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<document id="B99AC9E167C07E08D8E9B00567C05B78" ID-DOI="10.5281/zenodo.12612328" ID-ISSN="2064-2474" ID-Zenodo-Dep="12612328" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719865400710" checkinUser="felipe" docAuthor="Oláh, J. &amp; Johanson, K." docDate="2011" docId="747987ACFF9664165D404DF9FD01FCBE" docLanguage="en" docName="ActaZoolAcadSciH.57.2.117-129.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 57 (2)" docStyle="DocumentStyle:4FD749C6DF7B87D8940F4D8C7B0A0514.3:ActaZoolAcadSciH.2002-2014.journal_article" docStyleId="4FD749C6DF7B87D8940F4D8C7B0A0514" docStyleName="ActaZoolAcadSciH.2002-2014.journal_article" docStyleVersion="3" docTitle="Madagocerum bhemi Oláh &amp; Johanson, 2011, sp. n." docType="treatment" docVersion="2" lastPageNumber="123" masterDocId="8840FFD4FF9264105F304934FFDAFFB8" masterDocTitle="Description Of A New Endemic Genus Of Trichoptera From Madagascar (Odontoceridae)" masterLastPageNumber="129" masterPageNumber="117" pageNumber="121" updateTime="1719866007046" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
<mods:mods id="4845FF97B1DEA8E8AA005C984A2609BF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CBEA3068B0E122ADF1CA5B4B550DA7FE">
<mods:title id="880C1ABD0CC7765D0C5F03FD19AFD994">Description Of A New Endemic Genus Of Trichoptera From Madagascar (Odontoceridae)</mods:title>
</mods:titleInfo>
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<mods:role id="BD7D61A6AEA0C230DEDBDB31AC8938DA">
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<mods:namePart id="42C0DCA3D9714E925BF2588EBAEF203D">Oláh, J.</mods:namePart>
<mods:affiliation id="89AFED5B970676B585226F9E6F45CD42">and J, K. A. &amp; Szent István University, Gödöllő, Centre of Environmental Health, Gyula, Hungary Residence postal address: Tarján u. 28, H- 4032 Debrecen, Hungary; E-mail: profolah @ gmail. com</mods:affiliation>
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<mods:namePart id="F7D1AC1A3F96C8444CC2752FE0DFA7A7">Johanson, K.</mods:namePart>
<mods:affiliation id="EB3FF7BE9CBC41B082C9C455C1AE464B">Swedish Museum of Natural History, Entomology Department Box 50007, SE- 10405 Stockholm, Sweden; E-mail: kjell. arne. johanson @ nrm. se</mods:affiliation>
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<mods:typeOfResource id="FF3B76EBF8C22FA2E4934E0CCDE93F0D">text</mods:typeOfResource>
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<mods:title id="E480E838FAEC4867DC7440393B5F0C37">Acta Zoologica Academiae Scientiarum Hungaricae</mods:title>
</mods:titleInfo>
<mods:part id="CD3DAE50EE0999996096BDC718AD50C8">
<mods:date id="393E935A20CEB3FFC4B8E1B1E915EA75">2011</mods:date>
<mods:detail id="9FCF48C85A5C445A134E2FD2297BDF89" type="pubDate">
<mods:number id="F4046189F1BBB320697B79D2168E583B">2011-06-10</mods:number>
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<mods:detail id="C1084F5D36C6AF3CAA70DF038DAC9858" type="volume">
<mods:number id="A72826DB3BF0F33068FEC72DC0DCD8B4">57</mods:number>
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<mods:identifier id="A7B49D3077C6F24B577819E13DBAD89B" type="DOI">10.5281/zenodo.12612328</mods:identifier>
<mods:identifier id="FE2136C991117FBC7E05095A2038893C" type="ISSN">2064-2474</mods:identifier>
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<treatment id="747987ACFF9664165D404DF9FD01FCBE" ID-DOI="http://doi.org/10.5281/zenodo.12612305" ID-Zenodo-Dep="12612305" LSID="urn:lsid:plazi:treatment:747987ACFF9664165D404DF9FD01FCBE" httpUri="http://treatment.plazi.org/id/747987ACFF9664165D404DF9FD01FCBE" lastPageId="6" lastPageNumber="123" pageId="4" pageNumber="121">
<subSubSection id="B4CA6531FF9664145D404DF9FC18FB50" box="[624,962,1229,1256]" pageId="4" pageNumber="121" type="nomenclature">
<paragraph id="FC6F36BAFF9664145D404DF9FC18FB50" blockId="4.[624,962,1229,1256]" box="[624,962,1229,1256]" pageId="4" pageNumber="121">
<heading id="A72781D6FF9664145D404DF9FC18FB50" box="[624,962,1229,1256]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="121" reason="2">
<taxonomicName id="3BD04D39FF9664145D404DF9FCA6FB5F" authorityName="Oláh &amp; Johanson" authorityYear="2011" box="[624,892,1229,1255]" class="Insecta" family="Odontoceridae" genus="Madagocerum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="121" phylum="Arthropoda" rank="species" species="bhemi" status="sp. nov.">
<emphasis id="CEA4EAA8FF9664145D404DF9FCA6FB5F" bold="true" box="[624,892,1229,1255]" pageId="4" pageNumber="121">Madagocerum bhemi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D59757D3FF9664145CB44DFAFC18FB50" box="[900,962,1230,1256]" pageId="4" pageNumber="121" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B4CA6531FF9664165DED4DC5FD01FCBE" lastPageId="6" lastPageNumber="123" pageId="4" pageNumber="121" type="description">
<paragraph id="FC6F36BAFF9664145DED4DC5FC8CFAB3" blockId="4.[733,854,1265,1291]" box="[733,854,1265,1291]" pageId="4" pageNumber="121">
<heading id="A72781D6FF9664145DED4DC5FC8CFAB3" box="[733,854,1265,1291]" centered="true" fontSize="11" level="3" pageId="4" pageNumber="121" reason="3">
(
<figureCitation id="64EB2A3FFF9664145DD54DC5FC97FAB3" box="[741,845,1265,1291]" captionStart="Figs 1–4" captionStartId="5.[317,360,1774,1795]" captionTargetBox="[324,1266,838,1742]" captionTargetId="figure-188@5.[317,1269,832,1744]" captionTargetPageId="5" captionText="Figs 1–4. Madagocerum bhemi gen. et sp. n., holotype:1 = right wings, 2 = genitalia, lateral view, 3 = genitalia, dorsal view, 4 = phallus, lateral view" figureDoi="http://doi.org/10.5281/zenodo.12612330" httpUri="https://zenodo.org/record/12612330/files/figure.png" pageId="4" pageNumber="121">Figs 1–4</figureCitation>
)
</heading>
</paragraph>
<paragraph id="FC6F36BAFF9664145E4E4C03FC50F9B9" blockId="4.[317,1270,1335,1537]" pageId="4" pageNumber="121">
Diagnosis –This species is most similar to
<taxonomicName id="3BD04D39FF9664145C5C4C0CFC30FAEA" authorityName="Oláh &amp; Johanson" authorityYear="2011" box="[876,1002,1336,1362]" class="Insecta" family="Odontoceridae" genus="Madagocerum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="121" phylum="Arthropoda" rank="species" species="idvigun" status="sp. n.">
<emphasis id="CEA4EAA8FF9664145C5C4C0CFC30FAEA" box="[876,1002,1336,1362]" italics="true" pageId="4" pageNumber="121">M. idvigun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D59757D3FF9664145CC04C03FBF4FAE9" box="[1008,1070,1335,1361]" pageId="4" pageNumber="121" rank="species">sp. n.</taxonomicNameLabel>
in having a short segment IX. It is distinguished from
<taxonomicName id="3BD04D39FF9664145DC04C6FFCADFACD" authorityName="Oláh &amp; Johanson" authorityYear="2011" box="[752,887,1371,1397]" class="Insecta" family="Odontoceridae" genus="Madagocerum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="121" phylum="Arthropoda" rank="species" species="idvigum">
<emphasis id="CEA4EAA8FF9664145DC04C6FFCADFACD" box="[752,887,1371,1397]" italics="true" pageId="4" pageNumber="121">M. idvigum</emphasis>
</taxonomicName>
by the broader, blunt and divergent preanal appendages in dorsal view, not narrow, tapering and tangential; segment X is shorter than the preanal appendages, not longer; the coxopodites have no apicodorsal spiny protuberance; the harpagones are narrow, not broad; and the phallic apparatus is broad, not narrowing apically.
</paragraph>
<paragraph id="FC6F36BAFF9664145E4E4F10FB1CF9EE" blockId="4.[317,1270,1572,1824]" pageId="4" pageNumber="121">Description – Male (in alcohol). Body dark brown, except paler ventrally; cephalic and thoracic appendages light brown; forewing membrane light brown, without pattern (in alcohol).</paragraph>
<paragraph id="FC6F36BAFF9664145E4E4F6AFDEDF95E" blockId="4.[317,1270,1572,1824]" pageId="4" pageNumber="121">
Wings (
<figureCitation id="64EB2A3FFF9664145EFB4F6AFDDCF9CB" box="[459,518,1630,1651]" captionStart="Figs 1–4" captionStartId="5.[317,360,1774,1795]" captionTargetBox="[324,1266,838,1742]" captionTargetId="figure-188@5.[317,1269,832,1744]" captionTargetPageId="5" captionText="Figs 1–4. Madagocerum bhemi gen. et sp. n., holotype:1 = right wings, 2 = genitalia, lateral view, 3 = genitalia, dorsal view, 4 = phallus, lateral view" figureDoi="http://doi.org/10.5281/zenodo.12612330" httpUri="https://zenodo.org/record/12612330/files/figure.png" pageId="4" pageNumber="121">Fig. 1</figureCitation>
): Forewing apex narrowly rounded; 5.0 mm; membrane light brown; termen slightly convex; basal lobe present covered by microtrichia; jugum absent. Forewing forks I, II, III, and V present; Sc running to R1, and R1 running to R2 before C, meeting at hypertrophied pterostigma; distally almost indiscernible; Cu2 apparently absent in forewings; postanal vein running closely to posterior wing margin.
</paragraph>
<paragraph id="FC6F36BAFF9664165E4E4FDAFDD2FDE1" blockId="4.[317,1270,1572,1824]" lastBlockId="6.[317,1270,436,601]" lastPageId="6" lastPageNumber="123" pageId="4" pageNumber="121">
Male genitalia (
<figureCitation id="64EB2A3FFF9664145D244FDAFDB7F8BB" box="[532,621,1774,1795]" captionStart="Figs 1–4" captionStartId="5.[317,360,1774,1795]" captionTargetBox="[324,1266,838,1742]" captionTargetId="figure-188@5.[317,1269,832,1744]" captionTargetPageId="5" captionText="Figs 1–4. Madagocerum bhemi gen. et sp. n., holotype:1 = right wings, 2 = genitalia, lateral view, 3 = genitalia, dorsal view, 4 = phallus, lateral view" figureDoi="http://doi.org/10.5281/zenodo.12612330" httpUri="https://zenodo.org/record/12612330/files/figure.png" pageId="4" pageNumber="121">Figs 2–4</figureCitation>
): Abdominal segment IX fused annularly, short in lateral view, particularly at dorsal 1/3rd; venter 3 times longer than dorsum; anterior margin convex in lateral view, with lateral concavity conspicuous in dorsal view; posterior margins concave, with small, subtriangular apical lobe at dorsal margin of coxopodites; each antecosta and antecostal suture narrow on anterior margin of segment IX, forming pigmented marginal rim running evenly along anterior margin; acrotergite absent; surface of segment IX glabrous, spine row absent on posterior margins. Intersegmental depression between segments IX and X very deep, stepwise; segment X sunk to upper 1/3rd of segment IX, visible in lateral view; intersegmental depression covered by fused base of preanal appendages. Segment X broadly rod-like, directed straight posterad. Apicoventral setose lobes represented by setose apex of segment. Apicodorsal setose lobes reduced to central areas with scattered setae. Segment X body deeply cleft, with tangential lobes. Dorsal interlobular gap filled, as seen in dorsal view. Large, deeply bilobed preanal appendages dominating above phallic apparatus; fused basally; horizontal, posterad directed lobes fused to dorsum of segment IX; fused seam, representing borderline between segment IX and preanal appendages indiscernible; in lateral view, dorsal and ventral margins straight, producing regular, parallel-sided rods. Preanal appendages divergent in dorsal view, parallel-sided; rod-like. Coxopodites robust, dorsal margins rounded, convex; ventral margins concave, ending in small apicoventral corners. Harpagones slender, constricted at mid-length; capitate, with mesad turning apex with few, dark, short, conical spines. Phallic apparatus with broad, ventrally directed, short basal part; horizontal apical part long, straight. Small phallotremal sclerites almost indiscernible.
</paragraph>
<caption id="A8AF6632FF9764155E0D4FDAFC20F898" ID-DOI="http://doi.org/10.5281/zenodo.12612330" ID-Zenodo-Dep="12612330" httpUri="https://zenodo.org/record/12612330/files/figure.png" pageId="5" pageNumber="122" startId="5.[317,360,1774,1795]" targetBox="[324,1266,838,1742]" targetPageId="5" targetType="figure">
<paragraph id="FC6F36BAFF9764155E0D4FDAFC20F898" blockId="5.[317,1270,1774,1824]" pageId="5" pageNumber="122">
<emphasis id="CEA4EAA8FF9764155E0D4FDAFE4CF8BB" bold="true" box="[317,406,1774,1795]" pageId="5" pageNumber="122">Figs 1–4.</emphasis>
<taxonomicName id="3BD04D39FF9764155EAA4FDBFDBCF8BC" authorityName="Oláh &amp; Johanson" authorityYear="2011" box="[410,614,1775,1796]" class="Insecta" family="Odontoceridae" genus="Madagocerum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="species" species="bhemi" status="gen. et sp. n.">
<emphasis id="CEA4EAA8FF9764155EAA4FDBFDBCF8BC" box="[410,614,1775,1796]" italics="true" pageId="5" pageNumber="122">Madagocerum bhemi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D59757D3FF9764155D5A4FDAFD3AF8BB" box="[618,736,1774,1795]" pageId="5" pageNumber="122" rank="species">gen. et sp. n.</taxonomicNameLabel>
, holotype: 1 = right wings, 2 = genitalia, lateral view, 3 = genitalia, dorsal view, 4 = phallus, lateral view
</paragraph>
</caption>
<paragraph id="FC6F36BAFF9464165E4E4B4AFB2CFD2B" blockId="6.[317,1270,638,688]" box="[382,1270,638,659]" pageId="6" pageNumber="123">
<materialsCitation id="4CB83CE7FF9464165E4E4B4AFB2CFD2B" box="[382,1270,638,659]" collectingDate="1954-01" country="Madagascar" elevation="1800" location="Material" municipality="Male" pageId="6" pageNumber="123" specimenCount="1" typeStatus="holotype">
<location id="F90F6061FF9464165E4E4B4AFE15FD2B" LSID="urn:lsid:plazi:treatment:747987ACFF9664165D404DF9FD01FCBE:F90F6061FF9464165E4E4B4AFE15FD2B" box="[382,463,638,659]" country="Madagascar" municipality="Male" name="Material" pageId="6" pageNumber="123">Material</location>
examined –
<typeStatus id="236B8818FF9464165D794B4AFD7CFD2B" box="[585,678,638,659]" pageId="6" pageNumber="123" type="holotype">Holotype</typeStatus>
:
<collectingMunicipality id="1C0BACC0FF9464165D9F4B4AFD3EFD2B" box="[687,740,638,659]" pageId="6" pageNumber="123">Male</collectingMunicipality>
: “
<collectingCountry id="84C7762AFF9464165DC54B4AFCB5FD2B" box="[757,879,638,659]" name="Madagascar" pageId="6" pageNumber="123">Madagascar</collectingCountry>
: Andohahela:
<quantity id="3B289B5FFF9464165CC74B4AFB98FD2B" box="[1015,1090,638,659]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="123" unit="m" value="1800.0">
<elevation id="77FDD189FF9464165CC74B4AFB98FD2B" box="[1015,1090,638,659]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="123" unit="m" value="1800.0">1800 m</elevation>
</quantity>
,
<date id="886E107AFF9464165B7B4B4AFB5DFD2B" box="[1099,1159,638,659]" pageId="6" pageNumber="123" value="1954-01">
<collectingDate id="982AE992FF9464165B7B4B4AFB5DFD2B" box="[1099,1159,638,659]" pageId="6" pageNumber="123" value="1954-01">i.1954</collectingDate>
</date>
[Paulian]”.
</materialsCitation>
</paragraph>
<paragraph id="FC6F36BAFF9464165E0D4BAFFD14FD08" blockId="6.[317,1270,638,688]" box="[317,718,667,688]" pageId="6" pageNumber="123">
<materialsCitation id="4CB83CE7FF9464165E0D4BAFFD10FD08" box="[317,714,667,688]" collectingDate="1954-01" country="Madagascar" elevation="1800" location="Material" municipality="Male" pageId="6" pageNumber="123" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="236B8818FF9464165E0D4BAFFE4FFD08" box="[317,405,667,688]" pageId="6" pageNumber="123" type="paratype">Paratype</typeStatus>
:
<specimenCount id="EAD6FD33FF9464165EAF4BAFFE3FFD08" box="[415,485,667,688]" count="1" pageId="6" pageNumber="123" type="male">1 male</specimenCount>
: same data as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="FC6F36BAFF9464165E4E4BE0FD01FCBE" blockId="6.[317,1269,724,774]" pageId="6" pageNumber="123">
Etymology:
<emphasis id="CEA4EAA8FF9464165EC74BE1FDEEFD52" box="[503,564,725,746]" italics="true" pageId="6" pageNumber="123">Bhemi</emphasis>
, slender in Sanskrit, named after the slender harpagones in this species compared to in other species in the genus.
</paragraph>
</subSubSection>
</treatment>
</document>
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