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<document id="C5F6CB77EF10E3E0F4705F248D07E22E" ID-DOI="10.5281/zenodo.12612328" ID-ISSN="2064-2474" ID-Zenodo-Dep="12612328" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719865400710" checkinUser="felipe" docAuthor="Oláh, J. &amp; Johanson, K." docDate="2011" docId="747987ACFF9464185D584A63FC08FD75" docLanguage="en" docName="ActaZoolAcadSciH.57.2.117-129.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 57 (2)" docStyle="DocumentStyle:4FD749C6DF7B87D8940F4D8C7B0A0514.3:ActaZoolAcadSciH.2002-2014.journal_article" docStyleId="4FD749C6DF7B87D8940F4D8C7B0A0514" docStyleName="ActaZoolAcadSciH.2002-2014.journal_article" docStyleVersion="3" docTitle="Madagocerum idvigun Oláh &amp; Johanson, 2011, sp. n." docType="treatment" docVersion="2" lastPageNumber="125" masterDocId="8840FFD4FF9264105F304934FFDAFFB8" masterDocTitle="Description Of A New Endemic Genus Of Trichoptera From Madagascar (Odontoceridae)" masterLastPageNumber="129" masterPageNumber="117" pageNumber="123" updateTime="1719866007046" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
<mods:mods id="462143BBD86DCA6AE75B4E48306C12C1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="29E965D77EFA4FACC4CF6363534C37B6">
<mods:title id="D940C446EAA1DB5371EBEC6C19C3E4B0">Description Of A New Endemic Genus Of Trichoptera From Madagascar (Odontoceridae)</mods:title>
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<mods:roleTerm id="A54578ED09E2F87BD9BD05A175F0B36A">Author</mods:roleTerm>
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<mods:namePart id="6B1D64B5F7E42532405E8FD7E4038F84">Oláh, J.</mods:namePart>
<mods:affiliation id="01D7925CE0FCE94064AB800C97A6946D">and J, K. A. &amp; Szent István University, Gödöllő, Centre of Environmental Health, Gyula, Hungary Residence postal address: Tarján u. 28, H- 4032 Debrecen, Hungary; E-mail: profolah @ gmail. com</mods:affiliation>
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<mods:namePart id="9D8D88D0F3DF3E4CC1D59DCA8D566A1C">Johanson, K.</mods:namePart>
<mods:affiliation id="3D9E6C159D0221848957A495F3B37265">Swedish Museum of Natural History, Entomology Department Box 50007, SE- 10405 Stockholm, Sweden; E-mail: kjell. arne. johanson @ nrm. se</mods:affiliation>
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<mods:typeOfResource id="DE173BADF49075E407A3283C0D078ED6">text</mods:typeOfResource>
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<mods:title id="CFDC43D300903DF9BD7F4CC565E8C592">Acta Zoologica Academiae Scientiarum Hungaricae</mods:title>
</mods:titleInfo>
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<mods:date id="642D796C0597DF5AC264AA006216FFC0">2011</mods:date>
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<mods:number id="EB0C0A894267141FD4455B1A72BDFC16">2011-06-10</mods:number>
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<mods:number id="073AA0A56940769F77C45CA36ADA7BAA">57</mods:number>
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<mods:classification id="F7C498B8ED092093938697396EA69220">journal article</mods:classification>
<mods:identifier id="81E60F9AAC2F4F0B9748470086885D4C" type="DOI">10.5281/zenodo.12612328</mods:identifier>
<mods:identifier id="8827F2714C774A6AB2C30792DB930830" type="ISSN">2064-2474</mods:identifier>
<mods:identifier id="8AD02B722D2DDB38E0ED5B8F3F94FC27" type="Zenodo-Dep">12612328</mods:identifier>
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<treatment id="747987ACFF9464185D584A63FC08FD75" LSID="urn:lsid:plazi:treatment:747987ACFF9464185D584A63FC08FD75" httpUri="http://treatment.plazi.org/id/747987ACFF9464185D584A63FC08FD75" lastPageId="8" lastPageNumber="125" pageId="6" pageNumber="123">
<subSubSection id="B4CA6531FF9464165D584A63FC10FCC9" box="[616,970,855,881]" pageId="6" pageNumber="123" type="nomenclature">
<paragraph id="FC6F36BAFF9464165D584A63FC10FCC9" blockId="6.[616,970,855,881]" box="[616,970,855,881]" pageId="6" pageNumber="123">
<heading id="A72781D6FF9464165D584A63FC10FCC9" box="[616,970,855,881]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="123" reason="2">
<taxonomicName id="3BD04D39FF9464165D584A63FC5FFCC9" authorityName="Oláh &amp; Johanson" authorityYear="2011" box="[616,901,855,881]" class="Insecta" family="Odontoceridae" genus="Madagocerum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="idvigun" status="sp. nov.">
<emphasis id="CEA4EAA8FF9464165D584A63FC5FFCC9" bold="true" box="[616,901,855,881]" pageId="6" pageNumber="123">Madagocerum idvigun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D59757D3FF9464165CBC4A63FC10FCC9" box="[908,970,855,881]" pageId="6" pageNumber="123" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B4CA6531FF9464185DE54A4FFC08FD75" lastPageId="8" lastPageNumber="125" pageId="6" pageNumber="123" type="description">
<paragraph id="FC6F36BAFF9464165DE54A4FFC84FC2D" blockId="6.[725,862,891,917]" box="[725,862,891,917]" pageId="6" pageNumber="123">
<heading id="A72781D6FF9464165DE54A4FFC84FC2D" box="[725,862,891,917]" centered="true" fontSize="11" level="3" pageId="6" pageNumber="123" reason="3">
(
<figureCitation id="64EB2A3FFF9464165DED4A4FFC8FFC2D" box="[733,853,891,917]" captionStart-0="Figs 5–12" captionStart-1="Figs 13–16" captionStartId-0="7.[317,360,1751,1772]" captionStartId-1="8.[317,360,1780,1801]" captionTargetBox-0="[321,1267,650,1718]" captionTargetBox-1="[322,1266,773,1741]" captionTargetId-0="figure-94@7.[317,1269,649,1722]" captionTargetId-1="figure-109@8.[317,1269,768,1743]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Figs 5–12. Madagocerum idvigum sp. n., holotype: 5 = head, frontal view, 6 = head dorsal view, 7 = right maxilary palp, 8 = tentorium, dorsal view, 9 = tentorium, lateral view, 10 = pronotum, dorsal" captionText-1="Figs 13–16. Madagocerum idvigum sp. n., holotype: 13 = genitalia, lateral view, 14 = genitalia, dorsal view, 15 = genitalia, ventral view, 16 – phallus, lateral view" figureDoi-0="http://doi.org/10.5281/zenodo.12612332" figureDoi-1="http://doi.org/10.5281/zenodo.12612335" httpUri-0="https://zenodo.org/record/12612332/files/figure.png" httpUri-1="https://zenodo.org/record/12612335/files/figure.png" pageId="6" pageNumber="123">Figs 5–16</figureCitation>
)
</heading>
</paragraph>
<paragraph id="FC6F36BAFF9464165E4E4AF5FD75FB16" blockId="6.[317,1270,961,1198]" pageId="6" pageNumber="123">
Diagnosis – This species resembles
<taxonomicName id="3BD04D39FF9464165C164AF5FC4FFC63" authorityName="Oláh &amp; Johanson" authorityYear="2011" box="[806,917,961,987]" class="Insecta" family="Odontoceridae" genus="Madagocerum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="bhemi" status="sp. n.">
<emphasis id="CEA4EAA8FF9464165C164AF5FC4FFC63" box="[806,917,961,987]" italics="true" pageId="6" pageNumber="123">M. bhemi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D59757D3FF9464165CAC4AF5FC01FC63" box="[924,987,961,987]" pageId="6" pageNumber="123" rank="species">sp. n.</taxonomicNameLabel>
in its short segment IX, but the remaining parts of the genitalia are different.
<taxonomicName id="3BD04D39FF9464165C964AD0FB6AFC46" authorityName="Oláh &amp; Johanson" authorityYear="2011" box="[934,1200,996,1022]" class="Insecta" family="Odontoceridae" genus="Madagocerum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="idvigun" status="sp. nov.">
<emphasis id="CEA4EAA8FF9464165C964AD0FB6AFC46" box="[934,1200,996,1022]" italics="true" pageId="6" pageNumber="123">Madagocerum idvigun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D59757D3FF9464165B874AD0FB2FFC46" box="[1207,1269,996,1022]" pageId="6" pageNumber="123" rank="species">sp. n.</taxonomicNameLabel>
has narrow, tapering and tangential preanal appendages, not broad, blunt and divergent as in
<taxonomicName id="3BD04D39FF9464165EE44D1FFD98FBFD" authorityName="Oláh &amp; Johanson" authorityYear="2011" box="[468,578,1067,1093]" class="Insecta" family="Odontoceridae" genus="Madagocerum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="bhemi">
<emphasis id="CEA4EAA8FF9464165EE44D1FFD98FBFD" box="[468,578,1067,1093]" italics="true" pageId="6" pageNumber="123">M. bhemi</emphasis>
</taxonomicName>
; segment X is as long as the preanal appendages, not shorter as in
<taxonomicName id="3BD04D39FF9464165E4A4D7AFE33FBD0" authorityName="Oláh &amp; Johanson" authorityYear="2011" box="[378,489,1102,1128]" class="Insecta" family="Odontoceridae" genus="Madagocerum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="bhemi">
<emphasis id="CEA4EAA8FF9464165E4A4D7AFE33FBD0" box="[378,489,1102,1128]" italics="true" pageId="6" pageNumber="123">M. bhemi</emphasis>
</taxonomicName>
; each coxopodite has an apicodorsal spiny protuberance; both harpagones are broad, not narrow as in
<taxonomicName id="3BD04D39FF9464165DDD4D45FC87FB33" authorityName="Oláh &amp; Johanson" authorityYear="2011" box="[749,861,1137,1163]" class="Insecta" family="Odontoceridae" genus="Madagocerum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="bhemi">
<emphasis id="CEA4EAA8FF9464165DDD4D45FC87FB33" box="[749,861,1137,1163]" italics="true" pageId="6" pageNumber="123">M. bhemi</emphasis>
</taxonomicName>
; and the phallic apparatus is more slender and narrowing apically.
</paragraph>
<paragraph id="FC6F36BAFF9464165E4E4DE5FE25FABB" blockId="6.[317,1271,1233,1801]" pageId="6" pageNumber="123">Description – Male (in alcohol). Body light brown; forewing membrane light brown, without pattern (in alcohol).</paragraph>
<paragraph id="FC6F36BAFF9464165E4E4C3FFE26FA2B" blockId="6.[317,1271,1233,1801]" pageId="6" pageNumber="123">
Wings (
<figureCitation id="64EB2A3FFF9464165EFA4C3FFDC9FA98" box="[458,531,1291,1312]" captionStart="Figs 5–12" captionStartId="7.[317,360,1751,1772]" captionTargetBox="[321,1267,650,1718]" captionTargetId="figure-94@7.[317,1269,649,1722]" captionTargetPageId="7" captionText="Figs 5–12. Madagocerum idvigum sp. n., holotype: 5 = head, frontal view, 6 = head dorsal view, 7 = right maxilary palp, 8 = tentorium, dorsal view, 9 = tentorium, lateral view, 10 = pronotum, dorsal" figureDoi="http://doi.org/10.5281/zenodo.12612332" httpUri="https://zenodo.org/record/12612332/files/figure.png" pageId="6" pageNumber="123">Fig. 12</figureCitation>
): Forewing apex narrowly rounded; 5.0 mm; membrane light brown; termen slightly convex; basal lobe covered by microtrichia; jugum absent. Forewing forks I, II, III, and V present; Sc fusing with R1; R1 fusing with R2 before C, meeting at hypertrophied pterostigma; distally almost indiscernible; Cu2 apparently absent in forewings; postanal vein running closely to posterior wing-margin.
</paragraph>
<paragraph id="FC6F36BAFF9464185E4E4CAFFC90FDBB" blockId="6.[317,1271,1233,1801]" lastBlockId="8.[317,1270,436,515]" lastPageId="8" lastPageNumber="125" pageId="6" pageNumber="123">
Male genitalia (
<figureCitation id="64EB2A3FFF9464165D294CAFFD5EFA08" box="[537,644,1435,1456]" captionStart="Figs 13–16" captionStartId="8.[317,360,1780,1801]" captionTargetBox="[322,1266,773,1741]" captionTargetId="figure-109@8.[317,1269,768,1743]" captionTargetPageId="8" captionText="Figs 13–16. Madagocerum idvigum sp. n., holotype: 13 = genitalia, lateral view, 14 = genitalia, dorsal view, 15 = genitalia, ventral view, 16 – phallus, lateral view" figureDoi="http://doi.org/10.5281/zenodo.12612335" httpUri="https://zenodo.org/record/12612335/files/figure.png" pageId="6" pageNumber="123">Figs13–16</figureCitation>
): Abdominal segment IX fused annularly, short in lateral view; tergum almost as long as venter; anterior margin convex, with triangular pleural plate bearing mesally depressed lateral concavity in dorsal view; posterior margin straight, with tiny subtriangular apical lobe below preanal appendages; antecosta and antecostal suture, on anterior margin of segment IX narrow, forming dark marginal rim running evenly along anterior margin; acrotergite well developed, with sclerotized rim; entire surface of segment IX glabrous, spine row absent on posterior margins of segment IX, setose areas absent from apicopleural and apicoventral regions. Intersegmental depression between segments IX and X very deep, stepwise. Segment X sunk deep to upper 1/3rd of segment IX in lateral view; intersegmental depression covered by fused base of preanal appendages. Segment X very shallow, slender; deeply bilobed; slightly arching mesad in dorsal view, dorsad in lateral view. Apicoventral setose lobes represented by apex; apicodorsal setose lobes reduced to small setal areas on middle; segment X deeply cleft; dorsal interlobular gap long, almost closed in ventral view. Large, deeply bilobed preanal appendages dominating over phallic apparatus; fused basally; horizontal, posterad directed lobes fused to dorsum of segment IX; fused seam, representing borderline between segment IX and preanal appendages invisible; in lateral view, dorsum straight horizontal, venter slightly tapering; in dorsal view with individual preanal appendages, or bilobed structures, more slender than in dorsal view, tapering, weakly constricted; mesal margins parallel sided in dorsal view; almost touching along their lengths. Coxopodites robust, with protuberance on apicodorsal corner covered by stout, medium-long spines; slightly directed mesad. Harpagones large, view, 11 = mesonotum, dorsal view, 12 = right wings broad, with slightly ventrad and mesad curving spiny apical surface, spines dark, short, conical. Phallic apparatus with broad, ventrally directed, short, basal part; apical part horizontal, straight, narrowing. Small phallotremal sclerites almost indiscernible.
</paragraph>
<caption id="A8AF6632FF9564175E0D4FE3FB2CF8B1" ID-DOI="http://doi.org/10.5281/zenodo.12612332" ID-Zenodo-Dep="12612332" httpUri="https://zenodo.org/record/12612332/files/figure.png" pageId="7" pageNumber="124" startId="7.[317,360,1751,1772]" targetBox="[321,1267,650,1718]" targetPageId="7" targetType="figure">
<paragraph id="FC6F36BAFF9564175E0D4FE3FB2CF8B1" blockId="7.[317,1270,1751,1830]" pageId="7" pageNumber="124">
<emphasis id="CEA4EAA8FF9564175E0D4FE3FE79F954" bold="true" box="[317,419,1751,1772]" pageId="7" pageNumber="124">Figs 5–12.</emphasis>
<taxonomicName id="3BD04D39FF9564175E984FECFD5DF955" authorityName="Oláh &amp; Johanson" authorityYear="2011" box="[424,647,1752,1773]" class="Insecta" family="Odontoceridae" genus="Madagocerum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="124" phylum="Arthropoda" rank="species" species="idvigum" status="sp. nov.">
<emphasis id="CEA4EAA8FF9564175E984FECFD5DF955" box="[424,647,1752,1773]" italics="true" pageId="7" pageNumber="124">Madagocerum idvigum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D59757D3FF9564175DBC4FECFD1AF955" box="[652,704,1752,1773]" pageId="7" pageNumber="124" rank="species">sp. n.</taxonomicNameLabel>
, holotype: 5 = head, frontal view, 6 = head dorsal view, 7 = right maxilary palp, 8 = tentorium, dorsal view, 9 = tentorium, lateral view, 10 = pronotum, dorsal
</paragraph>
</caption>
<paragraph id="FC6F36BAFF9A64185E4E4B1CFB2FFD85" blockId="8.[317,1269,552,601]" box="[382,1269,552,573]" pageId="8" pageNumber="125">
<materialsCitation id="4CB83CE7FF9A64185E4E4B1CFB2FFD85" box="[382,1269,552,573]" collectingDate="1958-01-22" country="Madagascar" county="Imatso" elevation="1550" location="Material" municipality="Foret" pageId="8" pageNumber="125" specimenCount="1" typeStatus="holotype">
<location id="F90F6061FF9A64185E4E4B1CFE14FD85" LSID="urn:lsid:plazi:treatment:747987ACFF9464185D584A63FC08FD75:F90F6061FF9A64185E4E4B1CFE14FD85" box="[382,462,552,573]" country="Madagascar" county="Imatso" municipality="Foret" name="Material" pageId="8" pageNumber="125">Material</location>
examined –
<typeStatus id="236B8818FF9A64185D754B1CFD7AFD85" box="[581,672,552,573]" pageId="8" pageNumber="125" type="holotype">Holotype</typeStatus>
:
<location id="F90F6061FF9A64185D984B1CFD06FD85" LSID="urn:lsid:plazi:treatment:747987ACFF9464185D584A63FC08FD75:F90F6061FF9A64185D984B1CFD06FD85" box="[680,732,552,573]" country="Madagascar" county="Imatso" municipality="Foret" name="Male" pageId="8" pageNumber="125">Male</location>
: “
<collectingCountry id="84C7762AFF9A64185DDC4B1CFCB9FD85" box="[748,867,552,573]" name="Madagascar" pageId="8" pageNumber="125">Madagascar</collectingCountry>
:
<location id="F90F6061FF9A64185C5C4B1CFC1BFD85" LSID="urn:lsid:plazi:treatment:747987ACFF9464185D584A63FC08FD75:F90F6061FF9A64185C5C4B1CFC1BFD85" box="[876,961,552,573]" country="Madagascar" county="Imatso" municipality="Foret" name="Vacoana" pageId="8" pageNumber="125">Vacoana</location>
,
<collectingMunicipality id="1C0BACC0FF9A64185CF94B1CFC27FD85" box="[969,1021,552,573]" pageId="8" pageNumber="125">Foret</collectingMunicipality>
,
<collectingCounty id="150E4E36FF9A64185B364B1CFB9CFD85" box="[1030,1094,552,573]" pageId="8" pageNumber="125">Imatso</collectingCounty>
<quantity id="3B289B5FFF9A64185B7B4B1CFB4EFD85" box="[1099,1172,552,573]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="8" pageNumber="125" unit="m" value="1550.0">
<elevation id="77FDD189FF9A64185B7B4B1CFB4EFD85" box="[1099,1172,552,573]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="8" pageNumber="125" unit="m" value="1550.0">1550 m</elevation>
</quantity>
,
<date id="886E107AFF9A64185BAD4B1CFB2FFD85" box="[1181,1269,552,573]" pageId="8" pageNumber="125" value="1958-01-22">
<collectingDate id="982AE992FF9A64185BAD4B1CFB2FFD85" box="[1181,1269,552,573]" pageId="8" pageNumber="125" value="1958-01-22">22.i.1958</collectingDate>
</date>
</materialsCitation>
</paragraph>
<paragraph id="FC6F36BAFF9A64185E0D4B70FE7DFDE1" blockId="8.[317,1269,552,601]" box="[317,423,580,601]" pageId="8" pageNumber="125">[Paulian]”.</paragraph>
<paragraph id="FC6F36BAFF9A64185E4E4B4AFC08FD75" blockId="8.[317,1270,638,717]" pageId="8" pageNumber="125">
Etymology:
<emphasis id="CEA4EAA8FF9A64185EC44B4AFDE7FD2B" box="[500,573,638,659]" italics="true" pageId="8" pageNumber="125">Idvigun</emphasis>
, from Sanskrit “idvigunn”, meaning double; referring to the similar shape of the preanal appendages and segment X in dorsal view; and the additional spiny hump on the dorsum of the coxopodites duplicate the spiny apex of the harpagones.
</paragraph>
</subSubSection>
</treatment>
</document>
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