194 lines
23 KiB
XML
194 lines
23 KiB
XML
<document id="C233B1F3873BCEBF9B5C06F8ADA1AE98" ID-DOI="10.11646/zootaxa.4044.2.1" ID-GBIF-Dataset="4be691ce-6649-4fc2-9c81-b54c72c2179d" ID-ISSN="1175-5326" ID-Zenodo-Dep="235030" ID-ZooBank="4ABB85F0-D989-4702-9962-BA6E4D26BDE2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461338415922" checkinUser="plazi" docAuthor="Kupriyanova, Elena K. & Ippolitov, Alexei P." docDate="2015" docId="741287F4FFF68C30FF25FA1D9683232C" docLanguage="en" docName="zt04044p200.pdf" docOrigin="Zootaxa 4044 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Spirodiscus Fauvel 1909" docType="treatment" docVersion="10" lastPageNumber="163" masterDocId="882BFF8CFFFD8C3CFFB2FFBD972B2071" masterDocTitle="Deep-sea serpulids (Annelida: Polychaeta) in tetragonal tubes: on a tube convergence path from the Mesozoic to Recent" masterLastPageNumber="200" masterPageNumber="151" pageNumber="162" updateTime="1698619758002" updateUser="plazi">
|
||
<mods:mods id="4A723CC493B19B365DC1B31D83CF4B68" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="EA9EBF395F77D48CD80546A5D591971E">
|
||
<mods:title id="9F81F6967D3A2D4F7C2386110503F3D5">Deep-sea serpulids (Annelida: Polychaeta) in tetragonal tubes: on a tube convergence path from the Mesozoic to Recent</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="C08EF43DD8C2FD20B99A78A9B8706601" type="personal">
|
||
<mods:role id="557A9C4E8625FC1D6F6ACD0E716CDB24">
|
||
<mods:roleTerm id="8091756FD03F4158E764DCF5B3060F54">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="82C49D492E64A5EB04557BC3A4AB981B">Kupriyanova, Elena K.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="D188C7A00E89AA3F33DDB21078921B8D" type="personal">
|
||
<mods:role id="D1FE48330E5B3F8C43F106511DD0029D">
|
||
<mods:roleTerm id="100949B4530463C1735EF92FEF6A0607">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="333C7E8C2C925233F3C8EA4B22FFCD66">Ippolitov, Alexei P.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="86C4CA40173F27E99B01B06EB2EA783F">text</mods:typeOfResource>
|
||
<mods:relatedItem id="A4C59C9A7A0B3E4CB0B6CE6B60EC5115" type="host">
|
||
<mods:titleInfo id="B8CCFA0937732B999CB9543337BF1AE0">
|
||
<mods:title id="D83B76B842264C5EB98029E0A32E616E">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="22E4BC67ED0FF5D2B599B8B1CF81F58A">
|
||
<mods:date id="B9398D089B14F4EEBE17A447826545EF">2015</mods:date>
|
||
<mods:detail id="18B3968DC858CA7423568C91089F9665" type="volume">
|
||
<mods:number id="1A6AD69123E92865488B9F43252D2E97">4044</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="A2F4BEB09621918775F615BD27A57A6D" type="issue">
|
||
<mods:number id="0E0DC4BA9D349FFD94BE9ABE70F4BE03">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="83968C1C5E956411F2BAF950B6D26F8E" unit="page">
|
||
<mods:start id="AEF2602A293D80B4EBC164C5479081C1">151</mods:start>
|
||
<mods:end id="C4D14922EA10275CBD361FAA6901E97D">200</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="5EE9A01040C4D936D7EA4614AEBC71EF">journal article</mods:classification>
|
||
<mods:identifier id="7833A7F3791BDFD5BB9C05AACCEB8255" type="DOI">10.11646/zootaxa.4044.2.1</mods:identifier>
|
||
<mods:identifier id="BFA65CC7A960C3E0C6989C40FE60D11B" type="GBIF-Dataset">4be691ce-6649-4fc2-9c81-b54c72c2179d</mods:identifier>
|
||
<mods:identifier id="8A965B569C65A4616B470C068DB71FFD" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="EB167EEAFA7035B3ECB7928072A80568" type="Zenodo-Dep">235030</mods:identifier>
|
||
<mods:identifier id="D21A944E59E85B391A6995D13DE1774A" type="ZooBank">4ABB85F0-D989-4702-9962-BA6E4D26BDE2</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="741287F4FFF68C30FF25FA1D9683232C" ID-DOI="http://doi.org/10.5281/zenodo.5324170" ID-GBIF-Taxon="119644900" ID-Zenodo-Dep="5324170" LSID="urn:lsid:plazi:treatment:741287F4FFF68C30FF25FA1D9683232C" httpUri="http://treatment.plazi.org/id/741287F4FFF68C30FF25FA1D9683232C" lastPageId="12" lastPageNumber="163" pageId="11" pageNumber="162">
|
||
<subSubSection id="B4A16569FFF68C37FF25FA1D950B25CB" box="[151,544,1440,1466]" pageId="11" pageNumber="162" type="nomenclature">
|
||
<paragraph id="FC0436E2FFF68C37FF25FA1D950B25CB" blockId="11.[151,544,1440,1466]" box="[151,544,1440,1466]" pageId="11" pageNumber="162">
|
||
<heading id="A74C818EFFF68C37FF25FA1D950B25CB" bold="true" box="[151,544,1440,1466]" fontSize="11" level="1" pageId="11" pageNumber="162" reason="1">
|
||
<emphasis id="CECFEAF0FFF68C37FF25FA1D950B25CB" bold="true" box="[151,544,1440,1466]" pageId="11" pageNumber="162">
|
||
Genus
|
||
<taxonomicName id="3BBB4D61FFF68C37FF5CFA1D950B25CB" ID-CoL="84TS9" authority="Fauvel, 1909" authorityName="Fauvel" authorityYear="1909" box="[238,544,1440,1466]" class="Polychaeta" family="Serpulidae" genus="Spirodiscus" higherTaxonomySource="CoL" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="162" phylum="Annelida" rank="genus">
|
||
<emphasis id="CECFEAF0FFF68C37FF5CFA1D965125CB" bold="true" box="[238,378,1440,1466]" italics="true" pageId="11" pageNumber="162">Spirodiscus</emphasis>
|
||
<bibRefCitation id="982A4B13FFF68C37FE33FA1D950B25CB" author="Fauvel" box="[385,544,1440,1466]" pageId="11" pageNumber="162" refString="Fauvel, P. (1909) Deuxieme note preliminaire sur les polychetes provenant des campagnes de l'Hirondelle et de la princesse Alice ou deposes dans le Musee oceanographique de Monaco. Bulletin de l'Institute Oceanographique, 142, 1 - 76." type="journal article" year="1909">Fauvel, 1909</bibRefCitation>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B4A16569FFF68C37FF25FA5B932B266A" pageId="11" pageNumber="162" type="reference_group">
|
||
<paragraph id="FC0436E2FFF68C37FF25FA5B95F8258D" blockId="11.[151,1024,1510,1563]" box="[151,723,1510,1532]" pageId="11" pageNumber="162">
|
||
<treatmentCitationGroup id="DCAB11CCFFF68C37FF25FA5B95F8258D" box="[151,723,1510,1532]" pageId="11" pageNumber="162">
|
||
<taxonomicName id="3BBB4D61FFF68C37FF25FA5B9696258D" ID-CoL="84TS9" authority="Fauvel, 1909: 56" authorityName="Fauvel" authorityPageNumber="56" authorityYear="1909" box="[151,445,1510,1532]" class="Polychaeta" family="Serpulidae" genus="Spirodiscus" higherTaxonomySource="CoL" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="162" phylum="Annelida" rank="genus">
|
||
<emphasis id="CECFEAF0FFF68C37FF25FA5B9627258D" box="[151,268,1510,1532]" italics="true" pageId="11" pageNumber="162">Spirodiscus</emphasis>
|
||
<treatmentCitation id="7D1A10F3FFF68C37FEA0FA5B9696258D" author="Fauvel" box="[274,445,1510,1532]" page="56" pageId="11" pageNumber="162" year="1909">
|
||
<bibRefCitation id="982A4B13FFF68C37FEA0FA5B96B1258D" author="Fauvel" box="[274,410,1510,1532]" pageId="11" pageNumber="162" refString="Fauvel, P. (1909) Deuxieme note preliminaire sur les polychetes provenant des campagnes de l'Hirondelle et de la princesse Alice ou deposes dans le Musee oceanographique de Monaco. Bulletin de l'Institute Oceanographique, 142, 1 - 76." type="journal article" year="1909">Fauvel, 1909</bibRefCitation>
|
||
: 56
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
–57.—
|
||
<treatmentCitation id="7D1A10F3FFF68C37FDB2FA5B95E4258D" author="Fauchald" box="[512,719,1510,1532]" page="147" pageId="11" pageNumber="162" year="1977">
|
||
<bibRefCitation id="982A4B13FFF68C37FDB2FA5B95B6258D" author="Fauchald" box="[512,669,1510,1532]" pageId="11" pageNumber="162" refString="Fauchald, K. (1977) The polychaete worms. Definitions and keys to the orders, families and genera. Natural History Museum of Los Angeles County Science Series, 28, 1 - 188." type="journal article" year="1977">Fauchald 1977</bibRefCitation>
|
||
: 147
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="FC0436E2FFF68C37FF25F9B8932B266A" blockId="11.[151,1024,1510,1563]" box="[151,1024,1541,1563]" pageId="11" pageNumber="162">
|
||
<treatmentCitationGroup id="DCAB11CCFFF68C37FF25F9B8932B266A" box="[151,1024,1541,1563]" pageId="11" pageNumber="162">
|
||
<taxonomicName id="3BBB4D61FFF68C37FF25F9B897C6266A" ID-CoL="87JGN" box="[151,237,1541,1563]" class="Polychaeta" family="Serpulidae" genus="Nogrobs" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="162" phylum="Annelida" rank="genus">
|
||
<emphasis id="CECFEAF0FFF68C37FF25F9B897C6266A" box="[151,237,1541,1563]" italics="true" pageId="11" pageNumber="162">Nogrobs</emphasis>
|
||
</taxonomicName>
|
||
.—ten
|
||
<treatmentCitation id="7D1A10F3FFF68C37FE80F9BB9540266A" author="Hove" box="[306,619,1541,1563]" httpUri="http://treatment.plazi.org/id/03BD87F8C93CFF8C7E93FF4F1A03136E" page="68" pageId="11" pageNumber="162" year="2009">
|
||
<bibRefCitation id="982A4B13FFF68C37FE80F9BB9561266A" author="Hove" box="[306,586,1541,1563]" pageId="11" pageNumber="162" refString="Kupriyanova, E. K., Hove, H. A. ten, Sket, B., Zaksek, V., Trontelj, P. & Rouse, G. W. (2009) Evolution of the unique freshwater cave-dwelling tube worm Marifugia cavatica (Annelida: Serpulidae). Systematics and Biodiversity, 7, 389 - 401. http: // dx. doi. org / 10.1017 / S 1477200009990168" type="journal article" year="2009">Hove & Kupriyanova 2009</bibRefCitation>
|
||
: 68
|
||
</treatmentCitation>
|
||
–69.—Kupriyanova & Nishi 2011: 1–2.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B4A16569FFF68C37FF25F9F595E82611" box="[151,707,1608,1633]" pageId="11" pageNumber="162" type="materials_examined">
|
||
<paragraph id="FC0436E2FFF68C37FF25F9F595E82611" blockId="11.[151,1437,1608,2029]" box="[151,707,1608,1633]" pageId="11" pageNumber="162">
|
||
<emphasis id="CECFEAF0FFF68C37FF25F9F596002610" bold="true" box="[151,299,1608,1633]" pageId="11" pageNumber="162">
|
||
<typeStatus id="23008840FFF68C37FF25F9F597F92610" box="[151,210,1608,1633]" pageId="11" pageNumber="162">Type</typeStatus>
|
||
species
|
||
</emphasis>
|
||
:
|
||
<taxonomicName id="3BBB4D61FFF68C37FE88F9F595E82611" authority="Fauvel, 1909" authorityName="Fauvel" authorityYear="1909" box="[314,707,1608,1633]" class="Polychaeta" family="Serpulidae" genus="Spirodiscus" higherTaxonomySource="CoL" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="162" phylum="Annelida" rank="species" species="grimaldii">
|
||
<emphasis id="CECFEAF0FFF68C37FE88F9F595012610" box="[314,554,1608,1633]" italics="true" pageId="11" pageNumber="162">Spirodiscus grimaldii</emphasis>
|
||
<bibRefCitation id="982A4B13FFF68C37FD83F9F595E82611" author="Fauvel" box="[561,707,1608,1633]" pageId="11" pageNumber="162" refString="Fauvel, P. (1909) Deuxieme note preliminaire sur les polychetes provenant des campagnes de l'Hirondelle et de la princesse Alice ou deposes dans le Musee oceanographique de Monaco. Bulletin de l'Institute Oceanographique, 142, 1 - 76." type="journal article" year="1909">Fauvel, 1909</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B4A16569FFF68C30FF75F9D196AD2089" lastPageId="12" lastPageNumber="163" pageId="11" pageNumber="162" type="diagnosis">
|
||
<paragraph id="FC0436E2FFF68C30FF75F9D196AD2089" blockId="11.[151,1437,1608,2029]" lastBlockId="12.[151,1437,151,861]" lastPageId="12" lastPageNumber="163" pageId="11" pageNumber="162">
|
||
<emphasis id="CECFEAF0FFF68C37FF75F9D1961126F4" bold="true" box="[199,314,1644,1669]" pageId="11" pageNumber="162">Diagnosis</emphasis>
|
||
(after Kupriyanova & Nishi 2011, emend.): Tube white, free-lying, polygonal in cross-section, quadrangular or octagonal (= modified quadrangular), straight to slightly curved (tusk-shaped), or initially cylindrical, then quadrangular in cross-section and coiled clockwise into a flat spiral, finally with short cylindrical straight distal part. Peristomes absent. Operculum inverse cone (ampulla) with chitinous endplate and central depression. Peduncle thick, pinnulated, without distal wings, with dorsal groove distally, with or without constriction beneath ampulla; inserted as second right radiole, up to 3 times as wide as other radioles. Pseudoperculum absent. Arrangement of radioles semi-circular, up to 8 per lobe. Inter-radiolar membrane, radiolar eyes, and stylodes absent. Mouth palps absent. 5 or 6 thoracic chaetigerous segments. Collar non-lobed with straight edge, no clear separation towards thoracic membranes that end at second chaetiger; tonguelets between ventral and lateral collar parts absent. No apron. Collar chaetae limbate.
|
||
<taxonomicName id="3BBB4D61FFF68C37FC76F80C931F27B9" box="[964,1076,1969,1992]" class="Polychaeta" family="Serpulidae" genus="Apomatus" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="162" phylum="Annelida" rank="genus">
|
||
<emphasis id="CECFEAF0FFF68C37FC76F80C931F27B9" box="[964,1076,1969,1992]" italics="true" pageId="11" pageNumber="162">Apomatus</emphasis>
|
||
</taxonomicName>
|
||
chaetae absent. Thoracic uncini saw-to-rasp-shaped with numerous teeth (> 12) in profile, 2–3 teeth per row; anterior peg gouged. Thoracic triangular depression absent. Abdominal chaetae short, with flat triangular denticulate blade; uncini similar to thoracic ones. Achaetous anterior abdominal zone absent. Long posterior capillary chaetae absent. Posterior glandular pad absent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B4A16569FFF18C30FF75FEB99683232C" pageId="12" pageNumber="163" type="discussion">
|
||
<paragraph id="FC0436E2FFF18C30FF75FEB992B6224C" blockId="12.[151,1437,151,861]" pageId="12" pageNumber="163">
|
||
<emphasis id="CECFEAF0FFF18C30FF75FEB99610216C" bold="true" box="[199,315,260,285]" pageId="12" pageNumber="163">Remarks.</emphasis>
|
||
The Recent genus
|
||
<taxonomicName id="3BBB4D61FFF18C30FD96FEB99460216D" authority="Fauvel, 1909" authorityName="Fauvel" authorityYear="1909" box="[548,843,260,285]" class="Polychaeta" family="Serpulidae" genus="Spirodiscus" higherTaxonomySource="CoL" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="163" phylum="Annelida" rank="genus">
|
||
<emphasis id="CECFEAF0FFF18C30FD96FEB9958D216C" box="[548,678,260,285]" italics="true" pageId="12" pageNumber="163">Spirodiscus</emphasis>
|
||
<bibRefCitation id="982A4B13FFF18C30FD03FEB99460216D" author="Fauvel" box="[689,843,260,285]" pageId="12" pageNumber="163" refString="Fauvel, P. (1909) Deuxieme note preliminaire sur les polychetes provenant des campagnes de l'Hirondelle et de la princesse Alice ou deposes dans le Musee oceanographique de Monaco. Bulletin de l'Institute Oceanographique, 142, 1 - 76." type="journal article" year="1909">Fauvel, 1909</bibRefCitation>
|
||
</taxonomicName>
|
||
, characterised by an unattached quadrangular tube coiled into a flat spiral (hence the generic name), was synonymised with the fossil
|
||
<taxonomicName id="3BBB4D61FFF18C30FBF0FE9A938A2131" box="[1090,1185,295,320]" class="Polychaeta" family="Serpulidae" genus="Nogrobs" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="163" phylum="Annelida" rank="genus">
|
||
<emphasis id="CECFEAF0FFF18C30FBF0FE9A938A2131" box="[1090,1185,295,320]" italics="true" pageId="12" pageNumber="163">Nogrobs</emphasis>
|
||
</taxonomicName>
|
||
de
|
||
<bibRefCitation id="982A4B13FFF18C30FB7EFE9A92532131" author="Montfort" box="[1228,1400,295,320]" pageId="12" pageNumber="163" refString="Montfort, D. de. (1808) Conchyliologie systematique et classification methodique des coquilles, 1. Coquilles univalves, cloisonees. Schoell, Paris, 409 pp." type="book" year="1808">Montfort, 1808</bibRefCitation>
|
||
by
|
||
<bibRefCitation id="982A4B13FFF18C30FF25FEF0960C2114" author="Jager" box="[151,295,332,357]" pageId="12" pageNumber="163" refString="Jager, M. (2005) Serpulidae und Spirorbidae (Polychaeta sedentaria) aus Campan und Maastricht von Norddeutschland, den Niederlanden, Belgien und angrenzenden Gebieten. Geologisches Jahrbuch, A 157 (for 2004), 121 - 249." type="journal article" year="2005">Jäger (2005)</bibRefCitation>
|
||
. This was supported by ten
|
||
<bibRefCitation id="982A4B13FFF18C30FDD3FEF0948D2115" author="Hove" box="[609,934,332,357]" pageId="12" pageNumber="163" refString="Kupriyanova, E. K., Hove, H. A. ten, Sket, B., Zaksek, V., Trontelj, P. & Rouse, G. W. (2009) Evolution of the unique freshwater cave-dwelling tube worm Marifugia cavatica (Annelida: Serpulidae). Systematics and Biodiversity, 7, 389 - 401. http: // dx. doi. org / 10.1017 / S 1477200009990168" type="journal article" year="2009">Hove & Kupriyanova (2009)</bibRefCitation>
|
||
and Kupriyanova & Nishi (2011). However, tubes of Recent
|
||
<taxonomicName id="3BBB4D61FFF18C30FEEAFED2956621F9" box="[344,589,367,392]" class="Polychaeta" family="Serpulidae" genus="Spirodiscus" higherTaxonomySource="CoL" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="163" phylum="Annelida" rank="species" species="grimaldii">
|
||
<emphasis id="CECFEAF0FFF18C30FEEAFED2956621F9" box="[344,589,367,392]" italics="true" pageId="12" pageNumber="163">Spirodiscus grimaldii</emphasis>
|
||
</taxonomicName>
|
||
are strictly clockwise coiled (when looking from the upper side), while
|
||
<taxonomicName id="3BBB4D61FFF18C30FF25FE2997C821DC" box="[151,227,404,429]" class="Polychaeta" family="Serpulidae" genus="Ditrypa" higherTaxonomySource="CoL" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="163" phylum="Annelida" rank="species" species="known">known</taxonomicName>
|
||
populations of fossil representatives of genus
|
||
<taxonomicName id="3BBB4D61FFF18C30FC84FE2994BE21DC" box="[822,917,404,429]" class="Polychaeta" family="Serpulidae" genus="Nogrobs" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="163" phylum="Annelida" rank="genus">
|
||
<emphasis id="CECFEAF0FFF18C30FC84FE2994BE21DC" box="[822,917,404,429]" italics="true" pageId="12" pageNumber="163">Nogrobs</emphasis>
|
||
</taxonomicName>
|
||
demonstrate a mixture of clockwise and anticlockwise coiled specimens (Ippolitov, unpubl.). At the same time, although coiling direction appears to be a morphological argument against Jager’s synonymy, this character is not very important taxonomically. The results of comparative SEM studies of tube wall ultrastructures (see below) show very different crystal arrangement in
|
||
<taxonomicName id="3BBB4D61FFF18C30FF25FD999632224C" box="[151,281,548,573]" class="Polychaeta" family="Serpulidae" genus="Spirodiscus" higherTaxonomySource="CoL" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="163" phylum="Annelida" rank="genus">
|
||
<emphasis id="CECFEAF0FFF18C30FF25FD999632224C" box="[151,281,548,573]" italics="true" pageId="12" pageNumber="163">Spirodiscus</emphasis>
|
||
</taxonomicName>
|
||
and in the
|
||
<typeStatus id="23008840FFF18C30FE26FD9896EF224C" box="[404,452,549,573]" pageId="12" pageNumber="163">type</typeStatus>
|
||
species of genus
|
||
<taxonomicName id="3BBB4D61FFF18C30FD3AFD9995CC224C" box="[648,743,548,573]" class="Polychaeta" family="Serpulidae" genus="Nogrobs" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="163" phylum="Annelida" rank="genus">
|
||
<emphasis id="CECFEAF0FFF18C30FD3AFD9995CC224C" box="[648,743,548,573]" italics="true" pageId="12" pageNumber="163">Nogrobs</emphasis>
|
||
</taxonomicName>
|
||
, thus indicating that these genera should not be synonymised.
|
||
</paragraph>
|
||
<paragraph id="FC0436E2FFF18C30FF75FDFA9683232C" blockId="12.[151,1437,151,861]" pageId="12" pageNumber="163">
|
||
<taxonomicName id="3BBB4D61FFF18C30FF75FDFA96972211" box="[199,444,583,608]" class="Polychaeta" family="Serpulidae" genus="Spirodiscus" higherTaxonomySource="CoL" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="163" phylum="Annelida" rank="species" species="grimaldii">
|
||
<emphasis id="CECFEAF0FFF18C30FF75FDFA96972211" box="[199,444,583,608]" italics="true" pageId="12" pageNumber="163">Spirodiscus grimaldii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3BBB4D61FFF18C30FE4EFDF595902211" box="[508,699,583,608]" class="Polychaeta" family="Serpulidae" genus="Spirodiscus" higherTaxonomySource="CoL" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="163" phylum="Annelida" rank="species" species="groenlandicus" status="comb. nov.">
|
||
<emphasis id="CECFEAF0FFF18C30FE4EFDF595902211" box="[508,699,583,608]" italics="true" pageId="12" pageNumber="163">S. groenlandicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CECFEAF0FFF18C30FD75FDFA94602211" bold="true" box="[711,843,583,608]" pageId="12" pageNumber="163">
|
||
<taxonomicNameLabel id="D5FC578BFFF18C30FD75FDFA94602211" box="[711,843,583,608]" pageId="12" pageNumber="163">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(formerly
|
||
<taxonomicName id="3BBB4D61FFF18C30FC7CFDF493ED2211" box="[974,1222,583,608]" class="Polychaeta" family="Serpulidae" genus="Ditrupa" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="163" phylum="Annelida" rank="species" species="groenlandica">
|
||
<emphasis id="CECFEAF0FFF18C30FC7CFDF493ED2211" box="[974,1222,583,608]" italics="true" pageId="12" pageNumber="163">Ditrupa groenlandica</emphasis>
|
||
</taxonomicName>
|
||
) described below, have very different tube morphologies: coiled tetragonal in the former and tusk-shaped octagonal in the latter. However, the animals inhabiting these tubes are very similar (except for five thoracic chaetigers in
|
||
<taxonomicName id="3BBB4D61FFF18C30FA35FD2D961D22BC" class="Polychaeta" family="Serpulidae" genus="Spirodiscus" higherTaxonomySource="CoL" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="163" phylum="Annelida" rank="species" species="groenlandicus" status="comb. nov.">
|
||
<emphasis id="CECFEAF0FFF18C30FA35FD2D961D22BC" italics="true" pageId="12" pageNumber="163">S. groenlandicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CECFEAF0FFF18C30FEF2FD0996EF22BC" bold="true" box="[320,452,692,717]" pageId="12" pageNumber="163">
|
||
<taxonomicNameLabel id="D5FC578BFFF18C30FEF2FD0996EF22BC" box="[320,452,692,717]" pageId="12" pageNumber="163">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and six in
|
||
<taxonomicName id="3BBB4D61FFF18C30FDF9FD0895F922BC" box="[587,722,692,717]" class="Polychaeta" family="Serpulidae" genus="Spirodiscus" higherTaxonomySource="CoL" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="163" phylum="Annelida" rank="species" species="grimaldii">
|
||
<emphasis id="CECFEAF0FFF18C30FDF9FD0895F922BC" box="[587,722,692,717]" italics="true" pageId="12" pageNumber="163">S. grimaldii</emphasis>
|
||
</taxonomicName>
|
||
) and have identical chaetation patterns. Their tube walls also show similar ultrastructures having a well-defined outer layer with characteristic crystal orientation (see below). All this indicates that
|
||
<taxonomicName id="3BBB4D61FFF18C30FE20FD4095332364" box="[402,536,764,789]" class="Polychaeta" family="Serpulidae" genus="Spirodiscus" higherTaxonomySource="CoL" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="163" phylum="Annelida" rank="species" species="grimaldii">
|
||
<emphasis id="CECFEAF0FFF18C30FE20FD4095332364" box="[402,536,764,789]" italics="true" pageId="12" pageNumber="163">S. grimaldii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3BBB4D61FFF18C30FDE3FD4094262364" box="[593,781,764,789]" class="Polychaeta" family="Serpulidae" genus="Spirodiscus" higherTaxonomySource="CoL" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="163" phylum="Annelida" rank="species" species="groenlandicus" status="comb. nov.">
|
||
<emphasis id="CECFEAF0FFF18C30FDE3FD4094262364" box="[593,781,764,789]" italics="true" pageId="12" pageNumber="163">S. groenlandicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CECFEAF0FFF18C30FCA4FD4194B32364" bold="true" box="[790,920,764,789]" pageId="12" pageNumber="163">
|
||
<taxonomicNameLabel id="D5FC578BFFF18C30FCA4FD4194B32364" box="[790,920,764,789]" pageId="12" pageNumber="163">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are closely related and justifies placing them into one genus. The generic diagnosis here has been emended to include species with both coiled tetragonal and straight octagonal tubes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |