160 lines
20 KiB
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160 lines
20 KiB
XML
<document ID-CLB-Dataset="7157" ID-DOI="http://dx.doi.org/10.3897/zookeys.1035.65767" ID-GBIF-Dataset="cbd813c3-228e-4bce-b30a-1a150d778c5b" ID-PMC="PMC8096803" ID-Pensoft-Pub="1313-2970-1035-145" ID-Pensoft-UUID="55CAE096B4A451D88EA433175454DA3F" ID-PubMed="33958933" ID-ZooBank="ADD73A162DB34873890B1E490C2C8841" ModsDocID="1313-2970-1035-145" checkinTime="1619617199447" checkinUser="pensoft" docAuthor="Zamani, Alireza & Marusik, Yuri M." docDate="2021" docId="75080B2F5CE955999C9274B037E542BE" docLanguage="en" docName="ZooKeys 1035: 145-193" docOrigin="ZooKeys 1035" docPubDate="2021-04-27" docSource="http://dx.doi.org/10.3897/zookeys.1035.65767" docTitle="Lachesana kavirensis Zamani & Marusik 2021, sp. nov." docType="treatment" docUuid="3BC489D8-1BD7-46C1-A4D2-BCB9A3BCDE58" docUuidSource="ZooBank" docVersion="6" id="55CAE096B4A451D88EA433175454DA3F" lastPageNumber="145" masterDocId="55CAE096B4A451D88EA433175454DA3F" masterDocTitle="Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species" masterLastPageNumber="193" masterPageNumber="145" pageNumber="145" updateTime="1732670661048" updateUser="ExternalLinkService">
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<mods:mods id="FA8F0C31AE2C7F39640E8443A6320517" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="F2D3F22853DAD3E31815317AC8E9AFB7">
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<mods:title id="ED62FFFDD789E12CDFCE54EE14C8CDEB">Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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</mods:titleInfo>
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<mods:name id="5F0FA5A7462537CCD6E16A7FEA15FDA9" type="personal">
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<mods:role id="E979AF12DFB584A841A2ABA55187AFFB">
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<mods:roleTerm id="47D26DE87A32964B885616985B133139">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F6CD8B773B0BB31C55D91A0AA31C437A">Zamani, Alireza</mods:namePart>
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<mods:nameIdentifier id="967513BD6597C1D6C9FE17D816EC22E8" type="ORCID">https://orcid.org/0000-0002-8084-9666</mods:nameIdentifier>
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<mods:affiliation id="61DA7F38BC09B404A13F293584B25DC6">Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland</mods:affiliation>
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<mods:nameIdentifier id="DDD9FD0457AC18880C958F211E5B8462" type="email">zamani.alireza5@gmail.com</mods:nameIdentifier>
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<mods:name id="0D6F0359EC66BA68B59A0250D9FC573E" type="personal">
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<mods:roleTerm id="06041ACD13D88011CCEEB77D7830B500">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8FD96A31F642E5A046443BC302396D92">Marusik, Yuri M.</mods:namePart>
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<mods:nameIdentifier id="E3AD3A73C1923F4D089BC5F328FBF785" type="ORCID">https://orcid.org/0000-0002-4499-5148</mods:nameIdentifier>
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<mods:affiliation id="D9C7C12EDC219E3DC0A33444BBE7E658">Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia & Department of Zoology & Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="C79B6713ED74FC67F14DAA6C8C658F25">text</mods:typeOfResource>
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<mods:title id="7000E0CAB5EE1F2BE33408F2F63E66A9">ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="AECD80157F9C216CDE63E820E7053AC4">
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<mods:date id="CB3DDD2068509F9DCFDF3C1446FF959C">2021</mods:date>
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<mods:number id="7D8EA48EFBF112AD95F77841C7D97B58">2021-04-27</mods:number>
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<mods:detail id="C68E695E5947395D3D81B2C00D7F76A6" type="volume">
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<mods:number id="04AE88B7447A0575AE29CD5BE331A56E">1035</mods:number>
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<mods:start id="AB3069608E6925C34A580035966A4D9E">145</mods:start>
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<mods:end id="ADD3FFA4366EF64EBC17C348FC67A5E9">193</mods:end>
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<mods:location id="F22668E86E3A0792A1D16311AEE440FD">
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<mods:url id="0470D8F81711388AD623ECFF4F4C2139">http://dx.doi.org/10.3897/zookeys.1035.65767</mods:url>
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</mods:location>
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<mods:classification id="EFDBF242DCB781ACEFD213EE73612C8D">journal article</mods:classification>
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<mods:identifier id="4DC7AB8201D5B129AB90355B3BDFBD27" type="DOI">http://dx.doi.org/10.3897/zookeys.1035.65767</mods:identifier>
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<mods:identifier id="109F797E5586B26B6162998DBEE6702D" type="Pensoft-Pub">1313-2970-1035-145</mods:identifier>
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<mods:identifier id="76AE88CC94317B08FFB447D65EED6EE4" type="ZooBank">ADD73A162DB34873890B1E490C2C8841</mods:identifier>
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<treatment id="75080B2F5CE955999C9274B037E542BE" ID-GBIF-Taxon="181753555" LSID="urn:lsid:zoobank.org:act:3BC489D8-1BD7-46C1-A4D2-BCB9A3BCDE58" httpUri="http://treatment.plazi.org/id/75080B2F5CE955999C9274B037E542BE" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Lachesaninae">
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<subSubSection id="604C2C99F30B1F1234E0A822F116B93D" pageId="0" pageNumber="145" type="nomenclature">
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<paragraph id="0652F93F8648253B38C99090BAA5A5F4" pageId="0" pageNumber="145">
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<taxonomicName id="FD2D5B0ACF69683FA3B9F0CD925AA485" ID-CoL="98YYV" LSID="http://zoobank.org/3BC489D8-1BD7-46C1-A4D2-BCB9A3BCDE58" authority="Zamani & Marusik, 2021" authorityName="Zamani & Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana kavirensis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="kavirensis" status="sp. nov.">Lachesana kavirensis</taxonomicName>
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<taxonomicNameLabel id="6F9266FD992B194DB159817F468C6961" pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
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<figureCitation id="1EBCAC53707311EA5F299A866C33C333" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">Figs 1A-C</figureCitation>
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<figureCitation id="7D8D8367E734BC9A24BF4370318190D5" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">, 3A-E</figureCitation>
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<figureCitation id="3620ED69DCD6F826F7E6B13D8DD38725" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">, 4I</figureCitation>
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<figureCitation id="4E0449A87159DED2C316FDAF4582E224" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="5C8A2B03B4B9250026BF42C87C0DBA01" pageId="0" pageNumber="145" type="type material">
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<paragraph id="40EEE7B48C74BE9FB6263AD1CBAC29F8" pageId="0" pageNumber="145">Type material.</paragraph>
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<paragraph id="362367AE0476F7FE6F33BA14E2207BA5" pageId="0" pageNumber="145">
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<emphasis id="5F294A06E0007CB5FDC8EFF7F5F77BAA" bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
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♂ (MHNG), Iran:
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<emphasis id="063C512755E185B0C5D495855D7160F1" italics="true" pageId="0" pageNumber="145">Qom Province</emphasis>
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: Dasht-e Masileh,
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<geoCoordinate id="729F8F592B18821BCE57EF5B16A2E7A7" degrees="34" direction="north" minutes="47" orientation="latitude" precision="925" value="34.783333">34°47'N</geoCoordinate>
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,
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<geoCoordinate id="3284D87A06E53983F4B8EA878A52C191" degrees="51" direction="east" minutes="11" orientation="longitude" precision="925" value="51.183334">51°11'E</geoCoordinate>
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, 11.2017 (M. Mirghazanfari).
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</paragraph>
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</subSubSection>
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<subSubSection id="43A9C13ED5E96F9550DE1616D3789377" pageId="0" pageNumber="145" type="etymology">
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<paragraph id="1E6E5A1FB6B6229D70CCDE963B31603C" pageId="0" pageNumber="145">Etymology.</paragraph>
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<paragraph id="ECD674A6220A7BF754CEACBB2AE0814D" pageId="0" pageNumber="145">
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The specific epithet is derived from
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<normalizedToken id="3320C6189828C831E44BB2136D4B772C" originalValue="‘Kavir’">'Kavir'</normalizedToken>
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, a Persian word for desert, referring to the occurrence of the species in the central deserts of Iran.
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</paragraph>
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</subSubSection>
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<subSubSection id="1F81D003D25F702331063A2445B67BBA" pageId="0" pageNumber="145" type="diagnosis">
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<paragraph id="C0899A83683D2FC5E6470B7DDE141D49" pageId="0" pageNumber="145">Diagnosis.</paragraph>
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<paragraph id="8D39B779133E88E64E924F91A5735C1E" pageId="0" pageNumber="145">
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The male of
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<taxonomicName id="AFE5CE602157642213A101E334CCD8C6" lsidName="L. kavirensis" pageId="0" pageNumber="145" rank="species" species="kavirensis">
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<emphasis id="1EF70A5F5F24B1B44FE46320B724BC43" italics="true" pageId="0" pageNumber="145">L. kavirensis</emphasis>
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</taxonomicName>
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sp. nov. can be distinguished from the similar
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<taxonomicName id="C1B7658179D4C32CA663F6574DF32EE3" lsidName="L. dyachkovi" pageId="0" pageNumber="145" rank="species" species="dyachkovi">
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<emphasis id="3A7180F9435650C08A0F9936F76B1254" italics="true" pageId="0" pageNumber="145">L. dyachkovi</emphasis>
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</taxonomicName>
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Fomichev & Marusik, 2019 by having a gently bent cheliceral fang (
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<emphasis id="9E6831C9F200F912EBC9329F42D2B5F0" italics="true" pageId="0" pageNumber="145">vs.</emphasis>
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bent over 90°, cf. Fig.
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<figureCitation id="B5D684590E0E33B03A34BE6FD95620C5" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1C</figureCitation>
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and fig. 15 in
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<bibRefCitation id="0301D11C0FA22D61052F96E758AE6C5E" DOI="https://doi.org/10.15298/arthsel.28.4.08" author="Fomichev, AA" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="145" pagination="556 - 561" refId="B5" refString="Fomichev, AA, Marusik, YM, 2019. A new species of Lachesana Strand, 1932 (Aranei: Zodariidae) from southern Kazakhstan. Arthropoda Selecta 28 (4): 556 - 561, DOI: https://doi.org/10.15298/arthsel.28.4.08" title="A new species of Lachesana Strand, 1932 (Aranei: Zodariidae) from southern Kazakhstan." url="https://doi.org/10.15298/arthsel.28.4.08" volume="28" year="2019">Fomichev and Marusik 2019</bibRefCitation>
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). The shape of the RTA is also different from other species of the region (stalk with a gentle curve ventrally and tip straightly pointed; cf. Fig.
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<figureCitation id="F3E50994048DE8ACA551316E63BF6DF9" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4I</figureCitation>
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and Fig.
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<figureCitation id="1E0695FF0E08863C57682C767F4B6717" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4F-H, J</figureCitation>
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).
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</paragraph>
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<caption id="6E02DF173E7CCAA75D1160FD8E3AE8BC" doi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145" start="Figure 1" startId="F1">
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<paragraph id="70A7462D885D40811A3429CF85B87B7E" pageId="0" pageNumber="145">
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<emphasis id="89C53FD05330D9D2077A9D7083CAF455" bold="true" pageId="0" pageNumber="145">Figure 1.</emphasis>
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Habitus and chelicerae of male
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<taxonomicName id="6F067D0A69C2959CB658B05CE5A33D62" authorityName="Zamani & Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana kavirensis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="kavirensis">
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<emphasis id="983FFCDC3921BDEFC278449D0B8D47F9" italics="true" pageId="0" pageNumber="145">Lachesana kavirensis</emphasis>
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</taxonomicName>
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sp. nov. (
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<emphasis id="F48042E2001767800412B211E5F7AA9A" bold="true" pageId="0" pageNumber="145">A-C</emphasis>
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) and
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<taxonomicName id="29138CFD20DAA6C163A6C315F04A635F" lsidName="L. perseus" pageId="0" pageNumber="145" rank="species" species="perseus">
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<emphasis id="E0445D307571C5E8230322D4C98CB5EB" italics="true" pageId="0" pageNumber="145">L. perseus</emphasis>
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</taxonomicName>
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sp. nov. (
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<emphasis id="4524C988A95A19F56DA00C5EF7023E96" bold="true" pageId="0" pageNumber="145">D-F</emphasis>
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)
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<emphasis id="94212D215E4FB0E23E79327BDEF48081" bold="true" pageId="0" pageNumber="145">A, B, D, E</emphasis>
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habitus, dorsal
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<emphasis id="F32153E33FD758F95B2139BEEA32BC42" bold="true" pageId="0" pageNumber="145">C, F</emphasis>
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chelicerae, ventral.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="CAADF2ECB101A1BC983F8F3B31A3CD1C" pageId="0" pageNumber="145" type="description">
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<paragraph id="13CDC11543B4E7C5463C451ABCD70CCC" pageId="0" pageNumber="145">Description.</paragraph>
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<paragraph id="2259A8733FDE5B904D5BF6F639983779" pageId="0" pageNumber="145">
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<emphasis id="C221BC61D6EDF4B20A3FF93023EFCA67" bold="true" pageId="0" pageNumber="145">Male.</emphasis>
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Habitus as in Fig.
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<figureCitation id="A679570090A25AC627FE334E723AF8DB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1A, B</figureCitation>
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. Total length 11.55. Carapace 5.80 long, 2.60 wide at pars cephalica, 3.75 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.23, ALE: 0.20, PME: 0.17, PLE: 0.23, PME-PME: 0.14. Carapace, sternum, labium, chelicerae and maxillae light brown. Chelicera (Fig.
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<figureCitation id="E0CE6D685589F3867E1F35B4E4479CBE" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1C</figureCitation>
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) with dense, black setae and one promarginal tooth; fangs gently curved, almost straight. Legs the same color as carapace, without annulations and with numerous spines. Abdomen pale, dorsally with a longitudinal dark stripe and several small spots. Spinnerets slightly lighter than abdomen, uniform in color. Measurements of legs: I: 15.49 (4.47, 1.88, 3.33, 3.29, 2.52), II: 15.03 (4.20, 1.79, 3.02, 3.50, 2.52), III: 15.78 (4.08, 1.96, 2.48, 4.71, 2.55), IV: 18.30 (4.58, 2.07, 3.20, 5.24, 3.21).
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</paragraph>
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<paragraph id="304E1DF7FAA0F3E0AA1213524791DAFC" pageId="0" pageNumber="145">
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Palp as in Figs
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<figureCitation id="81D0B86A58E887F9B50E48105E1A8DA9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">3A-E</figureCitation>
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,
|
||
<figureCitation id="E48303C1CB0CE47198597D0A987CE857" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4I</figureCitation>
|
||
. RTA long, over 1.2 times longer than tibia, its widest part less than 2 times wider than stalk; cymbium with 2 spines in mesal proximal part; bulb elongate, 1.3 times longer than wide; base of embolus large, 1/3 of the tegulum's length; embolus filamentous, lying in chute formed by conductor.
|
||
</paragraph>
|
||
<paragraph id="8D0F520E623C595BDB6A4BF97F263548" pageId="0" pageNumber="145">
|
||
<emphasis id="3E3506B042F14DB6D86A3B4879E5FC4D" bold="true" pageId="0" pageNumber="145">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="93138FA7613F60626D020CBFA0315E7F" pageId="0" pageNumber="145" type="distribution">
|
||
<paragraph id="349676EA43DCC9D6FE7D20BC5FDAF437" pageId="0" pageNumber="145">Distribution.</paragraph>
|
||
<paragraph id="F8D01D058C16801AC1AB2064130704D7" pageId="0" pageNumber="145">
|
||
Known only from the type locality in Qom Province, northern Iran (Fig.
|
||
<figureCitation id="4821E25AC0408CC2280B594D11AC825A" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |