNumber="55">FOL/SVL ratio</th>
|
||
<td id="00464A44FF9197BBFEDCFBF7F549FB81" box="[280,379,1038,1083]" gridcol="1" gridrow="5" pageId="3" pageNumber="55">0.48 (0.46–0.49)</td>
|
||
<td id="00464A44FF9197BBFE50FBF7F5CAFB81" box="[404,504,1038,1083]" gridcol="2" gridrow="5" pageId="3" pageNumber="55">0.56 ± 0.03 (0.51–0.60)</td>
|
||
<td id="00464A44FF9197BBFDD7FBF7F64DFB81" box="[531,639,1038,1083]" gridcol="3" gridrow="5" pageId="3" pageNumber="55">0.51 ± 0.05 (0.46–0.63)</td>
|
||
<td id="00464A44FF9197BBFD52FBF7F73DFB81" box="[662,783,1038,1083]" gridcol="4" gridrow="5" pageId="3" pageNumber="55">0.66 ± 0.04 (0.60–0.71)</td>
|
||
<td id="00464A44FF9197BBFCE1FBF7F7A1FB81" box="[805,915,1038,1083]" gridcol="5" gridrow="5" pageId="3" pageNumber="55">0.49 ± 0.03 (0.45–0.56)</td>
|
||
<td id="00464A44FF9197BBFC63FBF7F7C1FB81" box="[935,1011,1038,1083]" gridcol="6" gridrow="5" pageId="3" pageNumber="55">0.64</td>
|
||
<td id="00464A44FF9197BBFBC8FBF7F0B5FB81" box="[1036,1159,1038,1083]" gridcol="7" gridrow="5" pageId="3" pageNumber="55">0.54 ± 0.06 (0.51–0.80)</td>
|
||
<td id="00464A44FF9197BBFB64FBF7F136FB81" box="[1184,1284,1038,1083]" gridcol="8" gridrow="5" pageId="3" pageNumber="55">0.54 ± 0.04 (0.49–0.62)</td>
|
||
<td id="00464A44FF9197BBFAD2FBF7F1A9FB81" box="[1302,1435,1038,1083]" gridcol="9" gridrow="5" pageId="3" pageNumber="55">0.52 ± 0.03 (0.47–0.55)</td>
|
||
</tr>
|
||
<tr id="43972338FF9197BBFF65FBB6F1A9FBC7" box="[161,1435,1103,1149]" gridrow="6" pageId="3" pageNumber="55">
|
||
<th id="00464A44FF9197BBFF65FBB6F536FBC7" box="[161,260,1103,1149]" gridcol="0" gridrow="6" pageId="3" pageNumber="55">HAL/SVL ratio</th>
|
||
<td id="00464A44FF9197BBFEDCFBB6F549FBC7" box="[280,379,1103,1149]" gridcol="1" gridrow="6" pageId="3" pageNumber="55">0.28 (0.26–0.29)</td>
|
||
<td id="00464A44FF9197BBFE50FBB6F5CAFBC7" box="[404,504,1103,1149]" gridcol="2" gridrow="6" pageId="3" pageNumber="55">0.29 ± 0.01 (0.27–0.32)</td>
|
||
<td id="00464A44FF9197BBFDD7FBB6F64DFBC7" box="[531,639,1103,1149]" gridcol="3" gridrow="6" pageId="3" pageNumber="55">0.29 ± 0.02 (0.27–0.31)</td>
|
||
<td id="00464A44FF9197BBFD52FBB6F73DFBC7" box="[662,783,1103,1149]" gridcol="4" gridrow="6" pageId="3" pageNumber="55">0.30 ± 0.02 (0.27–0.32)</td>
|
||
<td id="00464A44FF9197BBFCE1FBB6F7A1FBC7" box="[805,915,1103,1149]" gridcol="5" gridrow="6" pageId="3" pageNumber="55">0.29 ± 0.01 (0.27–0.33)</td>
|
||
<td id="00464A44FF9197BBFC63FBB6F7C1FBC7" box="[935,1011,1103,1149]" gridcol="6" gridrow="6" pageId="3" pageNumber="55">0.30</td>
|
||
<td id="00464A44FF9197BBFBC8FBB6F0B5FBC7" box="[1036,1159,1103,1149]" gridcol="7" gridrow="6" pageId="3" pageNumber="55">0.27 ± 0.01 (0.25–0.31)</td>
|
||
<td id="00464A44FF9197BBFB64FBB6F136FBC7" box="[1184,1284,1103,1149]" gridcol="8" gridrow="6" pageId="3" pageNumber="55">0.29 ± 0.02 (0.25–0.32)</td>
|
||
<td id="00464A44FF9197BBFAD2FBB6F1A9FBC7" box="[1302,1435,1103,1149]" gridcol="9" gridrow="6" pageId="3" pageNumber="55">0.30 ± 0.02 (0.27–0.33)</td>
|
||
</tr>
|
||
<tr id="43972338FF9197BBFF65FB6BF1A9FB60" box="[161,1435,1170,1242]" gridrow="7" pageId="3" pageNumber="55">
|
||
<th id="00464A44FF9197BBFF65FB6BF536FB60" box="[161,260,1170,1242]" gridcol="0" gridrow="7" pageId="3" pageNumber="55">RHL of most specimens</th>
|
||
<td id="00464A44FF9197BBFEDCFB6BF549FB60" box="[280,379,1170,1242]" gridcol="1" gridrow="7" pageId="3" pageNumber="55">TT reaches at most nostril</td>
|
||
<td id="00464A44FF9197BBFE50FB6BF5CAFB60" box="[404,504,1170,1242]" gridcol="2" gridrow="7" pageId="3" pageNumber="55">TT reaches snout tip or beyond</td>
|
||
<td id="00464A44FF9197BBFDD7FB6BF64DFB60" box="[531,639,1170,1242]" gridcol="3" gridrow="7" pageId="3" pageNumber="55">TT reaches at most nostril</td>
|
||
<td id="00464A44FF9197BBFD52FB6BF73DFB60" box="[662,783,1170,1242]" gridcol="4" gridrow="7" pageId="3" pageNumber="55">TT reaches beyond snout tip</td>
|
||
<td id="00464A44FF9197BBFCE1FB6BF7A1FB60" box="[805,915,1170,1242]" gridcol="5" gridrow="7" pageId="3" pageNumber="55">TT reaches at most nostril</td>
|
||
<td id="00464A44FF9197BBFC63FB6BF7C1FB60" box="[935,1011,1170,1242]" gridcol="6" gridrow="7" pageId="3" pageNumber="55">--</td>
|
||
<td id="00464A44FF9197BBFBC8FB6BF0B5FB60" box="[1036,1159,1170,1242]" gridcol="7" gridrow="7" pageId="3" pageNumber="55">TT reaches beyond snout tip</td>
|
||
<td id="00464A44FF9197BBFB64FB6BF136FB60" box="[1184,1284,1170,1242]" gridcol="8" gridrow="7" pageId="3" pageNumber="55">TT reaches snout tip or beyond</td>
|
||
<td id="00464A44FF9197BBFAD2FB6BF1A9FB60" box="[1302,1435,1170,1242]" gridcol="9" gridrow="7" pageId="3" pageNumber="55">TT does not reach snout tip</td>
|
||
</tr>
|
||
<tr id="43972338FF9197BBFF65FB17F1A9FAA6" box="[161,1435,1262,1308]" gridrow="8" pageId="3" pageNumber="55">
|
||
<th id="00464A44FF9197BBFF65FB17F536FAA6" box="[161,260,1262,1308]" gridcol="0" gridrow="8" pageId="3" pageNumber="55">Webbing of toe 5</th>
|
||
<td id="00464A44FF9197BBFEDCFB17F549FAA6" box="[280,379,1262,1308]" gridcol="1" gridrow="8" pageId="3" pageNumber="55">2</td>
|
||
<td id="00464A44FF9197BBFE50FB17F5CAFAA6" box="[404,504,1262,1308]" gridcol="2" gridrow="8" pageId="3" pageNumber="55">0.75–1</td>
|
||
<td id="00464A44FF9197BBFDD7FB17F64DFAA6" box="[531,639,1262,1308]" gridcol="3" gridrow="8" pageId="3" pageNumber="55">1.5–2</td>
|
||
<td id="00464A44FF9197BBFD52FB17F73DFAA6" box="[662,783,1262,1308]" gridcol="4" gridrow="8" pageId="3" pageNumber="55">2</td>
|
||
<td id="00464A44FF9197BBFCE1FB17F7A1FAA6" box="[805,915,1262,1308]" gridcol="5" gridrow="8" pageId="3" pageNumber="55">1.25</td>
|
||
<td id="00464A44FF9197BBFC63FB17F7C1FAA6" box="[935,1011,1262,1308]" gridcol="6" gridrow="8" pageId="3" pageNumber="55">--</td>
|
||
<td id="00464A44FF9197BBFBC8FB17F0B5FAA6" box="[1036,1159,1262,1308]" gridcol="7" gridrow="8" pageId="3" pageNumber="55">1</td>
|
||
<td id="00464A44FF9197BBFB64FB17F136FAA6" box="[1184,1284,1262,1308]" gridcol="8" gridrow="8" pageId="3" pageNumber="55">1</td>
|
||
<td id="00464A44FF9197BBFAD2FB17F1A9FAA6" box="[1302,1435,1262,1308]" gridcol="9" gridrow="8" pageId="3" pageNumber="55">1.5</td>
|
||
</tr>
|
||
<tr id="43972338FF9197BBFF65FAC8F1A9FA2E" box="[161,1435,1329,1428]" gridrow="9" pageId="3" pageNumber="55">
|
||
<th id="00464A44FF9197BBFF65FAC8F536FA2E" box="[161,260,1329,1428]" gridcol="0" gridrow="9" pageId="3" pageNumber="55">Femoral glands in males</th>
|
||
<td id="00464A44FF9197BBFEDCFAC8F549FA2E" box="[280,379,1329,1428]" gridcol="1" gridrow="9" pageId="3" pageNumber="55">large, prominent</td>
|
||
<td id="00464A44FF9197BBFE50FAC8F5CAFA2E" box="[404,504,1329,1428]" gridcol="2" gridrow="9" pageId="3" pageNumber="55">large, prominent</td>
|
||
<td id="00464A44FF9197BBFDD7FAC8F64DFA2E" box="[531,639,1329,1428]" gridcol="3" gridrow="9" pageId="3" pageNumber="55">large, prominent</td>
|
||
<td id="00464A44FF9197BBFD52FAC8F73DFA2E" box="[662,783,1329,1428]" gridcol="4" gridrow="9" pageId="3" pageNumber="55">large, very prominent</td>
|
||
<td id="00464A44FF9197BBFCE1FAC8F7A1FA2E" box="[805,915,1329,1428]" gridcol="5" gridrow="9" pageId="3" pageNumber="55">small to large, not very prominent</td>
|
||
<td id="00464A44FF9197BBFC63FAC8F7C1FA2E" box="[935,1011,1329,1428]" gridcol="6" gridrow="9" pageId="3" pageNumber="55">--</td>
|
||
<td id="00464A44FF9197BBFBC8FAC8F0B5FA2E" box="[1036,1159,1329,1428]" gridcol="7" gridrow="9" pageId="3" pageNumber="55">small, i ndistinct</td>
|
||
<td id="00464A44FF9197BBFB64FAC8F136FA2E" box="[1184,1284,1329,1428]" gridcol="8" gridrow="9" pageId="3" pageNumber="55">small, indistinct</td>
|
||
<td id="00464A44FF9197BBFAD2FAC8F1A9FA2E" box="[1302,1435,1329,1428]" gridcol="9" gridrow="9" pageId="3" pageNumber="55">large, not very prominent</td>
|
||
</tr>
|
||
<tr id="43972338FF9197BBFF65FA51F1A9F99C" box="[161,1435,1448,1574]" gridrow="10" pageId="3" pageNumber="55">
|
||
<th id="00464A44FF9197BBFF65FA51F536F99C" box="[161,260,1448,1574]" gridcol="0" gridrow="10" pageId="3" pageNumber="55">Frenal stripe between forelimb and eye</th>
|
||
<td id="00464A44FF9197BBFEDCFA51F549F99C" box="[280,379,1448,1574]" gridcol="1" gridrow="10" pageId="3" pageNumber="55">present</td>
|
||
<td id="00464A44FF9197BBFE50FA51F5CAF99C" box="[404,504,1448,1574]" gridcol="2" gridrow="10" pageId="3" pageNumber="55">absent</td>
|
||
<td id="00464A44FF9197BBFDD7FA51F64DF99C" box="[531,639,1448,1574]" gridcol="3" gridrow="10" pageId="3" pageNumber="55">present</td>
|
||
<td id="00464A44FF9197BBFD52FA51F73DF99C" box="[662,783,1448,1574]" gridcol="4" gridrow="10" pageId="3" pageNumber="55">present</td>
|
||
<td id="00464A44FF9197BBFCE1FA51F7A1F99C" box="[805,915,1448,1574]" gridcol="5" gridrow="10" pageId="3" pageNumber="55">present</td>
|
||
<td id="00464A44FF9197BBFC63FA51F7C1F99C" box="[935,1011,1448,1574]" gridcol="6" gridrow="10" pageId="3" pageNumber="55">present</td>
|
||
<td id="00464A44FF9197BBFBC8FA51F0B5F99C" box="[1036,1159,1448,1574]" gridcol="7" gridrow="10" pageId="3" pageNumber="55">present</td>
|
||
<td id="00464A44FF9197BBFB64FA51F136F99C" box="[1184,1284,1448,1574]" gridcol="8" gridrow="10" pageId="3" pageNumber="55">present</td>
|
||
<td id="00464A44FF9197BBFAD2FA51F1A9F99C" box="[1302,1435,1448,1574]" gridcol="9" gridrow="10" pageId="3" pageNumber="55">present</td>
|
||
</tr>
|
||
<tr id="43972338FF9197BBFF65F9C3F1A9F902" box="[161,1435,1594,1720]" gridrow="11" pageId="3" pageNumber="55">
|
||
<th id="00464A44FF9197BBFF65F9C3F536F902" box="[161,260,1594,1720]" gridcol="0" gridrow="11" pageId="3" pageNumber="55">Frenal stripe between eye and nostril</th>
|
||
<td id="00464A44FF9197BBFEDCF9C3F549F902" box="[280,379,1594,1720]" gridcol="1" gridrow="11" pageId="3" pageNumber="55">present, reaches close to nostril</td>
|
||
<td id="00464A44FF9197BBFE50F9C3F5CAF902" box="[404,504,1594,1720]" gridcol="2" gridrow="11" pageId="3" pageNumber="55">absent</td>
|
||
<td id="00464A44FF9197BBFDD7F9C3F64DF902" box="[531,639,1594,1720]" gridcol="3" gridrow="11" pageId="3" pageNumber="55">present, reaches nostril or close to it</td>
|
||
<td id="00464A44FF9197BBFD52F9C3F73DF902" box="[662,783,1594,1720]" gridcol="4" gridrow="11" pageId="3" pageNumber="55">present, reaches close to nostril</td>
|
||
<td id="00464A44FF9197BBFCE1F9C3F7A1F902" box="[805,915,1594,1720]" gridcol="5" gridrow="11" pageId="3" pageNumber="55">usually stops anterior to the eye</td>
|
||
<td id="00464A44FF9197BBFC63F9C3F7C1F902" box="[935,1011,1594,1720]" gridcol="6" gridrow="11" pageId="3" pageNumber="55">slightly fading anterior to eye</td>
|
||
<td id="00464A44FF9197BBFBC8F9C3F0B5F902" box="[1036,1159,1594,1720]" gridcol="7" gridrow="11" pageId="3" pageNumber="55">present, reaches nostril or close to it</td>
|
||
<td id="00464A44FF9197BBFB64F9C3F136F902" box="[1184,1284,1594,1720]" gridcol="8" gridrow="11" pageId="3" pageNumber="55">fades ante- rior to eye</td>
|
||
<td id="00464A44FF9197BBFAD2F9C3F1A9F902" box="[1302,1435,1594,1720]" gridcol="9" gridrow="11" pageId="3" pageNumber="55">fades anterior to eye</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="FD18217AFF916847FF65F935F141F965" blockId="3.[161,1435,1740,2011]" box="[161,1395,1740,1759]" pageId="3" pageNumber="55">Throat black or black with a grey with light with black, with light black or dark black or light grey–</paragraph>
|
||
<paragraph id="FD18217AFF916847FF65F91EF766F80F" blockId="3.[161,1435,1740,2011]" pageId="3" pageNumber="55">coloration dark brown, distinct large white indistinct, small white brown, with a dark yellowish, with with a white stripe spots weak darker spots which median line or brown, a rather broad median line which is forming a marbling form a row of spots with a light median and spots continued median row median row, median line stripe which onto the ven- sometimes or row of begins as dister fusing to a spots tinct white spot median close to the lip stripe</paragraph>
|
||
<paragraph id="FD18217AFF916840FAE6F830F120FCA0" blockId="3.[161,1435,1740,2011]" lastBlockId="4.[151,1436,698,2019]" lastPageId="4" lastPageNumber="56" pageId="3" pageNumber="55">
|
||
<emphasis id="CFD3FD68FF916847FAE6F830F1A9F861" box="[1314,1435,1993,2011]" italics="true" pageId="3" pageNumber="55">...... continued</emphasis>
|
||
<emphasis id="CFD3FD68FF966840FF03FD42F57CFD69" bold="true" box="[199,334,699,723]" pageId="4" pageNumber="56">Etymology.</emphasis>
|
||
The specific name—paidroa—refers to the typical advertisement call of the species, and derives from
|
||
<collectingCountry id="85B061EAFF966840FF11FD27F570FD4C" box="[213,322,734,758]" name="Madagascar" pageId="4" pageNumber="56">Malagasy</collectingCountry>
|
||
words paika (meaning notes) and roa (double or two) (according to
|
||
<collectingCountry id="85B061EAFF966840FB84FD27F09FFD4C" box="[1088,1197,734,758]" name="Madagascar" pageId="4" pageNumber="56">Malagasy</collectingCountry>
|
||
grammar, the ending —ka is replaced by —d in the composite word). The name is used as an invariable noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="A9D871F2FF966840FF65FF5BF5BDFF03" ID-Table-UUID="A9D871F2FF966840FF65FF5BF5BDFF03" box="[161,399,162,186]" httpUri="http://table.plazi.org/id/A9D871F2FF966840FF65FF5BF5BDFF03" pageId="4" pageNumber="56" targetBox="[161,1361,207,638]" targetIsTable="true" targetPageId="4">
|
||
<paragraph id="FD18217AFF966840FF65FF5BF5BDFF03" blockId="4.[161,399,162,186]" box="[161,399,162,186]" pageId="4" pageNumber="56">
|
||
<emphasis id="CFD3FD68FF966840FF65FF5BF53FFF00" bold="true" box="[161,269,162,186]" pageId="4" pageNumber="56">TABLE 1</emphasis>
|
||
(continued)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FD18217AFF966840FEDCFF36F10AFDD9" pageId="4" pageNumber="56">
|
||
<table id="8FA7D3DAFF9697BBFF65FF36F163FDC4" box="[161,1361,207,638]" gridcols="9" gridrows="4" pageId="4" pageNumber="56">
|
||
<tr id="43972338FF9697BBFF65FF36F163FF46" box="[161,1361,207,252]" gridrow="0" pageId="4" pageNumber="56">
|
||
<th id="00464A44FF9697BBFF65FF36F545FF46" box="[161,375,207,252]" gridcol="0" gridrow="0" pageId="4" pageNumber="56">
|
||
<taxonomicName id="3AA75AF9FF966840FEDCFF36F545FF5B" box="[280,375,207,225]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="4" pageNumber="56" phylum="Chordata" rank="species" species="paidroa" status="sp. nov.">
|
||
<emphasis id="CFD3FD68FF966840FEDCFF36F545FF5B" box="[280,375,207,225]" italics="true" pageId="4" pageNumber="56">M. paidroa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D4E04013FF966840FEDCFF10F569FF46" box="[280,347,233,252]" pageId="4" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</th>
|
||
<th id="00464A44FF9697BBFE50FF36F5C6FF46" box="[404,500,207,252]" gridcol="1" gridrow="0" pageId="4" pageNumber="56">
|
||
<taxonomicName id="3AA75AF9FF966840FE50FF36F5C6FF46" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="4" pageNumber="56" phylum="Chordata" rank="species" species="aerumnalis">
|
||
<emphasis id="CFD3FD68FF966840FE50FF36F5C6FF46" italics="true" pageId="4" pageNumber="56">M. aerumnalis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="00464A44FF9697BBFDD0FF36F64DFF46" box="[532,639,207,252]" gridcol="2" gridrow="0" pageId="4" pageNumber="56">
|
||
<taxonomicName id="3AA75AF9FF966840FDD0FF36F64DFF46" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="4" pageNumber="56" phylum="Chordata" rank="species" species="albofrenatus">
|
||
<emphasis id="CFD3FD68FF966840FDD0FF36F64DFF46" italics="true" pageId="4" pageNumber="56">M. albofrenatus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="00464A44FF9697BBFD52FF36F73DFF46" box="[662,783,207,252]" gridcol="3" gridrow="0" pageId="4" pageNumber="56">
|
||
<taxonomicName id="3AA75AF9FF966840FD52FF36F73DFF46" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="4" pageNumber="56" phylum="Chordata" rank="species" species="brevipalmatus">
|
||
<emphasis id="CFD3FD68FF966840FD52FF36F73DFF46" italics="true" pageId="4" pageNumber="56">M. brevipalmatus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="00464A44FF9697BBFCE1FF36F7B4FF46" box="[805,902,207,252]" gridcol="4" gridrow="0" pageId="4" pageNumber="56">
|
||
<taxonomicName id="3AA75AF9FF966840FCE1FF36F7B4FF46" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="4" pageNumber="56" phylum="Chordata" rank="species" species="charlotteae">
|
||
<emphasis id="CFD3FD68FF966840FCE1FF36F7B4FF46" italics="true" pageId="4" pageNumber="56">M. charlotteae</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="00464A44FF9697BBFC63FF36F7C4FF46" box="[935,1014,207,252]" gridcol="5" gridrow="0" pageId="4" pageNumber="56">
|
||
<taxonomicName id="3AA75AF9FF966840FC63FF36F7C1FF46" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="4" pageNumber="56" phylum="Chordata" rank="species" species="delormei">
|
||
<emphasis id="CFD3FD68FF966840FC63FF36F7C1FF46" italics="true" pageId="4" pageNumber="56">M. delormei</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="00464A44FF9697BBFBC8FF36F0B0FF46" box="[1036,1154,207,252]" gridcol="6" gridrow="0" pageId="4" pageNumber="56">
|
||
<taxonomicName id="3AA75AF9FF966840FBC8FF36F0B0FF46" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="4" pageNumber="56" phylum="Chordata" rank="species" species="melanopleura">
|
||
<emphasis id="CFD3FD68FF966840FBC8FF36F0B0FF46" italics="true" pageId="4" pageNumber="56">M. melanopleura</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="00464A44FF9697BBFB64FF36F0D5FF46" box="[1184,1255,207,252]" gridcol="7" gridrow="0" pageId="4" pageNumber="56">
|
||
<taxonomicName id="3AA75AF9FF966840FB64FF36F0D5FF46" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="4" pageNumber="56" phylum="Chordata" rank="species" species="opiparis">
|
||
<emphasis id="CFD3FD68FF966840FB64FF36F0D5FF46" italics="true" pageId="4" pageNumber="56">M. opiparis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="00464A44FF9697BBFAD2FF36F163FF46" box="[1302,1361,207,252]" gridcol="8" gridrow="0" pageId="4" pageNumber="56">
|
||
<taxonomicName id="3AA75AF9FF966840FAD2FF36F163FF46" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="4" pageNumber="56" phylum="Chordata" rank="species" species="zipperi">
|
||
<emphasis id="CFD3FD68FF966840FAD2FF36F163FF46" italics="true" pageId="4" pageNumber="56">M. zipperi</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="43972338FF9697BBFF65FEE8F163FE35" box="[161,1361,273,399]" gridrow="1" pageId="4" pageNumber="56">
|
||
<th id="00464A44FF9697BBFF65FEE8F545FE35" box="[161,375,273,399]" gridcol="0" gridrow="1" pageId="4" pageNumber="56">Duration of 8–14 single notes in adver- tisement calls (ms)</th>
|
||
<td id="00464A44FF9697BBFE50FEE8F5C6FE35" box="[404,500,273,399]" gridcol="1" gridrow="1" pageId="4" pageNumber="56">unknown</td>
|
||
<td id="00464A44FF9697BBFDD0FEE8F64DFE35" box="[532,639,273,399]" gridcol="2" gridrow="1" pageId="4" pageNumber="56">56–80</td>
|
||
<td id="00464A44FF9697BBFD52FEE8F73DFE35" box="[662,783,273,399]" gridcol="3" gridrow="1" pageId="4" pageNumber="56">9–21</td>
|
||
<td id="00464A44FF9697BBFCE1FEE8F7B4FE35" box="[805,902,273,399]" gridcol="4" gridrow="1" pageId="4" pageNumber="56">91–304</td>
|
||
<td id="00464A44FF9697BBFC63FEE8F7C4FE35" box="[935,1014,273,399]" gridcol="5" gridrow="1" pageId="4" pageNumber="56">unknown</td>
|
||
<td id="00464A44FF9697BBFBC8FEE8F0B0FE35" box="[1036,1154,273,399]" gridcol="6" gridrow="1" pageId="4" pageNumber="56">19–54</td>
|
||
<td id="00464A44FF9697BBFB64FEE8F0D5FE35" box="[1184,1255,273,399]" gridcol="7" gridrow="1" pageId="4" pageNumber="56">69–126</td>
|
||
<td id="00464A44FF9697BBFAD2FEE8F163FE35" box="[1302,1361,273,399]" gridcol="8" gridrow="1" pageId="4" pageNumber="56">21–31</td>
|
||
</tr>
|
||
<tr id="43972338FF9697BBFF65FE5AF163FD81" box="[161,1361,419,571]" gridrow="2" pageId="4" pageNumber="56">
|
||
<th id="00464A44FF9697BBFF65FE5AF545FD81" box="[161,375,419,571]" gridcol="0" gridrow="2" pageId="4" pageNumber="56">Duration of 12–16 double notes in advertise- ment calls (ms)</th>
|
||
<td id="00464A44FF9697BBFE50FE5AF5C6FD81" box="[404,500,419,571]" gridcol="1" gridrow="2" pageId="4" pageNumber="56">unknown</td>
|
||
<td id="00464A44FF9697BBFDD0FE5AF64DFD81" box="[532,639,419,571]" gridcol="2" gridrow="2" pageId="4" pageNumber="56">absent</td>
|
||
<td id="00464A44FF9697BBFD52FE5AF73DFD81" box="[662,783,419,571]" gridcol="3" gridrow="2" pageId="4" pageNumber="56">absent</td>
|
||
<td id="00464A44FF9697BBFCE1FE5AF7B4FD81" box="[805,902,419,571]" gridcol="4" gridrow="2" pageId="4" pageNumber="56">absent</td>
|
||
<td id="00464A44FF9697BBFC63FE5AF7C4FD81" box="[935,1014,419,571]" gridcol="5" gridrow="2" pageId="4" pageNumber="56">unknown</td>
|
||
<td id="00464A44FF9697BBFBC8FE5AF0B0FD81" box="[1036,1154,419,571]" gridcol="6" gridrow="2" pageId="4" pageNumber="56">absent</td>
|
||
<td id="00464A44FF9697BBFB64FE5AF0D5FD81" box="[1184,1255,419,571]" gridcol="7" gridrow="2" pageId="4" pageNumber="56">absent</td>
|
||
<td id="00464A44FF9697BBFAD2FE5AF163FD81" box="[1302,1361,419,571]" gridcol="8" gridrow="2" pageId="4" pageNumber="56">absent</td>
|
||
</tr>
|
||
<tr id="43972338FF9697BBFF65FDA9F163FDC4" box="[161,1361,592,638]" gridrow="3" pageId="4" pageNumber="56">
|
||
<th id="00464A44FF9697BBFF65FDA9F545FDC4" box="[161,375,592,638]" gridcol="0" gridrow="3" pageId="4" pageNumber="56">Relativetoe 5<3 length</th>
|
||
<td id="00464A44FF9697BBFE50FDA9F5C6FDC4" box="[404,500,592,638]" gridcol="1" gridrow="3" pageId="4" pageNumber="56">5=3</td>
|
||
<td id="00464A44FF9697BBFDD0FDA9F64DFDC4" box="[532,639,592,638]" gridcol="2" gridrow="3" pageId="4" pageNumber="56">5<3</td>
|
||
<td id="00464A44FF9697BBFD52FDA9F73DFDC4" box="[662,783,592,638]" gridcol="3" gridrow="3" pageId="4" pageNumber="56">5>3</td>
|
||
<td id="00464A44FF9697BBFCE1FDA9F7B4FDC4" box="[805,902,592,638]" gridcol="4" gridrow="3" pageId="4" pageNumber="56">5<3</td>
|
||
<td id="00464A44FF9697BBFC63FDA9F7C4FDC4" box="[935,1014,592,638]" gridcol="5" gridrow="3" pageId="4" pageNumber="56">5>3</td>
|
||
<td id="00464A44FF9697BBFBC8FDA9F0B0FDC4" box="[1036,1154,592,638]" gridcol="6" gridrow="3" pageId="4" pageNumber="56">5>3</td>
|
||
<td id="00464A44FF9697BBFB64FDA9F0D5FDC4" box="[1184,1255,592,638]" gridcol="7" gridrow="3" pageId="4" pageNumber="56">5>3</td>
|
||
<td id="00464A44FF9697BBFAD2FDA9F163FDC4" box="[1302,1361,592,638]" gridcol="8" gridrow="3" pageId="4" pageNumber="56">5<3</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<subSubSection id="B5BD72F1FF966842FF03FCDFF0D7FB22" lastPageId="6" lastPageNumber="58" pageId="4" pageNumber="56" type="description">
|
||
<paragraph id="FD18217AFF966840FF03FCDFF6B5FAC4" blockId="4.[151,1436,698,2019]" pageId="4" pageNumber="56">
|
||
<emphasis id="CFD3FD68FF966840FF03FCDFF623FC85" bold="true" box="[199,529,806,831]" pageId="4" pageNumber="56">
|
||
Description of the
|
||
<typeStatus id="221C9FD8FF966840FE67FCDEF63EFC85" box="[419,524,807,831]" pageId="4" pageNumber="56" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
SVL
|
||
<quantity id="3A5F8C9FFF966840FD97FCDFF688FC84" box="[595,698,806,830]" metricMagnitude="-2" metricUnit="m" metricValue="2.23" pageId="4" pageNumber="56" unit="mm" value="22.3">22.3 mm</quantity>
|
||
. Body relatively slender; head longer than wide, of same width as body; snout rounded in dorsal view, slightly pointed in lateral view; nostrils directed laterally, slightly protuberant, nearer to tip of snout than to eye; canthus rostralis rather indistinct, straight; loreal region concave; tympanum distinct, oval, wider horizontally than vertically, its horizontal diameter 143% of eye diameter; supratympanic fold indistinct; tongue absent, taken for tissue sampling; vomerine teeth distinct, one rounded aggregation on each side of buccal roof, positioned posteromedian to choana; choanae small, rounded. Arms slender, subarticular tubercles distinct, single; inner metacarpal tubercle distinct and ovoid, outer metacarpal tubercle dinstinct but smaller and rounded; fingers without webbing; comparative finger length 1<2<4<3, second finger distinctly shorter than fourth finger; finger disks slightly enlarged; nuptial pads absent. Hindlimbs slender; tibiotarsal articulation reaches the anterior eye corner; lateral metatarsalia not connected; comparative toe length 1<2<5<3<4; third toe slightly longer than fifth toe; inner metatarsal tubercle distinct, outer metatarsal tubercle not recognizable; webbing between toes weakly expressed, formula 1 (1),
|
||
<date id="891907BAFF966840FDC8FB4BF610FB70" box="[524,546,1202,1226]" pageId="4" pageNumber="56">2i</date>
|
||
(>>1), 2e (1),
|
||
<date id="891907BAFF966840FD02FB4BF6E9FB70" box="[710,731,1202,1226]" pageId="4" pageNumber="56">3i</date>
|
||
(2.25), 3e (2),
|
||
<date id="891907BAFF966840FC47FB4BF7AAFB70" box="[899,920,1202,1226]" pageId="4" pageNumber="56">4i</date>
|
||
(3.5), 4e (3.25), 5 (2). Exact extension of web on
|
||
<date id="891907BAFF966840FF7AFB2FF4E6FB54" box="[190,212,1238,1262]" pageId="4" pageNumber="56">2i</date>
|
||
cannot be determined more in detail because only one subarticular tubercle is recognizable. Dorsal skin smooth; dorsum with distinct dorsolateral folds; ventral skin smooth, slightly granular in the cloacal region; no distinct tubercles in the cloacal region. Femoral glands large and distinct (length 5.0, width
|
||
<quantity id="3A5F8C9FFF966840FBB3FAE7F0E3FA8C" box="[1143,1233,1310,1334]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="4" pageNumber="56" unit="mm" value="3.5">3.5 mm</quantity>
|
||
, distance between internal borders of glands on opposite thighs
|
||
<quantity id="3A5F8C9FFF966840FD49FABBF6D0FAE0" box="[653,738,1346,1370]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="56" unit="mm" value="0.7">0.7 mm</quantity>
|
||
(almost in contact in the cloacal region), of
|
||
<typeStatus id="221C9FD8FF966840FB09FABAF0CFFAE1" box="[1229,1277,1347,1371]" pageId="4" pageNumber="56">type</typeStatus>
|
||
3 with central depression as defined by
|
||
<bibRefCitation id="99365C8BFF966840FE76FA9FF6B0FAC4" box="[434,642,1382,1407]" pageId="4" pageNumber="60" refString="Glaw, F., Vences, M. & Gossmann, V. (2000) A new species of Mantidactylus from Madagascar, with a comparative survey of internal femoral gland structure in the genus (Amphibia: Ranidae: Mantellinae). Journal of Natural History, 34, 1135 - 1154." type="journal article">
|
||
Glaw
|
||
<emphasis id="CFD3FD68FF966840FE31FA91F61CFAC4" box="[501,558,1382,1407]" italics="true" pageId="4" pageNumber="56">et al.</emphasis>
|
||
(2000)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="FD18217AFF966840FF03FA73F77EF95D" blockId="4.[151,1436,698,2019]" pageId="4" pageNumber="56">
|
||
<emphasis id="CFD3FD68FF966840FF03FA73F5E9FA19" bold="true" box="[199,475,1418,1443]" pageId="4" pageNumber="56">
|
||
Colour of the
|
||
<typeStatus id="221C9FD8FF966840FEA9FA72F5E4FA19" box="[365,470,1419,1443]" pageId="4" pageNumber="56" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Colour in life (
|
||
<figureCitation id="659C3DFFFF966840FD4CFA73F6F5FA18" box="[648,711,1418,1442]" captionStart="FIGURE 1" captionStartId="2.[151,250,1959,1982]" captionTargetBox="[151,1436,337,1938]" captionTargetId="figure@2.[151,1436,337,1938]" captionTargetPageId="2" captionText="FIGURE 1. (a), (b) and (c), Photographs of male holotype of Mantidactylus paidroa (ZSM 1777 / 2008) in life (SVL 22.3 mm), in dorsolateral and ventral views." httpUri="https://zenodo.org/record/202729/files/figure.png" pageId="4" pageNumber="56">Fig. 1</figureCitation>
|
||
a–c) is dorsally light brown with irregular black blotches, with a conspicuous dorsolateral bold black border that separates the dorsum from the dark brown flanks, which have a few irregular bronze and white spots. From the insertion of forelimbs there is a white frenal stripe running under the tympanum and the eye, bending upward anterior to the eye and reaching the nostril. In the area between upper lip and nostril there are small bronze to light brown flecks. The iris is light brown in its upper and lower parts, with reddish brown colour in its anterior and posterior parts. The hindlimbs are light brown with dark brown transversal stripes. The ventral side is dark brown on the throat and greyish on the venter, with rounded white spots and a discontinuous median narrow white stripe running from the snout tip to the venter. The limbs are ventrally blackish with dark grey blotches. After two years in preservative, the colour tones are much less contrasted and especially the white and bronze elements have partly faded to dirty grey.
|
||
</paragraph>
|
||
<paragraph id="FD18217AFF966840FF03F90BF78CF8E8" blockId="4.[151,1436,698,2019]" pageId="4" pageNumber="56">
|
||
<emphasis id="CFD3FD68FF966840FF03F90BF509F8B1" bold="true" box="[199,315,1778,1803]" pageId="4" pageNumber="56">Variation.</emphasis>
|
||
The male
|
||
<typeStatus id="221C9FD8FF966840FE7CF90AF62AF8B1" box="[440,536,1779,1803]" pageId="4" pageNumber="56" type="paratype">paratype</typeStatus>
|
||
is morphologically similar to the
|
||
<typeStatus id="221C9FD8FF966840FC5DF90BF7CDF8B0" box="[921,1023,1778,1802]" pageId="4" pageNumber="56" type="holotype">holotype</typeStatus>
|
||
. The female
|
||
<typeStatus id="221C9FD8FF966840FB56F90AF0C7F8B1" box="[1170,1269,1779,1803]" pageId="4" pageNumber="56" type="paratype">paratype</typeStatus>
|
||
, in contrast, in preservative has a much lighter dorsal and ventral colouration with only indistinct white ventral spots and an indistinct white midventral line. The female has rudimentary femoral glands.
|
||
</paragraph>
|
||
<paragraph id="FD18217AFF966840FF03F8A6F1B6F858" blockId="4.[151,1436,698,2019]" pageId="4" pageNumber="56">
|
||
<emphasis id="CFD3FD68FF966840FF03F8A6F54FF8CD" bold="true" box="[199,381,1887,1911]" pageId="4" pageNumber="56">Measurements.</emphasis>
|
||
All given in mm. The first value refers to the
|
||
<typeStatus id="221C9FD8FF966840FC40F8A7F7DBF8CC" box="[900,1001,1886,1910]" pageId="4" pageNumber="56" type="holotype">holotype</typeStatus>
|
||
, the second and third values (in parentheses, male and female) to the
|
||
<typeStatus id="221C9FD8FF966840FDC0F87AF641F821" box="[516,627,1923,1947]" pageId="4" pageNumber="56" type="paratype">paratypes</typeStatus>
|
||
. SVL 22.3 (22.0; 27.0), HW 7.3 (7.4; 8.0), HL 9.5 (9.0; 10.0), TD 3.3 (3.3; 2.4), ED 2.3 (2.2; 2.3),
|
||
<collectingCountry id="85B061EAFF966840FE19F85EF625F804" box="[477,535,1959,1982]" name="India" pageId="4" pageNumber="56">END</collectingCountry>
|
||
2.4 (2.2; 2.4), NSD 1.7 (1.7; 1.8), NND 3.0 (3.3; 3.3), FORL 14.3 (13.3; 16.0), HAL 6.2 (6.3; 7.0), HIL 35.0 (36.0; 41.5), FOTL 15.5 (16.0; 18.3), FOL 10.7 (10.8; 12.5), TIL 11.1 (11.1; 13.0).
|
||
</paragraph>
|
||
<caption id="A9D871F2FF976841FF53F838F13FF84C" httpUri="https://zenodo.org/record/202730/files/figure.png" pageId="5" pageNumber="57" targetBox="[164,1427,178,1970]" targetPageId="5">
|
||
<paragraph id="FD18217AFF976841FF53F838F13FF84C" blockId="5.[151,1436,1985,2038]" pageId="5" pageNumber="57">
|
||
<emphasis id="CFD3FD68FF976841FF53F838F527F86C" bold="true" box="[151,277,1985,2008]" pageId="5" pageNumber="57">FIGURE 2.</emphasis>
|
||
Sonagrams and oscillograms of parts of a series of double notes, recorded from the holotype of
|
||
<taxonomicName id="3AA75AF9FF976841FACFF838F4D4F84C" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="57" phylum="Chordata" rank="species" species="paidroa">
|
||
<emphasis id="CFD3FD68FF976841FACFF838F4D4F84C" italics="true" pageId="5" pageNumber="57">Mantidactylus paidroa</emphasis>
|
||
</taxonomicName>
|
||
on 24 January 2008; (a) fragment of the call with 8 double notes; (b) enlarged view of two double notes.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FD18217AFF946842FF03FF61F6ECFEA1" blockId="6.[151,1437,151,1177]" pageId="6" pageNumber="58">
|
||
<emphasis id="CFD3FD68FF946842FF03FF61F5B6FF0A" bold="true" box="[199,388,151,176]" pageId="6" pageNumber="58">Natural History</emphasis>
|
||
. Calls of
|
||
<taxonomicName id="3AA75AF9FF946842FE31FF61F6C7FF0A" box="[501,757,152,176]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="paidroa">
|
||
<emphasis id="CFD3FD68FF946842FE31FF61F6C7FF0A" box="[501,757,152,176]" italics="true" pageId="6" pageNumber="58">Mantidactylus paidroa</emphasis>
|
||
</taxonomicName>
|
||
were heard only at the
|
||
<typeStatus id="221C9FD8FF946842FBC6FF61F000FF0A" box="[1026,1074,152,176]" pageId="6" pageNumber="58">type</typeStatus>
|
||
locality, Ankerana, in Ranomafana National Park, which is also the only locality where adults were collected. Other syntopic anurans at this site included two
|
||
<taxonomicName id="3AA75AF9FF946842FEF7FF26F5FAFF4D" box="[307,456,223,247]" class="Amphibia" family="Ranidae" genus="Chonomantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="58" phylum="Chordata" rank="genus">
|
||
<emphasis id="CFD3FD68FF946842FEF7FF26F5FAFF4D" box="[307,456,223,247]" italics="true" pageId="6" pageNumber="58">Chonomantis</emphasis>
|
||
</taxonomicName>
|
||
species (
|
||
<taxonomicName id="3AA75AF9FF946842FDF7FF26F70BFF4D" box="[563,825,223,247]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="opiparis">
|
||
<emphasis id="CFD3FD68FF946842FDF7FF26F70BFF4D" box="[563,825,223,247]" italics="true" pageId="6" pageNumber="58">Mantidactylus opiparis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="3AA75AF9FF946842FC8DFF19F03EFF4D" box="[841,1036,223,247]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="melanopleura">
|
||
<emphasis id="CFD3FD68FF946842FC8DFF19F03EFF4D" box="[841,1036,223,247]" italics="true" pageId="6" pageNumber="58">M. melanopleura</emphasis>
|
||
</taxonomicName>
|
||
) as well as
|
||
<taxonomicName id="3AA75AF9FF946842FB52FF26F1A6FF4D" box="[1174,1428,223,247]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="biporus">
|
||
<emphasis id="CFD3FD68FF946842FB52FF26F1A6FF4D" box="[1174,1428,223,247]" italics="true" pageId="6" pageNumber="58">Mantidactylus biporus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="3AA75AF9FF946842FF53FEFAF5B4FEA1" box="[151,390,259,283]" class="Amphibia" family="Mantellidae" genus="Gephyromantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="asper">
|
||
<emphasis id="CFD3FD68FF946842FF53FEFAF5B4FEA1" box="[151,390,259,283]" italics="true" pageId="6" pageNumber="58">Gephyromantis asper</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="CFD3FD68FF946842FE50FEFDF627FEA1" box="[404,533,259,283]" italics="true" pageId="6" pageNumber="58">
|
||
<taxonomicName id="3AA75AF9FF946842FE50FEFDF623FEA1" box="[404,529,259,283]" class="Amphibia" family="Mantellidae" genus="Gephyromantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="tschenki">G. tschenki</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="3AA75AF9FF946842FD88FEFAF685FEA1" box="[588,695,259,283]" class="Amphibia" family="Microhylidae" genus="Stumpffia" kingdom="Animalia" order="Anura" pageId="6" pageNumber="58" phylum="Chordata" rank="genus">
|
||
<emphasis id="CFD3FD68FF946842FD88FEFAF685FEA1" box="[588,695,259,283]" italics="true" pageId="6" pageNumber="58">Stumpffia</emphasis>
|
||
</taxonomicName>
|
||
sp.
|
||
</paragraph>
|
||
<paragraph id="FD18217AFF946842FF03FEDEF544FE74" blockId="6.[151,1437,151,1177]" pageId="6" pageNumber="58">
|
||
The
|
||
<typeStatus id="221C9FD8FF946842FF3DFED1F51BFEFA" box="[249,297,296,320]" pageId="6" pageNumber="58">type</typeStatus>
|
||
locality is characterized by the presence of various rocks of different size along a small, slow-running stream in rainforest. Specimens were found in humid sites under big rocks. Calls were emitted by the
|
||
<typeStatus id="221C9FD8FF946842FAC1FEB3F155FED8" box="[1285,1383,330,354]" pageId="6" pageNumber="58" type="holotype">holotype</typeStatus>
|
||
during the day, on
|
||
<date id="891907BAFF946842FE84FE97F5CBFE3C" box="[320,505,366,391]" pageId="6" pageNumber="58" value="2008-01-24">24 January 2008</date>
|
||
between 13:30–15:30. The two
|
||
<typeStatus id="221C9FD8FF946842FCA5FE96F7FFFE3D" box="[865,973,367,391]" pageId="6" pageNumber="58" type="paratype">paratypes</typeStatus>
|
||
were collected close to the
|
||
<typeStatus id="221C9FD8FF946842FAC5FE97F155FE3C" box="[1281,1383,366,390]" pageId="6" pageNumber="58" type="holotype">holotype</typeStatus>
|
||
. The female was found next to a rock under which the second male was calling, at about
|
||
<quantity id="3A5F8C9FFF946842FBF2FE6BF052FE10" box="[1078,1120,402,426]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="58" unit="m" value="5.0">5 m</quantity>
|
||
distance from the collecting site of the
|
||
<typeStatus id="221C9FD8FF946842FEC8FE4FF540FE74" box="[268,370,438,462]" pageId="6" pageNumber="58" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="FD18217AFF946842FF03FE20F4D6FD83" blockId="6.[151,1437,151,1177]" pageId="6" pageNumber="58">
|
||
The tadpole of
|
||
<taxonomicName id="3AA75AF9FF946842FEB0FE23F5C1FE4B" box="[372,499,473,497]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="paidroa">
|
||
<emphasis id="CFD3FD68FF946842FEB0FE23F5C1FE4B" box="[372,499,473,497]" italics="true" pageId="6" pageNumber="58">M. paidroa</emphasis>
|
||
</taxonomicName>
|
||
is described in detail in Grosjean
|
||
<emphasis id="CFD3FD68FF946842FCBCFE23F780FE4B" box="[888,946,473,497]" italics="true" pageId="6" pageNumber="58">et al.</emphasis>
|
||
(in press). It is of the funnel-mouthed
|
||
<typeStatus id="221C9FD8FF946842FAAFFE23F1AEFE48" box="[1387,1436,474,498]" pageId="6" pageNumber="58">type</typeStatus>
|
||
typical for
|
||
<emphasis id="CFD3FD68FF946842FED2FE04F580FDAF" box="[278,434,509,533]" italics="true" pageId="6" pageNumber="58">
|
||
<taxonomicName id="3AA75AF9FF946842FED2FE04F59CFDAF" box="[278,430,509,533]" class="Amphibia" family="Ranidae" genus="Chonomantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="58" phylum="Chordata" rank="genus">Chonomantis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
but as a character state unique in the subgenus, it has rudimentary labial teeth (keratodonts).
|
||
</paragraph>
|
||
<paragraph id="FD18217AFF946842FF03FDBDF1AEFCED" blockId="6.[151,1437,151,1177]" pageId="6" pageNumber="58">
|
||
<emphasis id="CFD3FD68FF946842FF03FDBDF58EFDE7" bold="true" box="[199,444,580,605]" pageId="6" pageNumber="58">Advertisement calls.</emphasis>
|
||
The call of
|
||
<taxonomicName id="3AA75AF9FF946842FD89FDBCF77DFDE7" box="[589,847,581,605]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="paidroa">
|
||
<emphasis id="CFD3FD68FF946842FD89FDBCF77DFDE7" box="[589,847,581,605]" italics="true" pageId="6" pageNumber="58">Mantidactylus paidroa</emphasis>
|
||
</taxonomicName>
|
||
consists of a series of pulsatile note pairs (
|
||
<figureCitation id="659C3DFFFF946842FA83FDBDF1BCFDE6" box="[1351,1422,580,604]" captionStart="FIGURE 2" captionStartId="5.[151,250,1985,2008]" captionTargetBox="[164,1427,178,1970]" captionTargetId="figure@5.[164,1429,178,1970]" captionTargetPageId="5" captionText="FIGURE 2. Sonagrams and oscillograms of parts of a series of double notes, recorded from the holotype of Mantidactylus paidroa on 24 January 2008; (a) fragment of the call with 8 double notes; (b) enlarged view of two double notes." httpUri="https://zenodo.org/record/202730/files/figure.png" pageId="6" pageNumber="58">Fig. 2</figureCitation>
|
||
). One completely recorded series had a duration of 47 seconds. At the beginning of the series, five single notes were emitted, followed by 33 note pairs, with a single note emitted in one case in-between two note pairs. Call frequency ranged from 1500–5900 Hz. Temporal call parameters are as follows: note length of single notes 8–14 ms (10.6±10.6 ms; n= 5); note length of first note in note pairs 12–14 ms (12.6±0.8 ms; n=11); note length of second note in note pairs 12–16 ms (13.4±1.1 ms; n=11); interval between single notes 908–1915 ms (1218.5±468.2 ms; n=4); interval between notes in note pairs 100–135 ms (116±8.6 ms; n=11); interval between note pairs 479–1471 ms (980.7±368.7 ms; n=11). For comparison of some call characters with other species in the subgenus see
|
||
<tableCitation id="B02514C1FF946842FA87FCC6F1A4FCED" box="[1347,1430,831,855]" captionStart="TABLE 1" captionStartId="3.[151,235,440,461]" captionTargetBox="[161,1435,653,1720]" captionTargetPageId="3" captionText="TABLE 1. Morphometric ratios and distinctive morphological and bioacoustic characters in Mantidactylus paidroa sp. nov. compared with other species of Chonomantis, based on data published by Vences & Glaw (2004). Abbreviations used: SVL, snout-vent length; TD, tympanum diameter; ED, eye diameter; FOL, foot length; HAL, hand length; RHL, relative hindlimb length; TT, tibiotarsal articulation. Measurement values are mean ± standard deviation, followed by minimum – maximum values in parentheses. Data for M. delormei are based on the female holotype and photos of additional specimens in Glaw & Vences (2007) for colour in life." httpUri="http://table.plazi.org/id/A9D871F2FF916847FF53FE41F5EEFDD0" pageId="6" pageNumber="58" tableUuid="A9D871F2FF916847FF53FE41F5EEFDD0">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="FD18217AFF946842FF03FC9BF0D7FB22" blockId="6.[151,1437,151,1177]" pageId="6" pageNumber="58">
|
||
<emphasis id="CFD3FD68FF946842FF03FC9BF580FCC1" bold="true" box="[199,434,866,891]" pageId="6" pageNumber="58">Genetic divergence.</emphasis>
|
||
In the alignment of 458 nucleotides of the 16S rRNA gene (after deleting portions at the beginning and the end of the DNA fragment that were missing in some taxa), the male
|
||
<typeStatus id="221C9FD8FF946842FBB0FC7EF0E6FC25" box="[1140,1236,903,927]" pageId="6" pageNumber="58" type="paratype">paratype</typeStatus>
|
||
of
|
||
<taxonomicName id="3AA75AF9FF946842FB38FC7FF4DDFC78" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="paidroa">
|
||
<emphasis id="CFD3FD68FF946842FB38FC7FF4DDFC78" italics="true" pageId="6" pageNumber="58">Mantidactylus paidroa</emphasis>
|
||
</taxonomicName>
|
||
differed by a single substitution from the
|
||
<typeStatus id="221C9FD8FF946842FD13FC53F70BFC78" box="[727,825,938,962]" pageId="6" pageNumber="58" type="holotype">holotype</typeStatus>
|
||
and female
|
||
<typeStatus id="221C9FD8FF946842FC0EFC52F01FFC79" box="[970,1069,939,963]" pageId="6" pageNumber="58" type="paratype">paratype</typeStatus>
|
||
, which had identical sequences. The new species was strongly differentiated from all other species of
|
||
<taxonomicName id="3AA75AF9FF946842FC62FC37F009FC5C" box="[934,1083,974,998]" class="Amphibia" family="Ranidae" genus="Chonomantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="58" phylum="Chordata" rank="genus">
|
||
<emphasis id="CFD3FD68FF946842FC62FC37F009FC5C" box="[934,1083,974,998]" italics="true" pageId="6" pageNumber="58">Chonomantis</emphasis>
|
||
</taxonomicName>
|
||
(sequences as reported by
|
||
<bibRefCitation id="99365C8BFF946842FAA8FC37F570FBB3" author="Vieites" pageId="6" pageNumber="60" refString="Vieites, D. R., Wollenberg, K. C., Andreone, F., Kohler, J., Glaw, F. & Vences, M. (2009) Vast underestimation of Madagascar's biodiversity evidenced by an integrative amphibian inventory. Proceedings of the National Academy of Sciences of the U. S. A., 106, 8267 - 8272." type="journal article" year="2009">
|
||
Vieites
|
||
<emphasis id="CFD3FD68FF946842FF01FC0BF4CCFBB3" box="[197,254,1009,1033]" italics="true" pageId="6" pageNumber="58">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
). We found 24–46 substitutions in pairwise comparisons to all other species, corresponding to 5.6– 10.8% uncorrected pairwise sequence divergence (p-distances). The lowest distances (24–25 substitutions; 5.6– 5.8% divergence) were found to
|
||
<emphasis id="CFD3FD68FF946842FDCDFBC0F76AFBEB" box="[521,856,1081,1105]" italics="true" pageId="6" pageNumber="58">
|
||
<taxonomicName id="3AA75AF9FF946842FDCDFBC0F766FBEB" box="[521,852,1081,1105]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="brevipalmatus">Mantidactylus brevipalmatus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
For a preliminary phylogenetic analysis of the 16S sequences that however could not satisfyingly resolve relationships among species of
|
||
<taxonomicName id="3AA75AF9FF946842FBB2FBA4F139FBCF" box="[1142,1291,1117,1141]" class="Amphibia" family="Ranidae" genus="Chonomantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="58" phylum="Chordata" rank="genus">
|
||
<emphasis id="CFD3FD68FF946842FBB2FBA4F139FBCF" box="[1142,1291,1117,1141]" italics="true" pageId="6" pageNumber="58">Chonomantis</emphasis>
|
||
</taxonomicName>
|
||
, see supplementary information of
|
||
<bibRefCitation id="99365C8BFF946842FE67FB79F6B3FB22" box="[419,641,1152,1176]" pageId="6" pageNumber="60" refString="Vieites, D. R., Wollenberg, K. C., Andreone, F., Kohler, J., Glaw, F. & Vences, M. (2009) Vast underestimation of Madagascar's biodiversity evidenced by an integrative amphibian inventory. Proceedings of the National Academy of Sciences of the U. S. A., 106, 8267 - 8272." type="journal article">
|
||
Vieites
|
||
<emphasis id="CFD3FD68FF946842FE3CFB78F602FB22" box="[504,560,1152,1176]" italics="true" pageId="6" pageNumber="58">et al.</emphasis>
|
||
(2009)
|
||
</bibRefCitation>
|
||
where the species is included as
|
||
<taxonomicName id="3AA75AF9FF946842FC30FB79F0A6FB22" box="[1012,1172,1152,1176]" class="Amphibia" family="Mantellidae" genus="Mantidactylus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="58" phylum="Chordata" rank="genus">
|
||
<emphasis id="CFD3FD68FF946842FC30FB79F0A6FB22" box="[1012,1172,1152,1176]" italics="true" pageId="6" pageNumber="58">Mantidactylus</emphasis>
|
||
</taxonomicName>
|
||
sp. 59.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |