treatments-xml/data/74/1E/87/741E87E4FFA3FD0764D835D17BCAFB57.xml
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<mods:title id="1E66FC34DBC260C6205251E3DFFA64A8">A new species of Pseudouroctonus from the Pinaleño Mountains southern Arizona (Scorpiones Vaejovidae)</mods:title>
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<taxonomicName id="3BB74D71FFA3FD0A64D835D17C25FB08" ID-CoL="992RH" authority="Ayrey &amp; Kovařík &amp; Myers, 2021" authorityName="Ayrey &amp; Kovařík &amp; Myers" authorityYear="2021" box="[255,556,1224,1251]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="moyeri" status="sp. nov.">Pseudouroctonus moyeri</taxonomicName>
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(
<figureCitation id="648C2A77FFA3FD0A651535F27FCBFAE8" box="[306,450,1259,1283]" captionStart-0="Figures 12" captionStart-1="Figures 36" captionStart-2="Figures 714" captionStart-3="Figures 1521" captionStart-4="Figures 2231" captionStart-5="Figure 3233" captionStart-6="Figure 3435" captionStartId-0="3.[113,190,2014,2036]" captionStartId-1="5.[115,192,1992,2014]" captionStartId-2="6.[115,192,1980,2002]" captionStartId-3="7.[115,192,1979,2001]" captionStartId-4="8.[115,192,1940,1962]" captionStartId-5="10.[115,183,2005,2027]" captionStartId-6="11.[114,182,1989,2011]" captionTargetBox-0="[113,1474,234,2001]" captionTargetBox-1="[165,1431,245,1975]" captionTargetBox-2="[113,1459,299,1907]" captionTargetBox-3="[113,1461,244,1949]" captionTargetBox-4="[120,1469,280,1879]" captionTargetBox-5="[113,1474,234,1973]" captionTargetBox-6="[113,1474,234,1974]" captionTargetId-0="figure-28@3.[113,1474,234,2003]" captionTargetId-1="figure-46@5.[113,1474,242,1980]" captionTargetId-2="figure-55@6.[113,1474,287,1955]" captionTargetId-3="figure-47@7.[113,1474,244,1949]" captionTargetId-4="figure-65@8.[113,1474,278,1892]" captionTargetId-5="figure-37@10.[113,1474,234,1975]" captionTargetId-6="figure-42@11.[113,1474,234,1977]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="10" captionTargetPageId-6="11" captionText-0="Figures 12. Pseudouroctonus moyeri sp. n., paratypes, female (1) and male shortly after last ecdysis (2)." captionText-1="Figures 36: Pseudouroctonus moyeri sp. n., paratypes. Figures 34. Female in dorsal (3) and ventral (4) views. Figures 56. Male in dorsal (5) and ventral (6) views." captionText-2="Figures 714. Pseudouroctonus moyeri sp. n., paratype male (FKCP), pedipalp. chela, dorsal (7), external (8), and ventral (9) views. Patella, dorsal (10), external (11) and ventral (12) views. Femur and trochanter dorsal (13). Movable finger (14)." captionText-3="Figures 1521. Pseudouroctonus moyeri sp. n., paratype male (FKCP), sternopectinal area (15), left chelicerae, ventral view (16), right legs IIIIV, retrolateral aspect (1718), metasoma and telson lateral (19), ventral (20) and dorsal (21) views." captionText-4="Figures 2231: Pseudouroctonus moyeri sp. n., paratype male (USNM). Figures 2225. Hemispermatophore internal (22), ventral (23), external (24), and dorsal (25) views. Figures 2631. Mating plug dorsal (26), distal (27), ventral (28), proximal (29), internal (30), and external (31) views." captionText-5="Figure 3233. Pseudouroctonus moyeri sp. n., paratype juvenile female 4th instar (32) and female with newborns (33)." captionText-6="Figure 3435: Figure 34. Map showing the type localities for the four species of Pseudouroctonus known from southern Arizona. Figure 35. Pseudouroctonus moyeri sp. n., type locality." figureDoi-0="http://doi.org/10.5281/zenodo.5742155" figureDoi-1="http://doi.org/10.5281/zenodo.5742157" figureDoi-2="http://doi.org/10.5281/zenodo.5742161" figureDoi-3="http://doi.org/10.5281/zenodo.5742165" figureDoi-4="http://doi.org/10.5281/zenodo.5742169" figureDoi-5="http://doi.org/10.5281/zenodo.5742173" figureDoi-6="http://doi.org/10.5281/zenodo.5742175" httpUri-0="https://zenodo.org/record/5742155/files/figure.png" httpUri-1="https://zenodo.org/record/5742157/files/figure.png" httpUri-2="https://zenodo.org/record/5742161/files/figure.png" httpUri-3="https://zenodo.org/record/5742165/files/figure.png" httpUri-4="https://zenodo.org/record/5742169/files/figure.png" httpUri-5="https://zenodo.org/record/5742173/files/figure.png" httpUri-6="https://zenodo.org/record/5742175/files/figure.png" pageId="4" pageNumber="3">Figures 135</figureCitation>
,
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, leg.
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<collectionCode id="9AA6AE37FFA3FD0A661337727C8CF968" box="[564,645,1643,1667]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="3" type="Museum">USNM</collectionCode>
);
<specimenCount id="EAB1FD7BFFA3FD0A66B237727CBFF968" box="[661,694,1643,1667]" pageId="4" pageNumber="3" type="female">1♀</specimenCount>
,
<date id="88091032FFA3FD0A66E637727EAEF948" pageId="4" pageNumber="3" value="2015-06-05">5 June 2015</date>
, leg.
<collectionCode id="9AA6AE37FFA3FD0A64C737927EFDF949" box="[224,244,1675,1698]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="3">R</collectionCode>
. Moyer, specimen #1167 (
<collectionCode id="9AA6AE37FFA3FD0A662037927C34F948" box="[519,573,1675,1699]" country="Brazil" lsid="urn:lsid:biocol.org:col:14341" name="Universidade Federal do Rio de Janeiro" pageId="4" pageNumber="3" type="Herbarium">RFA</collectionCode>
);
<specimenCount id="EAB1FD7BFFA3FD0A667737927C78F948" box="[592,625,1675,1699]" pageId="4" pageNumber="3" type="female">1♀</specimenCount>
,
<date id="88091032FFA3FD0A665937927D09F948" box="[638,768,1675,1699]" pageId="4" pageNumber="3" value="2015-06-05">5 June 2015</date>
, leg.
<collectionCode id="9AA6AE37FFA3FD0A648737B27EBDF929" box="[160,180,1707,1730]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="3">R</collectionCode>
. Moyer, specimen #1168 (
<collectionCode id="9AA6AE37FFA3FD0A65F737B27C0FF928" box="[464,518,1707,1731]" country="Brazil" lsid="urn:lsid:biocol.org:col:14341" name="Universidade Federal do Rio de Janeiro" pageId="4" pageNumber="3" type="Herbarium">RFA</collectionCode>
);
<specimenCount id="EAB1FD7BFFA3FD0A663D37B27C35F928" box="[538,572,1707,1731]" pageId="4" pageNumber="3" type="male">1♂</specimenCount>
,
<date id="88091032FFA3FD0A666D37B27CD9F928" box="[586,720,1707,1731]" pageId="4" pageNumber="3" value="2010-05-05">5 May 2010</date>
, leg.
<collectionCode id="9AA6AE37FFA3FD0A645637D27E8CF909" box="[113,133,1739,1762]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="3">R</collectionCode>
. F. Ayrey, specimen #1174 (
<collectionCode id="9AA6AE37FFA3FD0A658137D27FE4F908" box="[422,493,1739,1763]" pageId="4" pageNumber="3">FKCP</collectionCode>
). The type specimens were found using a blacklight at night. The vegetation type is pine oak woodland (see
<figureCitation id="648C2A77FFA3FD0A651236127F89F8C8" box="[309,384,1803,1827]" captionStart="Figure 3435" captionStartId="11.[114,182,1989,2011]" captionTargetBox="[113,1474,234,1974]" captionTargetId="figure-42@11.[113,1474,234,1977]" captionTargetPageId="11" captionText="Figure 3435: Figure 34. Map showing the type localities for the four species of Pseudouroctonus known from southern Arizona. Figure 35. Pseudouroctonus moyeri sp. n., type locality." figureDoi="http://doi.org/10.5281/zenodo.5742175" httpUri="https://zenodo.org/record/5742175/files/figure.png" pageId="4" pageNumber="3">Fig. 35</figureCitation>
).
<taxonomicName id="3BB74D71FFA3FD0A65B636127CD1F8C8" authority="Hughes, 2011" authorityName="Hughes" authorityYear="2011" box="[401,728,1803,1827]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="electrum">
<emphasis id="CEC3EAE0FFA3FD0A65B636127C4FF8C8" box="[401,582,1803,1827]" italics="true" pageId="4" pageNumber="3">Vaejovis electrum</emphasis>
Hughes, 2011
</taxonomicName>
and
<taxonomicName id="3BB74D71FFA3FD0A645636327C0FF8A8" authority="Ewing, 1928" authorityName="Ewing" authorityYear="1928" box="[113,518,1835,1859]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="sculpturatus">
<emphasis id="CEC3EAE0FFA3FD0A645636327F74F8A8" box="[113,381,1835,1859]" italics="true" pageId="4" pageNumber="3">Centruroides sculpturatus</emphasis>
Ewing, 1928
</taxonomicName>
were found syntopically with
<taxonomicName id="3BB74D71FFA3FD0A648236527F0FF889" authorityName="Ayrey &amp; Kovařík &amp; Myers" authorityYear="2021" box="[165,262,1867,1890]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="moyeri">
<emphasis id="CEC3EAE0FFA3FD0A648236527F0FF889" box="[165,262,1867,1890]" italics="true" pageId="4" pageNumber="3">P. moyeri</emphasis>
</taxonomicName>
during five field trips to Mount Graham.
</paragraph>
<paragraph id="FC0836F2FFA3FD0A645636727C5AF7E8" blockId="4.[113,772,1514,2051]" pageId="4" pageNumber="3">
<materialsCitation id="4CDF3CAFFFA3FD0A645636727F5BF848" collectingDate="2010-05-05" collectionCode="OTHER, MATERIAL, STUDIED" pageId="4" pageNumber="3" specimenCount="5" specimenCount-female="3" specimenCount-male="2">
<collectionCode id="9AA6AE37FFA3FD0A645636727F88F868" box="[113,385,1899,1923]" pageId="4" pageNumber="3">OTHER MATERIAL STUDIED</collectionCode>
.
<specimenCount id="EAB1FD7BFFA3FD0A65AA36727FB9F868" box="[397,432,1899,1923]" pageId="4" pageNumber="3" type="male">1♂</specimenCount>
<specimenCount id="EAB1FD7BFFA3FD0A659736727FD9F868" box="[432,464,1899,1923]" pageId="4" pageNumber="3" type="female">3♀</specimenCount>
(same locality as paratypes):
<specimenCount id="EAB1FD7BFFA3FD0A645636927E9DF848" box="[113,148,1931,1955]" pageId="4" pageNumber="3" type="male">1♂</specimenCount>
,
<date id="88091032FFA3FD0A648736927F2DF848" box="[160,292,1931,1955]" pageId="4" pageNumber="3" value="2010-05-05">
<collectingDate id="984DE9DAFFA3FD0A648736927F2DF848" box="[160,292,1931,1955]" pageId="4" pageNumber="3" value="2010-05-05">5 May 2010</collectingDate>
</date>
, leg
</materialsCitation>
.
<collectionCode id="9AA6AE37FFA3FD0A657B36927F78F849" box="[348,369,1931,1954]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="3">R</collectionCode>
.
<materialsCitation id="4CDF3CAFFFA3FD0A655C36927F27F828" collectingDate="2015-06-05" collectionCode="RFA" pageId="4" pageNumber="3" specimenCount="1" specimenCount-female="1">
Moyer, specimen #1166, (
<collectionCode id="9AA6AE37FFA3FD0A66B636927CC0F848" box="[657,713,1931,1955]" country="Brazil" lsid="urn:lsid:biocol.org:col:14341" name="Universidade Federal do Rio de Janeiro" pageId="4" pageNumber="3" type="Herbarium">RFA</collectionCode>
);
<specimenCount id="EAB1FD7BFFA3FD0A66FC36927CF5F848" box="[731,764,1931,1955]" pageId="4" pageNumber="3" type="female">1♀</specimenCount>
,
<date id="88091032FFA3FD0A645636B27EF5F828" box="[113,252,1963,1987]" pageId="4" pageNumber="3" value="2015-06-05">
<collectingDate id="984DE9DAFFA3FD0A645636B27EF5F828" box="[113,252,1963,1987]" pageId="4" pageNumber="3" value="2015-06-05">5 June 2015</collectingDate>
</date>
, leg
</materialsCitation>
.
<collectionCode id="9AA6AE37FFA3FD0A651C36B27F59F829" box="[315,336,1963,1986]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="3">R</collectionCode>
.
<materialsCitation id="4CDF3CAFFFA3FD0A657A36B27F06F808" collectingDate="2015-06-05" collectionCode="RFA" pageId="4" pageNumber="3" specimenCount="1" specimenCount-female="1">
Moyer, specimen #1165 (
<collectionCode id="9AA6AE37FFA3FD0A665136B27CA7F828" box="[630,686,1963,1987]" country="Brazil" lsid="urn:lsid:biocol.org:col:14341" name="Universidade Federal do Rio de Janeiro" pageId="4" pageNumber="3" type="Herbarium">RFA</collectionCode>
);
<specimenCount id="EAB1FD7BFFA3FD0A66E336B27CECF828" box="[708,741,1963,1987]" pageId="4" pageNumber="3" type="female">1♀</specimenCount>
,
<date id="88091032FFA3FD0A66D136B27EE8F808" pageId="4" pageNumber="3" value="2015-06-05">
<collectingDate id="984DE9DAFFA3FD0A66D136B27EE8F808" pageId="4" pageNumber="3" value="2015-06-05">5 June 2015</collectingDate>
</date>
, leg
</materialsCitation>
.
<collectionCode id="9AA6AE37FFA3FD0A653E36D27F27F809" box="[281,302,1995,2018]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="3">R</collectionCode>
.
<materialsCitation id="4CDF3CAFFFA3FD0A651F36D27ED1F7E8" collectingDate="2015-06-05" collectionCode="RFA" pageId="4" pageNumber="3" specimenCount="1" specimenCount-female="1">
Moyer, specimen #1169 (
<collectionCode id="9AA6AE37FFA3FD0A666136D27C77F808" box="[582,638,1995,2019]" country="Brazil" lsid="urn:lsid:biocol.org:col:14341" name="Universidade Federal do Rio de Janeiro" pageId="4" pageNumber="3" type="Herbarium">RFA</collectionCode>
);
<specimenCount id="EAB1FD7BFFA3FD0A66B736D27CB8F808" box="[656,689,1995,2019]" pageId="4" pageNumber="3" type="female">1♀</specimenCount>
,
<date id="88091032FFA3FD0A669936D27EA0F7E8" pageId="4" pageNumber="3" value="2015-06-05">
<collectingDate id="984DE9DAFFA3FD0A669936D27EA0F7E8" pageId="4" pageNumber="3" value="2015-06-05">5 June 2015</collectingDate>
</date>
, leg
</materialsCitation>
.
<collectionCode id="9AA6AE37FFA3FD0A64C536F27EFEF7E9" box="[226,247,2027,2050]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="3">R</collectionCode>
. Moyer, specimen #1170 (
<collectionCode id="9AA6AE37FFA3FD0A663736F27C41F7E8" box="[528,584,2027,2051]" country="Brazil" lsid="urn:lsid:biocol.org:col:14341" name="Universidade Federal do Rio de Janeiro" pageId="4" pageNumber="3" type="Herbarium">RFA</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="B4AD6579FFA3FD0A671735D07ACAFAEA" pageId="4" pageNumber="3" type="etymology">
<paragraph id="FC0836F2FFA3FD0A671735D07ACAFAEA" blockId="4.[816,1474,1225,1281]" pageId="4" pageNumber="3">
ETYMOLOGY. This species is named in honor of Ryan Moyer who collected most of the
<typeStatus id="230C8850FFA3FD0A606C35F07A70FAEA" box="[1099,1145,1257,1281]" pageId="4" pageNumber="3">type</typeStatus>
series.
</paragraph>
</subSubSection>
<subSubSection id="B4AD6579FFA3FD0A671734307A08F86A" pageId="4" pageNumber="3" type="diagnosis">
<paragraph id="FC0836F2FFA3FD0A671734307B3CFA0A" blockId="4.[816,1475,1321,1505]" pageId="4" pageNumber="3">
DIAGNOSIS. Large (
<quantity id="3B4F9B17FFA3FD0A602B34307A82FAAA" box="[1036,1163,1321,1345]" metricMagnitude="-2" metricUnit="m" metricValue="3.45" metricValueMax="3.7" metricValueMin="3.2" pageId="4" pageNumber="3" unit="mm" value="34.5" valueMax="37.0" valueMin="32.0">3237 mm</quantity>
) scorpions. Color is dark, reddish brown, lighter on legs (see
<figureCitation id="648C2A77FFA3FD0A608C34507B18FA8A" box="[1195,1297,1353,1377]" captionStart="Figures 12" captionStartId="3.[113,190,2014,2036]" captionTargetBox="[113,1474,234,2001]" captionTargetId="figure-28@3.[113,1474,234,2003]" captionTargetPageId="3" captionText="Figures 12. Pseudouroctonus moyeri sp. n., paratypes, female (1) and male shortly after last ecdysis (2)." figureDoi="http://doi.org/10.5281/zenodo.5742155" httpUri="https://zenodo.org/record/5742155/files/figure.png" pageId="4" pageNumber="3">Figs. 12</figureCitation>
), telson reddish. Pedipalp movable finger usually with 7 ID denticles and fixed finger usually with 6. Carapace of female is the same size or longer than the metasomal segment V. Mean pectinal tooth count: females 8.916 [n=12], males 11.00 [n=6].
</paragraph>
<paragraph id="FC0836F2FFA3FD0A671737107A08F86A" blockId="4.[816,1475,1545,2049]" pageId="4" pageNumber="3">
<emphasis id="CEC3EAE0FFA3FD0A671737107A12F9CA" bold="true" box="[816,1051,1545,1569]" pageId="4" pageNumber="3">Hemispermatophore</emphasis>
(
<figureCitation id="648C2A77FFA3FD0A601C37107AC7F9CA" box="[1083,1230,1545,1569]" captionStart="Figures 2231" captionStartId="8.[115,192,1940,1962]" captionTargetBox="[120,1469,280,1879]" captionTargetId="figure-65@8.[113,1474,278,1892]" captionTargetPageId="8" captionText="Figures 2231: Pseudouroctonus moyeri sp. n., paratype male (USNM). Figures 2225. Hemispermatophore internal (22), ventral (23), external (24), and dorsal (25) views. Figures 2631. Mating plug dorsal (26), distal (27), ventral (28), proximal (29), internal (30), and external (31) views." figureDoi="http://doi.org/10.5281/zenodo.5742169" httpUri="https://zenodo.org/record/5742169/files/figure.png" pageId="4" pageNumber="3">Figs. 2225</figureCitation>
). Description based on left hemispermatophore. All measurements in mm. Hemispermatophore with wide base, deep dorsal trough. Lamina edges nearly parallel with no obvious basal constriction. Distinct distal crest on dorsal side of lamina, on posterior edge. Strongly bifurcated lamellar hook, originating from dorsal trough. Measurements as follows (mm): Total length = 4.47, lamina length = 2.74, lamellar hook length = 0.96, trough difference = 0.43, base width = 0.98, lamina width = 0.44. Morphometric ratios are as follows: lamellar hook length / lamina length = 0.35, trough difference / lamellar hook length = 0.45.
</paragraph>
</subSubSection>
<subSubSection id="B4AD6579FFA3FD07671736907D82FBB7" lastPageId="9" lastPageNumber="8" pageId="4" pageNumber="3" type="description">
<paragraph id="FC0836F2FFA3FD0A671736907A71F7EB" blockId="4.[816,1475,1545,2049]" pageId="4" pageNumber="3">
<emphasis id="CEC3EAE0FFA3FD0A671736907DB5F84A" bold="true" box="[816,956,1929,1953]" pageId="4" pageNumber="3">Mating Plug</emphasis>
(
<figureCitation id="648C2A77FFA3FD0A67EE36907A43F84A" box="[969,1098,1929,1953]" captionStart="Figures 2231" captionStartId="8.[115,192,1940,1962]" captionTargetBox="[120,1469,280,1879]" captionTargetId="figure-65@8.[113,1474,278,1892]" captionTargetPageId="8" captionText="Figures 2231: Pseudouroctonus moyeri sp. n., paratype male (USNM). Figures 2225. Hemispermatophore internal (22), ventral (23), external (24), and dorsal (25) views. Figures 2631. Mating plug dorsal (26), distal (27), ventral (28), proximal (29), internal (30), and external (31) views." figureDoi="http://doi.org/10.5281/zenodo.5742169" httpUri="https://zenodo.org/record/5742169/files/figure.png" pageId="4" pageNumber="3">Figs. 2631</figureCitation>
). Sclerotized mating plug with wide base, wide stem. Smooth barb, with a slight roundedness, but not crescent-shaped. One tine of barb significantly longer than other. Stem projection present.
</paragraph>
<caption id="A8C8667AFFA2FD0B645436D17F6EF810" ID-DOI="http://doi.org/10.5281/zenodo.5742157" ID-Zenodo-Dep="5742157" httpUri="https://zenodo.org/record/5742157/files/figure.png" pageId="5" pageNumber="4" startId="5.[115,192,1992,2014]" targetBox="[165,1431,245,1975]" targetPageId="5">
<paragraph id="FC0836F2FFA2FD0B645436D17F6EF810" blockId="5.[115,1473,1992,2043]" pageId="5" pageNumber="4">
<emphasis id="CEC3EAE0FFA2FD0B645436D17EE0F836" bold="true" box="[115,233,1992,2014]" pageId="5" pageNumber="4">Figures 36</emphasis>
:
<taxonomicName id="3BB74D71FFA2FD0B64D136D17FEDF835" authority="Ayrey &amp; Kovařík &amp; Myers, 2021" authorityName="Ayrey &amp; Kovařík &amp; Myers" authorityYear="2021" box="[246,484,1992,2014]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="moyeri" status="sp. nov.">
<emphasis id="CEC3EAE0FFA2FD0B64D136D17FEDF835" box="[246,484,1992,2014]" italics="true" pageId="5" pageNumber="4">Pseudouroctonus moyeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D5F0579BFFA2FD0B65CD36D17C16F835" box="[490,543,1992,2014]" pageId="5" pageNumber="4" rank="species">
<emphasis id="CEC3EAE0FFA2FD0B65CD36D17C08F836" bold="true" box="[490,513,1992,2013]" pageId="5" pageNumber="4">sp</emphasis>
.
<emphasis id="CEC3EAE0FFA2FD0B662B36D17C10F836" bold="true" box="[524,537,1992,2013]" pageId="5" pageNumber="4">n</emphasis>
.
</taxonomicNameLabel>
, paratypes.
<emphasis id="CEC3EAE0FFA2FD0B66B436D17D03F836" bold="true" box="[659,778,1992,2014]" pageId="5" pageNumber="4">Figures 34</emphasis>
. Female in dorsal (3) and ventral (4) views.
<emphasis id="CEC3EAE0FFA2FD0B609336D17B22F836" bold="true" box="[1204,1323,1992,2014]" pageId="5" pageNumber="4">Figures 56</emphasis>
. Male in dorsal (5) and ventral (6) views.
</paragraph>
</caption>
<caption id="A8C8667AFFA1FD08645436A57A55F804" ID-DOI="http://doi.org/10.5281/zenodo.5742161" ID-Zenodo-Dep="5742161" httpUri="https://zenodo.org/record/5742161/files/figure.png" pageId="6" pageNumber="5" startId="6.[115,192,1980,2002]" targetBox="[113,1459,299,1907]" targetPageId="6">
<paragraph id="FC0836F2FFA1FD08645436A57A55F804" blockId="6.[115,1473,1980,2031]" pageId="6" pageNumber="5">
<emphasis id="CEC3EAE0FFA1FD08645436A57EFFF83A" bold="true" box="[115,246,1980,2002]" pageId="6" pageNumber="5">Figures 714</emphasis>
.
<taxonomicName id="3BB74D71FFA1FD08652536A57FF8F83A" authority="Ayrey &amp; Kovařík &amp; Myers, 2021" authorityName="Ayrey &amp; Kovařík &amp; Myers" authorityYear="2021" box="[258,497,1980,2001]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="moyeri" status="sp. nov.">
<emphasis id="CEC3EAE0FFA1FD08652536A57FF8F83A" box="[258,497,1980,2001]" italics="true" pageId="6" pageNumber="5">Pseudouroctonus moyeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D5F0579BFFA1FD0865D036A57C25F83A" box="[503,556,1980,2001]" pageId="6" pageNumber="5" rank="species">
<emphasis id="CEC3EAE0FFA1FD0865D036A57C07F83A" bold="true" box="[503,526,1980,2001]" pageId="6" pageNumber="5">sp</emphasis>
.
<emphasis id="CEC3EAE0FFA1FD08663D36A57C2EF83A" bold="true" box="[538,551,1980,2001]" pageId="6" pageNumber="5">n</emphasis>
.
</taxonomicNameLabel>
, paratype male (FKCP), pedipalp. chela, dorsal (7), external (8), and ventral (9) views. Patella, dorsal (10), external (11) and ventral (12) views. Femur and trochanter dorsal (13). Movable finger (14).
</paragraph>
</caption>
<caption id="A8C8667AFFA0FD09645436A27A5DF805" ID-DOI="http://doi.org/10.5281/zenodo.5742165" ID-Zenodo-Dep="5742165" httpUri="https://zenodo.org/record/5742165/files/figure.png" pageId="7" pageNumber="6" startId="7.[115,192,1979,2001]" targetBox="[113,1461,244,1949]" targetPageId="7">
<paragraph id="FC0836F2FFA0FD09645436A27A5DF805" blockId="7.[115,1473,1979,2030]" pageId="7" pageNumber="6">
<emphasis id="CEC3EAE0FFA0FD09645436A27F0BF83B" bold="true" box="[115,258,1979,2001]" pageId="7" pageNumber="6">Figures 1521</emphasis>
.
<taxonomicName id="3BB74D71FFA0FD09652836A27FF7F83A" authority="Ayrey &amp; Kovařík &amp; Myers, 2021" authorityName="Ayrey &amp; Kovařík &amp; Myers" authorityYear="2021" box="[271,510,1979,2001]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="moyeri" status="sp. nov.">
<emphasis id="CEC3EAE0FFA0FD09652836A27FF7F83A" box="[271,510,1979,2001]" italics="true" pageId="7" pageNumber="6">Pseudouroctonus moyeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D5F0579BFFA0FD09662336A27C33F83B" box="[516,570,1979,2000]" pageId="7" pageNumber="6" rank="species">
<emphasis id="CEC3EAE0FFA0FD09662336A27C12F83B" bold="true" box="[516,539,1979,2000]" pageId="7" pageNumber="6">sp</emphasis>
.
<emphasis id="CEC3EAE0FFA0FD09660F36A27C3CF83B" bold="true" box="[552,565,1979,2000]" pageId="7" pageNumber="6">n</emphasis>
.
</taxonomicNameLabel>
, paratype male (FKCP), sternopectinal area (15), left chelicerae, ventral view (16), right legs IIIIV, retrolateral aspect (1718), metasoma and telson lateral (19), ventral (20) and dorsal (21) views.
</paragraph>
</caption>
<caption id="A8C8667AFFAFFD066454368D7EE9F80F" ID-DOI="http://doi.org/10.5281/zenodo.5742169" ID-Zenodo-Dep="5742169" httpUri="https://zenodo.org/record/5742169/files/figure.png" pageId="8" pageNumber="7" startId="8.[115,192,1940,1962]" targetBox="[120,1469,280,1879]" targetPageId="8">
<paragraph id="FC0836F2FFAFFD066454368D7EE9F80F" blockId="8.[115,1473,1940,2020]" pageId="8" pageNumber="7">
<emphasis id="CEC3EAE0FFAFFD066454368D7F0CF842" bold="true" box="[115,261,1940,1962]" pageId="8" pageNumber="7">Figures 2231</emphasis>
:
<taxonomicName id="3BB74D71FFAFFD066533368D7C0FF841" authority="Ayrey &amp; Kovařík &amp; Myers, 2021" authorityName="Ayrey &amp; Kovařík &amp; Myers" authorityYear="2021" box="[276,518,1940,1962]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="moyeri" status="sp. nov.">
<emphasis id="CEC3EAE0FFAFFD066533368D7C0FF841" box="[276,518,1940,1962]" italics="true" pageId="8" pageNumber="7">Pseudouroctonus moyeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D5F0579BFFAFFD066628368D7C4EF841" box="[527,583,1940,1962]" pageId="8" pageNumber="7" rank="species">
<emphasis id="CEC3EAE0FFAFFD066628368D7C2FF842" bold="true" box="[527,550,1940,1961]" pageId="8" pageNumber="7">sp</emphasis>
.
<emphasis id="CEC3EAE0FFAFFD066612368D7C4BF842" bold="true" box="[565,578,1940,1961]" pageId="8" pageNumber="7">n</emphasis>
.
</taxonomicNameLabel>
, paratype male (USNM).
<emphasis id="CEC3EAE0FFAFFD066768368D7DE8F842" bold="true" box="[847,993,1940,1962]" pageId="8" pageNumber="7">Figures 2225</emphasis>
. Hemispermatophore internal (22), ventral (23), external (24), and dorsal (25) views.
<emphasis id="CEC3EAE0FFAFFD0665E836A87C54F82D" bold="true" box="[463,605,1969,1991]" pageId="8" pageNumber="7">Figures 2631</emphasis>
. Mating plug dorsal (26), distal (27), ventral (28), proximal (29), internal (30), and external (31) views.
</paragraph>
</caption>
<paragraph id="FC0836F2FFAEFD07645631F27EBDFEC8" blockId="9.[113,772,235,2019]" pageId="9" pageNumber="8">
DESCRIPTION. Based on
<typeStatus id="230C8850FFAEFD0765A231F27FEBFEE8" box="[389,482,235,259]" pageId="9" pageNumber="8" type="holotype">holotype</typeStatus>
female, unless otherwise noted.
</paragraph>
<paragraph id="FC0836F2FFAEFD07645630337ECEFE88" blockId="9.[113,772,235,2019]" pageId="9" pageNumber="8">
<emphasis id="CEC3EAE0FFAEFD07645630337EBBFEA9" bold="true" box="[113,178,298,322]" pageId="9" pageNumber="8">Color</emphasis>
. Color is dark, reddish brown, lighter on the legs, telson reddish.
</paragraph>
<paragraph id="FC0836F2FFAEFD07645630737EB0FD28" blockId="9.[113,772,235,2019]" pageId="9" pageNumber="8">
<emphasis id="CEC3EAE0FFAEFD07645630737ED6FE69" bold="true" box="[113,223,362,386]" pageId="9" pageNumber="8">Carapace</emphasis>
. Carapace of female is longer than the metasomal segment V. Anterior edge with a conspicuous median indentation, providing a ratio of 0.066 when its depth is compared to the carapaces length; edge with six setae visible; entire surface moderately covered with medium sized granules. Three lateral eyes are present. Median eyes and tubercle of medium size, positioned anterior of middle with the following length and width ratios: 0.344 (anterior edge to medium tubercle middle / carapace length) and 0.129 (width of median tubercle including eyes / width of carapace at that point).
</paragraph>
<paragraph id="FC0836F2FFAEFD07645633D27C4CFB08" blockId="9.[113,772,235,2019]" pageId="9" pageNumber="8">
<emphasis id="CEC3EAE0FFAEFD07645633D27EEEFD08" bold="true" box="[113,231,715,739]" pageId="9" pageNumber="8">Metasoma</emphasis>
. Length/width ratio of segment I 0.76; segment II 0.95; segment III 1.02; segment IV 1.42; segment V 2.30. Segments IIV: dorsolateral carinae moderate, granular with distal denticle of I-IV enlarged and spinoid. Lateral supramedian carinae IIV strong and granular with enlarged spinoid distal denticle. Lateral inframedian carinae moderately granular on segment I, posterior half of segment II, one-third of segment III, and obsolete on distal one-fourth of segment IV. Ventrolateral carinae on segments III strong, granular; on IIIIV moderate, granular. Ventral submedian carinae weak on segment I and II, weak to moderate on III, moderate, granular on IV. Dorsal and lateral intercarinal spaces finely granular. Segment IIV ventral submedian setae 3/3. Segment V dorsolateral carinae strong. Lateromedian carinae moderate and granular on basal 3/5, obsolete on distal 2/5. Ventrolateral and ventromedian carinae strong. Intercarinal spaces finely granular. Segment V ventrolateral setae 3/3.
</paragraph>
<paragraph id="FC0836F2FFAEFD07645635F27F2AFA89" blockId="9.[113,772,235,2019]" pageId="9" pageNumber="8">
<emphasis id="CEC3EAE0FFAEFD07645635F27EB3FAE8" bold="true" box="[113,186,1259,1283]" pageId="9" pageNumber="8">Telson</emphasis>
. Smooth with 4 pairs of large setae on the ventral surface, 3 large setae along both lateral edges of the vesicle and numerous smaller setae. Small, spinoid subaculear tubercle present.
</paragraph>
<paragraph id="FC0836F2FFAEFD07645634727C22F8E8" blockId="9.[113,772,235,2019]" pageId="9" pageNumber="8">
<emphasis id="CEC3EAE0FFAEFD07645634727EE0FA68" bold="true" box="[113,233,1387,1411]" pageId="9" pageNumber="8">Mesosoma</emphasis>
. Tergites finely granular, with vestigial median carina on tergites IVI. Tergite VII with weak median carina on anterior one-third and strong dorsal lateral and lateral supramedian granular carinae. Sternites IIIVI very finely granular and without carinae. Sternite VII with vestigial ventral lateral carinae on middle third. Presternites smooth. Spiracles ovoid with median side rotated 35 degrees from posterior sternite margin. Sternites with variable number of microsetae. Pectinal tooth count 08/08 for
<typeStatus id="230C8850FFAEFD07666B37727CA0F968" box="[588,681,1643,1667]" pageId="9" pageNumber="8" type="holotype">holotype</typeStatus>
female. All pectinal teeth have exterodistal angling with large sensorial area. Middle lamellae 6/6. Fulcra are present. Each fulcra with 13 central setae. Sternum is Type 2. Genital Operculum is sclerites separated on posterior one-third.
</paragraph>
<paragraph id="FC0836F2FFAEFD07645636137FC1F888" blockId="9.[113,772,235,2019]" pageId="9" pageNumber="8">
<emphasis id="CEC3EAE0FFAEFD07645636137EE3F8C9" bold="true" box="[113,234,1802,1826]" pageId="9" pageNumber="8">Chelicerae</emphasis>
. Dorsal edge of movable cheliceral finger with two subdistal (
<emphasis id="CEC3EAE0FFAEFD07652936327F2FF8A8" box="[270,294,1835,1859]" italics="true" pageId="9" pageNumber="8">sd</emphasis>
) denticles. Ventral edge is smooth, with well developed serrula on distal half.
</paragraph>
<paragraph id="FC0836F2FFAEFD07645636737D70FE28" blockId="9.[113,772,235,2019]" lastBlockId="9.[816,1475,235,515]" pageId="9" pageNumber="8">
<emphasis id="CEC3EAE0FFAEFD07645636737EE9F869" bold="true" box="[113,224,1898,1922]" pageId="9" pageNumber="8">Pedipalps</emphasis>
. Trichobothrial pattern
<typeStatus id="230C8850FFAEFD0765FB36727C03F868" box="[476,522,1899,1923]" pageId="9" pageNumber="8">type</typeStatus>
C (
<bibRefCitation id="98264B03FFAEFD07660936727CCCF868" author="VACHON" box="[558,709,1899,1923]" pageId="9" pageNumber="8" pagination="857 - 958" refId="ref4635" refString="VACHON, M. 1974. Etude des caracteres utilizes pour classer les familles et les genres de Scorpions (Arachnides). 1. La trichobothriotaxie enArachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les Scorpions. Bulletin du Museum National d'Historie Naturelle, Paris, (Ser. 3), 140 (Zool. 104): 857 - 958." type="journal article" year="1974">Vachon, 1974</bibRefCitation>
,
<figureCitation id="648C2A77FFAEFD0766E936727EA2F848" captionStart="Figures 714" captionStartId="6.[115,192,1980,2002]" captionTargetBox="[113,1459,299,1907]" captionTargetId="figure-55@6.[113,1474,287,1955]" captionTargetPageId="6" captionText="Figures 714. Pseudouroctonus moyeri sp. n., paratype male (FKCP), pedipalp. chela, dorsal (7), external (8), and ventral (9) views. Patella, dorsal (10), external (11) and ventral (12) views. Femur and trochanter dorsal (13). Movable finger (14)." figureDoi="http://doi.org/10.5281/zenodo.5742161" httpUri="https://zenodo.org/record/5742161/files/figure.png" pageId="9" pageNumber="8">Figs. 713</figureCitation>
). Trichobothria
<emphasis id="CEC3EAE0FFAEFD07657E36927F67F848" box="[345,366,1931,1955]" italics="true" pageId="9" pageNumber="8">ib</emphasis>
and
<emphasis id="CEC3EAE0FFAEFD07658236927FBDF849" box="[421,436,1931,1954]" italics="true" pageId="9" pageNumber="8">it</emphasis>
are at the base of fixed finger. Pedipalp ratios: chela length/width 3.22; femur length/width 2.73; patella length/width 2.45; fixed finger length/carapace length 0.67. Chela with carinae moderate. Fixed finger median denticles (MD) aligned and divided into 6 subrows by 5 outer denticles (OD) and 6 inner denticles (ID). Movable finger with 6 subrows of MD, 5 OD and usually 7 ID (Soleglad &amp; Sissom, 2001). Femur with carinae moderate. Patella with carinae strong, internal surface with very large granules on the
<emphasis id="CEC3EAE0FFAEFD0761A230927BCBFE49" box="[1413,1474,395,418]" italics="true" pageId="9" pageNumber="8">DPSc</emphasis>
carina.
</paragraph>
<paragraph id="FC0836F2FFAEFD07671730D27B60FDE8" blockId="9.[816,1475,235,515]" pageId="9" pageNumber="8">
<emphasis id="CEC3EAE0FFAEFD07671730D27D6CFE08" bold="true" box="[816,869,459,483]" pageId="9" pageNumber="8">Legs</emphasis>
. Ventral surface of tarsomere II with single median row of spinules terminating distally with one spinule pair.
</paragraph>
<paragraph id="FC0836F2FFAEFD07671733317A95FD74" blockId="9.[816,1474,551,672]" pageId="9" pageNumber="8">VARIABILITY. Pectinal tooth count 8/8 [n=1], 9/9 [n=4], 9/10 [n=1] with a mean of 8.916 [n=12], standard deviation 0.493 for females and 11/11[n=3] with a mean of 11.00 [n=6], standard deviation 0.00 for males.</paragraph>
<paragraph id="FC0836F2FFAEFD07671733DD7D82FBB7" blockId="9.[816,1475,708,1116]" pageId="9" pageNumber="8">
REPRODUCTION. Four females were kept alive in order to observe them giving birth and to count the number of first instar juveniles (see
<figureCitation id="648C2A77FFAEFD07603A321D7A7BFCF7" box="[1053,1138,772,796]" captionStart="Figure 3233" captionStartId="10.[115,183,2005,2027]" captionTargetBox="[113,1474,234,1973]" captionTargetId="figure-37@10.[113,1474,234,1975]" captionTargetPageId="10" captionText="Figure 3233. Pseudouroctonus moyeri sp. n., paratype juvenile female 4th instar (32) and female with newborns (33)." figureDoi="http://doi.org/10.5281/zenodo.5742173" httpUri="https://zenodo.org/record/5742173/files/figure.png" pageId="9" pageNumber="8">Fig. 33</figureCitation>
). All
<specimenCount id="EAB1FD7BFFAEFD07609F321D7B4DFCF7" box="[1208,1348,772,796]" pageId="9" pageNumber="8" type="female">four females</specimenCount>
gave birth between the 28th of August and the 2nd of September, 2015. The juvenile counts were 34, 32, 35, and 35; mean = 34.0 (n=4), SD 1.2247. The 1
<superScript id="0BC29BBAFFAEFD076067327A7A43FC9A" attach="left" box="[1088,1098,867,881]" fontSize="6" pageId="9" pageNumber="8">st</superScript>
instar orientation on the mothers back was “non-random”, as reported by
<bibRefCitation id="98264B03FFAEFD0760C2329D7B67FC77" author="AYREY" box="[1253,1390,900,924]" pageId="9" pageNumber="8" pagination="1 - 12" refId="ref4021" refString="AYREY, R. F. 2009. Sky Island Vaejovis: A new species (Scorpiones: Vaejovidae). Euscorpius, 86: 1 - 12." type="journal article" year="2009">Ayrey (2009</bibRefCitation>
,
<bibRefCitation id="98264B03FFAEFD07615D329D7BB7FC77" author="AYREY" box="[1402,1470,900,924]" pageId="9" pageNumber="8" pagination="1 - 15" refId="ref4050" refString="AYREY, R. F. 2013 a. Reproduction and birth in the vorhiesi group of the genus Vaejovis (Scorpiones: Vaejovidae). Part 1. Clutch size. Euscorpius, 166: 1 - 15." type="journal article" year="2013">2013a</bibRefCitation>
,
<bibRefCitation id="98264B03FFAEFD07671732BD7D7FFC57" author="AYREY" box="[816,886,932,956]" pageId="9" pageNumber="8" pagination="1 - 13" refId="ref4090" refString="AYREY, R. F. 2013 b. A new species of Vaejovis from the Mogollon Rim of northern Arizona (Scorpiones: Vaejovidae). Euscorpius, 176: 1 - 13." type="journal article" year="2013">2013b</bibRefCitation>
,
<bibRefCitation id="98264B03FFAEFD0767A532BD7DB5FC57" author="AYREY" box="[898,956,932,956]" pageId="9" pageNumber="8" pagination="1 - 12" refId="ref4125" refString="AYREY, R. F. 2020. A new species of Vaejovis from Mingus Mountain, northern Arizona (Scorpiones: Vaejovidae). Euscorpius, 303: 1 - 12." type="journal article" year="2020">2020</bibRefCitation>
). They were facing anteriorly with the prosoma down and the metasoma raised over the prosoma of the juvenile immediately posterior to them, the same orientation as in
<taxonomicName id="3BB74D71FFAEFD076754351D7A91FBF7" authorityName="Ayrey &amp; Soleglad" authorityYear="2015" box="[883,1176,1028,1052]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="santarita">
<emphasis id="CEC3EAE0FFAEFD076754351D7A91FBF7" box="[883,1176,1028,1052]" italics="true" pageId="9" pageNumber="8">Pseudouroctonus santarita</emphasis>
</taxonomicName>
and
<taxonomicName id="3BB74D71FFAEFD0760F1351C7DD5FBD7" authority="(Ayrey &amp; Soleglad, 2015)" baseAuthorityName="Ayrey &amp; Soleglad" baseAuthorityYear="2015" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="kremani">
<emphasis id="CEC3EAE0FFAEFD0760F1351C7B45FBF7" box="[1238,1356,1028,1052]" italics="true" pageId="9" pageNumber="8">P. kremani</emphasis>
(Ayrey &amp; Soleglad, 2015)
</taxonomicName>
. Postpartum behavior is as described in
<bibRefCitation id="98264B03FFAEFD0761A5353D7D8EFBB7" author="AYREY" pageId="9" pageNumber="8" pagination="1 - 15" refId="ref4050" refString="AYREY, R. F. 2013 a. Reproduction and birth in the vorhiesi group of the genus Vaejovis (Scorpiones: Vaejovidae). Part 1. Clutch size. Euscorpius, 166: 1 - 15." type="journal article" year="2013">Ayrey (2013a)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B4AD6579FFAEFD076717359D7BCAFB57" pageId="9" pageNumber="8" type="distribution">
<paragraph id="FC0836F2FFAEFD076717359D7BCAFB57" blockId="9.[816,1475,1156,1212]" pageId="9" pageNumber="8">
<materialsCitation id="4CDF3CAFFFAEFD076717359D7BB6FB57" country="United States of America" county="Graham County" location="Mount Graham" municipality="Pinaleno Mountains" pageId="9" pageNumber="8" specimenCount="1" stateProvince="Arizona">
DISTRIBUTION. Known only from the
<typeStatus id="230C8850FFAEFD0760F9359C7B05FB76" box="[1246,1292,1157,1181]" pageId="9" pageNumber="8">type</typeStatus>
locality,
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,
<collectingMunicipality id="1C6CAC88FFAEFD0767B535BD7A60FB57" box="[914,1129,1188,1212]" pageId="9" pageNumber="8">Pinaleño Mountains</collectingMunicipality>
,
<collectingCounty id="15694E7EFFAEFD07605535BD7B14FB57" box="[1138,1309,1188,1212]" pageId="9" pageNumber="8">Graham County</collectingCounty>
,
<collectingRegion id="3E73F810FFAEFD07610335BD7B74FB57" box="[1316,1405,1188,1212]" country="United States of America" name="Arizona" pageId="9" pageNumber="8">Arizona</collectingRegion>
,
<collectingCountry id="84A07662FFAEFD0761A135BD7BB6FB57" box="[1414,1471,1188,1212]" name="United States of America" pageId="9" pageNumber="8">USA</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>